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<mods:title id="0A0943BABB56A0A9AEAB38A1A0A33BEA">The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera</mods:title>
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<mods:namePart id="3ACA9C9207EB38ED749699D4F52E57AE">Harzhauser, Mathias</mods:namePart>
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<mods:namePart id="8E34D6755A874ED4B49D3FA3DD5A46F7">Landau, Bernard</mods:namePart>
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<mods:namePart id="F1279C19CFC5DB9D481D2EC0F69C0A88">Janssen, Ronald</mods:namePart>
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<subSubSection id="C327654CFFA5FF8FFFBAFE0769CFFE06" box="[151,445,427,454]" pageId="39" pageNumber="40" type="nomenclature">
<paragraph id="8B8236C7FFA5FF8FFFBAFE0769CFFE06" blockId="39.[151,445,427,454]" box="[151,445,427,454]" pageId="39" pageNumber="40">
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<taxonomicName id="4C3D4D44FFA5FF8FFF8CFE006964FE06" ID-CoL="7NZRZ" box="[161,278,427,454]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FFA5FF8FFF8CFE006964FE06" bold="true" box="[161,278,427,454]" italics="true" pageId="39" pageNumber="40">Clavatula</emphasis>
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<taxonomicName id="4C3D4D44FFA5FF8FFE0AFE006919FE06" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1891" box="[295,363,427,454]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="juliae">
<emphasis id="B949EAD5FFA5FF8FFE0AFE006919FE06" bold="true" box="[295,363,427,454]" italics="true" pageId="39" pageNumber="40">juliae</emphasis>
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group
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We unite here two moderately large species, which are characterized by a broad conical coronate spire, a strongly constricted and strongly concave base leading to a long siphonal canal. Early teleoconch sculpture is bipartite with faint (
<emphasis id="B949EAD5FFA5FF8FFFCBFD97688DFD96" box="[230,255,572,598]" italics="true" pageId="39" pageNumber="40">C.</emphasis>
<emphasis id="B949EAD5FFA5FF8FFE3CFD97690CFD96" box="[273,382,572,598]" italics="true" pageId="39" pageNumber="40">jarzynkae</emphasis>
) or prominent (
<emphasis id="B949EAD5FFA5FF8FFD13FD976A2AFD96" box="[574,600,572,598]" italics="true" pageId="39" pageNumber="40">C.</emphasis>
<taxonomicName id="4C3D4D44FFA5FF8FFD44FD976ADBFD96" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1891" box="[617,681,572,598]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="juliae">
<emphasis id="B949EAD5FFA5FF8FFD44FD976ADBFD96" box="[617,681,572,598]" italics="true" pageId="39" pageNumber="40">juliae</emphasis>
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) opisthocline riblets on broad mid-portion. The columella is more broadly excavated than in most other clavatulids. The bipartite early teleoconch whorls distinguish this group from
<taxonomicName id="4C3D4D44FFA5FF8FFFBAFD2F694DFD5E" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[151,319,644,670]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FFA5FF8FFFBAFD2F694DFD5E" box="[151,319,644,670]" italics="true" pageId="39" pageNumber="40">Megaclavatula</emphasis>
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. The long siphonal canal and the opisthocline riblets on early teleoconch whorls allow a separation from
<taxonomicName id="4C3D4D44FFA5FF8FFFF9FD036930FD02" box="[212,322,680,706]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FFA5FF8FFFF9FD036930FD02" box="[212,322,680,706]" italics="true" pageId="39" pageNumber="40">Clavatula</emphasis>
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. The last whorl profile separates the
<taxonomicName id="4C3D4D44FFA5FF8FFDC4FD036B25FD02" box="[745,855,680,706]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FFA5FF8FFDC4FD036B25FD02" box="[745,855,680,706]" italics="true" pageId="39" pageNumber="40">Clavatula</emphasis>
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<taxonomicName id="4C3D4D44FFA5FF8FFC45FD036BDAFD02" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1891" box="[872,936,680,706]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="juliae">
<emphasis id="B949EAD5FFA5FF8FFC45FD036BDAFD02" box="[872,936,680,706]" italics="true" pageId="39" pageNumber="40">juliae</emphasis>
</taxonomicName>
group from other bipatite groups such as the
<emphasis id="B949EAD5FFA5FF8FFFBAFD676952FD26" box="[151,288,716,742]" italics="true" pageId="39" pageNumber="40">bimarginata</emphasis>
-clade and the
<taxonomicName id="4C3D4D44FFA5FF8FFEE3FD676A4EFD26" box="[462,572,716,742]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FFA5FF8FFEE3FD676A4EFD26" box="[462,572,716,742]" italics="true" pageId="39" pageNumber="40">Clavatula</emphasis>
</taxonomicName>
<taxonomicName id="4C3D4D44FFA5FF8FFD66FD676AD9FD26" baseAuthorityName="Hoernes R. &amp; Auinger" baseAuthorityYear="1891" box="[587,683,716,742]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="camillae">
<emphasis id="B949EAD5FFA5FF8FFD66FD676AD9FD26" box="[587,683,716,742]" italics="true" pageId="39" pageNumber="40">camillae</emphasis>
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group, which differ in their slender fusiform profile.
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