227 lines
26 KiB
XML
227 lines
26 KiB
XML
<document ID-CLB-Dataset="94519" ID-DOI="http://dx.doi.org/10.3897/zookeys.1112.85752" ID-GBIF-Dataset="5b4d920a-898b-49da-b958-8ee8a0774e8a" ID-PMC="PMC9848642" ID-Pensoft-Pub="1313-2970-1112-11" ID-Pensoft-UUID="59936DE5F02E5E1D8346772863977AEA" ID-PubMed="36760626" ID-ZooBank="F9F1E9A2CB9249CB8D4CFB3FAC592F96" ModsDocID="1313-2970-1112-11" checkinTime="1657650131167" checkinUser="pensoft" docAuthor="Sanchez, Enrico Gerard S., Delocado, Emmanuel D. & Freitag, Hendrik" docDate="2022" docId="1F1266B781235291B023B3190488AEC5" docLanguage="en" docName="ZooKeys 1112: 11-25" docOrigin="ZooKeys 1112" docPubDate="2022-07-12" docSource="http://dx.doi.org/10.3897/zookeys.1112.85752" docTitle="Anacaena auxilium Sanchez & Delocado & Freitag 2022, sp. nov." docType="treatment" docUuid="E3BC4B40-3228-4317-8213-CF2BE3A13748" docUuidSource="ZooBank" docVersion="5" id="59936DE5F02E5E1D8346772863977AEA" lastPageNumber="11" masterDocId="59936DE5F02E5E1D8346772863977AEA" masterDocTitle="Two new species of Anacaena Thomson, 1859 (Coleoptera, Hydrophilidae) from Northern Luzon, Philippines" masterLastPageNumber="25" masterPageNumber="11" pageNumber="11" updateTime="1732844981149" updateUser="ExternalLinkService">
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<mods:mods id="4CD9551718DB77FBCEC0BB8392DE94AD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0B81A8A564F6875D9F94AD947F66AB7E">
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<mods:title id="FE3428B08B523223DCD7AB6CCD812FD5">Two new species of Anacaena Thomson, 1859 (Coleoptera, Hydrophilidae) from Northern Luzon, Philippines</mods:title>
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</mods:titleInfo>
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<mods:name id="D9643AE1986938A3E14F13C19E3C963E" type="personal">
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<mods:roleTerm id="AB84CF0E7AF41D086B9BD5050908C67A">Author</mods:roleTerm>
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<mods:namePart id="ED9E60F4B009DB08639215CE64DBA2DB">Sanchez, Enrico Gerard S.</mods:namePart>
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<mods:affiliation id="A7DF5E8B887FDA33315935F5F7E2E8B8">Ateneo Biodiversity Research Laboratory, Department of Biology, School of Science and Engineering, Ateneo de Manila University, Quezon City, 1108 Philippines</mods:affiliation>
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<mods:nameIdentifier id="6623DB877767B08262BBDBB75605D6B7" type="email">enrico.sanchez@obf.ateneo.edu</mods:nameIdentifier>
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<mods:roleTerm id="04C2FF97348EA642099FABFB35D25143">Author</mods:roleTerm>
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<mods:namePart id="AFB888ADA51F233A651FD420A78401CD">Delocado, Emmanuel D.</mods:namePart>
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<mods:nameIdentifier id="2B514C5047BA986D84DBDC39A4B7E9A4" type="ORCID">https://orcid.org/0000-0001-7731-3476</mods:nameIdentifier>
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<mods:affiliation id="A9C57530CC588EF4045EACDCEA57E7DA">Ateneo Biodiversity Research Laboratory, Department of Biology, School of Science and Engineering, Ateneo de Manila University, Quezon City, 1108 Philippines</mods:affiliation>
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<mods:roleTerm id="A78E8D68A6A5BE8E0D5F31124D68820C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E83D7B478118A08AF352D483156DEBEF">Freitag, Hendrik</mods:namePart>
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<mods:nameIdentifier id="0A6C4EC3C5CF26250331AE56121D71F8" type="ORCID">https://orcid.org/0000-0002-1325-0979</mods:nameIdentifier>
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<mods:affiliation id="53F5F7C9C5939DA1C4B21F2834C039E7">Ateneo Biodiversity Research Laboratory, Department of Biology, School of Science and Engineering, Ateneo de Manila University, Quezon City, 1108 Philippines</mods:affiliation>
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<mods:title id="DB63F11C0941FFA3E595E2FB56119CB8">ZooKeys</mods:title>
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<mods:part id="634FC80718A22C33EC6732260B7D0CB2">
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<mods:date id="195566C6946CBA3F4EED1787304F391D">2022</mods:date>
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<mods:number id="4A226B7F1DBCBC3F31730B91CC7684A9">2022-07-12</mods:number>
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<mods:number id="A4C3D4357B225A278E2C078B7F46ABBC">1112</mods:number>
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<mods:start id="2AE379A42D313808FCEE01A4A228D9D6">11</mods:start>
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<mods:end id="7F822442F273A0CB637B971825976A5F">25</mods:end>
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<mods:url id="0B39FF441F573135BAB4FB1A0F9987B7">http://dx.doi.org/10.3897/zookeys.1112.85752</mods:url>
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<mods:classification id="9F90B4B924EF432297B04D93B76A7326">journal article</mods:classification>
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<mods:identifier id="BB60FAC9A39E6E92ADE656D35B1B7F1A" type="DOI">http://dx.doi.org/10.3897/zookeys.1112.85752</mods:identifier>
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<treatment id="1F1266B781235291B023B3190488AEC5" ID-GBIF-Taxon="197228275" LSID="urn:lsid:zoobank.org:act:E3BC4B40-3228-4317-8213-CF2BE3A13748" httpUri="http://treatment.plazi.org/id/1F1266B781235291B023B3190488AEC5" lastPageNumber="11" pageId="0" pageNumber="11" scope_family="Hydrophilidae" scope_order="Coleoptera">
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<subSubSection id="3E7F70289B869AF1F1394CC30FF68693" pageId="0" pageNumber="11" type="nomenclature">
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<paragraph id="0CA8AE545A2CD3BDC72E7AECEBA8595D" pageId="0" pageNumber="11">
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<taxonomicName id="B2CD457E8E95DC58ECD1F3ECE341D50A" ID-CoL="9GDLB" LSID="http://zoobank.org/E3BC4B40-3228-4317-8213-CF2BE3A13748" authority="Sanchez & Delocado & Freitag, 2022" authorityName="Sanchez & Delocado & Freitag" authorityYear="2022" class="Insecta" family="Hydrophilidae" genus="Anacaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anacaena auxilium" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="auxilium" status="sp. nov.">Anacaena auxilium</taxonomicName>
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<taxonomicNameLabel id="0FF13DACE4A4161098A694557F43F47F" pageId="0" pageNumber="11">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="A12B00E582AAE1C43D785DFFA06A7B2A" pageId="0" pageNumber="11" type="description">
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<paragraph id="C5F63E87198FE45C1BCA10B4CF10AF43" pageId="0" pageNumber="11">
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<figureCitation id="BFDC73766CEF13F6FBFEE4FE09FECD97" captionStart="Figure 1–4" captionStartId="F1" captionText="Figure 1 - 4. Habitus of new Anacaena species 1, 2 Anacaena angatbuhay sp. nov. 3, 4 Anacaena auxilium sp. nov. in (1, 3) dorsal and (2, 4) lateral views. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1112.85752.figure1-4" httpUri="https://binary.pensoft.net/fig/713647" pageId="0" pageNumber="11">Figs 3-4</figureCitation>
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<figureCitation id="087A6199C4C1321CF4CE4C94EDBC2E67" captionStart="Figure 5–6" captionStartId="F2" captionText="Figure 5 - 6. Aedeagi of 5 Anacaena angatbuhay sp. nov. holotype (GS 076) 6 Anacaena auxilium sp. nov. holotype (GS 066) in dorsal view. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1112.85752.figure5-6" httpUri="https://binary.pensoft.net/fig/713648" pageId="0" pageNumber="11">, 6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="622ED7EB3CB7370DC57B4F39BD6B9C7E" pageId="0" pageNumber="11" type="type locality">
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<paragraph id="507697C0F7AA313CFF6C364FB75931DA" pageId="0" pageNumber="11">Type locality.</paragraph>
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<paragraph id="188C5C8A39B81A14F9B105634EC6AF99" pageId="0" pageNumber="11">
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Philippines • Luzon, Mountain Province, Bauko; Besang mountain river, Chico River tributary; rural/agricultural area; boulders;
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<geoCoordinate id="818614F2B99F0E9A151E94F7CC9BFE82" degrees="17" direction="north" minutes="00" orientation="latitude" precision="15" seconds="14" value="17.00389">17°00'14"N</geoCoordinate>
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,
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<geoCoordinate id="74FFAE2653B28BCC80E178E43919E761" degrees="120" direction="east" minutes="53" orientation="longitude" precision="15" seconds="49" value="120.89694">120°53'49"E</geoCoordinate>
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; ca. 1080 m a.s.l.
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</paragraph>
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</subSubSection>
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<subSubSection id="D827C419710FF6832DA5809170A41F09" pageId="0" pageNumber="11" type="type material">
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<paragraph id="E62154D52BCF515F975BEE96CF4685EA" pageId="0" pageNumber="11">Type material.</paragraph>
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<paragraph id="ADD9E5B100249A8F19651690F8C507D6" pageId="0" pageNumber="11">
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<emphasis id="491FDA282B2F7B9E9856F3B730758B4B" bold="true" italics="true" pageId="0" pageNumber="11">Holotype</emphasis>
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: Philippines • ♂ (PNM: GS066), "PHIL.: Luzon, Mountain Province, \ Bauko; Besang mt. river, Chico River tributary; \ rural/agricultural area; boulders, \
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<geoCoordinate id="A9AA1F5DD685A2DA82C809ECA2ACFF34" degrees="17" direction="north" minutes="00" orientation="latitude" precision="15" seconds="14" value="17.00389">17°00'14"N</geoCoordinate>
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;
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<geoCoordinate id="C606ACE66430D1269946C120C3AE4C08" degrees="120" direction="east" minutes="53" orientation="longitude" precision="15" seconds="49" value="120.89694">120°53'49"E</geoCoordinate>
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; ca. 1080 m a.s.l.; \ Nov. 1997, leg. Mey (451)L"; GS066, specimen and terminal parts of the abdomen, including genitalia, were glued separately on the entomological card.
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<emphasis id="097C3259AA18F990C2D5294A535D63E2" bold="true" italics="true" pageId="0" pageNumber="11">Paratypes</emphasis>
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: Philippines • 1♂, 2♀♀ (ADMU: GS009, GS067; ZMB: GS010): same data as holotype. • 1♂, 2♀♀ (ADMU: GS034, GS035, GS038): Luzon, Ilocos Sur, Suyo Municipality, Tagudin-Cervantes-Sabangan Road, Besang Pass Area; secondary forest, mountain creek;
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<geoCoordinate id="A47EA34ADEA2FE6A1A3805EA7E145B11" degrees="16" direction="north" minutes="57" orientation="latitude" precision="15" seconds="17" value="16.954723">16°57'17"N</geoCoordinate>
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;
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<geoCoordinate id="27B84BD2633DECF6A25C967B517B2073" degrees="120" direction="east" minutes="38" orientation="longitude" precision="15" seconds="57" value="120.64916">120°38'57"E</geoCoordinate>
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; ca. 1200 m a.s.l.; 15 Apr. 2019, leg. Freitag, Garces, Pangantihon
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<normalizedToken id="ED89D451542F844274665FE778339B6E" originalValue="“(448)L”">"(448)L"</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="22FBB7696E12F82BDB26CCC388FA6A53" pageId="0" pageNumber="11" type="description">
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<paragraph id="689BBD089BE3BD1915882F0F9D7E32CE" pageId="0" pageNumber="11">Description.</paragraph>
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<paragraph id="B6D87AA5F079B83A12215C54F317A238" pageId="0" pageNumber="11">
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(Fig.
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<figureCitation id="5D5D90D76801D39C72778EF4FC5959EF" captionStart="Figure 1–4" captionStartId="F1" captionText="Figure 1 - 4. Habitus of new Anacaena species 1, 2 Anacaena angatbuhay sp. nov. 3, 4 Anacaena auxilium sp. nov. in (1, 3) dorsal and (2, 4) lateral views. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1112.85752.figure1-4" httpUri="https://binary.pensoft.net/fig/713647" pageId="0" pageNumber="11">3</figureCitation>
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). TL 2.8 mm (2.3-2.8 mm); TW 1.7 mm (1.4-1.7 mm); EL 1.8 mm (1.5-1.9 mm). Body form oval, moderately convex (Fig.
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<figureCitation id="C18CEA0F119E9FB3CBE9AAB717ECBAB3" captionStart="Figure 1–4" captionStartId="F1" captionText="Figure 1 - 4. Habitus of new Anacaena species 1, 2 Anacaena angatbuhay sp. nov. 3, 4 Anacaena auxilium sp. nov. in (1, 3) dorsal and (2, 4) lateral views. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1112.85752.figure1-4" httpUri="https://binary.pensoft.net/fig/713647" pageId="0" pageNumber="11">4</figureCitation>
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); elytra about 3.4 times as long as pronotum (dorsal view), greatest width at anterior one-third.
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</paragraph>
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<paragraph id="8AB1799662E9DAD283DAC5EFB178E592" pageId="0" pageNumber="11">
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<emphasis id="0A01E06B1A5C434DACF12E97B673680C" bold="true" italics="true" pageId="0" pageNumber="11">Head</emphasis>
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: Clypeus black, moderately large, straight anteriorly, with distinct antero-lateral angle. Frons black; frontoclypeal suture visible laterally. Labrum black. Puncturation regular, coarse, shallowly impressed; interstices as wide as diameter of one puncture mediad, approximately twice as wide as diameter of punctures laterad; series of densely arranged punctures along inner margin of eyes absent. Ventral punctures obsolete. Maxillary palpomeres yellow to brown; palpomeres 1-3 dark yellow; palpomere 2 moderately inflated. Palpomere 4 almost entirely infuscated, lighter towards palpomere 3, widest towards mid-length; inner margin slightly convex; outer margin distinctly convex. Mentum with fine setae on lateral margins; anterior margin with median incision. Labial palpi stout, not longer than lateral edge of mentum; palpomere 3 slightly longer than palpomere 2. Eyes not constricted anteriorly, dorsal and ventral portions of almost equal size. Antennae capitate, 9-segmented; scape parallel-sided; pedicel narrowest towards mid-length; antennomere 1 (segment 3) not distinctly elongated, broadest at base, narrows distally; antennomeres 2-4 smallest in length and width, lighter than adjacent segments, smaller than antennomere 1 (segment 3), increasing size distally; antennomere 5-7 darkest of all antennomeres; antennomeres 5 and 6 of equal length; setae along lateral margins black, long, thick, erect. Segment 7 irregularly shaped, asymmetrical; segment 8 globular. Terminal segment as long as segments 7 and 8, widest at base, subparallel on lateral margins, asymmetrical apicad with apex present on distal side; setae distributed throughout, black, long, thick, erect, but setae on apical one-third longer. Preocular patches absent.
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</paragraph>
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<paragraph id="6786DC2AE50DF3B9AEF9D2F4C252CFD7" pageId="0" pageNumber="11">
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<emphasis id="996A9C65A55AB2CE16F6F969FDD08C76" bold="true" italics="true" pageId="0" pageNumber="11">Thorax</emphasis>
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: PW/PL = 2.62; PL/PW = 0.38. Pronotum largely black, with narrow light brown or yellow lateral margins not wider than diameter of eye. Puncturation of two series: larger, coarser series evenly spaced, with interstices as wide as diameter of one puncture; smaller, fine series near postero-medial margin, irregularly scattered, one-fourth as wide as larger series. Anterior margin curving inwards on both sides from antero-lateral angle, with deepest point on 0.2 laterad and curving outwards mediad. Lateral margins slightly curved, without setae. Postero-lateral angle ca. 70-80°. Posterior margin almost straight with curvature laterad. Prosternum flat. EL/EW = 1.13; EL/PL = 4.13; EW/PW = 1.11; TL/EW = 1.69. Elytra black, with very narrow brown lateral margins and posterior area; setae absent. Shoulder regions not accentuated. Punctures coarse, strongly impressed especially near the disc, arrangement dense, spacing as wide as punctures near the disc, wider than punctures laterad; depressions or rows of coarser punctures on lateral portion absent. Mesoventrite distinctly elevated medially, with slightly pointed protuberance.
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</paragraph>
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<paragraph id="D5DA97BAB3491E1343D7282CB600FB1A" pageId="0" pageNumber="11">
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<emphasis id="7DA44CF811405CBC1C1A7A6A8E8BD2F2" bold="true" italics="true" pageId="0" pageNumber="11">Legs</emphasis>
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: Dark brown, but tarsi lighter. Spine-like setae on procoxa present. Metafemur pubescent on proximal one-fourth and near outer margin, hairline direction horizontal and recumbent. Tibia with long, thick, black, spine-like setae along lateral margins, pointing distally; terminal pair of setae at least 1.5 times as long as adjacent setae; tibiae length ratio (protibia: mesotibia: metatibia) 1: 1.3: 1.5. Metatarsus size as long as metatibia or slightly shorter. Tarsi length ratio (protarsus: mesotarsus: metatarsus) 1.0: 1.4: 2.2.
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</paragraph>
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<paragraph id="A9F3DD9F1F496C6D4822C1C19E71BD90" pageId="0" pageNumber="11">
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<emphasis id="5EB5692715E82EC5827A1324F64541DA" bold="true" italics="true" pageId="0" pageNumber="11">Aedeagus</emphasis>
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: (Fig.
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<figureCitation id="54D357681666C1BBF964BF55EA952FCE" captionStart="Figure 5–6" captionStartId="F2" captionText="Figure 5 - 6. Aedeagi of 5 Anacaena angatbuhay sp. nov. holotype (GS 076) 6 Anacaena auxilium sp. nov. holotype (GS 066) in dorsal view. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1112.85752.figure5-6" httpUri="https://binary.pensoft.net/fig/713648" pageId="0" pageNumber="11">6</figureCitation>
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) Parameres apical region blunt, broadly pointing medially, slightly asymmetrical; inner margins almost straight on anterior half, widening medially on posterior half, mesal gap tapering to a narrow opening about 0.25 times as long as parameres; outer margins convex, bulging at anterior 0.5; combined width of parameres approximately the same or slightly broader as diameter of phallobase; basal portion 4 times as wide as apical part. Median lobe with apex blunt, apical portion subparallel, broadening starting in anterior 0.25, broadest before basal apophyses; base of median lobe not distinctly connected with parameres, about 2 times as wide as the width of the apex of median lobe; basal apophyses barely extending beyond the anterior portion of phallobase, narrow and subparallel. Phallobase longer than parameres, longer than wide; median reinforcement or pigmented line absent.
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</paragraph>
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</subSubSection>
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<subSubSection id="40AA3C3563038D424B39FF1227C9A518" pageId="0" pageNumber="11" type="differential diagnosis">
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<paragraph id="6FDD13F952EAF5E5FB72B9FD94DA1EE5" pageId="0" pageNumber="11">Differential diagnosis.</paragraph>
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<paragraph id="DFA27CACE62FD322F7DEA8940B363FE7" pageId="0" pageNumber="11">
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<taxonomicName id="94E0942A8596523B0CF4867556933094" lsidName="A. auxilium" pageId="0" pageNumber="11" rank="species" species="auxilium">
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<emphasis id="ECD5A22408981411EF3CB98B7A6AC820" italics="true" pageId="0" pageNumber="11">A. auxilium</emphasis>
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</taxonomicName>
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sp. nov. (Fig.
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<figureCitation id="C6E3C9A6A3CB08067DEE54944A143223" captionStart="Figure 1–4" captionStartId="F1" captionText="Figure 1 - 4. Habitus of new Anacaena species 1, 2 Anacaena angatbuhay sp. nov. 3, 4 Anacaena auxilium sp. nov. in (1, 3) dorsal and (2, 4) lateral views. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1112.85752.figure1-4" httpUri="https://binary.pensoft.net/fig/713647" pageId="0" pageNumber="11">3</figureCitation>
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) is remarkably larger than most Philippine congeners. In length,
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<taxonomicName id="B3BFEA49BFDE7031E6C811ECF7A6F741" lsidName="A. auxilium" pageId="0" pageNumber="11" rank="species" species="auxilium">
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<emphasis id="73CF22372318BA6FC951ABBB0A603DC0" italics="true" pageId="0" pageNumber="11">A. auxilium</emphasis>
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</taxonomicName>
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sp. nov. (2.3-2.8 mm) overlaps with
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<taxonomicName id="5FD3AA19BC6735A80A73FD7C5735A590" lsidName="A. cordillera" pageId="0" pageNumber="11" rank="species" species="cordillera">
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||
<emphasis id="13713398C03931509FD43720252B2DDD" italics="true" pageId="0" pageNumber="11">A. cordillera</emphasis>
|
||
</taxonomicName>
|
||
Komarek & Freitag, 2014 (2.6-3.1 mm) and
|
||
<taxonomicName id="A2AA566437E087BB5F10B796296306FB" lsidName="A. levistriata" pageId="0" pageNumber="11" rank="species" species="levistriata">
|
||
<emphasis id="2A9FF8870434A6B4CBC7A56A326B345B" italics="true" pageId="0" pageNumber="11">A. levistriata</emphasis>
|
||
</taxonomicName>
|
||
Komarek & Freitag, 2014 (2.1-2.7 mm). However, the new species has a smaller body width than
|
||
<taxonomicName id="482EF3ABEEA38BC8EE226C3BC24BE069" lsidName="A. cordillera" pageId="0" pageNumber="11" rank="species" species="cordillera">
|
||
<emphasis id="C2371BB4D3BF73E1257D7D00FF254284" italics="true" pageId="0" pageNumber="11">A. cordillera</emphasis>
|
||
</taxonomicName>
|
||
(1.3 mm vs.> 1.5-1.8 mm) and a longer elytral length (1.5 mm vs. <1.20-1.27 mm) than both congeners. Additionally, the new species has stout labial palpi which are not longer than the lateral edge of the mentum, while both
|
||
<emphasis id="3F399EC4CD028D916C943800A2FC1025" italics="true" pageId="0" pageNumber="11">
|
||
<taxonomicName id="72DC924873A136300975C7F33B918EFA" lsidName="A. cordillera" pageId="0" pageNumber="11" rank="species" species="cordillera">A. cordillera</taxonomicName>
|
||
and
|
||
<taxonomicName id="25EE7BC6A4868304DB3843627A46C137" lsidName="A. levistriata" pageId="0" pageNumber="11" rank="species" species="levistriata">A. levistriata</taxonomicName>
|
||
</emphasis>
|
||
have labial palpi longer than the lateral edge of the mentum. Additionally, the labial palpi of
|
||
<taxonomicName id="16068BAFE325CA048262930C911AA88B" lsidName="A. auxilium" pageId="0" pageNumber="11" rank="species" species="auxilium">
|
||
<emphasis id="9F6C1F3A1CF2803DC992388E709C58B4" italics="true" pageId="0" pageNumber="11">A. auxilium</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. are stout, while the labial palpi of
|
||
<taxonomicName id="7FBD31D38502D7EFEF1D4AE9FF68A774" lsidName="A. cordillera" pageId="0" pageNumber="11" rank="species" species="cordillera">
|
||
<emphasis id="33FCCCC8F20D03482C95678F50267E3D" italics="true" pageId="0" pageNumber="11">A. cordillera</emphasis>
|
||
</taxonomicName>
|
||
are slender. Moreover, pre-ocular patches are absent in
|
||
<taxonomicName id="91FB4C61F97387C798C6FD0DB605AA6F" lsidName="A. auxilium" pageId="0" pageNumber="11" rank="species" species="auxilium">
|
||
<emphasis id="07920F2500B93712E666A3136D613999" italics="true" pageId="0" pageNumber="11">A. auxilium</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., while these are distinct in
|
||
<taxonomicName id="E176E9DC03FB4D359850FCC63CC1C2CF" lsidName="A. cordillera" pageId="0" pageNumber="11" rank="species" species="cordillera">
|
||
<emphasis id="84D77666ECB7840D349585992710D1D6" italics="true" pageId="0" pageNumber="11">A. cordillera</emphasis>
|
||
</taxonomicName>
|
||
. Remarkably, the new species has a bi-punctuate elytra with large, coarse series and small, fine series, while
|
||
<taxonomicName id="48D54394FE35B498288546B8E1755D65" lsidName="A. cordillera" pageId="0" pageNumber="11" rank="species" species="cordillera">
|
||
<emphasis id="7006655537536922F4A11FF744A10E21" italics="true" pageId="0" pageNumber="11">A. cordillera</emphasis>
|
||
</taxonomicName>
|
||
only exhibits the latter and
|
||
<taxonomicName id="81D27D21FACCB00B6F477DFBD81C162C" lsidName="A. levistriata" pageId="0" pageNumber="11" rank="species" species="levistriata">
|
||
<emphasis id="45CA83B0FE0C997FCEFC1539AE06262C" italics="true" pageId="0" pageNumber="11">A. levistriata</emphasis>
|
||
</taxonomicName>
|
||
only exhibits the former.
|
||
</paragraph>
|
||
<paragraph id="B324D55FFA50C75F0B06AB7374842F9B" pageId="0" pageNumber="11">
|
||
While the parameres of the new species and
|
||
<taxonomicName id="2CA5812324C1B64E004ED8E48820577B" lsidName="A. levistriata" pageId="0" pageNumber="11" rank="species" species="levistriata">
|
||
<emphasis id="F75BA8DD496A31720FD37E30CBDCA8F4" italics="true" pageId="0" pageNumber="11">A. levistriata</emphasis>
|
||
</taxonomicName>
|
||
look similar,
|
||
<taxonomicName id="648A9773AAA9ECCB48055F3AE6628996" lsidName="A. auxilium" pageId="0" pageNumber="11" rank="species" species="auxilium">
|
||
<emphasis id="50B7915FDBA891820602029D611F7F06" italics="true" pageId="0" pageNumber="11">A. auxilium</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be differentiated in that its median lobe is distinctly longer (ca. 0.4
|
||
<normalizedToken id="E14508AE5DA8B2DA0F3EDB829BB8E362" originalValue="×">x</normalizedToken>
|
||
) than its parameres. Meanwhile, the apical portions of the median lobe and parameres of
|
||
<taxonomicName id="CBC730E32BD43995C7098FED53FA63A0" lsidName="A. cordillera" pageId="0" pageNumber="11" rank="species" species="cordillera">
|
||
<emphasis id="35A2C7F10B0443CE4A8FE24828A259D7" italics="true" pageId="0" pageNumber="11">A. cordillera</emphasis>
|
||
</taxonomicName>
|
||
are clearly different from those of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CB7578103FD157CDC053611083879AAA" pageId="0" pageNumber="11" type="distribution">
|
||
<paragraph id="1C86080DAE6F613CF98CD0A3ECB6D55A" pageId="0" pageNumber="11">Distribution.</paragraph>
|
||
<paragraph id="C311517FEDD224761A7DF65A66E3230A" pageId="0" pageNumber="11">
|
||
The species is documented from two nearby localities, namely Ilocos Sur and Mountain Province (Fig.
|
||
<figureCitation id="E1573E1F13565197EF50263BB310BF21" captionStart="Figure 7" captionStartId="F3" captionText="Figure 7. Map showing the updated distribution of Anacaena in Northern Luzon, including Komarek & Freitag (2014) data." figureDoi="10.3897/zookeys.1112.85752.figure7" httpUri="https://binary.pensoft.net/fig/713649" pageId="0" pageNumber="11">7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6B457FE0F92A46E7EA8A15C7F868A73E" pageId="0" pageNumber="11" type="remarks">
|
||
<paragraph id="9A1EFE18BF921CCB681D15C5EFC66BB4" pageId="0" pageNumber="11">Remarks.</paragraph>
|
||
<paragraph id="AB262657CE2466694772657AD8E31B16" pageId="0" pageNumber="11">Specimens from a preserved collection during 1997 fieldwork showed no pertinent phenotypic plasticity to specimens collected in a field expedition 22 years later. No external sexual dimorphism is observed.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="18ECF4D332745C8A243C78F53BCE8CFE" pageId="0" pageNumber="11" type="etymology">
|
||
<paragraph id="11064DAAB5E28F7DEC17E54C7969EB77" pageId="0" pageNumber="11">Etymology.</paragraph>
|
||
<paragraph id="82756D77C8724C118D2766D9F6101DB2" pageId="0" pageNumber="11">
|
||
The species epithet
|
||
<normalizedToken id="D9072F6619E2B0B94D90807FCBD4FB49" originalValue="‘auxilium’">'auxilium'</normalizedToken>
|
||
is Latin for
|
||
<normalizedToken id="C0E18D6089DE41A22E169E3319C70DEA" originalValue="‘help’">'help'</normalizedToken>
|
||
. The new species is dedicated to the Mary Help of Christians Seminary system to which the first author expresses gratitude for constant inspiration. Additionally,
|
||
<normalizedToken id="1E74034F96DB8BA1D5E74A7FA95629C4" originalValue="‘help’">'help'</normalizedToken>
|
||
alludes to the vulnerable state of freshwater biodiversity in the tropics.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |