245 lines
32 KiB
XML
245 lines
32 KiB
XML
<document id="40F95D2E33C16C1829F201F523D1EFAD" ID-CLB-Dataset="20492" ID-DOI="10.5169/seals-787049" ID-GBIF-Dataset="ce0756e2-5184-4b74-8472-ab69fed7951c" ID-ISSN="2624-9170" ID-Zenodo-Dep="6285683" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637935326602" checkinUser="carolina" docAuthor="Huber, Charles & Schmidt, Joachim" docDate="2017" docId="03948799FFF7FFEFFF41FCD2FE06FC89" docLanguage="en" docName="ContribNatHist.36.1-85.pdf.imf" docOrigin="Contributions to Natural History 36" docStyle="DocumentStyle:D319E9290ED630111D2A11900DAFC11F.4:ContribNatHist.2011-.journal_article" docStyleId="D319E9290ED630111D2A11900DAFC11F" docStyleName="ContribNatHist.2011-.journal_article" docStyleVersion="4" docTitle="Nebria (Epinebriola) impunctata Huber & Schmidt, 2017, sp. nov." docType="treatment" docVersion="12" lastPageNumber="48" masterDocId="FFADFFE1FFDBFFC0FF9CFFE9FFB0FFA7" masterDocTitle="Notes on the Nebria subgenus Epinebriola K. DANIEL, 1904 with the description of Barbonebriola subgen. nov. and 13 new species from the Himalaya-Tibet orogen (Coleoptera, Carabidae, Nebriini)" masterLastPageNumber="85" masterPageNumber="1" pageNumber="45" updateTime="1699470525451" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="1D3F8CFAFAB0CE889C5A46990D7B7061">Notes on the Nebria subgenus Epinebriola K. DANIEL, 1904 with the description of Barbonebriola subgen. nov. and 13 new species from the Himalaya-Tibet orogen (Coleoptera, Carabidae, Nebriini)</mods:title>
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<mods:namePart id="AD710D709B55CA8BFF599CA492B0A62A">Huber, Charles</mods:namePart>
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<mods:roleTerm id="BC24D6706DE4D5D913B1D89CBD0774C5">Author</mods:roleTerm>
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<mods:namePart id="01B860A08F9C43D406318EE17662281A">Schmidt, Joachim</mods:namePart>
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<mods:typeOfResource id="B025593698898E69C323C0787C53545F">text</mods:typeOfResource>
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<mods:title id="CEF8044B4E4CB6F34D2F758E46D05C6D">Contributions to Natural History</mods:title>
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<mods:part id="1DBE4888DB4152275B68C9914928D868">
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<mods:date id="265A6CBC45ADF6C89690F0A27708F353">2017</mods:date>
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<mods:number id="3A38F10B1496FE474768CFC01AF17690">2017-07-17</mods:number>
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<mods:detail id="D31B4336BCC431C051713754FA41E790" type="volume">
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<mods:number id="243ECE5ED840E008544BBAFF1939B91D">36</mods:number>
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<mods:classification id="ED0D65EB2CAD7097D7CC68771AE2FA75">journal article</mods:classification>
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<mods:identifier id="1BDC59371B2CD66BCC971EAB79AD0064" type="CLB-Dataset">20492</mods:identifier>
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<mods:identifier id="3DFE68127C134121BF2B9491CFFD38A7" type="DOI">10.5169/seals-787049</mods:identifier>
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<mods:identifier id="AE3222BF8E7EFC38D583DD339656F854" type="ISSN">2624-9170</mods:identifier>
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<treatment id="03948799FFF7FFEFFF41FCD2FE06FC89" ID-DOI="http://doi.org/10.5281/zenodo.6315666" ID-GBIF-Taxon="193820246" ID-Zenodo-Dep="6315666" LSID="urn:lsid:plazi:treatment:03948799FFF7FFEFFF41FCD2FE06FC89" httpUri="http://treatment.plazi.org/id/03948799FFF7FFEFFF41FCD2FE06FC89" lastPageId="47" lastPageNumber="48" pageId="44" pageNumber="45">
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<subSubSection id="C3276504FFF7FFECFF41FCD2FCBDFCF3" box="[221,781,827,852]" pageId="44" pageNumber="45" type="nomenclature">
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<paragraph id="8B82368FFFF7FFECFF41FCD2FCBDFCF3" blockId="44.[221,781,827,852]" box="[221,781,827,852]" pageId="44" pageNumber="45">
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<heading id="D0CA81E3FFF7FFECFF41FCD2FCBDFCF3" box="[221,781,827,852]" centered="true" fontSize="12" level="2" pageId="44" pageNumber="45" reason="2">
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<emphasis id="B949EA9DFFF7FFECFF41FCD2FD19FCF3" bold="true" box="[221,681,827,852]" pageId="44" pageNumber="45">
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<taxonomicName id="4C3D4D0CFFF7FFECFF41FCD2FDFBFCF3" ID-CoL="8MLNC" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[221,587,827,852]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="impunctata" status="sp. nov." subGenus="Epinebriola">
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<emphasis id="B949EA9DFFF7FFECFF41FCD2FE99FCF3" bold="true" box="[221,297,827,852]" italics="true" pageId="44" pageNumber="45">Nebria</emphasis>
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(
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<emphasis id="B949EA9DFFF7FFECFEA5FCD2FE0BFCF3" bold="true" box="[313,443,827,852]" italics="true" pageId="44" pageNumber="45">Epinebriola</emphasis>
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)
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<emphasis id="B949EA9DFFF7FFECFE57FCD2FDFBFCF3" bold="true" box="[459,587,827,852]" italics="true" pageId="44" pageNumber="45">impunctata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27A57E6FFF7FFECFDCDFCD2FD19FCF3" box="[593,681,827,852]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="13062A0AFFF7FFECFD2AFCD5FCB3FCF3" box="[694,771,828,852]" captionStart="Fig" captionStartId="46.[221,249,161,179]" captionTargetBox="[463,1106,153,1196]" captionTargetId="figure-108@46.[463,1106,153,1196]" captionTargetPageId="46" captionText="Fig. 23: Nebria (Epinebriola) impunctata sp. nov. Paratype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6285778" httpUri="https://zenodo.org/record/6285778/files/figure.png" pageId="44" pageNumber="45">Fig. 23</figureCitation>
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)
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3276504FFF7FFECFF41FC6FFD3AFB93" pageId="44" pageNumber="45" type="materials_examined">
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||
<paragraph id="8B82368FFFF7FFECFF41FC6FFBF6FC63" blockId="44.[221,1107,899,1598]" pageId="44" pageNumber="45">
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<materialsCitation id="3B553CD2FFF7FFECFF41FC6FFBF6FC63" ID-GBIF-Occurrence="3497686303" collectingDate="2000-06-13" collectingDateMax="2000-06-16" collectingDateMin="2000-06-13" collectionCode="cSCHM" collectorName="Iman Ghale & Santos Tamang & Ram & Santa & Sante Gurung" country="Nepal" elevation="4400" location="Ganesh Himal" municipality="Jaisuli Kund" pageId="44" pageNumber="45" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="5486882DFFF7FFECFF41FC6FFEF5FC39" box="[221,325,902,926]" pageId="44" pageNumber="45" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D3BFD06FFF7FFECFED1FC6AFED1FC39" box="[333,353,899,926]" pageId="44" pageNumber="45" type="male">♂</specimenCount>
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:
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<collectingCountry id="F32A761FFFF7FFECFEECFC6FFE07FC39" box="[368,439,902,926]" name="Nepal" pageId="44" pageNumber="45">Nepal</collectingCountry>
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,
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<location id="8EE26054FFF7FFECFE5EFC6FFDD4FC39" LSID="urn:lsid:plazi:treatment:03948799FFF7FFEFFF41FCD2FE06FC89:8EE26054FFF7FFECFE5EFC6FFDD4FC39" box="[450,612,902,926]" country="Nepal" municipality="Jaisuli Kund" name="Ganesh Himal" pageId="44" pageNumber="45">Ganesh Himal</location>
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,
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<location id="8EE26054FFF7FFECFDF3FC6FFC80FC39" LSID="urn:lsid:plazi:treatment:03948799FFF7FFEFFF41FCD2FE06FC89:8EE26054FFF7FFECFDF3FC6FFC80FC39" box="[623,816,902,926]" country="Nepal" municipality="Jaisuli Kund" name="Jaisuli Kund env." pageId="44" pageNumber="45">Jaisuli Kund env.</location>
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,
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||
<quantity id="4CC59B6AFFF7FFECFCA6FC6FFC6EFC39" box="[826,990,902,926]" metricMagnitude="3" metricUnit="m" metricValue="4.4" metricValueMax="4.5" metricValueMin="4.3" pageId="44" pageNumber="45" unit="m" value="4400.0" valueMax="4500.0" valueMin="4300.0">
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<elevation id="0010D1BCFFF7FFECFCA6FC6FFC6EFC39" box="[826,990,902,926]" metricMagnitude="3" metricUnit="m" metricValue="4.4" metricValueMax="4.5" metricValueMin="4.3" pageId="44" pageNumber="45" unit="m" value="4400.0" valueMax="4500.0" valueMin="4300.0">4300–4500 m</elevation>
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||
</quantity>
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||
,
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||
<date id="FF83104FFFF7FFECFC75FC6FFEA8FC63" pageId="44" pageNumber="45" value="2000-06-13" valueMax="2000-06-16" valueMin="2000-06-13">
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<collectingDate id="EFC7E9A7FFF7FFECFC75FC6FFEA8FC63" pageId="44" pageNumber="45" value="2000-06-13" valueMax="2000-06-16" valueMin="2000-06-13">13.–16.6. 2000</collectingDate>
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</date>
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||
, leg.
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||
<collectorName id="26C85359FFF7FFECFEC8FC45FE67FC63" box="[340,471,940,964]" pageId="44" pageNumber="45">Iman Ghalé</collectorName>
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||
,
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||
<collectorName id="26C85359FFF7FFECFE7DFC45FD21FC63" box="[481,657,940,964]" pageId="44" pageNumber="45">Santos Tamang</collectorName>
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||
,
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||
<collectorName id="26C85359FFF7FFECFD06FC45FD60FC63" box="[666,720,940,964]" pageId="44" pageNumber="45">Ram</collectorName>
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,
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||
<collectorName id="26C85359FFF7FFECFD45FC45FCABFC63" box="[729,795,940,964]" pageId="44" pageNumber="45">Santa</collectorName>
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&
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||
<collectorName id="26C85359FFF7FFECFCA5FC45FC63FC63" box="[825,979,940,964]" pageId="44" pageNumber="45">Santé Gurung</collectorName>
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(
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||
<collectionCode id="ED2CAE4AFFF7FFECFC43FC45FB8CFC63" box="[991,1084,940,964]" pageId="44" pageNumber="45">cSCHM</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8B82368FFFF7FFECFE9FFC38FD3AFB93" blockId="44.[221,1107,899,1598]" pageId="44" pageNumber="45">
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<materialsCitation id="3B553CD2FFF7FFECFE9FFC38FC93FC4E" ID-GBIF-Occurrence="3497686336" box="[259,803,974,1001]" collectionCode="NMBE, cSCHM" pageId="44" pageNumber="45" specimenCount="9" specimenCount-female="5" specimenCount-male="4" typeStatus="paratype">
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<typeStatus id="5486882DFFF7FFECFE9FFC38FEC9FC4E" box="[259,377,977,1001]" pageId="44" pageNumber="45" type="paratype">Paratypes</typeStatus>
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||
:
|
||
<specimenCount id="9D3BFD06FFF7FFECFE1FFC38FE1CFC4E" box="[387,428,974,1001]" pageId="44" pageNumber="45" type="male">4 ♂</specimenCount>
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||
,
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||
<specimenCount id="9D3BFD06FFF7FFECFE24FC38FE51FC4E" box="[440,481,974,1001]" pageId="44" pageNumber="45" type="female">5 ♀</specimenCount>
|
||
, same data (
|
||
<collectionCode id="ED2CAE4AFFF7FFECFDF3FC38FD0BFC4E" box="[623,699,977,1001]" collectionName="Switzerland, Bern, Naturhistorische Museums" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="44" pageNumber="45">NMBE</collectionCode>
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||
,
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||
<collectionCode id="ED2CAE4AFFF7FFECFD58FC38FCADFC4E" box="[708,797,977,1001]" pageId="44" pageNumber="45">cSCHM</collectionCode>
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)
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B553CD2FFF7FFECFCB0FC38FD35FB93" ID-GBIF-Occurrence="3497686327" collectingDate="2000-06-14" collectionCode="cSCHM" collectorName="Iman Ghale & Santos Tamang & Ram & Santa & Sante Gurung" country="Nepal" elevation="4100" location="Ganesh Himal" municipality="Jaisuli Kund" pageId="44" pageNumber="45" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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||
<specimenCount id="9D3BFD06FFF7FFECFCB0FC38FCE5FC4E" box="[812,853,974,1001]" pageId="44" pageNumber="45" type="male">1 ♂</specimenCount>
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,
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||
<collectingCountry id="F32A761FFFF7FFECFCFDFC38FC16FC4E" box="[865,934,977,1001]" name="Nepal" pageId="44" pageNumber="45">Nepal</collectingCountry>
|
||
,
|
||
<location id="8EE26054FFF7FFECFC2CFC38FBFEFC4E" LSID="urn:lsid:plazi:treatment:03948799FFF7FFEFFF41FCD2FE06FC89:8EE26054FFF7FFECFC2CFC38FBFEFC4E" box="[944,1102,977,1001]" country="Nepal" municipality="Jaisuli Kund" name="Ganesh Himal" pageId="44" pageNumber="45">Ganesh Himal</location>
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||
,
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||
<location id="8EE26054FFF7FFECFF41FC1FFE5BFBA9" LSID="urn:lsid:plazi:treatment:03948799FFF7FFEFFF41FCD2FE06FC89:8EE26054FFF7FFECFF41FC1FFE5BFBA9" box="[221,491,1014,1038]" country="Nepal" municipality="Jaisuli Kund" name="Jaisuli Kund base camp" pageId="44" pageNumber="45">Jaisuli Kund base camp</location>
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,
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<quantity id="4CC59B6AFFF7FFECFE69FC1FFDFEFBA9" box="[501,590,1014,1038]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="44" pageNumber="45" unit="m" value="4100.0">
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<elevation id="0010D1BCFFF7FFECFE69FC1FFDFEFBA9" box="[501,590,1014,1038]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="44" pageNumber="45" unit="m" value="4100.0">4100 m</elevation>
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||
</quantity>
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||
,
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||
<date id="FF83104FFFF7FFECFDC5FC1FFD7DFBA9" box="[601,717,1014,1038]" pageId="44" pageNumber="45" value="2000-06-14">
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||
<collectingDate id="EFC7E9A7FFF7FFECFDC5FC1FFD7DFBA9" box="[601,717,1014,1038]" pageId="44" pageNumber="45" value="2000-06-14">14.6.2000</collectingDate>
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||
</date>
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||
, leg.
|
||
<collectorName id="26C85359FFF7FFECFC91FC1FFC22FBA9" box="[781,914,1014,1038]" pageId="44" pageNumber="45">Iman Ghalé</collectorName>
|
||
,
|
||
<collectorName id="26C85359FFF7FFECFC01FC1FFBFFFBA9" box="[925,1103,1014,1038]" pageId="44" pageNumber="45">Santos Tamang</collectorName>
|
||
,
|
||
<collectorName id="26C85359FFF7FFECFF41FBF5FEA3FB93" box="[221,275,1052,1076]" pageId="44" pageNumber="45">Ram</collectorName>
|
||
,
|
||
<collectorName id="26C85359FFF7FFECFE80FBF5FEEEFB93" box="[284,350,1052,1076]" pageId="44" pageNumber="45">Santa</collectorName>
|
||
&
|
||
<collectorName id="26C85359FFF7FFECFEE0FBF5FDA6FB93" box="[380,534,1052,1076]" pageId="44" pageNumber="45">Santé Gurung</collectorName>
|
||
(
|
||
<collectionCode id="ED2CAE4AFFF7FFECFDBFFBF5FD30FB93" box="[547,640,1052,1076]" pageId="44" pageNumber="45">cSCHM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFF7FFEFFE9FFBA8FC9AFE79" lastPageId="47" lastPageNumber="48" pageId="44" pageNumber="45" type="description">
|
||
<paragraph id="8B82368FFFF7FFECFE9FFBA8FD08FB03" blockId="44.[221,1107,899,1598]" pageId="44" pageNumber="45">
|
||
Body length
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||
<quantity id="4CC59B6AFFF7FFECFE0EFBA8FDBBFBFE" box="[402,523,1089,1113]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="44" pageNumber="45" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">10–11 mm</quantity>
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||
. Colour black, mandibulae and appendages of head piceous. Basal antennomere darkened. Femura black, trochanters lightened, tibiae and knees dark brown, tarsi brown.
|
||
</paragraph>
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||
<paragraph id="8B82368FFFF7FFEDFE9FFB58FEFCFF59" blockId="44.[221,1107,899,1598]" lastBlockId="45.[184,1069,156,1598]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
|
||
Head with a shallow transverse impression behind the prominent eyes. Labrum with anterior margin straight, bearing six setae. Apical margin of clypeus straight. One supraorbital seta, rarely asymmetrically bisetose. Longitudinal supraorbital furrows in a shallow impression. Vertex impunctate, distinctly transversally wrinkled. Antennae long and slender extending to the middle of the elytra. Antennal scape elongate, a little longer than the eye's diameter, basally narrowed, cylindrical apically, with one dorsal seta. 2
|
||
<superScript id="7C489BC7FFF7FFECFC02FA67FC05FA06" attach="left" box="[926,949,1422,1441]" fontSize="8" pageId="44" pageNumber="45">nd</superScript>
|
||
antennomere with one seta ventroapically. Maxillary stipes normal, laterally with 7–8 long setae (
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||
<figureCitation id="13062A0AFFF7FFECFEB0FA35FEC5FA53" box="[300,373,1500,1524]" captionStart="Fig" captionStartId="10.[221,248,535,552]" captionTargetBox="[464,1080,503,1146]" captionTargetId="figure-230@10.[465,1116,512,1070]" captionTargetPageId="10" captionText="Fig. 2: Right maxilla of N. (Epinebriola) impunctata sp. nov. A: ventral. B: dorsal. bpm: basal palpomere; ca: cardo; ga: galea; la: lacinia; mp: maxillar palpus; st: stipes. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6285690" httpUri="https://zenodo.org/record/6285690/files/figure.png" pageId="44" pageNumber="45">Figs 2</figureCitation>
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||
,
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||
<figureCitation id="13062A0AFFF7FFECFE19FA35FE24FA53" box="[389,404,1500,1524]" captionStart="Fig" captionStartId="12.[221,250,1427,1445]" captionTargetBox="[221,1105,882,1411]" captionTargetId="figure-85@12.[221,1108,882,1411]" captionTargetPageId="12" captionText="Fig. 4: SEM micrograph of the stipes of the right maxilla of N. (Epinebriola) impunctata sp. nov. A: Detail of the right maxillary stipes (lateral part, ventral view). B: Finely acuminate tip of a mechanosensitive seta. cS = campaniform sensillum; Sm = mechanosensitive seta; SS = membrane of seta socket; Tu = sunken chaetiphorous tubercle." figureDoi="http://doi.org/10.5281/zenodo.6285698" httpUri="https://zenodo.org/record/6285698/files/figure.png" pageId="44" pageNumber="45">4</figureCitation>
|
||
). Characters only visible on SEM micrograph: Cuticle of the stipes only with a few cone-shaped campaniform sensilla but without pores, and the stipal setae long and finely acuminate). Penultimate labial palpomere trisetose. Ligula long, spiny, with two lateral setae. Mentum with bifid medial tooth. Submentum with a row of 11–15 long setae. Microreticulation of the head isodiametric.
|
||
</paragraph>
|
||
<paragraph id="8B82368FFFF6FFEDFF41FEE5FCC7FC63" blockId="45.[184,1069,156,1598]" pageId="45" pageNumber="46">
|
||
Pronotum transverse, distinctly convex, narrowed basally. Ratio length/ width of the pronotum = 1.38. Lateral margin straight in basal half, widely rounded to the anterior angles, convexly narrowed to the posterior angle, no concave sinuation in front of the posterior angles (
|
||
<figureCitation id="13062A0AFFF6FFEDFD60FE95FCD0FE33" box="[764,864,380,404]" captionStart="Fig" captionStartId="61.[184,213,1317,1335]" captionTargetBox="[185,1068,153,1276]" captionTargetId="figure-106@61.[551,733,151,716]" captionTargetPageId="61" captionText="Fig. 31: Left part of the pronotum of Epinebriola species. A: Nebria (Epinebriola) triseriata sp. nov.; B: N. (E.) rupina sp. nov.; C: N. (E.) montisanimae sp. nov.; D: N. (E.) impunctata sp. nov.; E: N. (E.) tuberculata sp. nov.; F: N. (E.) numburica sp. nov.; G: N. (E.) retingensis sp. nov.; H: N. (E.) delicata sp. nov.; I: N. (E.) incarinata sp. nov.; Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6285816" httpUri="https://zenodo.org/record/6285816/files/figure.png" pageId="45" pageNumber="46">Fig. 31D</figureCitation>
|
||
). Lateral explanation very narrow with a conspicuous groove, broadened basally. Lateral margin basally blade-like, obliquely or vertically upturned. Posterior angles acute, projected posteriorly. Anterior angles wide, rounded, weakly protruding. Basal margin bisinuate, before the posterior angle with a distinct incision towards the basal fovea. Pronotal disc weakly convex. Basal fovea deep, anterior and posterior transverse sulci deep, median line shallow. Basal fovea, anterior and posterior transverse sulci sparsely and faintly punctate or almost impunctate; the narrow lateral groove impunctate. Apical margination of the pronotum restricted to lateral one-fourths, merges with a faint ridge at the anterior angle; the ridge continues along the lateral groove to the lateral seta. Base of the pronotum not margined. Basolateral seta absent. One midlateral seta in the apical half of the pronotum, rarely with two setae; midlateral setae inserted in distinct grooves at the inside of the slightly upturned margin. Microreticulation of the pronotum isodiametric, impunctate on disc. Proepisternum smooth. Prosternal process elongate, unmargined, not flattened, rounded apically.
|
||
</paragraph>
|
||
<paragraph id="8B82368FFFF6FFEDFF41FC38FB97FA9E" blockId="45.[184,1069,156,1598]" pageId="45" pageNumber="46">Elytra short, ratio length/width of the elytra = 1.50. Elytral silhouette regularly ovoid, compact, widest in or slightly behind the middle; no subapical sinuation. Elytral apex acutely rounded. Basal margination straight, smoothly merged, or joined at an obtuse angle, with the lateral margination. Humeral carina faintly developed. Striae faint, impunctate. Striae obliterate towards the apex; striae 7 and 8 scarcely visible. Intervals rather convex, more convex on disc and towards the basal margin; interval 3 with 1–4 long setae adjoining stria 3. Scutellar seta present, rarely two setae. Microreticulation oblong. Mesepisterna smooth. Metepisterna 1.6 times as long as wide, smooth. Metacoxa with 2–4 basal and one apical setae. Hindwings present as a strap-like vestige.</paragraph>
|
||
<paragraph id="8B82368FFFF6FFEDFF41FAAFFCA4FA69" blockId="45.[184,1069,156,1598]" pageId="45" pageNumber="46">Sternum 3 medially asetose. Sterna 4–6 each with 3–5 very long posterior paramedial setae, which extend the sternal length; Anal sternum with 2–4 paramedial setae in males and in females. Sterna 4–5 with a faint longitudinal posterolateral impression of half of the sternit length.</paragraph>
|
||
<paragraph id="8B82368FFFF6FFEEFF41FA35FBB4FA23" blockId="45.[184,1069,156,1598]" lastBlockId="46.[221,1107,1313,1598]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">Legs long and slender. All tarsi dorsally glabrous. Protarsus of male with tarsomeres 1–3 distinctly broadened, ventrally with pads of adhesive setae. Ventroapical tooth of the metatarsomere 4 long, markedly projected, longer than half of the dorsal tarsomere lenght. Meso- and metatarsomeres 2–4 ventroapically with long setae. All tarsomeres 5 ventrally with two rows of 4–6 short setae. Metatarsomere 5 distinctly longer than the metatarsomeres 3+4.</paragraph>
|
||
<caption id="DF426607FFF5FFEEFF41FF48FECAFEB2" ID-DOI="http://doi.org/10.5281/zenodo.6285778" ID-Zenodo-Dep="6285778" httpUri="https://zenodo.org/record/6285778/files/figure.png" pageId="46" pageNumber="47" startId="46.[221,249,161,179]" targetBox="[463,1106,153,1196]" targetPageId="46">
|
||
<paragraph id="8B82368FFFF5FFEEFF41FF48FECAFEB2" blockId="46.[221,390,161,277]" pageId="46" pageNumber="47">
|
||
Fig. 23:
|
||
<taxonomicName id="4C3D4D0CFFF5FFEEFEBDFF48FEFEFF6C" authority="(Epinebriola)" authorityName="LATREILLE" authorityYear="1802" baseAuthorityName="Epinebriola" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
Nebria
|
||
<emphasis id="B949EA9DFFF5FFEEFF41FF50FEFEFF6C" bold="true" box="[221,334,185,203]" pageId="46" pageNumber="47">
|
||
(
|
||
<emphasis id="B949EA9DFFF5FFEEFF78FF50FEF7FF6C" bold="true" box="[228,327,185,203]" italics="true" pageId="46" pageNumber="47">Epinebriola</emphasis>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B949EA9DFFF5FFEEFF41FF38FECAFEB2" bold="true" pageId="46" pageNumber="47">
|
||
<taxonomicName id="4C3D4D0CFFF5FFEEFF41FF38FE8DFF44" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[221,317,209,227]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="impunctata" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFF5FFEEFF41FF38FE8DFF44" bold="true" box="[221,317,209,227]" italics="true" pageId="46" pageNumber="47">impunctata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFF5FFEEFEDEFF38FE36FF44" box="[322,390,209,227]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<typeStatus id="5486882DFFF5FFEEFF41FF00FE9CFF5C" box="[221,300,233,251]" pageId="46" pageNumber="47" type="paratype">Paratype</typeStatus>
|
||
. Scale bar =
|
||
<quantity id="4CC59B6AFFF5FFEEFED9FEEAFEC6FEB2" box="[325,374,259,277]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="46" pageNumber="47" unit="mm" value="5.0">5 mm</quantity>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B82368FFFF5FFEEFE9FFA78FE44F999" blockId="46.[221,1107,1313,1598]" pageId="46" pageNumber="47">
|
||
Male genitalia: Edeagus (
|
||
<figureCitation id="13062A0AFFF5FFEEFDB1FA78FD26FA0E" box="[557,662,1425,1449]" captionStart="Fig" captionStartId="38.[221,250,1317,1335]" captionTargetBox="[221,1106,153,1296]" captionTargetId="figure-70@38.[221,704,627,867]" captionTargetPageId="38" captionText="Fig. 20: Male edeagi of new Nebria species of the subgenus Epinebriola K. DANIEL, 1904. A: Nebria (Epinebriola) triseriata sp. nov.; B: N. (E.) rupina sp. nov.; C: N. (E.) montisanimae sp. nov.; D: N. (E.) impunctata sp. nov.; E: N. (E.) numburica sp. nov., with parameres; F: N. (E.) retingensis sp. nov.; G: N. (E.) incarinata sp. nov.; H: N. (E.) delicata sp. nov.; I: N. (E.) incarinata sp. nov., apex in dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6285764" httpUri="https://zenodo.org/record/6285764/files/figure.png" pageId="46" pageNumber="47">Fig. 20D</figureCitation>
|
||
): Basal part of the median lobe wide, with prominent triangular basolateral lobes. Mid-shaft thin, strongly curved at base, moderately curved and progressively narrowed to the apex. Apex short, trianglular, faintly deflected to the left. Mid-shaft of the endophallus densely packed with short setae.
|
||
</paragraph>
|
||
<paragraph id="8B82368FFFF4FFEFFF41FF75FD46FEEE" blockId="47.[183,1069,156,814]" pageId="47" pageNumber="48">
|
||
Female genitalia: Gonocoxa (
|
||
<figureCitation id="13062A0AFFF4FFEFFDA8FF75FD2EFF13" box="[564,670,156,180]" captionStart="Fig" captionStartId="20.[221,250,1430,1448]" captionTargetBox="[228,1106,153,1365]" captionTargetId="figure-16@20.[574,757,751,1260]" captionTargetPageId="20" captionText="Fig. 10: Right gonocoxae of Epinebriola species, in ventral view. A: Nebria (Epinebriola) triseriata sp. nov.; B: N. (E.) rupina sp. nov.; C: N. (E.) montisanimae sp. nov.; D: N. (E.) impunctata sp. nov.; E: N. (E.) tuberculata sp. nov.; F: N. (E.) numburica sp. nov.; G: N. (E.) numburica sp. nov. in lateral view with the distale gonocoxite 2 dorsally grooved (arrow); H: N. (E.) christinae HUBER & SCHMIDT, 2007; I: N. (E.) pseudorestias sp. nov.; J: N. (E.) retingensis sp. nov.; Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6285726" httpUri="https://zenodo.org/record/6285726/files/figure.png" pageId="47" pageNumber="48">Fig. 10D</figureCitation>
|
||
): Gonocoxite 2 stocky, triangular, only half the length of gonocoxite 1. Apex broadly rounded, dorsally arcuate, dorsally grooved. Ventral preapical insertion furrow elongate-oval, with two nematiforme setae. Gonocoxites ventrally unjointed and continuously sclerotized, dorsally separated by a membranous area.
|
||
</paragraph>
|
||
<paragraph id="8B82368FFFF4FFEFFF41FEBFFC9AFE79" blockId="47.[183,1069,156,814]" pageId="47" pageNumber="48">Body ratios: hea.w/fro.w = 1.43± 0.020 (1.40–1.47); prm.w/hea.w = 1.33± 0.031 (1.29–1.39); prm.w/prp.w = 1.61±0.048 (1.52–1.69); pra.w/prp.w = 1.14± 0.028 (1.09–1.17); ely.w/prm.w = 1.61±0.038 (1.55–1.68); prm.w/prm.l = 1.38± 0.031 (1.35–1.43); ely.l/ely.w = 1.50±0.038 (1.44–1.54).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFF4FFEFFF41FE05FBBAFDA3" box="[221,1034,492,516]" pageId="47" pageNumber="48" type="etymology">
|
||
<paragraph id="8B82368FFFF4FFEFFF41FE05FBBAFDA3" blockId="47.[183,1069,156,814]" box="[221,1034,492,516]" pageId="47" pageNumber="48">Etymology: The specific epithet refers to impunctate striae of the elytra.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFF4FFEFFF41FDF8FE08FD19" pageId="47" pageNumber="48" type="diagnosis">
|
||
<paragraph id="8B82368FFFF4FFEFFF41FDF8FE08FD19" blockId="47.[183,1069,156,814]" pageId="47" pageNumber="48">
|
||
Diagnosis: Close to
|
||
<taxonomicName id="4C3D4D0CFFF4FFEFFE21FDF8FDCEFD8E" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[445,638,529,553]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="montisanimae" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFF4FFEFFE21FDF8FDCEFD8E" box="[445,638,529,553]" italics="true" pageId="47" pageNumber="48">N. montisanimae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFF4FFEFFD1AFDF8FD6EFD8E" box="[646,734,529,553]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from the Manaslu Himal. The posterior angles of the pronotum are distinctly and acutely projecting in
|
||
<taxonomicName id="4C3D4D0CFFF4FFEFFB8FFDDFFE88FDD3" authorityName="C.Huber & J.Schmidt" authorityYear="2017" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="impunctata" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFF4FFEFFB8FFDDFFE88FDD3" italics="true" pageId="47" pageNumber="48">N. impunctata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFF4FFEFFEA3FDB5FE27FDD3" box="[319,407,604,628]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in contrast to
|
||
<taxonomicName id="4C3D4D0CFFF4FFEFFDA7FDB5FD4CFDD3" authorityName="C.Huber & J.Schmidt" authorityYear="2017" box="[571,764,604,628]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="montisanimae" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFF4FFEFFDA7FDB5FD4CFDD3" box="[571,764,604,628]" italics="true" pageId="47" pageNumber="48">N. montisanimae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFF4FFEFFC9FFDB5FCEBFDD3" box="[771,859,604,628]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
|
||
with the posterior angles not or only faintly protruding. The elytra are less elongate than in
|
||
<taxonomicName id="4C3D4D0CFFF4FFEFFB8FFD68FEE9FD19" authorityName="C.Huber & J.Schmidt" authorityYear="2017" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="montisanimae" status="sp. nov.">
|
||
<emphasis id="B949EA9DFFF4FFEFFB8FFD68FEE9FD19" italics="true" pageId="47" pageNumber="48">N. montisanimae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57E6FFF4FFEFFEFCFD4FFE08FD19" box="[352,440,678,702]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFF4FFEFFF41FD25FE64FCAE" pageId="47" pageNumber="48" type="distribution">
|
||
<paragraph id="8B82368FFFF4FFEFFF41FD25FE64FCAE" blockId="47.[183,1069,156,814]" pageId="47" pageNumber="48">
|
||
Distribution (
|
||
<figureCitation id="13062A0AFFF4FFEFFEF1FD25FE09FD43" box="[365,441,716,740]" captionStart="Fig" captionStartId="76.[221,250,160,178]" captionTargetId="figure-8@75.[184,1068,152,1645]" captionTargetPageId="75" captionText="Fig. 41: Locations with occurrences of Nebria (Epinebriola) species from the Central and Eastern Himalaya and from South Tibet." figureDoi="http://doi.org/10.5281/zenodo.6285892" httpUri="https://zenodo.org/record/6285892/files/figure.png" pageId="47" pageNumber="48">Fig. 41</figureCitation>
|
||
): Known only from the
|
||
<typeStatus id="5486882DFFF4FFEFFD20FD25FD5EFD43" box="[700,750,716,740]" pageId="47" pageNumber="48">type</typeStatus>
|
||
locality in the Ganesh Himal in western Central
|
||
<collectingCountry id="F32A761FFFF4FFEFFE16FD18FE60FCAE" box="[394,464,753,777]" name="Nepal" pageId="47" pageNumber="48">Nepal</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3276504FFF4FFEFFF41FCFFFE06FC89" box="[221,438,790,814]" pageId="47" pageNumber="48" type="biology_ecology">
|
||
<paragraph id="8B82368FFFF4FFEFFF41FCFFFE06FC89" blockId="47.[183,1069,156,814]" box="[221,438,790,814]" pageId="47" pageNumber="48">Habitat: Not noted.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |