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<document id="ACF6CEC2DCA449BE891A91B7CC99D38D" ID-DOI="10.5281/zenodo.182262" ID-GBIF-Dataset="63642e86-410e-4b94-a2cb-4787f3c52296" ID-ISSN="1175-5326" ID-Zenodo-Dep="182262" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459857368870" checkinUser="plazi" docAuthor="Fontoura, Paulo, Pilato, Giovanni &amp; Lisi, Oscar" docDate="2008" docId="038D5952FD38E155188C0C24FCFE08E6" docLanguage="en" docName="zt01778p047.pdf" docOrigin="Zootaxa 1778" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Macrobiotus almadai Fontoura, Pilato &amp; Lisi, 2008, sp. nov." docType="treatment" docVersion="10" lastPageNumber="40" masterDocId="FFB4212AFD39E156181B0F2EFF910E34" masterDocTitle="New records of eutardigrades (Tardigrada) from Faial and Pico Islands, the Azores, with the description of two new species" masterLastPageNumber="47" masterPageNumber="37" pageNumber="38" updateTime="1698225734066" updateUser="plazi">
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<mods:title id="CC3BAEF9B72DE48A95F6D6469B0D1EE6">New records of eutardigrades (Tardigrada) from Faial and Pico Islands, the Azores, with the description of two new species</mods:title>
</mods:titleInfo>
<mods:name id="1B37475CCD56CE6A8199D2BE7210EE84" type="personal">
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<mods:namePart id="24C711AFD535BD2F075382608977520A">Fontoura, Paulo</mods:namePart>
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<mods:namePart id="B95EFBAAFB57F4F6C18B06A12DEE262B">Pilato, Giovanni</mods:namePart>
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<mods:namePart id="4181CA146170EF246C14C54910C8B78A">Lisi, Oscar</mods:namePart>
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<mods:title id="D24CDF620918AE1E65D6309B04562FF3">Zootaxa</mods:title>
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<mods:part id="EB5D065F73CFBFEDEB32791EFE76B35F">
<mods:date id="3C8B1C99A49CC285B878ED207259256B">2008</mods:date>
<mods:detail id="51510A190DC3F8987727525948C408CD" type="volume">
<mods:number id="FB896367B1544ECD59EA3A0A69355646">1778</mods:number>
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<mods:identifier id="2CB7C2707542940523AA04D98F0D7688" type="DOI">10.5281/zenodo.182262</mods:identifier>
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<treatment id="038D5952FD38E155188C0C24FCFE08E6" ID-DOI="http://doi.org/10.5281/zenodo.5626475" ID-GBIF-Taxon="119363113" ID-Zenodo-Dep="5626475" LSID="urn:lsid:plazi:treatment:038D5952FD38E155188C0C24FCFE08E6" httpUri="http://treatment.plazi.org/id/038D5952FD38E155188C0C24FCFE08E6" lastPageId="3" lastPageNumber="40" pageId="1" pageNumber="38">
<subSubSection id="C33EBBCFFD38E157188C0C24FF4A0D73" pageId="1" pageNumber="38" type="nomenclature">
<paragraph id="8B9BE844FD38E157188C0C24FE6C0D10" blockId="1.[151,509,778,839]" box="[151,509,778,804]" pageId="1" pageNumber="38">
<heading id="D0D35F28FD38E157188C0C24FE6C0D10" bold="true" box="[151,509,778,804]" fontSize="11" level="1" pageId="1" pageNumber="38" reason="1">
<emphasis id="B9503456FD38E157188C0C24FE6C0D10" bold="true" box="[151,509,778,804]" pageId="1" pageNumber="38">
<taxonomicName id="4C2493C7FD38E157188C0C24FE080D10" ID-CoL="92c56c82-35cc-4638-bad2-f339b60932f7" box="[151,409,778,804]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="1" pageNumber="38" phylum="Tardigrada" rank="species" species="almadai" status="sp. nov.">
<emphasis id="B9503456FD38E157188C0C24FE080D10" bold="true" box="[151,409,778,804]" italics="true" pageId="1" pageNumber="38">Macrobiotus almadai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A263892DFD38E15719BB0C24FE6C0D10" box="[416,509,778,804]" pageId="1" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9BE844FD38E157188C0C03FF4A0D73" blockId="1.[151,509,778,839]" box="[151,219,813,839]" pageId="1" pageNumber="38">
<figureCitation id="131FF4C1FD38E157188C0C03FF4A0D73" box="[151,219,813,839]" captionStart="FIGURE 1. A G" captionStartId="2.[151,255,1339,1363]" captionTargetBox="[268,1285,318,1338]" captionTargetId="figure@2.[255,1333,261,1373]" captionTargetPageId="2" captionText="FIGURE 1. A G, Macrobiotus almadai sp. nov. (holotype): A, bucco-pharyngeal apparatus; B and C, claws of the third pair of legs; D, claws of the hind legs; E, egg; F and G, details of the egg shell (arrow indicates the small teeth of the terminal disc). (Scale bar = 10 µm)." httpUri="https://zenodo.org/record/182263/files/figure.png" pageId="1" pageNumber="38">Fig. 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33EBBCFFD38E157188C0C56FC8A0BF6" pageId="1" pageNumber="38" type="materials_examined">
<paragraph id="8B9BE844FD38E157188C0C56FEA40A06" blockId="1.[151,1437,888,2034]" pageId="1" pageNumber="38">
<emphasis id="B9503456FD38E157188C0C56FE1F0DA6" bold="true" box="[151,398,888,914]" pageId="1" pageNumber="38">Material examined:</emphasis>
<specimenCount id="9D2223CDFD38E157198D0C56FDA70DA6" box="[406,566,888,914]" pageId="1" pageNumber="38" type="generic">53 specimens</specimenCount>
and
<specimenCount id="9D2223CDFD38E1571A6A0C56FD410DA6" box="[625,720,888,914]" pageId="1" pageNumber="38" type="egg">15 eggs</specimenCount>
:
<specimenCount id="9D2223CDFD38E1571AC70C56FCFC0DA6" box="[732,877,888,914]" pageId="1" pageNumber="38" type="generic">3 specimens</specimenCount>
mounted in polyvinyl lactophenol collected on Túnel da Caldeira (Faial Island);
<specimenCount id="9D2223CDFD38E1571A3C0C8EFD3F0D8E" box="[551,686,928,954]" pageId="1" pageNumber="38" type="generic">1 specimen</specimenCount>
and
<specimenCount id="9D2223CDFD38E1571AF00C8EFCBC0D8E" box="[747,813,928,954]" pageId="1" pageNumber="38" type="egg">1 egg</specimenCount>
mounted in Hoyers medium from Parque Florestal (Faial Island);
<specimenCount id="9D2223CDFD38E15719530CE6FE7B0DD6" box="[328,490,968,994]" pageId="1" pageNumber="38" type="generic">33 specimens</specimenCount>
(30 mounted in Hoyers medium and
<quantity id="4CDC45A1FD38E1571BA70CE6FC7A0DD6" box="[956,1003,968,994]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="1" pageNumber="38" unit="in" value="3.0">3 in</quantity>
polyvinyl lactophenol) and
<specimenCount id="9D2223CDFD38E1571D240CE6FA0C0DD6" box="[1343,1437,968,994]" pageId="1" pageNumber="38" type="egg">12 eggs</specimenCount>
from Lages do Pico (Pico Island);
<specimenCount id="9D2223CDFD38E1571A370CDEFD5A0A3E" box="[556,715,1008,1034]" pageId="1" pageNumber="38" type="generic">16 specimens</specimenCount>
and
<specimenCount id="9D2223CDFD38E1571B1E0CDEFCC00A3E" box="[773,849,1008,1034]" pageId="1" pageNumber="38" type="egg">2 eggs</specimenCount>
mounted in Hoyers medium from Terras do Pico (Pico Island).
</paragraph>
<paragraph id="8B9BE844FD38E15718DE0B6EFE1A0AE6" blockId="1.[151,1437,888,2034]" pageId="1" pageNumber="38">
<emphasis id="B9503456FD38E15718DE0B6EFE060A6E" bold="true" box="[197,407,1088,1114]" pageId="1" pageNumber="38">
<typeStatus id="549F56E6FD38E15718DE0B6EFE930A6E" box="[197,258,1088,1114]" pageId="1" pageNumber="38">Type</typeStatus>
repository:
</emphasis>
<typeStatus id="549F56E6FD38E15719B80B6EFD810A6E" box="[419,528,1088,1114]" pageId="1" pageNumber="38" type="holotype">Holotype</typeStatus>
(slide No. 5220) and two eggs are deposited in the collection of Binda &amp; Pilato (Museo del Dipartimento di Biologia Animale “Marcello La Greca”, Università di Catania).
<typeStatus id="549F56E6FD38E1571D310B46FA0D0AB6" box="[1322,1436,1128,1154]" pageId="1" pageNumber="38" type="paratype">Paratypes</typeStatus>
are deposited in the collection of P. Fontoura (Department of Zoology and Anthropology, Faculty of Sciences, University of Porto).
</paragraph>
<paragraph id="8B9BE844FD38E15718DE0BCEFC8A0BF6" blockId="1.[151,1437,888,2034]" pageId="1" pageNumber="38">
<emphasis id="B9503456FD38E15718DE0BCEFE3F0ACE" bold="true" box="[197,430,1248,1274]" pageId="1" pageNumber="38">Specific diagnosis:</emphasis>
Species belonging to
<taxonomicName id="4C2493C7FD38E1571AA60BCEFCBD0ACD" box="[701,812,1248,1273]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="1" pageNumber="38" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="B9503456FD38E1571AA60BCEFCBD0ACD" box="[701,812,1248,1273]" italics="true" pageId="1" pageNumber="38">hufelandi</emphasis>
</taxonomicName>
group. Cuticle smooth with some small pores (fine granulation present only on hind legs), eyes present; buccal tube narrow; buccal armature very simple with peribuccal lamellae and three dorsal and three ventral thin transversal ridges; bands of teeth not visible; pharyngeal bulb with apophyses, two macroplacoids and microplacoid; eggs with reticulated shell and processes shaped in the form of inverted goblets; reticular design delicate and very regular; terminal disc of processes with very small teeth visible only in some orientations.
</paragraph>
</subSubSection>
<subSubSection id="C33EBBCFFD38E15418DE0AFEFB070F36" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38" type="description">
<paragraph id="8B9BE844FD38E15718DE0AFEFC3B0936" blockId="1.[151,1437,888,2034]" pageId="1" pageNumber="38">
<emphasis id="B9503456FD38E15718DE0AFEFDB00BDE" bold="true" box="[197,545,1488,1514]" pageId="1" pageNumber="38">
Description of the
<typeStatus id="549F56E6FD38E15719AD0AFEFDB00BDE" box="[438,545,1488,1514]" pageId="1" pageNumber="38" type="holotype">holotype</typeStatus>
</emphasis>
: Body length 340 μm, colorless, eyes present; cuticle smooth with some small circular and elliptical pores (fine granulation present only on hind legs). Buccal armature simple: only 10 peribuccal lamellae and a system of three thin dorsal and three ventral transversal ridges present. Bands of teeth not visible. Rigid buccal tube narrow, 32.4 µm long and 3.6 µm (
<emphasis id="B9503456FD38E1571BFD0966FBC20855" box="[998,1107,1608,1633]" italics="true" pageId="1" pageNumber="38">pt = 11.1</emphasis>
) wide externally (
<figureCitation id="131FF4C1FD38E1571D340966FA1C0856" box="[1327,1421,1608,1634]" captionStart="FIGURE 1. A G" captionStartId="2.[151,255,1339,1363]" captionTargetBox="[268,1285,318,1338]" captionTargetId="figure@2.[255,1333,261,1373]" captionTargetPageId="2" captionText="FIGURE 1. A G, Macrobiotus almadai sp. nov. (holotype): A, bucco-pharyngeal apparatus; B and C, claws of the third pair of legs; D, claws of the hind legs; E, egg; F and G, details of the egg shell (arrow indicates the small teeth of the terminal disc). (Scale bar = 10 µm)." httpUri="https://zenodo.org/record/182263/files/figure.png" pageId="1" pageNumber="38">Fig. 1A</figureCitation>
). Stylet supports inserted on the buccal tube wall at 76.5 % of its length (
<emphasis id="B9503456FD38E1571BC4095EFBDD08BD" box="[991,1100,1648,1673]" italics="true" pageId="1" pageNumber="38">pt = 76.5</emphasis>
). Pharyngeal bulb with apophyses, two rod-shaped macroplacoids and microplacoid. First macroplacoid, with central constriction, 8.2 µm long (
<emphasis id="B9503456FD38E157191009EEFEE708ED" box="[267,374,1728,1753]" italics="true" pageId="1" pageNumber="38">pt = 25.3</emphasis>
), second 5.5 µm (
<emphasis id="B9503456FD38E1571A5709EEFDF308ED" box="[588,610,1728,1753]" italics="true" pageId="1" pageNumber="38">pt</emphasis>
=
<emphasis id="B9503456FD38E1571A9909EEFD2408ED" box="[642,693,1728,1753]" italics="true" pageId="1" pageNumber="38">17.0</emphasis>
), microplacoid 2.7 µm long (
<emphasis id="B9503456FD38E1571C1509EEFBFA08ED" box="[1038,1131,1728,1753]" italics="true" pageId="1" pageNumber="38">pt = 8.3</emphasis>
). Placoid row length 16.6 µm (
<emphasis id="B9503456FD38E15718D409C6FF770935" box="[207,230,1768,1793]" italics="true" pageId="1" pageNumber="38">pt</emphasis>
=
<emphasis id="B9503456FD38E157191E09C6FEA90935" box="[261,312,1768,1793]" italics="true" pageId="1" pageNumber="38">51.2</emphasis>
). Macroplacoid row length 14.0 µm long (
<emphasis id="B9503456FD38E1571B3409C6FCD70935" box="[815,838,1768,1793]" italics="true" pageId="1" pageNumber="38">pt</emphasis>
=
<emphasis id="B9503456FD38E1571B7E09C6FC090935" box="[869,920,1768,1793]" italics="true" pageId="1" pageNumber="38">43.2</emphasis>
).
</paragraph>
<paragraph id="8B9BE844FD38E15718DE083EFCF1094E" blockId="1.[151,1437,888,2034]" pageId="1" pageNumber="38">
Claws of
<taxonomicName id="4C2493C7FD38E1571921083EFE38091D" box="[314,425,1808,1833]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="1" pageNumber="38" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="B9503456FD38E1571921083EFE38091D" box="[314,425,1808,1833]" italics="true" pageId="1" pageNumber="38">hufelandi</emphasis>
</taxonomicName>
<typeStatus id="549F56E6FD38E15719A8083EFE74091E" box="[435,485,1808,1834]" pageId="1" pageNumber="38">type</typeStatus>
with accessory points on the main branches and smooth lunules (
<figureCitation id="131FF4C1FD38E1571CE5083EFA1F091E" box="[1278,1422,1808,1834]" captionStart="FIGURE 1. A G" captionStartId="2.[151,255,1339,1363]" captionTargetBox="[268,1285,318,1338]" captionTargetId="figure@2.[255,1333,261,1373]" captionTargetPageId="2" captionText="FIGURE 1. A G, Macrobiotus almadai sp. nov. (holotype): A, bucco-pharyngeal apparatus; B and C, claws of the third pair of legs; D, claws of the hind legs; E, egg; F and G, details of the egg shell (arrow indicates the small teeth of the terminal disc). (Scale bar = 10 µm)." httpUri="https://zenodo.org/record/182263/files/figure.png" pageId="1" pageNumber="38">Figs. 1B, D</figureCitation>
). External and internal claws of the third pair of legs 9.6 µm (
<emphasis id="B9503456FD38E1571B770816FC4D0965" box="[876,988,1848,1873]" italics="true" pageId="1" pageNumber="38">pt = 29.6</emphasis>
) long and 9.5 µm (
<emphasis id="B9503456FD38E1571CD30816FB4E0965" box="[1224,1247,1848,1873]" italics="true" pageId="1" pageNumber="38">pt</emphasis>
=
<emphasis id="B9503456FD38E1571D1A0816FAA50965" box="[1281,1332,1848,1873]" italics="true" pageId="1" pageNumber="38">29.3</emphasis>
) respectively. Posterior claws of hind legs 10.9 µm (
<emphasis id="B9503456FD38E1571ABD084EFC82094D" box="[678,787,1888,1913]" italics="true" pageId="1" pageNumber="38">pt = 33.6</emphasis>
) long.
</paragraph>
<paragraph id="8B9BE844FD38E15418DE08A6FB070F36" blockId="1.[151,1437,888,2034]" lastBlockId="2.[151,1436,152,258]" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38">
Eggs laid free, spherical, with reticulated shell (
<figureCitation id="131FF4C1FD38E1571B1B08A6FC160996" box="[768,903,1928,1954]" captionStart="FIGURE 1. A G" captionStartId="2.[151,255,1339,1363]" captionTargetBox="[268,1285,318,1338]" captionTargetId="figure@2.[255,1333,261,1373]" captionTargetPageId="2" captionText="FIGURE 1. A G, Macrobiotus almadai sp. nov. (holotype): A, bucco-pharyngeal apparatus; B and C, claws of the third pair of legs; D, claws of the hind legs; E, egg; F and G, details of the egg shell (arrow indicates the small teeth of the terminal disc). (Scale bar = 10 µm)." httpUri="https://zenodo.org/record/182263/files/figure.png" pageId="1" pageNumber="38">Figs. 1E, F</figureCitation>
) and with processes in the shape of inverted goblets (
<figureCitation id="131FF4C1FD38E15718E5089EFE1C09FE" box="[254,397,1968,1994]" captionStart="FIGURE 1. A G" captionStartId="2.[151,255,1339,1363]" captionTargetBox="[268,1285,318,1338]" captionTargetId="figure@2.[255,1333,261,1373]" captionTargetPageId="2" captionText="FIGURE 1. A G, Macrobiotus almadai sp. nov. (holotype): A, bucco-pharyngeal apparatus; B and C, claws of the third pair of legs; D, claws of the hind legs; E, egg; F and G, details of the egg shell (arrow indicates the small teeth of the terminal disc). (Scale bar = 10 µm)." httpUri="https://zenodo.org/record/182263/files/figure.png" pageId="1" pageNumber="38">Figs. 1E, G</figureCitation>
). Diameter without processes 6978 µm, including them 7486 µm. On circumference 2327 processes were present (
<quantity id="4CDC45A1FD38E1571A1708F6FD1409C6" box="[524,645,2008,2034]" metricMagnitude="0" metricUnit="m" metricValue="2.4892" metricValueMax="2.7939999999999996" metricValueMin="2.1844" pageId="1" pageNumber="38" unit="in" value="98.0" valueMax="110.0" valueMin="86.0">86110 in</quantity>
hemisphere). Processes up to 5.9 µm high with basal diameter of 5.15.5 µm. Terminal disc, 3.85.5 µm in diameter, may appear smooth (
<figureCitation id="131FF4C1FD3BE1541BF00FB6FBFF0E86" box="[1003,1134,152,178]" captionStart="FIGURE 1. A G" captionStartId="2.[151,255,1339,1363]" captionTargetBox="[268,1285,318,1338]" captionTargetId="figure@2.[255,1333,261,1373]" captionTargetPageId="2" captionText="FIGURE 1. A G, Macrobiotus almadai sp. nov. (holotype): A, bucco-pharyngeal apparatus; B and C, claws of the third pair of legs; D, claws of the hind legs; E, egg; F and G, details of the egg shell (arrow indicates the small teeth of the terminal disc). (Scale bar = 10 µm)." httpUri="https://zenodo.org/record/182263/files/figure.png" pageId="2" pageNumber="39">Figs. 1E, F</figureCitation>
), but in some orientations it shows very small teeth (
<figureCitation id="131FF4C1FD3BE15419CF0FEEFDA30EEE" box="[468,562,192,218]" captionStart="FIGURE 1. A G" captionStartId="2.[151,255,1339,1363]" captionTargetBox="[268,1285,318,1338]" captionTargetId="figure@2.[255,1333,261,1373]" captionTargetPageId="2" captionText="FIGURE 1. A G, Macrobiotus almadai sp. nov. (holotype): A, bucco-pharyngeal apparatus; B and C, claws of the third pair of legs; D, claws of the hind legs; E, egg; F and G, details of the egg shell (arrow indicates the small teeth of the terminal disc). (Scale bar = 10 µm)." httpUri="https://zenodo.org/record/182263/files/figure.png" pageId="2" pageNumber="39">Fig. 1G</figureCitation>
). Egg shell with very regular, elegant, and homogeneous reticular design (
<figureCitation id="131FF4C1FD3BE15418840FC6FEB40F36" box="[159,293,232,258]" captionStart="FIGURE 1. A G" captionStartId="2.[151,255,1339,1363]" captionTargetBox="[268,1285,318,1338]" captionTargetId="figure@2.[255,1333,261,1373]" captionTargetPageId="2" captionText="FIGURE 1. A G, Macrobiotus almadai sp. nov. (holotype): A, bucco-pharyngeal apparatus; B and C, claws of the third pair of legs; D, claws of the hind legs; E, egg; F and G, details of the egg shell (arrow indicates the small teeth of the terminal disc). (Scale bar = 10 µm)." httpUri="https://zenodo.org/record/182263/files/figure.png" pageId="2" pageNumber="39">Figs. 1E, F</figureCitation>
). Margins of meshes very thin and basal meshes slightly larger than others.
</paragraph>
</subSubSection>
<caption id="DF5BB8CCFD3BE154188C0A15FE690BAC" httpUri="https://zenodo.org/record/182263/files/figure.png" pageId="2" pageNumber="39" targetBox="[268,1285,318,1338]" targetPageId="2">
<paragraph id="8B9BE844FD3BE154188C0A15FE690BAC" blockId="2.[151,1436,1339,1432]" pageId="2" pageNumber="39">
<emphasis id="B9503456FD3BE154188C0A15FE880B67" bold="true" box="[151,281,1339,1363]" pageId="2" pageNumber="39">FIGURE 1.</emphasis>
AG,
<taxonomicName id="4C2493C7FD3BE15419450A15FDD20B66" box="[350,579,1339,1362]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="39" phylum="Tardigrada" rank="species" species="almadai" status="sp. nov.">
<emphasis id="B9503456FD3BE15419450A15FDD20B66" box="[350,579,1339,1362]" italics="true" pageId="2" pageNumber="39">Macrobiotus almadai</emphasis>
</taxonomicName>
<emphasis id="B9503456FD3BE1541A510A15FD0E0B67" bold="true" box="[586,671,1339,1363]" pageId="2" pageNumber="39">
<taxonomicNameLabel id="A263892DFD3BE1541A510A15FD0E0B67" box="[586,671,1339,1363]" pageId="2" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype): A, bucco-pharyngeal apparatus; B and C, claws of the third pair of legs; D, claws of the hind legs; E, egg; F and G, details of the egg shell (arrow indicates the small teeth of the terminal disc). (Scale bar = 10 µm).
</paragraph>
</caption>
<subSubSection id="C33EBBCFFD3BE15418DE0AE5FDC508B1" pageId="2" pageNumber="39" type="discussion">
<paragraph id="8B9BE844FD3BE15418DE0AE5FDCE0839" blockId="2.[151,1436,1483,2029]" pageId="2" pageNumber="39">
<emphasis id="B9503456FD3BE15418DE0AE5FED00BD1" bold="true" box="[197,321,1483,1509]" pageId="2" pageNumber="39">Remarks:</emphasis>
<typeStatus id="549F56E6FD3BE15419520AE5FE2A0BD1" box="[329,443,1483,1509]" pageId="2" pageNumber="39" type="paratype">Paratypes</typeStatus>
similar to
<typeStatus id="549F56E6FD3BE1541A220AE5FD310BD1" box="[569,672,1483,1509]" pageId="2" pageNumber="39" type="holotype">holotype</typeStatus>
in both qualitative and morphometric characters. Smallest specimen 161 µm long, largest one 394 µm.
</paragraph>
<paragraph id="8B9BE844FD3BE15418DE0935FDC508B1" blockId="2.[151,1436,1483,2029]" pageId="2" pageNumber="39">
Re-examination of some specimens and eggs collected on Pico and St. Maria Islands (
<bibRefCitation id="EFB595B5FD3BE1541CC10935FA020801" author="Fontoura" box="[1242,1427,1563,1589]" pageId="2" pageNumber="47" refString="Fontoura, P. (1985) Primeiro contributo para o conhecimento da fauna de tardigrados dos Acores. Arquipelago, 4, 45 - 56." type="journal article" year="1985">Fontoura, 1985</bibRefCitation>
) showed that they do not belong to
<taxonomicName id="4C2493C7FD3BE1541A22096AFD400869" box="[569,721,1604,1629]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="39" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="B9503456FD3BE1541A22096AFD400869" box="[569,721,1604,1629]" italics="true" pageId="2" pageNumber="39">M. hufelandi</emphasis>
</taxonomicName>
, as previously identified, and must be attributed to the new species
<taxonomicName id="4C2493C7FD3BE15418E80942FE7F08B1" box="[243,494,1644,1669]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="39" phylum="Tardigrada" rank="species" species="almadai" status="sp. nov.">
<emphasis id="B9503456FD3BE15418E80942FE7F08B1" box="[243,494,1644,1669]" italics="true" pageId="2" pageNumber="39">Macrobiotus almadai</emphasis>
</taxonomicName>
<emphasis id="B9503456FD3BE15419ED0945FDC508B1" bold="true" box="[502,596,1643,1669]" pageId="2" pageNumber="39">
<taxonomicNameLabel id="A263892DFD3BE15419ED0945FDC508B1" box="[502,596,1643,1669]" pageId="2" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33EBBCFFD3BE15518DE09BDFCFE08E6" lastPageId="3" lastPageNumber="40" pageId="2" pageNumber="39" type="etymology">
<paragraph id="8B9BE844FD3BE15418DE09BDFBA808E1" blockId="2.[151,1436,1483,2029]" pageId="2" pageNumber="39">
<emphasis id="B9503456FD3BE15418DE09BDFEC70899" bold="true" box="[197,342,1683,1709]" pageId="2" pageNumber="39">Etymology:</emphasis>
The new species is dedicated in honor to the coordinator of the Eco-Ethology Research Unit, Prof. Vitor Almada, in appreciation of his support to our studies on tardigrades.
</paragraph>
<paragraph id="8B9BE844FD3BE15418DE09CDFD0A09D9" blockId="2.[151,1436,1483,2029]" pageId="2" pageNumber="39">
<emphasis id="B9503456FD3BE15418DE09CDFE4D08C9" bold="true" box="[197,476,1763,1789]" pageId="2" pageNumber="39">Differential diagnosis:</emphasis>
<taxonomicName id="4C2493C7FD3BE15419FD09CAFD7408C9" box="[486,741,1764,1789]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="39" phylum="Tardigrada" rank="species" species="almadai" status="sp. nov.">
<emphasis id="B9503456FD3BE15419FD09CAFD7408C9" box="[486,741,1764,1789]" italics="true" pageId="2" pageNumber="39">Macrobiotus almadai</emphasis>
</taxonomicName>
<emphasis id="B9503456FD3BE1541AF409CDFCDF08C9" bold="true" box="[751,846,1763,1789]" pageId="2" pageNumber="39">
<taxonomicNameLabel id="A263892DFD3BE1541AF409CDFCDF08C9" box="[751,846,1763,1789]" pageId="2" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to the
<taxonomicName id="4C2493C7FD3BE1541C1009CAFBEB08C9" box="[1035,1146,1764,1789]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="39" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="B9503456FD3BE1541C1009CAFBEB08C9" box="[1035,1146,1764,1789]" italics="true" pageId="2" pageNumber="39">hufelandi</emphasis>
</taxonomicName>
group of species. Some species of this group have narrow buccal tube and no (or almost not visible) bands of teeth in the buccal cavity (
<taxonomicName id="4C2493C7FD3BE15418BA081AFE700979" authority="Biserov, 1990" authorityName="Biserov" authorityYear="1990" box="[161,481,1843,1869]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="39" phylum="Tardigrada" rank="species" species="diversus">
<emphasis id="B9503456FD3BE15418BA081AFEBC0979" box="[161,301,1844,1869]" italics="true" pageId="2" pageNumber="39">M. diversus</emphasis>
<bibRefCitation id="EFB595B5FD3BE154192C081DFE700979" author="Biserov" box="[311,481,1843,1869]" pageId="2" pageNumber="47" refString="Biserov, V. I. (1990) On the revision of the genus Macrobiotus, the subgenus Macrobiotus s. str.: A new systematic status of the group hufelandi (Tardigrada, Macrobiotidae). Communication I. Zoologichesky Zhurnal, 69, 5 - 17. (in Russian with English summary)" type="book chapter" year="1990">Biserov, 1990</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C2493C7FD3BE15419EB081AFD390979" box="[496,680,1844,1869]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="39" phylum="Tardigrada" rank="species" species="madegassus">
<emphasis id="B9503456FD3BE15419EB081AFD390979" box="[496,680,1844,1869]" italics="true" pageId="2" pageNumber="39">M. madegassus</emphasis>
</taxonomicName>
;
<taxonomicName id="4C2493C7FD3BE1541AA0081AFCAA0979" box="[699,827,1844,1869]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="39" phylum="Tardigrada" rank="species" species="humilis">
<emphasis id="B9503456FD3BE1541AA0081AFCAA0979" box="[699,827,1844,1869]" italics="true" pageId="2" pageNumber="39">M. humilis</emphasis>
</taxonomicName>
;
<taxonomicName id="4C2493C7FD3BE1541B55081AFC600979" box="[846,1009,1844,1869]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="39" phylum="Tardigrada" rank="species" species="denticulus">
<emphasis id="B9503456FD3BE1541B55081AFC600979" box="[846,1009,1844,1869]" italics="true" pageId="2" pageNumber="39">M. denticulus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2493C7FD3BE1541C2A081AFAF40979" authority="Dastych, 2005" authorityName="Dastych" authorityYear="2005" box="[1073,1381,1843,1869]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="39" phylum="Tardigrada" rank="species" species="ramoli">
<emphasis id="B9503456FD3BE1541C2A081AFB380979" box="[1073,1193,1844,1869]" italics="true" pageId="2" pageNumber="39">M. ramoli</emphasis>
<bibRefCitation id="EFB595B5FD3BE1541CAF081DFAF40979" author="Dastych" box="[1204,1381,1843,1869]" pageId="2" pageNumber="47" refString="Dastych, H. (2005) Macrobiotus ramoli sp. nov., a new tardigrade species from the nival zone of the Otztal Alps, Austria (Tardigrada). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 102, 21 - 35." type="journal article" year="2005">Dastych, 2005</bibRefCitation>
</taxonomicName>
).
<taxonomicName id="4C2493C7FD3BE1541D67081AFF660941" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="39" phylum="Tardigrada" rank="species" species="almadai" status="sp. nov.">
<emphasis id="B9503456FD3BE1541D67081AFF660941" italics="true" pageId="2" pageNumber="39">M. almadai</emphasis>
</taxonomicName>
<emphasis id="B9503456FD3BE15419190875FEF30941" bold="true" box="[258,354,1883,1909]" pageId="2" pageNumber="39">
<taxonomicNameLabel id="A263892DFD3BE15419190875FEF30941" box="[258,354,1883,1909]" pageId="2" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from them in some characters of the eggs and, in addition, from
<taxonomicName id="4C2493C7FD3BE1541CC50872FAF80941" box="[1246,1385,1884,1909]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="39" phylum="Tardigrada" rank="species" species="diversus">
<emphasis id="B9503456FD3BE1541CC50872FAF80941" box="[1246,1385,1884,1909]" italics="true" pageId="2" pageNumber="39">M. diversus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2493C7FD3BE1541D670872FEB409A9" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="39" phylum="Tardigrada" rank="species" species="madegassus">
<emphasis id="B9503456FD3BE1541D670872FEB409A9" italics="true" pageId="2" pageNumber="39">M. madegassus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2493C7FD3BE154194608AAFE4B09A9" box="[349,474,1924,1949]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="39" phylum="Tardigrada" rank="species" species="humilis">
<emphasis id="B9503456FD3BE154194608AAFE4B09A9" box="[349,474,1924,1949]" italics="true" pageId="2" pageNumber="39">M. humilis</emphasis>
</taxonomicName>
in having three separate dorsal ridges instead of a long single ridge; from
<taxonomicName id="4C2493C7FD3BE1541D5A08AAFF7409F1" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="39" phylum="Tardigrada" rank="species" species="denticulus">
<emphasis id="B9503456FD3BE1541D5A08AAFF7409F1" italics="true" pageId="2" pageNumber="39">M. denticulus</emphasis>
</taxonomicName>
in lacking a tooth in the anterior part of the buccal tube, and from
<taxonomicName id="4C2493C7FD3BE1541C3F0882FB0C09F1" box="[1060,1181,1964,1989]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="39" phylum="Tardigrada" rank="species" species="ramoli">
<emphasis id="B9503456FD3BE1541C3F0882FB0C09F1" box="[1060,1181,1964,1989]" italics="true" pageId="2" pageNumber="39">M. ramoli</emphasis>
</taxonomicName>
in having the dorsomedian ridge very thin, and smooth lunules.
</paragraph>
<paragraph id="8B9BE844FD3AE15518DE0FB6FD930F66" blockId="3.[151,1436,152,338]" pageId="3" pageNumber="40">
The new species differs from
<taxonomicName id="4C2493C7FD3AE1551A3C0FB6FD2E0E85" box="[551,703,152,177]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="40" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="B9503456FD3AE1551A3C0FB6FD2E0E85" box="[551,703,152,177]" italics="true" pageId="3" pageNumber="40">M. hufelandi</emphasis>
</taxonomicName>
in having narrower buccal tube (
<tableCitation id="C6A6DDFFFD3AE1551C530FB6FB020E86" box="[1096,1171,152,178]" captionStart="TABLE 1" captionStartId="3.[151,239,391,415]" captionTargetBox="[159,1380,637,1519]" captionTargetPageId="3" captionText="TABLE 1. Measurements (in µm) of some structures of the holotype, the smallest and the largest measured specimens of Macrobiotus almadai sp. nov., of a paratype of Macrobiotus sandrae and of a specimen of Macrobiotus hufelandi collected in the locus typicus." httpUri="http://table.plazi.org/id/DF5BB8CCFD3AE155188C0EA9FE230FD0" pageId="3" pageNumber="40" tableUuid="DF5BB8CCFD3AE155188C0EA9FE230FD0">Tab. 1</tableCitation>
;
<figureCitation id="131FF4C1FD3AE1551CBF0FB6FA9D0E86" box="[1188,1292,152,178]" captionStart="FIGURE 1. A G" captionStartId="2.[151,255,1339,1363]" captionTargetBox="[268,1285,318,1338]" captionTargetId="figure@2.[255,1333,261,1373]" captionTargetPageId="2" captionText="FIGURE 1. A G, Macrobiotus almadai sp. nov. (holotype): A, bucco-pharyngeal apparatus; B and C, claws of the third pair of legs; D, claws of the hind legs; E, egg; F and G, details of the egg shell (arrow indicates the small teeth of the terminal disc). (Scale bar = 10 µm)." httpUri="https://zenodo.org/record/182263/files/figure.png" pageId="3" pageNumber="40">Figs. 1A</figureCitation>
,
<figureCitation id="131FF4C1FD3AE1551D020FB6FAD00E86" box="[1305,1345,152,178]" captionStart="FIGURE 2. A and B" captionStartId="4.[151,255,1191,1215]" captionTargetBox="[351,1233,318,1190]" captionTargetId="figure@4.[330,1258,277,1210]" captionTargetPageId="4" captionText="FIGURE 2. A and B, Macrobiotus hufelandi: A, bucco-pharyngeal apparatus; B, detail of the egg (Scale bar = 10 µm). C and D, Macrobiotus sandrae (paratype): C, bucco-pharyngeal apparatus; D, detail of the egg (Scale bar = 10 µm)." httpUri="https://zenodo.org/record/182264/files/figure.png" pageId="3" pageNumber="40">2A</figureCitation>
); bands of teeth in the buccal cavity not visible; stylet supports inserted on buccal tube in more anterior position (
<emphasis id="B9503456FD3AE1551D760FEEFA120EED" box="[1389,1411,192,217]" italics="true" pageId="3" pageNumber="40">pt</emphasis>
=
<quantity id="4CDC45A1FD3AE155188C0FC6FF7D0F36" box="[151,236,232,258]" metricMagnitude="0" metricUnit="m" metricValue="1.9431" pageId="3" pageNumber="40" unit="in" value="76.5">
<emphasis id="B9503456FD3AE155188C0FC6FF5B0F35" box="[151,202,232,257]" italics="true" pageId="3" pageNumber="40">76.5</emphasis>
in
</quantity>
<taxonomicName id="4C2493C7FD3AE15518EC0FC6FE130F35" box="[247,386,232,257]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="40" phylum="Tardigrada" rank="species" species="almadai" status="sp. nov.">
<emphasis id="B9503456FD3AE15518EC0FC6FE130F35" box="[247,386,232,257]" italics="true" pageId="3" pageNumber="40">M. almadai</emphasis>
</taxonomicName>
<emphasis id="B9503456FD3AE15519960FC9FE7F0F35" bold="true" box="[397,494,231,257]" pageId="3" pageNumber="40">
<taxonomicNameLabel id="A263892DFD3AE15519960FC9FE7F0F35" box="[397,494,231,257]" pageId="3" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<quantity id="4CDC45A1FD3AE1551A340FC6FD0B0F36" box="[559,666,232,258]" metricMagnitude="0" metricUnit="m" metricValue="2.0446999999999997" metricValueMax="2.0574" metricValueMin="2.032" pageId="3" pageNumber="40" unit="in" value="80.5" valueMax="81.0" valueMin="80.0">
<emphasis id="B9503456FD3AE1551A340FC6FDE90F35" box="[559,632,232,257]" italics="true" pageId="3" pageNumber="40">8081</emphasis>
in
</quantity>
<taxonomicName id="4C2493C7FD3AE1551ABE0FC6FCAE0F35" box="[677,831,232,257]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="40" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="B9503456FD3AE1551ABE0FC6FCAE0F35" box="[677,831,232,257]" italics="true" pageId="3" pageNumber="40">M. hufelandi</emphasis>
</taxonomicName>
); slightly shorter macroplacoids (
<tableCitation id="C6A6DDFFFD3AE1551CC20FC6FAB70F36" box="[1241,1318,232,258]" captionStart="TABLE 1" captionStartId="3.[151,239,391,415]" captionTargetBox="[159,1380,637,1519]" captionTargetPageId="3" captionText="TABLE 1. Measurements (in µm) of some structures of the holotype, the smallest and the largest measured specimens of Macrobiotus almadai sp. nov., of a paratype of Macrobiotus sandrae and of a specimen of Macrobiotus hufelandi collected in the locus typicus." httpUri="http://table.plazi.org/id/DF5BB8CCFD3AE155188C0EA9FE230FD0" pageId="3" pageNumber="40" tableUuid="DF5BB8CCFD3AE155188C0EA9FE230FD0">Tab. 1</tableCitation>
); slightly longer claws (
<tableCitation id="C6A6DDFFFD3AE15519200E3EFE150F1E" box="[315,388,272,298]" captionStart="TABLE 1" captionStartId="3.[151,239,391,415]" captionTargetBox="[159,1380,637,1519]" captionTargetPageId="3" captionText="TABLE 1. Measurements (in µm) of some structures of the holotype, the smallest and the largest measured specimens of Macrobiotus almadai sp. nov., of a paratype of Macrobiotus sandrae and of a specimen of Macrobiotus hufelandi collected in the locus typicus." httpUri="http://table.plazi.org/id/DF5BB8CCFD3AE155188C0EA9FE230FD0" pageId="3" pageNumber="40" tableUuid="DF5BB8CCFD3AE155188C0EA9FE230FD0">Tab. 1</tableCitation>
); eggs with more delicate and regular reticular design and terminal disc of processes with smaller teeth (
<figureCitation id="131FF4C1FD3AE15519260E16FE550F66" box="[317,452,312,338]" captionStart="FIGURE 1. A G" captionStartId="2.[151,255,1339,1363]" captionTargetBox="[268,1285,318,1338]" captionTargetId="figure@2.[255,1333,261,1373]" captionTargetPageId="2" captionText="FIGURE 1. A G, Macrobiotus almadai sp. nov. (holotype): A, bucco-pharyngeal apparatus; B and C, claws of the third pair of legs; D, claws of the hind legs; E, egg; F and G, details of the egg shell (arrow indicates the small teeth of the terminal disc). (Scale bar = 10 µm)." httpUri="https://zenodo.org/record/182263/files/figure.png" pageId="3" pageNumber="40">Figs. 1E, G</figureCitation>
,
<figureCitation id="131FF4C1FD3AE15519D40E16FE4E0F66" box="[463,479,312,338]" captionStart="FIGURE 2. A and B" captionStartId="4.[151,255,1191,1215]" captionTargetBox="[351,1233,318,1190]" captionTargetId="figure@4.[330,1258,277,1210]" captionTargetPageId="4" captionText="FIGURE 2. A and B, Macrobiotus hufelandi: A, bucco-pharyngeal apparatus; B, detail of the egg (Scale bar = 10 µm). C and D, Macrobiotus sandrae (paratype): C, bucco-pharyngeal apparatus; D, detail of the egg (Scale bar = 10 µm)." httpUri="https://zenodo.org/record/182264/files/figure.png" pageId="3" pageNumber="40">2</figureCitation>
B).
</paragraph>
<caption id="DF5BB8CCFD3AE155188C0EA9FE230FD0" ID-Table-UUID="DF5BB8CCFD3AE155188C0EA9FE230FD0" httpUri="http://table.plazi.org/id/DF5BB8CCFD3AE155188C0EA9FE230FD0" pageId="3" pageNumber="40" targetBox="[159,1380,637,1519]" targetIsTable="true" targetPageId="3">
<paragraph id="8B9BE844FD3AE155188C0EA9FE230FD0" blockId="3.[151,1436,391,484]" pageId="3" pageNumber="40">
<emphasis id="B9503456FD3AE155188C0EA9FE980FAB" bold="true" box="[151,265,391,415]" pageId="3" pageNumber="40">TABLE 1.</emphasis>
Measurements (in µm) of some structures of the holotype, the smallest and the largest measured specimens of
<taxonomicName id="4C2493C7FD3AE155188C0E84FEEF0FF5" box="[151,382,426,449]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="40" phylum="Tardigrada" rank="species" species="almadai" status="sp. nov.">
<emphasis id="B9503456FD3AE155188C0E84FEEF0FF5" box="[151,382,426,449]" italics="true" pageId="3" pageNumber="40">Macrobiotus almadai</emphasis>
</taxonomicName>
<emphasis id="B9503456FD3AE155199E0E84FE4D0FF6" bold="true" box="[389,476,426,450]" pageId="3" pageNumber="40">
<taxonomicNameLabel id="A263892DFD3AE155199E0E84FE4D0FF6" box="[389,476,426,450]" pageId="3" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, of a paratype of
<taxonomicName id="4C2493C7FD3AE1551A850E84FC150FF5" box="[670,900,426,449]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="40" phylum="Tardigrada" rank="species" species="sandrae">
<emphasis id="B9503456FD3AE1551A850E84FC150FF5" box="[670,900,426,449]" italics="true" pageId="3" pageNumber="40">Macrobiotus sandrae</emphasis>
</taxonomicName>
and of a specimen of
<taxonomicName id="4C2493C7FD3AE1551C6D0E84FAFB0FF5" box="[1142,1386,426,449]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="40" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="B9503456FD3AE1551C6D0E84FAFB0FF5" box="[1142,1386,426,449]" italics="true" pageId="3" pageNumber="40">Macrobiotus hufelandi</emphasis>
</taxonomicName>
collected in the locus typicus.
</paragraph>
</caption>
<paragraph id="8B9BE844FD3AE15518840D2BFAF50C28" blockId="3.[159,1380,516,617]" box="[159,1380,516,540]" pageId="3" pageNumber="40">
Characters
<taxonomicName id="4C2493C7FD3AE1551A540D2BFD5B0C28" box="[591,714,517,540]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="40" phylum="Tardigrada" rank="species" species="almadai" status="sp. nov.">
<emphasis id="B9503456FD3AE1551A540D2BFD5B0C28" box="[591,714,517,540]" italics="true" pageId="3" pageNumber="40">M. almadai</emphasis>
</taxonomicName>
<emphasis id="B9503456FD3AE1551ACB0D2AFCB70C28" bold="true" box="[720,806,516,540]" pageId="3" pageNumber="40">
<taxonomicNameLabel id="A263892DFD3AE1551ACB0D2AFCB70C28" box="[720,806,516,540]" pageId="3" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="B9503456FD3AE1551C270D2BFAF50C28" box="[1084,1380,517,540]" italics="true" pageId="3" pageNumber="40">
<taxonomicName id="4C2493C7FD3AE1551C270D2BFB270C28" box="[1084,1206,517,540]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="40" phylum="Tardigrada" rank="species" species="sandrae">M. sandrae</taxonomicName>
<taxonomicName id="4C2493C7FD3AE1551CC00D2BFAF50C28" box="[1243,1380,517,540]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="40" phylum="Tardigrada" rank="species" species="hufelandi">M. hufelandi</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8B9BE844FD3AE1551A540D1EFC620C73" blockId="3.[159,1380,516,617]" box="[591,1011,560,583]" pageId="3" pageNumber="40">
<typeStatus id="549F56E6FD3AE1551A540D1EFD230C73" box="[591,690,560,583]" pageId="3" pageNumber="40" type="holotype">Holotype</typeStatus>
Smallest Largest
</paragraph>
<paragraph id="8B9BE844FD3AE1551AE10D7CFB980C5D" blockId="3.[159,1380,516,617]" box="[762,1033,594,617]" pageId="3" pageNumber="40">specimen specimen</paragraph>
<paragraph id="8B9BE844FD3AE15518840D53FB790BDB" pageId="3" pageNumber="40">
<table id="F9241AE4FD3A1EA918840D53FAF50BDB" box="[159,1380,637,1519]" gridcols="6" gridrows="13" pageId="3" pageNumber="40">
<tr id="3514EA06FD3A1EA918840D53FAF50CA0" box="[159,1380,637,660]" gridrow="0" pageId="3" pageNumber="40">
<th id="76C5837AFD3A1EA918840D53FE5F0CA0" box="[159,462,637,660]" gridcol="0" gridrow="0" pageId="3" pageNumber="40">Body length</th>
<th id="76C5837AFD3A1EA91A540D53FDEF0CA0" box="[591,638,637,660]" gridcol="1" gridrow="0" pageId="3" pageNumber="40">340</th>
<th id="76C5837AFD3A1EA91AE10D53FCB80CA0" box="[762,809,637,660]" gridcol="2" gridrow="0" pageId="3" pageNumber="40">195</th>
<th id="76C5837AFD3A1EA91BBF0D53FC450CA0" box="[932,980,637,660]" gridcol="3" gridrow="0" pageId="3" pageNumber="40">394</th>
<th id="76C5837AFD3A1EA91C270D53FBFA0CA0" box="[1084,1131,637,660]" gridcol="4" gridrow="0" pageId="3" pageNumber="40">376</th>
<th id="76C5837AFD3A1EA91CC70D53FAF50CA0" box="[1244,1380,637,660]" gridcol="5" gridrow="0" pageId="3" pageNumber="40">?</th>
</tr>
<tr id="3514EA06FD3A1EA918840D86FAF50C8B" box="[159,1380,680,703]" gridrow="1" pageId="3" pageNumber="40">
<th id="76C5837AFD3A1EA918840D86FE5F0C8B" box="[159,462,680,703]" gridcol="0" gridrow="1" pageId="3" pageNumber="40">Buccal tube length</th>
<td id="76C5837AFD3A1EA91A540D86FDEF0C8B" box="[591,638,680,703]" gridcol="1" gridrow="1" pageId="3" pageNumber="40">32.4</td>
<td id="76C5837AFD3A1EA91AE10D86FCB80C8B" box="[762,809,680,703]" gridcol="2" gridrow="1" pageId="3" pageNumber="40">20.8</td>
<td id="76C5837AFD3A1EA91BBF0D86FC450C8B" box="[932,980,680,703]" gridcol="3" gridrow="1" pageId="3" pageNumber="40">34.9</td>
<td id="76C5837AFD3A1EA91C270D86FBFA0C8B" box="[1084,1131,680,703]" gridcol="4" gridrow="1" pageId="3" pageNumber="40">37.3</td>
<td id="76C5837AFD3A1EA91CC70D86FAF50C8B" box="[1244,1380,680,703]" gridcol="5" gridrow="1" pageId="3" pageNumber="40">34.5</td>
</tr>
<tr id="3514EA06FD3A1EA918840DFCFAF50D38" box="[159,1380,722,780]" gridrow="2" pageId="3" pageNumber="40">
<th id="76C5837AFD3A1EA918840DFCFE5F0D38" box="[159,462,722,780]" gridcol="0" gridrow="2" pageId="3" pageNumber="40">Buccal tube external width pt</th>
<td id="76C5837AFD3A1EA91A540DFCFDEF0D38" box="[591,638,722,780]" gridcol="1" gridrow="2" pageId="3" pageNumber="40">
3.6
<emphasis id="B9503456FD3AE1551A540DDBFDEB0D38" box="[591,634,757,780]" italics="true" pageId="3" pageNumber="40">11.1</emphasis>
</td>
<td id="76C5837AFD3A1EA91AE10DFCFCB80D38" box="[762,809,722,780]" gridcol="2" gridrow="2" pageId="3" pageNumber="40">
2.1
<emphasis id="B9503456FD3AE1551AE10DDBFCB80D38" box="[762,809,757,780]" italics="true" pageId="3" pageNumber="40">10.1</emphasis>
</td>
<td id="76C5837AFD3A1EA91BBF0DFCFC450D38" box="[932,980,722,780]" gridcol="3" gridrow="2" pageId="3" pageNumber="40">
4.0
<emphasis id="B9503456FD3AE1551BBF0DDBFC5E0D38" box="[932,975,757,780]" italics="true" pageId="3" pageNumber="40">11.5</emphasis>
</td>
<td id="76C5837AFD3A1EA91C270DFCFBFA0D38" box="[1084,1131,722,780]" gridcol="4" gridrow="2" pageId="3" pageNumber="40">
5.3
<emphasis id="B9503456FD3AE1551C270DDBFBFA0D38" box="[1084,1131,757,780]" italics="true" pageId="3" pageNumber="40">14.2</emphasis>
</td>
<td id="76C5837AFD3A1EA91CC70DFCFAF50D38" box="[1244,1380,722,780]" gridcol="5" gridrow="2" pageId="3" pageNumber="40">
4.8
<emphasis id="B9503456FD3AE1551CC70DDBFA9A0D38" box="[1244,1291,757,780]" italics="true" pageId="3" pageNumber="40">13.9</emphasis>
</td>
</tr>
<tr id="3514EA06FD3A1EA918840C0EFAF50D03" box="[159,1380,800,823]" gridrow="3" pageId="3" pageNumber="40">
<th id="76C5837AFD3A1EA918840C0EFE5F0D03" box="[159,462,800,823]" gridcol="0" gridrow="3" pageId="3" pageNumber="40">
Stylets support insertion
<emphasis id="B9503456FD3AE15519B30C0EFE2C0D03" box="[424,445,800,823]" italics="true" pageId="3" pageNumber="40">pt</emphasis>
</th>
<td id="76C5837AFD3A1EA91A540C0EFDEF0D03" box="[591,638,800,823]" gridcol="1" gridrow="3" pageId="3" pageNumber="40">
<emphasis id="B9503456FD3AE1551A540C0EFDEF0D03" box="[591,638,800,823]" italics="true" pageId="3" pageNumber="40">76.5</emphasis>
</td>
<td id="76C5837AFD3A1EA91AE10C0EFCB80D03" box="[762,809,800,823]" gridcol="2" gridrow="3" pageId="3" pageNumber="40">
<emphasis id="B9503456FD3AE1551AE10C0EFCB80D03" box="[762,809,800,823]" italics="true" pageId="3" pageNumber="40">74.0</emphasis>
</td>
<td id="76C5837AFD3A1EA91BBF0C0EFC450D03" box="[932,980,800,823]" gridcol="3" gridrow="3" pageId="3" pageNumber="40">
<emphasis id="B9503456FD3AE1551BBF0C0EFC420D03" box="[932,979,800,823]" italics="true" pageId="3" pageNumber="40">76.8</emphasis>
</td>
<td id="76C5837AFD3A1EA91C270C0EFBFA0D03" box="[1084,1131,800,823]" gridcol="4" gridrow="3" pageId="3" pageNumber="40">
<emphasis id="B9503456FD3AE1551C270C0EFBFA0D03" box="[1084,1131,800,823]" italics="true" pageId="3" pageNumber="40">76.4</emphasis>
</td>
<td id="76C5837AFD3A1EA91CC70C0EFAF50D03" box="[1244,1380,800,823]" gridcol="5" gridrow="3" pageId="3" pageNumber="40">
<emphasis id="B9503456FD3AE1551CC70C0EFA9A0D03" box="[1244,1291,800,823]" italics="true" pageId="3" pageNumber="40">81.1</emphasis>
</td>
</tr>
<tr id="3514EA06FD3A1EA918840C64FAF50DB0" box="[159,1380,842,900]" gridrow="4" pageId="3" pageNumber="40">
<th id="76C5837AFD3A1EA918840C64FE5F0DB0" box="[159,462,842,900]" gridcol="0" gridrow="4" pageId="3" pageNumber="40">First macroplacoid length pt</th>
<td id="76C5837AFD3A1EA91A540C64FDEF0DB0" box="[591,638,842,900]" gridcol="1" gridrow="4" pageId="3" pageNumber="40">
8.2
<emphasis id="B9503456FD3AE1551A540C43FDEF0DB0" box="[591,638,877,900]" italics="true" pageId="3" pageNumber="40">25.3</emphasis>
</td>
<td id="76C5837AFD3A1EA91AE10C64FCB80DB0" box="[762,809,842,900]" gridcol="2" gridrow="4" pageId="3" pageNumber="40">
4.6
<emphasis id="B9503456FD3AE1551AE10C43FCB80DB0" box="[762,809,877,900]" italics="true" pageId="3" pageNumber="40">22.1</emphasis>
</td>
<td id="76C5837AFD3A1EA91BBF0C64FC450DB0" box="[932,980,842,900]" gridcol="3" gridrow="4" pageId="3" pageNumber="40">
8.9
<emphasis id="B9503456FD3AE1551BBF0C43FC420DB0" box="[932,979,877,900]" italics="true" pageId="3" pageNumber="40">25.5</emphasis>
</td>
<td id="76C5837AFD3A1EA91C270C64FBFA0DB0" box="[1084,1131,842,900]" gridcol="4" gridrow="4" pageId="3" pageNumber="40">
10.5
<emphasis id="B9503456FD3AE1551C270C43FBFA0DB0" box="[1084,1131,877,900]" italics="true" pageId="3" pageNumber="40">28.2</emphasis>
</td>
<td id="76C5837AFD3A1EA91CC70C64FAF50DB0" box="[1244,1380,842,900]" gridcol="5" gridrow="4" pageId="3" pageNumber="40">
9.3
<emphasis id="B9503456FD3AE1551CC70C43FA9A0DB0" box="[1244,1291,877,900]" italics="true" pageId="3" pageNumber="40">27.0</emphasis>
</td>
</tr>
<tr id="3514EA06FD3A1EA918840CB6FAF50DE5" box="[159,1380,920,977]" gridrow="5" pageId="3" pageNumber="40">
<th id="76C5837AFD3A1EA918840CB6FE5F0DE5" box="[159,462,920,977]" gridcol="0" gridrow="5" pageId="3" pageNumber="40">Second macroplacoid length pt</th>
<td id="76C5837AFD3A1EA91A540CB6FDEF0DE5" box="[591,638,920,977]" gridcol="1" gridrow="5" pageId="3" pageNumber="40">
5.5
<emphasis id="B9503456FD3AE1551A540C94FDEF0DE5" box="[591,638,954,977]" italics="true" pageId="3" pageNumber="40">17.0</emphasis>
</td>
<td id="76C5837AFD3A1EA91AE10CB6FCB80DE5" box="[762,809,920,977]" gridcol="2" gridrow="5" pageId="3" pageNumber="40">
3.1
<emphasis id="B9503456FD3AE1551AE10C94FCB80DE5" box="[762,809,954,977]" italics="true" pageId="3" pageNumber="40">14.9</emphasis>
</td>
<td id="76C5837AFD3A1EA91BBF0CB6FC450DE5" box="[932,980,920,977]" gridcol="3" gridrow="5" pageId="3" pageNumber="40">
6.2
<emphasis id="B9503456FD3AE1551BBF0C94FC420DE5" box="[932,979,954,977]" italics="true" pageId="3" pageNumber="40">17.8</emphasis>
</td>
<td id="76C5837AFD3A1EA91C270CB6FBFA0DE5" box="[1084,1131,920,977]" gridcol="4" gridrow="5" pageId="3" pageNumber="40">
7.3
<emphasis id="B9503456FD3AE1551C270C94FBFA0DE5" box="[1084,1131,954,977]" italics="true" pageId="3" pageNumber="40">19.6</emphasis>
</td>
<td id="76C5837AFD3A1EA91CC70CB6FAF50DE5" box="[1244,1380,920,977]" gridcol="5" gridrow="5" pageId="3" pageNumber="40">
6.7
<emphasis id="B9503456FD3AE1551CC70C94FA9A0DE5" box="[1244,1291,954,977]" italics="true" pageId="3" pageNumber="40">19.4</emphasis>
</td>
</tr>
<tr id="3514EA06FD3A1EA918840CCBFAF50A2B" box="[159,1380,997,1055]" gridrow="6" pageId="3" pageNumber="40">
<th id="76C5837AFD3A1EA918840CCBFE5F0A2B" box="[159,462,997,1055]" gridcol="0" gridrow="6" pageId="3" pageNumber="40">Microplacoid length pt</th>
<td id="76C5837AFD3A1EA91A540CCBFDEF0A2B" box="[591,638,997,1055]" gridcol="1" gridrow="6" pageId="3" pageNumber="40">
2.7
<emphasis id="B9503456FD3AE1551A540B26FDE00A2B" box="[591,625,1032,1055]" italics="true" pageId="3" pageNumber="40">8.3</emphasis>
</td>
<td id="76C5837AFD3A1EA91AE10CCBFCB80A2B" box="[762,809,997,1055]" gridcol="2" gridrow="6" pageId="3" pageNumber="40">
1.5
<emphasis id="B9503456FD3AE1551AE10B26FC8D0A2B" box="[762,796,1032,1055]" italics="true" pageId="3" pageNumber="40">7.2</emphasis>
</td>
<td id="76C5837AFD3A1EA91BBF0CCBFC450A2B" box="[932,980,997,1055]" gridcol="3" gridrow="6" pageId="3" pageNumber="40">
2.8
<emphasis id="B9503456FD3AE1551BBF0B26FC570A2B" box="[932,966,1032,1055]" italics="true" pageId="3" pageNumber="40">8.0</emphasis>
</td>
<td id="76C5837AFD3A1EA91C270CCBFBFA0A2B" box="[1084,1131,997,1055]" gridcol="4" gridrow="6" pageId="3" pageNumber="40">
2.6
<emphasis id="B9503456FD3AE1551C270B26FBCF0A2B" box="[1084,1118,1032,1055]" italics="true" pageId="3" pageNumber="40">7.0</emphasis>
</td>
<td id="76C5837AFD3A1EA91CC70CCBFAF50A2B" box="[1244,1380,997,1055]" gridcol="5" gridrow="6" pageId="3" pageNumber="40">
2.4
<emphasis id="B9503456FD3AE1551CC70B26FB6F0A2B" box="[1244,1278,1032,1055]" italics="true" pageId="3" pageNumber="40">7.0</emphasis>
</td>
</tr>
<tr id="3514EA06FD3A1EA918840B1CFAF50A58" box="[159,1380,1074,1132]" gridrow="7" pageId="3" pageNumber="40">
<th id="76C5837AFD3A1EA918840B1CFE5F0A58" box="[159,462,1074,1132]" gridcol="0" gridrow="7" pageId="3" pageNumber="40">Placoid row length pt</th>
<td id="76C5837AFD3A1EA91A540B1CFDEF0A58" box="[591,638,1074,1132]" gridcol="1" gridrow="7" pageId="3" pageNumber="40">
16.6
<emphasis id="B9503456FD3AE1551A540B7BFDEF0A58" box="[591,638,1109,1132]" italics="true" pageId="3" pageNumber="40">51.2</emphasis>
</td>
<td id="76C5837AFD3A1EA91AE10B1CFCB80A58" box="[762,809,1074,1132]" gridcol="2" gridrow="7" pageId="3" pageNumber="40">
10.2
<emphasis id="B9503456FD3AE1551AE10B7BFCB80A58" box="[762,809,1109,1132]" italics="true" pageId="3" pageNumber="40">49.0</emphasis>
</td>
<td id="76C5837AFD3A1EA91BBF0B1CFC450A58" box="[932,980,1074,1132]" gridcol="3" gridrow="7" pageId="3" pageNumber="40">
18.6
<emphasis id="B9503456FD3AE1551BBF0B7BFC420A58" box="[932,979,1109,1132]" italics="true" pageId="3" pageNumber="40">53.3</emphasis>
</td>
<td id="76C5837AFD3A1EA91C270B1CFBFA0A58" box="[1084,1131,1074,1132]" gridcol="4" gridrow="7" pageId="3" pageNumber="40">
22.5
<emphasis id="B9503456FD3AE1551C270B7BFBFA0A58" box="[1084,1131,1109,1132]" italics="true" pageId="3" pageNumber="40">60.3</emphasis>
</td>
<td id="76C5837AFD3A1EA91CC70B1CFAF50A58" box="[1244,1380,1074,1132]" gridcol="5" gridrow="7" pageId="3" pageNumber="40">
19.2
<emphasis id="B9503456FD3AE1551CC70B7BFA9A0A58" box="[1244,1291,1109,1132]" italics="true" pageId="3" pageNumber="40">55.7</emphasis>
</td>
</tr>
<tr id="3514EA06FD3A1EA918840BAEFAF50A8E" box="[159,1380,1152,1210]" gridrow="8" pageId="3" pageNumber="40">
<th id="76C5837AFD3A1EA918840BAEFE5F0A8E" box="[159,462,1152,1210]" gridcol="0" gridrow="8" pageId="3" pageNumber="40">Macroplacoid row length pt</th>
<td id="76C5837AFD3A1EA91A540BAEFDEF0A8E" box="[591,638,1152,1210]" gridcol="1" gridrow="8" pageId="3" pageNumber="40">
14.0
<emphasis id="B9503456FD3AE1551A540B8DFDEF0A8E" box="[591,638,1187,1210]" italics="true" pageId="3" pageNumber="40">43.2</emphasis>
</td>
<td id="76C5837AFD3A1EA91AE10BAEFCB80A8E" box="[762,809,1152,1210]" gridcol="2" gridrow="8" pageId="3" pageNumber="40">
8.4
<emphasis id="B9503456FD3AE1551AE10B8DFCB80A8E" box="[762,809,1187,1210]" italics="true" pageId="3" pageNumber="40">40.4</emphasis>
</td>
<td id="76C5837AFD3A1EA91BBF0BAEFC450A8E" box="[932,980,1152,1210]" gridcol="3" gridrow="8" pageId="3" pageNumber="40">
15.8
<emphasis id="B9503456FD3AE1551BBF0B8DFC420A8E" box="[932,979,1187,1210]" italics="true" pageId="3" pageNumber="40">45.3</emphasis>
</td>
<td id="76C5837AFD3A1EA91C270BAEFBFA0A8E" box="[1084,1131,1152,1210]" gridcol="4" gridrow="8" pageId="3" pageNumber="40">
19.1
<emphasis id="B9503456FD3AE1551C270B8DFBFA0A8E" box="[1084,1131,1187,1210]" italics="true" pageId="3" pageNumber="40">51.2</emphasis>
</td>
<td id="76C5837AFD3A1EA91CC70BAEFAF50A8E" box="[1244,1380,1152,1210]" gridcol="5" gridrow="8" pageId="3" pageNumber="40">
16.4
<emphasis id="B9503456FD3AE1551CC70B8DFA9A0A8E" box="[1244,1291,1187,1210]" italics="true" pageId="3" pageNumber="40">47.5</emphasis>
</td>
</tr>
<tr id="3514EA06FD3A1EA918840BE3FAF50B33" box="[159,1380,1229,1287]" gridrow="9" pageId="3" pageNumber="40">
<th id="76C5837AFD3A1EA918840BE3FE5F0B33" box="[159,462,1229,1287]" gridcol="0" gridrow="9" pageId="3" pageNumber="40">External claws II, III pt</th>
<td id="76C5837AFD3A1EA91A540BE3FDEF0B33" box="[591,638,1229,1287]" gridcol="1" gridrow="9" pageId="3" pageNumber="40">
9.6
<emphasis id="B9503456FD3AE1551A540BDEFDEF0B33" box="[591,638,1264,1287]" italics="true" pageId="3" pageNumber="40">29.6</emphasis>
</td>
<td id="76C5837AFD3A1EA91AE10BE3FCB80B33" box="[762,809,1229,1287]" gridcol="2" gridrow="9" pageId="3" pageNumber="40">
6.0
<emphasis id="B9503456FD3AE1551AE10BDEFCB80B33" box="[762,809,1264,1287]" italics="true" pageId="3" pageNumber="40">28.8</emphasis>
</td>
<td id="76C5837AFD3A1EA91BBF0BE3FC450B33" box="[932,980,1229,1287]" gridcol="3" gridrow="9" pageId="3" pageNumber="40">
9.7
<emphasis id="B9503456FD3AE1551BBF0BDEFC420B33" box="[932,979,1264,1287]" italics="true" pageId="3" pageNumber="40">27.8</emphasis>
</td>
<td id="76C5837AFD3A1EA91C270BE3FBFA0B33" box="[1084,1131,1229,1287]" gridcol="4" gridrow="9" pageId="3" pageNumber="40">
9.2
<emphasis id="B9503456FD3AE1551C270BDEFBFA0B33" box="[1084,1131,1264,1287]" italics="true" pageId="3" pageNumber="40">24.7</emphasis>
</td>
<td id="76C5837AFD3A1EA91CC70BE3FAF50B33" box="[1244,1380,1229,1287]" gridcol="5" gridrow="9" pageId="3" pageNumber="40">
7.8
<emphasis id="B9503456FD3AE1551CC70BDEFA9A0B33" box="[1244,1291,1264,1287]" italics="true" pageId="3" pageNumber="40">22.6</emphasis>
</td>
</tr>
<tr id="3514EA06FD3A1EA918840A34FAF50B60" box="[159,1380,1306,1364]" gridrow="10" pageId="3" pageNumber="40">
<th id="76C5837AFD3A1EA918840A34FE5F0B60" box="[159,462,1306,1364]" gridcol="0" gridrow="10" pageId="3" pageNumber="40">Internal claws II, III pt</th>
<td id="76C5837AFD3A1EA91A540A34FDEF0B60" box="[591,638,1306,1364]" gridcol="1" gridrow="10" pageId="3" pageNumber="40">
9.5
<emphasis id="B9503456FD3AE1551A540A13FDEF0B60" box="[591,638,1341,1364]" italics="true" pageId="3" pageNumber="40">29.3</emphasis>
</td>
<td id="76C5837AFD3A1EA91AE10A34FCB80B60" box="[762,809,1306,1364]" gridcol="2" gridrow="10" pageId="3" pageNumber="40">
5.9
<emphasis id="B9503456FD3AE1551AE10A13FCB80B60" box="[762,809,1341,1364]" italics="true" pageId="3" pageNumber="40">28.4</emphasis>
</td>
<td id="76C5837AFD3A1EA91BBF0A34FC450B60" box="[932,980,1306,1364]" gridcol="3" gridrow="10" pageId="3" pageNumber="40">
9.6
<emphasis id="B9503456FD3AE1551BBF0A13FC420B60" box="[932,979,1341,1364]" italics="true" pageId="3" pageNumber="40">27.5</emphasis>
</td>
<td id="76C5837AFD3A1EA91C270A34FBFA0B60" box="[1084,1131,1306,1364]" gridcol="4" gridrow="10" pageId="3" pageNumber="40">
8.0
<emphasis id="B9503456FD3AE1551C270A13FBFA0B60" box="[1084,1131,1341,1364]" italics="true" pageId="3" pageNumber="40">21.5</emphasis>
</td>
<td id="76C5837AFD3A1EA91CC70A34FAF50B60" box="[1244,1380,1306,1364]" gridcol="5" gridrow="10" pageId="3" pageNumber="40">??</td>
</tr>
<tr id="3514EA06FD3A1EA918840A46FAF50B95" box="[159,1380,1384,1441]" gridrow="11" pageId="3" pageNumber="40">
<th id="76C5837AFD3A1EA918840A46FE5F0B95" box="[159,462,1384,1441]" gridcol="0" gridrow="11" pageId="3" pageNumber="40">Anterior claws IV pt</th>
<td id="76C5837AFD3A1EA91A540A46FDEF0B95" box="[591,638,1384,1441]" gridcol="1" gridrow="11" pageId="3" pageNumber="40">??</td>
<td id="76C5837AFD3A1EA91AE10A46FCB80B95" box="[762,809,1384,1441]" gridcol="2" gridrow="11" pageId="3" pageNumber="40">
6.3
<emphasis id="B9503456FD3AE1551AE10AA4FCB80B95" box="[762,809,1418,1441]" italics="true" pageId="3" pageNumber="40">30.3</emphasis>
</td>
<td id="76C5837AFD3A1EA91BBF0A46FC450B95" box="[932,980,1384,1441]" gridcol="3" gridrow="11" pageId="3" pageNumber="40">
11.0
<emphasis id="B9503456FD3AE1551BBF0AA4FC420B95" box="[932,979,1418,1441]" italics="true" pageId="3" pageNumber="40">31.5</emphasis>
</td>
<td id="76C5837AFD3A1EA91C270A46FBFA0B95" box="[1084,1131,1384,1441]" gridcol="4" gridrow="11" pageId="3" pageNumber="40">??</td>
<td id="76C5837AFD3A1EA91CC70A46FAF50B95" box="[1244,1380,1384,1441]" gridcol="5" gridrow="11" pageId="3" pageNumber="40">??</td>
</tr>
<tr id="3514EA06FD3A1EA918840A9BFAF50BDB" box="[159,1380,1461,1519]" gridrow="12" pageId="3" pageNumber="40">
<th id="76C5837AFD3A1EA918840A9BFE5F0BDB" box="[159,462,1461,1519]" gridcol="0" gridrow="12" pageId="3" pageNumber="40">Posterior claws IV pt</th>
<td id="76C5837AFD3A1EA91A540A9BFDEF0BDB" box="[591,638,1461,1519]" gridcol="1" gridrow="12" pageId="3" pageNumber="40">
10.9
<emphasis id="B9503456FD3AE1551A540AF6FDEF0BDB" box="[591,638,1496,1519]" italics="true" pageId="3" pageNumber="40">33.6</emphasis>
</td>
<td id="76C5837AFD3A1EA91AE10A9BFCB80BDB" box="[762,809,1461,1519]" gridcol="2" gridrow="12" pageId="3" pageNumber="40">??</td>
<td id="76C5837AFD3A1EA91BBF0A9BFC450BDB" box="[932,980,1461,1519]" gridcol="3" gridrow="12" pageId="3" pageNumber="40">??</td>
<td id="76C5837AFD3A1EA91C270A9BFBFA0BDB" box="[1084,1131,1461,1519]" gridcol="4" gridrow="12" pageId="3" pageNumber="40">??</td>
<td id="76C5837AFD3A1EA91CC70A9BFAF50BDB" box="[1244,1380,1461,1519]" gridcol="5" gridrow="12" pageId="3" pageNumber="40">??</td>
</tr>
</table>
</paragraph>
<paragraph id="8B9BE844FD3AE15518DE096EFCFE08E6" blockId="3.[151,1436,1599,1746]" pageId="3" pageNumber="40">
<taxonomicName id="4C2493C7FD3AE15518DE096EFEDD086D" box="[197,332,1600,1626]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="40" phylum="Tardigrada" rank="species" species="almadai" status="sp. nov.">
<emphasis id="B9503456FD3AE15518DE096EFF4C086D" box="[197,221,1600,1625]" italics="true" pageId="3" pageNumber="40">M</emphasis>
.
<emphasis id="B9503456FD3AE15518F7096EFEDD086D" box="[236,332,1600,1625]" italics="true" pageId="3" pageNumber="40">almadai</emphasis>
</taxonomicName>
<emphasis id="B9503456FD3AE15519480911FE21086D" bold="true" box="[339,432,1599,1625]" pageId="3" pageNumber="40">
<taxonomicNameLabel id="A263892DFD3AE15519480911FE21086D" box="[339,432,1599,1625]" pageId="3" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C2493C7FD3AE1551A51096EFD5E086D" box="[586,719,1600,1625]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="40" phylum="Tardigrada" rank="species" species="sandrae">
<emphasis id="B9503456FD3AE1551A51096EFD5E086D" box="[586,719,1600,1625]" italics="true" pageId="3" pageNumber="40">M. sandrae</emphasis>
</taxonomicName>
in having narrower buccal tube (
<tableCitation id="C6A6DDFFFD3AE1551C4B096EFB0A086E" box="[1104,1179,1600,1626]" captionStart="TABLE 1" captionStartId="3.[151,239,391,415]" captionTargetBox="[159,1380,637,1519]" captionTargetPageId="3" captionText="TABLE 1. Measurements (in µm) of some structures of the holotype, the smallest and the largest measured specimens of Macrobiotus almadai sp. nov., of a paratype of Macrobiotus sandrae and of a specimen of Macrobiotus hufelandi collected in the locus typicus." httpUri="http://table.plazi.org/id/DF5BB8CCFD3AE155188C0EA9FE230FD0" pageId="3" pageNumber="40" tableUuid="DF5BB8CCFD3AE155188C0EA9FE230FD0">Tab. 1</tableCitation>
;
<figureCitation id="131FF4C1FD3AE1551CB2096EFA81086E" box="[1193,1296,1600,1626]" captionStart="FIGURE 1. A G" captionStartId="2.[151,255,1339,1363]" captionTargetBox="[268,1285,318,1338]" captionTargetId="figure@2.[255,1333,261,1373]" captionTargetPageId="2" captionText="FIGURE 1. A G, Macrobiotus almadai sp. nov. (holotype): A, bucco-pharyngeal apparatus; B and C, claws of the third pair of legs; D, claws of the hind legs; E, egg; F and G, details of the egg shell (arrow indicates the small teeth of the terminal disc). (Scale bar = 10 µm)." httpUri="https://zenodo.org/record/182263/files/figure.png" pageId="3" pageNumber="40">Figs. 1A</figureCitation>
,
<figureCitation id="131FF4C1FD3AE1551D07096EFABD086E" box="[1308,1324,1600,1626]" captionStart="FIGURE 2. A and B" captionStartId="4.[151,255,1191,1215]" captionTargetBox="[351,1233,318,1190]" captionTargetId="figure@4.[330,1258,277,1210]" captionTargetPageId="4" captionText="FIGURE 2. A and B, Macrobiotus hufelandi: A, bucco-pharyngeal apparatus; B, detail of the egg (Scale bar = 10 µm). C and D, Macrobiotus sandrae (paratype): C, bucco-pharyngeal apparatus; D, detail of the egg (Scale bar = 10 µm)." httpUri="https://zenodo.org/record/182264/files/figure.png" pageId="3" pageNumber="40">2</figureCitation>
C); bands of teeth in buccal cavity not visible; slightly shorter macroplacoids (
<tableCitation id="C6A6DDFFFD3AE1551BCE0946FBB208B6" box="[981,1059,1640,1666]" captionStart="TABLE 1" captionStartId="3.[151,239,391,415]" captionTargetBox="[159,1380,637,1519]" captionTargetPageId="3" captionText="TABLE 1. Measurements (in µm) of some structures of the holotype, the smallest and the largest measured specimens of Macrobiotus almadai sp. nov., of a paratype of Macrobiotus sandrae and of a specimen of Macrobiotus hufelandi collected in the locus typicus." httpUri="http://table.plazi.org/id/DF5BB8CCFD3AE155188C0EA9FE230FD0" pageId="3" pageNumber="40" tableUuid="DF5BB8CCFD3AE155188C0EA9FE230FD0">Tab. 1</tableCitation>
;
<figureCitation id="131FF4C1FD3AE1551C2D0946FB3108B6" box="[1078,1184,1640,1666]" captionStart="FIGURE 1. A G" captionStartId="2.[151,255,1339,1363]" captionTargetBox="[268,1285,318,1338]" captionTargetId="figure@2.[255,1333,261,1373]" captionTargetPageId="2" captionText="FIGURE 1. A G, Macrobiotus almadai sp. nov. (holotype): A, bucco-pharyngeal apparatus; B and C, claws of the third pair of legs; D, claws of the hind legs; E, egg; F and G, details of the egg shell (arrow indicates the small teeth of the terminal disc). (Scale bar = 10 µm)." httpUri="https://zenodo.org/record/182263/files/figure.png" pageId="3" pageNumber="40">Figs. 1A</figureCitation>
,
<figureCitation id="131FF4C1FD3AE1551CAB0946FB5108B6" box="[1200,1216,1640,1666]" captionStart="FIGURE 2. A and B" captionStartId="4.[151,255,1191,1215]" captionTargetBox="[351,1233,318,1190]" captionTargetId="figure@4.[330,1258,277,1210]" captionTargetPageId="4" captionText="FIGURE 2. A and B, Macrobiotus hufelandi: A, bucco-pharyngeal apparatus; B, detail of the egg (Scale bar = 10 µm). C and D, Macrobiotus sandrae (paratype): C, bucco-pharyngeal apparatus; D, detail of the egg (Scale bar = 10 µm)." httpUri="https://zenodo.org/record/182264/files/figure.png" pageId="3" pageNumber="40">2</figureCitation>
C); slightly longer claws (
<tableCitation id="C6A6DDFFFD3AE15518F009BEFEA6089E" box="[235,311,1680,1706]" captionStart="TABLE 1" captionStartId="3.[151,239,391,415]" captionTargetBox="[159,1380,637,1519]" captionTargetPageId="3" captionText="TABLE 1. Measurements (in µm) of some structures of the holotype, the smallest and the largest measured specimens of Macrobiotus almadai sp. nov., of a paratype of Macrobiotus sandrae and of a specimen of Macrobiotus hufelandi collected in the locus typicus." httpUri="http://table.plazi.org/id/DF5BB8CCFD3AE155188C0EA9FE230FD0" pageId="3" pageNumber="40" tableUuid="DF5BB8CCFD3AE155188C0EA9FE230FD0">Tab. 1</tableCitation>
); egg shell with more delicate and regular reticular design (
<figureCitation id="131FF4C1FD3AE1551C1009BEFB03089E" box="[1035,1170,1680,1706]" captionStart="FIGURE 1. A G" captionStartId="2.[151,255,1339,1363]" captionTargetBox="[268,1285,318,1338]" captionTargetId="figure@2.[255,1333,261,1373]" captionTargetPageId="2" captionText="FIGURE 1. A G, Macrobiotus almadai sp. nov. (holotype): A, bucco-pharyngeal apparatus; B and C, claws of the third pair of legs; D, claws of the hind legs; E, egg; F and G, details of the egg shell (arrow indicates the small teeth of the terminal disc). (Scale bar = 10 µm)." httpUri="https://zenodo.org/record/182263/files/figure.png" pageId="3" pageNumber="40">Figs. 1F, G</figureCitation>
,
<figureCitation id="131FF4C1FD3AE1551CBB09BEFB21089E" box="[1184,1200,1680,1706]" captionStart="FIGURE 2. A and B" captionStartId="4.[151,255,1191,1215]" captionTargetBox="[351,1233,318,1190]" captionTargetId="figure@4.[330,1258,277,1210]" captionTargetPageId="4" captionText="FIGURE 2. A and B, Macrobiotus hufelandi: A, bucco-pharyngeal apparatus; B, detail of the egg (Scale bar = 10 µm). C and D, Macrobiotus sandrae (paratype): C, bucco-pharyngeal apparatus; D, detail of the egg (Scale bar = 10 µm)." httpUri="https://zenodo.org/record/182264/files/figure.png" pageId="3" pageNumber="40">2</figureCitation>
D); terminal disc of processes with small teeth instead of jagged margin (
<figureCitation id="131FF4C1FD3AE1551B180996FCD808E6" box="[771,841,1720,1746]" captionStart="FIGURE 2. A and B" captionStartId="4.[151,255,1191,1215]" captionTargetBox="[351,1233,318,1190]" captionTargetId="figure@4.[330,1258,277,1210]" captionTargetPageId="4" captionText="FIGURE 2. A and B, Macrobiotus hufelandi: A, bucco-pharyngeal apparatus; B, detail of the egg (Scale bar = 10 µm). C and D, Macrobiotus sandrae (paratype): C, bucco-pharyngeal apparatus; D, detail of the egg (Scale bar = 10 µm)." httpUri="https://zenodo.org/record/182264/files/figure.png" pageId="3" pageNumber="40">Fig. 2</figureCitation>
D).
</paragraph>
</subSubSection>
</treatment>
</document>