239 lines
27 KiB
XML
239 lines
27 KiB
XML
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<mods:title id="22C5C960C2802B06BC79A2265F6DFB1C">Revision of the higher taxonomy of Neotropical freshwater crabs of the family Pseudothelphusidae, based on multigene and morphological analyses</mods:title>
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<mods:namePart id="2CCF1B68502683890090D0BD08C7FE5E">Ojeda, Juan Carlos</mods:namePart>
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<mods:namePart id="4AAE3A2E7B165AFB2E629CD51E6A7E10">Villalobos, José Luis</mods:namePart>
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<mods:namePart id="6B846D2BA8175988AFEE9F41A2E78A6E">Magalhães, Célio</mods:namePart>
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<mods:namePart id="94DEDEFD1790587B74BA150341696198">Wehrtmann, Ingo S.</mods:namePart>
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<mods:namePart id="2AFAD875A4B722F5433623F4D8101100">Mantelatto, Fernando L.</mods:namePart>
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<treatment id="03EE0A71FFFFFFBEFC81FDA1C53923D3" ID-DOI="http://doi.org/10.5281/zenodo.5638841" ID-GBIF-Taxon="190216110" ID-Zenodo-Dep="5638841" LSID="urn:lsid:plazi:treatment:03EE0A71FFFFFFBEFC81FDA1C53923D3" httpUri="http://treatment.plazi.org/id/03EE0A71FFFFFFBEFC81FDA1C53923D3" lastPageId="16" lastPageNumber="989" pageId="15" pageNumber="988" scope_family="Pseudothelphusidae" scope_superFamily="Pseudothelphusoidea">
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<subSubSection id="C35DE8ECFFFFFFA1FC81FDA1C06122EB" pageId="15" pageNumber="988" type="nomenclature">
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<paragraph id="8BF8BB67FFFFFFA1FC81FDA1C06122EB" blockId="15.[853,1416,531,586]" lastBlockId="15.[827,1443,624,676]" pageId="15" pageNumber="988">
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SUBFAMILY
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<taxonomicName id="4C47C0E4FFFFFFA1FC08FDA1C1B62263" ID-CoL="8H862" authority="RODRIGUEZ, 1982" authorityName="Rodriguez" authorityYear="1982" box="[988,1410,531,556]" class="Malacostraca" family="Pseudothelphusidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="988" phylum="Arthropoda" rank="subFamily" subFamily="Strengerianinae">
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STRENGERIANINAE
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<bibRefCitation id="EFD6C696FFFFFFA1FB6DFDA1C1B62263" author="Rodriguez G" box="[1209,1410,531,556]" pageId="15" pageNumber="988" pagination="224" refId="ref21823" refString="Rodriguez G. 1982. Les crabes d'eau douce d'Amerique. Famille des Pseudothelphusidae. Collection Faune Tropicale 22. Paris: Editions ORSTOM, 224." type="journal article" year="1982">RODRIGUEZ, 1982</bibRefCitation>
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</taxonomicName>
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, NEW RANK
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<emphasis id="B9336775FFFFFFA1FCEFFDC2C75322CA" box="[827,871,624,645]" italics="true" pageId="15" pageNumber="988">lsid</emphasis>
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||
:
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<uri id="FFD6B765FFFFFFA1FCBEFDC2C06122EB" pageId="15" pageNumber="988">
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urn:lsid:zoobank.org:act:
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<uuid id="FFE181B2FFFFFFA1FB5CFDC2C06122EB" pageId="15" pageNumber="988">C8D16D88-2E2E-4177- BBE2-CBDCD68CCBFB</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35DE8ECFFFFFFA1FCEFFD71C154267D" pageId="15" pageNumber="988" type="diagnosis">
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<paragraph id="8BF8BB67FFFFFFA1FCEFFD71C154267D" blockId="15.[827,1444,706,1587]" pageId="15" pageNumber="988">
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<emphasis id="B9336775FFFFFFA1FCEFFD71C7B622B9" italics="true" pageId="15" pageNumber="988">
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Diagnosis (modified from
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<bibRefCitation id="EFD6C696FFFFFFA1FBA5FD70C1042297" author="Rodriguez G" box="[1137,1328,706,728]" pageId="15" pageNumber="988" pagination="224" refId="ref21823" refString="Rodriguez G. 1982. Les crabes d'eau douce d'Amerique. Famille des Pseudothelphusidae. Collection Faune Tropicale 22. Paris: Editions ORSTOM, 224." type="journal article" year="1982">Rodriguez, 1982</bibRefCitation>
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;
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<bibRefCitation id="EFD6C696FFFFFFA1FA94FD71C74722B9" author="Campos MR" pageId="15" pageNumber="988" refId="ref19132" refString="Campos MR. 2005. Freshwater crabs from Colombia: a taxonomic and distributional study. Coleccion Jorge Alvarez Lleras No. 24. Bogota: Academia Colombiana de Ciencias Exactas, Fisicas y Naturales." type="book" year="2005">Campos, 2005</bibRefCitation>
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):
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</emphasis>
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Anterolateral margin of carapace often finely serrate to unarmed, lateral armature indistinct. Exopod of third maxilliped not as long as, or slightly longer than, distal border of ischium; ratio exopod/ischium length ranging from 0.58 to 1.17. Opening of branchial efferent channel with jugal and lateral angles separated, in contact or almost in contact, closing aperture orifice. Gonopod long, slightly or evidently directed cephalically, lateromesially compressed, without lateral lobes on principal axis. Apical cavity rounded, next to mesial crest (
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<taxonomicName id="4C47C0E4FFFFFFA1FC97FBA6C7962466" box="[835,930,1044,1065]" class="Malacostraca" family="Pseudothelphusidae" genus="Chaceus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="988" phylum="Arthropoda" rank="genus">
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<emphasis id="B9336775FFFFFFA1FC97FBA6C7962466" box="[835,930,1044,1065]" italics="true" pageId="15" pageNumber="988">Chaceus</emphasis>
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</taxonomicName>
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), or narrow and caudocephalically extended, surrounded by distal crests of three plates: mesial, lateral and marginal (
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<emphasis id="B9336775FFFFFFA1FC33FBE3C08C2429" box="[999,1208,1105,1126]" italics="true" pageId="15" pageNumber="988">Eidocamtophallus</emphasis>
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,
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<emphasis id="B9336775FFFFFFA1FB12FBE3C1A82429" box="[1222,1436,1105,1126]" italics="true" pageId="15" pageNumber="988">Pallangothelphusa</emphasis>
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,
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<taxonomicName id="4C47C0E4FFFFFFA1FCEFFBC2C7EC24CA" box="[827,984,1136,1157]" class="Malacostraca" family="Pseudothelphusidae" genus="Strengeriana" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="988" phylum="Arthropoda" rank="genus">
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<emphasis id="B9336775FFFFFFA1FCEFFBC2C7EC24CA" box="[827,984,1136,1157]" italics="true" pageId="15" pageNumber="988">Strengeriana</emphasis>
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</taxonomicName>
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); field of apical spines in transversal position relative to principal axis, in distal or cephalic position; aperture of spermatic channel in mesocaudal position. Mesial process well developed, directed distally, acute or digitiform (
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<taxonomicName id="4C47C0E4FFFFFFA1FBC0FB58C04424B0" box="[1044,1136,1258,1279]" class="Malacostraca" family="Pseudothelphusidae" genus="Chaceus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="988" phylum="Arthropoda" rank="genus">
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<emphasis id="B9336775FFFFFFA1FBC0FB58C04424B0" box="[1044,1136,1258,1279]" italics="true" pageId="15" pageNumber="988">Chaceus</emphasis>
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</taxonomicName>
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), or expanded and projected cephalically forming triangular or a complex dentate projection (
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<taxonomicName id="4C47C0E4FFFFFFA1FC6CFA9AC07E2572" box="[952,1098,1320,1341]" class="Malacostraca" family="Pseudothelphusidae" genus="Strengeriana" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="988" phylum="Arthropoda" rank="genus">
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<emphasis id="B9336775FFFFFFA1FC6CFA9AC07E2572" box="[952,1098,1320,1341]" italics="true" pageId="15" pageNumber="988">Strengeriana</emphasis>
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</taxonomicName>
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), or divided into two lobes and projected cephalically (
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<emphasis id="B9336775FFFFFFA1FBE1FAF4C1362514" box="[1077,1282,1350,1371]" italics="true" pageId="15" pageNumber="988">Pallangothelphusa</emphasis>
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), or as conical, acute tooth, projected cephalically (
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<emphasis id="B9336775FFFFFFA1FB1CFAD7C1A12535" box="[1224,1429,1381,1402]" italics="true" pageId="15" pageNumber="988">Eidocamtophallus</emphasis>
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). Marginal plate simple, overlapping caudal portion of mesialandlateralsurfaces,endingdistallyinapicalcavity; distal border straight or rounded (
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<emphasis id="B9336775FFFFFFA1FB1FFA73C1A92599" box="[1227,1437,1473,1494]" italics="true" pageId="15" pageNumber="988">Eidocamtophallus</emphasis>
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,
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<emphasis id="B9336775FFFFFFA1FCEFFA52C03325BA" box="[827,1031,1504,1525]" italics="true" pageId="15" pageNumber="988">Pallangothelphusa</emphasis>
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,
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<taxonomicName id="4C47C0E4FFFFFFA1FBC6FA52C09525BA" box="[1042,1185,1504,1525]" class="Malacostraca" family="Pseudothelphusidae" genus="Strengeriana" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="988" phylum="Arthropoda" rank="genus">
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<emphasis id="B9336775FFFFFFA1FBC6FA52C09525BA" box="[1042,1185,1504,1525]" italics="true" pageId="15" pageNumber="988">Strengeriana</emphasis>
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</taxonomicName>
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), or projected over apex to form one of three apical elements (
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<taxonomicName id="4C47C0E4FFFFFFA1FB36FA4CC174265C" box="[1250,1344,1534,1555]" class="Malacostraca" family="Pseudothelphusidae" genus="Chaceus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="988" phylum="Arthropoda" rank="genus">
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<emphasis id="B9336775FFFFFFA1FB36FA4CC174265C" box="[1250,1344,1534,1555]" italics="true" pageId="15" pageNumber="988">Chaceus</emphasis>
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</taxonomicName>
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). Row of long setae on proximal portion of marginal suture.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35DE8ECFFFFFFA1FCEFF9EEC0972743" pageId="15" pageNumber="988" type="reference_group">
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||
<paragraph id="8BF8BB67FFFFFFA1FCEFF9EEC104263E" blockId="15.[827,1328,1628,1649]" box="[827,1328,1628,1649]" pageId="15" pageNumber="988">
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||
<emphasis id="B9336775FFFFFFA1FCEFF9EEC78B263E" box="[827,959,1628,1649]" italics="true" pageId="15" pageNumber="988">
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||
<typeStatus id="54FC05C5FFFFFFA1FCEFF9EEC744263E" box="[827,880,1628,1649]" pageId="15" pageNumber="988">Type</typeStatus>
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genus:
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</emphasis>
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||
<taxonomicName id="4C47C0E4FFFFFFA1FC1FF9EEC11E263E" ID-CoL="7Q34Z" authority="Pretzmann, 1971" authorityName="Pretzmann" authorityYear="1971" box="[971,1322,1628,1649]" class="Malacostraca" family="Pseudothelphusidae" genus="Strengeriana" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="988" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B9336775FFFFFFA1FC1FF9EEC054263E" box="[971,1120,1628,1649]" italics="true" pageId="15" pageNumber="988">Strengeriana</emphasis>
|
||
<bibRefCitation id="EFD6C696FFFFFFA1FBB3F9EEC11E263E" author="Pretzmann G" box="[1127,1322,1628,1649]" pageId="15" pageNumber="988" refId="ref21558" refString="Pretzmann G. 1971. Fortschritte in der Klassifizierung der Pseudothelphusidae. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften," type="book" year="1971">Pretzmann, 1971</bibRefCitation>
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||
</taxonomicName>
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||
.
|
||
</paragraph>
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||
<paragraph id="8BF8BB67FFFFFFA1FCEFF929C0972743" blockId="15.[827,1444,1690,1804]" pageId="15" pageNumber="988">
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||
<emphasis id="B9336775FFFFFFA1FCEFF929C09726FF" box="[827,1187,1690,1712]" italics="true" pageId="15" pageNumber="988">G e n e r a i n c l u d e d: C h a c e u s</emphasis>
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P r e t z m a n n, 1 9 7 1,
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||
<taxonomicName id="4C47C0E4FFFFFFA1FCEFF90BC1AF2681" ID-CoL="7PCW2" authority="Rodriguez & Hobbs, 1989" authorityName="Rodriguez & Hobbs" authorityYear="1989" box="[827,1435,1721,1743]" class="Malacostraca" family="Pseudothelphusidae" genus="Eidocamptophallus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="988" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9336775FFFFFFA1FCEFF90BC00C2681" box="[827,1080,1721,1742]" italics="true" pageId="15" pageNumber="988">Eidocamptophallus</emphasis>
|
||
<bibRefCitation id="EFD6C696FFFFFFA1FB92F90BC1AF2681" author="Rodriguez G & Hobbs HH Jr." box="[1094,1435,1721,1743]" pageId="15" pageNumber="988" pagination="183 - 192" refId="ref22066" refString="Rodriguez G, Hobbs HH Jr. 1989. A new cavernicolous crab, Zilchia falcata, from Guatemala, with notes on the genera of the Potamocarcinini (Crustacea: Decapoda: Pseudothelphusidae). Bulletin du Museum national d'Histoire naturelle (Paris), serie 4, section A 1: 183 - 192." type="journal article" year="1989">Rodríguez & Hobbs, 1989</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B9336775FFFFFFA1FCEFF96AC06326A2" box="[827,1111,1752,1773]" italics="true" pageId="15" pageNumber="988">P h a l l a n g o t h e l p h u s a</emphasis>
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P r e t z m a n n, 1 9 6 5 a n d
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||
<taxonomicName id="4C47C0E4FFFFFFA1FCEFF945C0A82744" ID-CoL="7Q34Z" authority="Pretzmann, 1971" authorityName="Pretzmann" authorityYear="1971" box="[827,1180,1782,1804]" class="Malacostraca" family="Pseudothelphusidae" genus="Strengeriana" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="988" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9336775FFFFFFA1FCEFF945C7E52743" box="[827,977,1783,1804]" italics="true" pageId="15" pageNumber="988">Strengeriana</emphasis>
|
||
<bibRefCitation id="EFD6C696FFFFFFA1FC0CF945C0A82744" author="Pretzmann G" box="[984,1180,1782,1804]" pageId="15" pageNumber="988" refId="ref21558" refString="Pretzmann G. 1971. Fortschritte in der Klassifizierung der Pseudothelphusidae. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften," type="book" year="1971">Pretzmann, 1971</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35DE8ECFFFFFFBEFCEFF887C53923D3" lastPageId="16" lastPageNumber="989" pageId="15" pageNumber="988" type="discussion">
|
||
<paragraph id="8BF8BB67FFFFFFBEFCEFF887C5FA22E8" blockId="15.[827,1443,1845,1898]" lastBlockId="16.[145,761,197,924]" lastPageId="16" lastPageNumber="989" pageId="15" pageNumber="988">
|
||
<emphasis id="B9336775FFFFFFA1FCEFF887C7842705" box="[827,944,1845,1866]" italics="true" pageId="15" pageNumber="988">Remarks:</emphasis>
|
||
The gonopod
|
||
<typeStatus id="54FC05C5FFFFFFA1FBBCF887C0A92705" box="[1128,1181,1845,1866]" pageId="15" pageNumber="988">type</typeStatus>
|
||
in this subfamily is based on those of the genera
|
||
<taxonomicName id="4C47C0E4FFFFFFA1FB78F8E6C1972726" authorityName="Pretzmann" authorityYear="1965" box="[1196,1443,1876,1897]" class="Malacostraca" family="Pseudothelphusidae" genus="Phallangothelphusa" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="988" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9336775FFFFFFA1FB78F8E6C1972726" box="[1196,1443,1876,1897]" italics="true" pageId="15" pageNumber="988">Phallangothelphusa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C47C0E4FFE0FFBEFF07FF74C5BC2094" authorityName="Pretzmann" authorityYear="1971" box="[211,392,198,219]" class="Malacostraca" family="Pseudothelphusidae" genus="Strengeriana" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9336775FFE0FFBEFF07FF74C5BC2094" box="[211,392,198,219]" italics="true" pageId="16" pageNumber="989">Strengeriana</emphasis>
|
||
</taxonomicName>
|
||
. Although no samples of
|
||
<taxonomicName id="4C47C0E4FFE0FFBEFF45FF57C59520B5" authorityName="Pretzmann" authorityYear="1965" box="[145,417,228,250]" class="Malacostraca" family="Pseudothelphusidae" genus="Phallangothelphusa" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9336775FFE0FFBEFF45FF57C59520B5" box="[145,417,228,250]" italics="true" pageId="16" pageNumber="989">Phallangothelphusa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C47C0E4FFE0FFBEFE26FF57C6CC20B5" authorityName="Rodriguez & Hobbs" authorityYear="1989" box="[498,760,228,250]" class="Malacostraca" family="Pseudothelphusidae" genus="Eidocamptophallus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9336775FFE0FFBEFE26FF57C6CC20B5" box="[498,760,228,250]" italics="true" pageId="16" pageNumber="989">Eidocamptophallus</emphasis>
|
||
</taxonomicName>
|
||
could be included in our molecular analysis, their gonopod morphology (see
|
||
<bibRefCitation id="EFD6C696FFE0FFBEFE3EFE93C6C72179" author="Rodriguez G" box="[490,755,289,311]" pageId="16" pageNumber="989" pagination="224" refId="ref21823" refString="Rodriguez G. 1982. Les crabes d'eau douce d'Amerique. Famille des Pseudothelphusidae. Collection Faune Tropicale 22. Paris: Editions ORSTOM, 224." type="journal article" year="1982">Rodriguez, 1982: 43</bibRefCitation>
|
||
, fig. 18; 121, fig. 78) clearly shows that they belong to this subfamily.
|
||
<bibRefCitation id="EFD6C696FFE0FFBEFE56FEEDC6CC213A" author="Campos MR & Magalhaes C" box="[386,760,350,373]" pageId="16" pageNumber="989" pagination="571 - 579" refId="ref19188" refString="Campos MR, Magalhaes C. 2014. Colombiathelphusa, a new genus of freshwater crab from Colombia, and the first location record of Eidocamptophallus chacei (Pretzmann, 1967) (Crustacea: Decapoda: Pseudothelphusidae). Zootaxa 3860: 571 - 579." type="journal article" year="2014">Campos & Magalhães (2014)</bibRefCitation>
|
||
provisionally included
|
||
<taxonomicName id="4C47C0E4FFE0FFBEFE75FECFC6ED21DD" baseAuthorityName="Pretzmann" baseAuthorityYear="1967" box="[417,729,381,402]" class="Malacostraca" family="Pseudothelphusidae" genus="Eidocamptophallus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="species" species="chacei">
|
||
<emphasis id="B9336775FFE0FFBEFE75FECFC6ED21DD" box="[417,729,381,402]" italics="true" pageId="16" pageNumber="989">Eidocamptophallus chacei</emphasis>
|
||
</taxonomicName>
|
||
in the
|
||
<taxonomicName id="4C47C0E4FFE0FFBEFF6BFE2EC54521FD" baseAuthorityName="Campos & Magalhaes" baseAuthorityYear="2014" box="[191,369,412,434]" class="Malacostraca" family="Pseudothelphusidae" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="994" phylum="Arthropoda" rank="tribe" tribe="Hypolobocerini">Hypolobocerini</taxonomicName>
|
||
after recognizing the difficulties in assessing their affinities with this or other tribes. It is herein suggested, however, placing this genus among the
|
||
<taxonomicName id="4C47C0E4FFE0FFBEFEC5FE4AC5F82241" authorityName="Rodriguez" authorityYear="1982" box="[273,460,504,526]" pageId="16" pageNumber="996" rank="subFamily" subFamily="Strengerianinae">Strengerianinae</taxonomicName>
|
||
due to the ornamentation of the apex of the gonopod bearing subapical spines on the lateral surface, in addition to the mesial process and the marginal tooth that are similar, although less developed, to the ornamentations of the gonopod of the species of
|
||
<taxonomicName id="4C47C0E4FFE0FFBEFEE5FD23C5F322E9" authorityName="Pretzmann" authorityYear="1971" box="[305,455,657,678]" class="Malacostraca" family="Pseudothelphusidae" genus="Strengeriana" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9336775FFE0FFBEFEE5FD23C5F322E9" box="[305,455,657,678]" italics="true" pageId="16" pageNumber="989">Strengeriana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF8BB67FFE0FFBEFF7DFD02C53923D3" blockId="16.[145,761,197,924]" pageId="16" pageNumber="989">
|
||
Pending a future analysis including more species,
|
||
<taxonomicName id="4C47C0E4FFE0FFBEFF45FD7CC4C422AC" box="[145,240,718,739]" class="Malacostraca" family="Pseudothelphusidae" genus="Chaceus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9336775FFE0FFBEFF45FD7CC4C422AC" box="[145,240,718,739]" italics="true" pageId="16" pageNumber="989">Chaceus</emphasis>
|
||
</taxonomicName>
|
||
is placed in the
|
||
<taxonomicName id="4C47C0E4FFE0FFBEFE7DFD7CC65622AB" authorityName="Rodriguez" authorityYear="1982" box="[425,610,718,740]" pageId="16" pageNumber="996" rank="subFamily" subFamily="Strengerianinae">Strengerianinae</taxonomicName>
|
||
based on the morphology of the gonopod of
|
||
<taxonomicName id="4C47C0E4FFE0FFBEFE33FD5FC641234D" box="[487,629,749,770]" class="Malacostraca" family="Pseudothelphusidae" genus="Chaceus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="species" species="turikensis">
|
||
<emphasis id="B9336775FFE0FFBEFE33FD5FC641234D" box="[487,629,749,770]" italics="true" pageId="16" pageNumber="989">C. turikensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFD6C696FFE0FFBEFD50FD5FC5B1236E" author="Rodriguez G & Herrera F" pageId="16" pageNumber="989" pagination="121 - 126" refId="ref22016" refString="Rodriguez G, Herrera F. 1994. A new troglobitic crab, Chaceus turikensis, from Venezuela, and additional notes on the stygobiont crab Chaceus caecus Rodriguez and Bosque, 1990 (Decapoda: Brachyura: Pseudothelphusidae). Memoires de Biospelogie 21: 121 - 126." type="journal article" year="1994">Rodriguez & Herrera, 1994: 123</bibRefCitation>
|
||
, fig. 2A–E) and on an additional BI analysis (majority rule routine) where
|
||
<taxonomicName id="4C47C0E4FFE0FFBEFDBDFC98C6FC2370" box="[617,712,810,831]" class="Malacostraca" family="Pseudothelphusidae" genus="Chaceus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9336775FFE0FFBEFDBDFC98C6FC2370" box="[617,712,810,831]" italics="true" pageId="16" pageNumber="989">Chaceus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C47C0E4FFE0FFBEFF45FCFBC51D2311" authorityName="Pretzmann" authorityYear="1971" box="[145,297,841,862]" class="Malacostraca" family="Pseudothelphusidae" genus="Strengeriana" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="989" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9336775FFE0FFBEFF45FCFBC51D2311" box="[145,297,841,862]" italics="true" pageId="16" pageNumber="989">Strengeriana</emphasis>
|
||
</taxonomicName>
|
||
are sister genera (
|
||
<figureCitation id="137CA7E2FFE0FFBEFDD1FCFBC6712311" box="[517,581,841,863]" captionStart="Figure 2" captionStartId="12.[144,222,1306,1328]" captionTargetBox="[148,1425,197,1265]" captionTargetId="figure-214@12.[145,1428,195,1266]" captionTargetPageId="12" captionText="Figure 2. Tree with divergence estimates for nodes of the main groups of the Pseudothelphusoidea (numbers at the nodes represent millions of years before present), with 95% confidence intervals represented by the grey bars. Three fossil crabs were incorporated into the analysis: (1) Hillius youngi Bishop, 1983, from the lower Cretaceous; (2) Tanzanonautes tuerkai Feldmann et al., 2007 in the Potamonautidae, from the Paleogene of Tanzania; and (3) an undetermined Trichodactylidae from the late Middle Paleocene of Peru (Klaus et al., 2017)." figureDoi="http://doi.org/10.5281/zenodo.5638801" httpUri="https://zenodo.org/record/5638801/files/figure.png" pageId="16" pageNumber="989">Fig. 2</figureCitation>
|
||
). The members of this subfamily are distributed in
|
||
<collectingCountry id="F350FBF7FFE0FFBEFD84FCDAC6F62331" box="[592,706,872,894]" name="Colombia" pageId="16" pageNumber="989">Colombia</collectingCountry>
|
||
and
|
||
<collectingCountry id="F350FBF7FFE0FFBEFF45FC34C53D23D3" box="[145,265,902,924]" name="Venezuela" pageId="16" pageNumber="989">Venezuela</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |