223 lines
29 KiB
XML
223 lines
29 KiB
XML
<document id="05847DDF566FF79D7E9692B3B4FCD5B3" ID-DOI="10.24199/j.mmv.2024.83.01" ID-ISSN="1447-2554" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718999175370" checkinUser="felipe" docAuthor="Mah, Christopher L." docDate="2023" docId="03889522DD7CFFA4FCE8FA01FC2A8A0F" docLanguage="en" docName="MemMusVictoria.83.1-35.pdf" docOrigin="Memoirs of Museum Victoria 83" docSource="http://dx.doi.org/10.24199/j.mmv.2024.83.01" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Atheraster pheos Mah, 2023, n. sp." docType="treatment" docVersion="1" lastPageNumber="9" masterDocId="FFB1ED5ADD79FFACFFD9FFE2FF8F8D61" masterDocTitle="New Goniasteridae and in situ observations significant to deep-sea coral predation" masterLastPageNumber="35" masterPageNumber="1" pageNumber="6" updateTime="1719015880144" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="086419D283A99F912455EFE8B7682267">
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<mods:title id="58FFEC7CA0058A60C4CB1D5C071E50C2">New Goniasteridae and in situ observations significant to deep-sea coral predation</mods:title>
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<mods:name id="B26FA545600C656466DD82EC3EC0C304" type="personal">
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<mods:namePart id="577DB45DAE4EC4474C1E84AA08C35C9B">Mah, Christopher L.</mods:namePart>
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<mods:typeOfResource id="FD13FC6F6665CBF95BDCC329F4B77864">text</mods:typeOfResource>
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<mods:title id="00BB6FCF2E7996C4AE9F83250D1EC333">Memoirs of Museum Victoria</mods:title>
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<mods:part id="3BD2DAE29FDD7B51466E72D670103B93">
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<mods:date id="5D7F6F4C28EF83591C4C245190926466">2023</mods:date>
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<mods:number id="5652BA25AA56D7FB8DE6C091E9F4CB39">2024-03-12</mods:number>
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<mods:number id="45618D217F1282499DF4357C0A7196EA">83</mods:number>
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<mods:url id="E654DE4F3FA272953D89AD930C6705B4">http://dx.doi.org/10.24199/j.mmv.2024.83.01</mods:url>
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<mods:classification id="01E4CF09CC71E7BDABE88BBB7838F730">journal article</mods:classification>
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<mods:identifier id="270306D1166048E2AA1286CD6305136A" type="DOI">10.24199/j.mmv.2024.83.01</mods:identifier>
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<mods:identifier id="AA44E1E23B1D304E0C05464822837F1C" type="ISSN">1447-2554</mods:identifier>
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</mods:mods>
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<treatment id="03889522DD7CFFA4FCE8FA01FC2A8A0F" LSID="urn:lsid:plazi:treatment:03889522DD7CFFA4FCE8FA01FC2A8A0F" httpUri="http://treatment.plazi.org/id/03889522DD7CFFA4FCE8FA01FC2A8A0F" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6">
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<subSubSection id="C33B77BFDD7CFFA9FCE8FA01FA278B43" pageId="5" pageNumber="6" type="nomenclature">
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<paragraph id="8B9E2434DD7CFFA9FCE8FA01FBAE8898" blockId="5.[817,1057,1507,1529]" box="[817,1057,1507,1529]" pageId="5" pageNumber="6">
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<heading id="D0D69358DD7CFFA9FCE8FA01FBAE8898" bold="true" box="[817,1057,1507,1529]" fontSize="9" level="2" pageId="5" pageNumber="6" reason="2">
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<emphasis id="B955F826DD7CFFA9FCE8FA01FBAE8898" bold="true" box="[817,1057,1507,1529]" pageId="5" pageNumber="6">
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<taxonomicName id="4C215FB7DD7CFFA9FCE8FA01FC6C8898" ID-CoL="C6D55" authorityName="Mah" authorityYear="2023" box="[817,995,1507,1529]" class="Asteroidea" family="Goniasteridae" genus="Atheraster" kingdom="Animalia" order="Valvatida" pageId="5" pageNumber="6" phylum="Echinodermata" rank="species" species="pheos" status="sp. nov.">
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<emphasis id="B955F826DD7CFFA9FCE8FA01FC6C8898" bold="true" box="[817,995,1507,1529]" italics="true" pageId="5" pageNumber="6">Atheraster pheos</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A266455DDD7CFFA9FC30FA06FBAE8898" box="[1001,1057,1508,1529]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9E2434DD7CFFA9FC8EF9EDFA278B43" blockId="5.[817,1448,1551,1596]" box="[855,1448,1551,1570]" pageId="5" pageNumber="6">urn:lsid:zoobank.org:act:4084989D-EA1D-4137-9D8B-</paragraph>
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</subSubSection>
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<subSubSection id="C33B77BFDD7CFFA4FCE8F9CBFC2A8A0F" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6" type="description">
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<paragraph id="8B9E2434DD7CFFA9FCE8F9CBFC4B8B5D" blockId="5.[817,1448,1551,1596]" box="[817,964,1577,1596]" pageId="5" pageNumber="6">944AA7547A94</paragraph>
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<paragraph id="8B9E2434DD7CFFA9FCE8F9B1FC258B09" blockId="5.[817,938,1619,1641]" box="[817,938,1619,1641]" pageId="5" pageNumber="6">
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<figureCitation id="131A38B1DD7CFFA9FCE8F9B1FC258B09" box="[817,938,1619,1641]" captionStart="Figure 3" captionStartId="7.[147,207,1830,1849]" captionTargetBox="[219,1376,227,1813]" captionTargetId="figure-7@7.[219,1376,227,1813]" captionTargetPageId="7" captionText="Figure 3. Atheraster pheos n. sp. holotype, WAM Z110039: a, abactinal surface; b, abactinal surface detail; c, abactinal arm surface; d, marginal plate surface showing spinelets; e, actinal surface; f, actinal surface, details; g; furrow spines, adambulacral spination. Pedicellariae circled. Scale: a, e=10 mm; b=3.0 mm; c, d, f=5.0 mm; g=3.0 mm." pageId="5" pageNumber="6">Figure 3a–g</figureCitation>
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</paragraph>
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<paragraph id="8B9E2434DD7CFFA9FCE8F99DFA048BD3" blockId="5.[817,1448,1663,1714]" pageId="5" pageNumber="6">
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<emphasis id="B955F826DD7CFFA9FCE8F99DFC2F8BF5" box="[817,928,1663,1684]" italics="true" pageId="5" pageNumber="6">Etymology.</emphasis>
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The species epithet
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<taxonomicName id="4C215FB7DD7CFFA9FBBDF99DFB238BF5" authorityName="Mah" authorityYear="2023" box="[1124,1196,1663,1684]" class="Asteroidea" family="Goniasteridae" genus="Atheraster" kingdom="Animalia" order="Valvatida" pageId="5" pageNumber="6" phylum="Echinodermata" rank="species" species="phaeos">
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<emphasis id="B955F826DD7CFFA9FBBDF99DFB238BF5" box="[1124,1196,1663,1684]" italics="true" pageId="5" pageNumber="6">phaeos</emphasis>
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</taxonomicName>
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is Greek for a spiny plant alluding to the prickly spines present on the marginal plates.
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</paragraph>
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<paragraph id="8B9E2434DD7CFFA4FCE8F92BFD198CEC" blockId="5.[817,1449,1736,1903]" lastBlockId="8.[140,771,229,397]" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6">
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<emphasis id="B955F826DD7CFFA9FCE8F92BFC198BBF" box="[817,918,1737,1758]" italics="true" pageId="5" pageNumber="6">Diagnosis</emphasis>
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. Body strongly stellate (R/r=3.4), arms elongate, arm tips strongly upturned. Abactinal plates bare with no accessories. Abactinal arm plates larger than those of disk, but gradually changing proximally on each arm. Marginal plates bearing 6–10 spines, interradially with a single prominent conical spine on arm spines. Pedicellariae paddle-shaped. Nearly all actinal plates with a prominent paddle-shaped pedicellaria, pointed granules. No actinal spines. Furrow spines, 6–9, spines blunt, quadrate. Proximal adambulacral plates, adjacent to furrow spines, large paddle-shaped pedicellariae, valves rectangular and edges smooth, succeeded distally by subambulacral spines, four to each plate.
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</paragraph>
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<caption id="DF5E74BCDD7FFFAAFF55F8C4FDCD8A0F" pageId="6" pageNumber="7" startId="6.[140,200,1830,1849]" targetBox="[191,1389,227,1813]" targetPageId="6" targetType="figure">
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<paragraph id="8B9E2434DD7FFFAAFF55F8C4FDCD8A0F" blockId="6.[140,1442,1830,1902]" pageId="6" pageNumber="7">
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Figure 2.
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<taxonomicName id="4C215FB7DD7FFFAAFF3FF8C4FEF98A58" authorityName="Mah" authorityYear="2023" box="[230,374,1830,1849]" class="Asteroidea" family="Goniasteridae" genus="Atheraster" kingdom="Animalia" order="Valvatida" pageId="6" pageNumber="7" phylum="Echinodermata" rank="species" species="luma" status="sp. nov.">
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<emphasis id="B955F826DD7FFFAAFF3FF8C4FEF98A58" box="[230,374,1830,1849]" italics="true" pageId="6" pageNumber="7">Atheraster luma</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A266455DDD7FFFAAFEA6F8C5FE3F8A5B" box="[383,432,1831,1850]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
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holotype, NMV F122778: a, abactinal surface; b, abactinal surface detail; c, abactinal arm surface; d, actinal surface; e, lateral surface, superomarginal plates; f, actinal surface, details; g, furrow spines and adjacent regions. Pedicellariae circled. Scale: a, d=10 mm; c, e=2.0 mm; c, g=5.0 mm; f=3.0 mm.
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</paragraph>
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</caption>
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<caption id="DF5E74BCDD7EFFABFF4AF8C4FD128A0F" pageId="7" pageNumber="8" startId="7.[147,207,1830,1849]" targetBox="[219,1376,227,1813]" targetPageId="7" targetType="figure">
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<paragraph id="8B9E2434DD7EFFABFF4AF8C4FD128A0F" blockId="7.[147,1449,1830,1902]" pageId="7" pageNumber="8">
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Figure 3.
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<taxonomicName id="4C215FB7DD7EFFABFF33F8C4FE0E8A58" authorityName="Mah" authorityYear="2023" box="[234,385,1830,1849]" class="Asteroidea" family="Goniasteridae" genus="Atheraster" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="species" species="pheos" status="sp. nov.">
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<emphasis id="B955F826DD7EFFABFF33F8C4FE0E8A58" box="[234,385,1830,1849]" italics="true" pageId="7" pageNumber="8">Atheraster pheos</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A266455DDD7EFFABFE5FF8C5FE398A5B" box="[390,438,1831,1850]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
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holotype, WAM Z110039: a, abactinal surface; b, abactinal surface detail; c, abactinal arm surface; d, marginal plate surface showing spinelets; e, actinal surface; f, actinal surface, details; g; furrow spines, adambulacral spination. Pedicellariae circled. Scale: a, e=10 mm; b=3.0 mm; c, d, f=5.0 mm; g=3.0 mm.
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||
</paragraph>
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||
</caption>
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||
<paragraph id="8B9E2434DD71FFA4FF55FE46FDEB8C95" blockId="8.[138,772,420,996]" pageId="8" pageNumber="9">
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<emphasis id="B955F826DD71FFA4FF55FE46FF748CD8" box="[140,251,420,441]" italics="true" pageId="8" pageNumber="9">Comments.</emphasis>
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Rather than a single large conical spine, most proximal marginal plates on
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<taxonomicName id="4C215FB7DD71FFA4FE7EFE23FE738CB7" authorityName="Mah" authorityYear="2023" box="[423,508,449,470]" class="Asteroidea" family="Goniasteridae" genus="Atheraster" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="pheos">
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<emphasis id="B955F826DD71FFA4FE7EFE23FE738CB7" box="[423,508,449,470]" italics="true" pageId="8" pageNumber="9">A. pheos</emphasis>
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</taxonomicName>
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variably display 3–4 short spines, with a single spine on more distal plates.
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</paragraph>
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<paragraph id="8B9E2434DD71FFA4FF68FE19FEB48E54" blockId="8.[138,772,420,996]" pageId="8" pageNumber="9">
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||
This species very closely resembles
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<taxonomicName id="4C215FB7DD71FFA4FDC0FE1EFD088F71" authorityName="Mah" authorityYear="2006" box="[537,647,507,529]" class="Asteroidea" family="Goniasteridae" genus="Circeaster" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="arandae">
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<emphasis id="B955F826DD71FFA4FDC0FE1EFD088F71" box="[537,647,507,529]" italics="true" pageId="8" pageNumber="9">A. arandae</emphasis>
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</taxonomicName>
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, with which it shares the presence of multiple spines on the interradial marginal plates in conjunction with more pronounced single spines on the distal marginal plates. It differs in possessing fewer furrow spines (6–9 versus 12–15 on
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<taxonomicName id="4C215FB7DD71FFA4FDA5FD92FD7A8FE4" authorityName="Mah" authorityYear="2006" box="[636,757,624,645]" class="Asteroidea" family="Goniasteridae" genus="Circeaster" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="arandae">
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<emphasis id="B955F826DD71FFA4FDA5FD92FD7A8FE4" box="[636,757,624,645]" italics="true" pageId="8" pageNumber="9">A. arandae</emphasis>
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</taxonomicName>
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), possessing thorny spinelets on the abactinal arm plates (lacking in
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<taxonomicName id="4C215FB7DD71FFA4FF7CFD49FE9D8FDE" authorityName="Mah" authorityYear="2006" box="[165,274,682,704]" class="Asteroidea" family="Goniasteridae" genus="Circeaster" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="arandae">
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<emphasis id="B955F826DD71FFA4FF7CFD49FE9D8FDE" box="[165,274,682,704]" italics="true" pageId="8" pageNumber="9">A. arandae</emphasis>
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</taxonomicName>
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), more weakly stellate body shape (R/r=3.5 versus>4.0 in
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<taxonomicName id="4C215FB7DD71FFA4FF38FD2AFEDC8FBC" authorityName="Mah" authorityYear="2006" box="[225,339,712,733]" class="Asteroidea" family="Goniasteridae" genus="Circeaster" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="arandae">
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<emphasis id="B955F826DD71FFA4FF38FD2AFEDC8FBC" box="[225,339,712,733]" italics="true" pageId="8" pageNumber="9">A. arandae</emphasis>
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</taxonomicName>
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), and much broader proximal arm width.
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<taxonomicName id="4C215FB7DD71FFA4FF55FD07FEDC8F9B" baseAuthorityName="Mah" baseAuthorityYear="2006" box="[140,339,741,762]" class="Asteroidea" family="Goniasteridae" genus="Atheraster" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="arandae">
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<emphasis id="B955F826DD71FFA4FF55FD07FEDC8F9B" box="[140,339,741,762]" italics="true" pageId="8" pageNumber="9">Atheraster arandae</emphasis>
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</taxonomicName>
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from
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<collectingCountry id="F33664A4DD71FFA4FE40FD07FD9E8F9A" box="[409,529,741,763]" name="Madagascar" pageId="8" pageNumber="9">Madagascar</collectingCountry>
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and
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<collectingCountry id="F33664A4DD71FFA4FD93FD07FD678F9A" box="[586,744,741,763]" name="New Caledonia" pageId="8" pageNumber="9">New Caledonia</collectingCountry>
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is found at similar depth,
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<quantity id="4CD989D1DD71FFA4FEAEFCE0FD8E8E79" box="[375,513,770,792]" metricMagnitude="3" metricUnit="m" metricValue="1.88" metricValueMax="2.16" metricValueMin="1.6" pageId="8" pageNumber="9" unit="m" value="1880.0" valueMax="2160.0" valueMin="1600.0">1600–2160 m</quantity>
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, to
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<taxonomicName id="4C215FB7DD71FFA4FDF1FCE0FDF08E76" authorityName="Mah" authorityYear="2023" box="[552,639,770,791]" class="Asteroidea" family="Goniasteridae" genus="Atheraster" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="pheos" status="n. sp.">
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||
<emphasis id="B955F826DD71FFA4FDF1FCE0FDF08E76" box="[552,639,770,791]" italics="true" pageId="8" pageNumber="9">A. pheos</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A266455DDD71FFA4FD5CFCE1FD348E79" box="[645,699,771,792]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
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, which occurs at
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<quantity id="4CD989D1DD71FFA4FF35FCFDFEB98E54" box="[236,310,799,821]" metricMagnitude="3" metricUnit="m" metricValue="1.8519999999999999" pageId="8" pageNumber="9" unit="m" value="1852.0">1852 m</quantity>
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||
.
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</paragraph>
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<paragraph id="8B9E2434DD71FFA4FF68FCDEFDEA8E85" blockId="8.[138,772,420,996]" pageId="8" pageNumber="9">
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||
The significance of the smaller-sized interradial spination is unclear. Other deep-sea goniasterids with similar morphology have been observed in complex, acrobatic positions on octocoral prey, suggesting that interradial spines are related to arm flexure. Alternatively, spination on these plates may represent phenotypic variation relative to differing
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<typeStatus id="549A9A96DD71FFA4FE74FC2DFE6D8E85" box="[429,482,975,996]" pageId="8" pageNumber="9">types</typeStatus>
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of predators.
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</paragraph>
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<paragraph id="8B9E2434DD71FFA4FF55FC19FE2D892A" blockId="8.[140,771,1019,1099]" pageId="8" pageNumber="9">
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||
<taxonomicName id="4C215FB7DD71FFA4FF55FC19FF708971" authorityName="Mah" authorityYear="2023" box="[140,255,1019,1040]" pageId="8" pageNumber="33" rank="subFamily" status="nov. subfamily" subFamily="Occurrence">
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||
<emphasis id="B955F826DD71FFA4FF55FC19FF708971" box="[140,255,1019,1040]" italics="true" pageId="8" pageNumber="9">Occurrence</emphasis>
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||
</taxonomicName>
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||
.
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<materialsCitation id="3B492E69DD71FFA4FED3FC19FE11892A" country="Australia" elevation="1852" latitude="-20.802267" location="Gascoyne Marine Preserve" longLatPrecision="1" longitude="111.61544" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Western Australia">
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||
<collectingCountry id="F33664A4DD71FFA4FED3FC19FEE78970" box="[266,360,1019,1041]" name="Australia" pageId="8" pageNumber="9">Australia</collectingCountry>
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||
.
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||
<location id="8EFE72EFDD71FFA4FEA9FC19FDF58971" LSID="urn:lsid:plazi:treatment:03889522DD7CFFA4FCE8FA01FC2A8A0F:8EFE72EFDD71FFA4FEA9FC19FDF58971" box="[368,634,1019,1041]" country="Australia" latitude="-20.802267" longLatPrecision="1" longitude="111.61544" name="Gascoyne Marine Preserve" pageId="8" pageNumber="9" stateProvince="Western Australia">Gascoyne Marine Preserve</location>
|
||
, circa
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||
<locationDeviation id="9A2C3D16DD71FFA4FD60FC19FEE5894F" location="Low Point" pageId="8" pageNumber="9">
|
||
<quantity id="4CD989D1DD71FFA4FD60FC19FC8C8970" box="[697,771,1019,1041]" metricMagnitude="5" metricUnit="m" metricValue="2.73" pageId="8" pageNumber="9" unit="km" value="273.0">273 km</quantity>
|
||
WNW of Low Point
|
||
</locationDeviation>
|
||
,
|
||
<collectingRegion id="49E5EAD6DD71FFA4FEA3FBFBFDBA894F" box="[378,565,1048,1070]" country="Australia" name="Western Australia" pageId="8" pageNumber="9">Western Australia</collectingRegion>
|
||
.
|
||
<geoCoordinate id="EE1542F3DD71FFA4FD9DFBFAFD71894C" box="[580,766,1048,1070]" degrees="20" direction="south" minutes="48" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="08.1612" value="-20.802267">20º 48'08.1612'' S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1542F3DD71FFA4FF55FBD7FEC5892A" box="[140,330,1077,1099]" degrees="111" direction="east" minutes="36" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="55.5984" value="111.61544">111º 36' 55.5984'' E</geoCoordinate>
|
||
,
|
||
<quantity id="4CD989D1DD71FFA4FE8DFBD7FE11892A" box="[340,414,1077,1099]" metricMagnitude="3" metricUnit="m" metricValue="1.8519999999999999" pageId="8" pageNumber="9" unit="m" value="1852.0">
|
||
<elevation id="000CC307DD71FFA4FE8DFBD7FE11892A" box="[340,414,1077,1099]" metricMagnitude="3" metricUnit="m" metricValue="1.8519999999999999" pageId="8" pageNumber="9" unit="m" value="1852.0">1852 m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9E2434DD71FFA4FF55FB80FD8589D3" blockId="8.[139,772,1122,1903]" pageId="8" pageNumber="9">
|
||
<emphasis id="B955F826DD71FFA4FF55FB80FE888916" box="[140,263,1122,1143]" italics="true" pageId="8" pageNumber="9">Description.</emphasis>
|
||
Body stout, strongly stellate (R/r=3.4), arms narrow, tapering, strongly upturned. Interradial arcs weakly curved to straight. Marginal plates lateral facing.
|
||
</paragraph>
|
||
<paragraph id="8B9E2434DD71FFA4FF68FB5BFD178BC3" blockId="8.[139,772,1122,1903]" pageId="8" pageNumber="9">Abactinal surface composed of two sizes of plates, disk plates abutted, round to irregularly polygonal, distinctly smaller than those on arms. Central surface on disk plates smooth and bare with no accessories. Arm plates gradually intergrading with those on disk to become larger distally, each becoming approximately 2–3 times the overall size of plates on the disk region. Disk plates homogeneous in size, shape relative to those on arm, which are larger but also more heterogeneous in size and shape. Disk plates each with peripheral granules, quadrate in shape, 6–16 per plate, homogeneous in size, peripheral granules forming approximately 20% of the total plate diameter. Granular periphery in contact with other granules forming doubled granule around plates on the abactinal surface. Papulae single, present most abundantly around radial regions, absent from arm surface. Pedicellariae paddle-shaped, present on a minority of plates. Madreporite large, irregular in shape, mound-like, flanked by 6–7 adjacent abactinal plates.</paragraph>
|
||
<paragraph id="8B9E2434DD71FFA4FF68F948FAAE8976" blockId="8.[139,772,1122,1903]" lastBlockId="8.[809,1442,229,1754]" pageId="8" pageNumber="9">Marginal plates, 27–30 per arm side, 54–60 per interradius (arm tip to arm tip). Marginal plates individually wide, predominantly with lateral-facing superomarginals forming narrow periphery when viewed from abactinal surface; inferomarginals prominent from actinal surface but not evident from abactinal surface. Superomarginal plate series most clearly quadrate in shape, with rounded abactinal-lateral angle distalmost on arm, plates interradially jumbled, arrangement irregular, some plates appearing fragmented and smaller than adjacent plates in series. Plates develop a more ordered and consistent appearance at arm base. Superomarginal plates with a pronounced armament, 6–10 short pointed spines, most strongly expressed interradially. More distally along arms 8–10 superomarginals away from the terminus, single, short thorn-like spines present on the plate surface. Superomarginals in the intervening distance along the arm with irregular alternation between single thorn-like spines and multiple smaller spines. Single large paddle-shaped pedicellaria present on a minority of superomarginal plates. Other than spines and pedicellariae, superomarginal plate surfaces are bare and smooth. Peripheral quadrate-shaped granules, 20–60, around superomarginal and inferomarginal plates. Inferomarginal plates with more pronounced facing on actinal surface. Surficial armament similar to that on superomarginal plates, irregularly alternating between six and ten spines proximally (especially interradially), becoming a single thorn-like spine distally along arm adjacent to terminus. Paddle-like pedicellariae on a minority of inferomarginal plates, but usually one per plate, especially numerous on those interradially. Inferomarginal plates similarly irregular in morphology and distribution interradially, with plate variability extending over midway along arm distance. A minority of superomarginal and inferomarginal plates distally along arm devoid of any armament, surface smooth. Spineless plates present distally on arms. Shallow fasciolar grooves present between superomarginal and actinal plates.</paragraph>
|
||
<paragraph id="8B9E2434DD71FFA4FC96FBFCFBF4885C" blockId="8.[809,1442,229,1754]" pageId="8" pageNumber="9">Actinal regions composed of two complete series, with irregular arrangement of plates adjacent to inferomarginal plate contact, these series limited to disk with no plates extending onto arms. Individual actinal plates weakly quadrate to irregularly polygonal in shape each with pointed granules, 5–20, widely spaced, present along each plate periphery. Most plates with broad paddle-shaped pedicellariae, each with valves bearing smooth to weakly developed “teeth”, four to five along top edge. Actinal plate surface otherwise bare and smooth. Shallow fasciolar grooves present.</paragraph>
|
||
<paragraph id="8B9E2434DD71FFA4FC96FAA7FB698BBB" blockId="8.[809,1442,229,1754]" pageId="8" pageNumber="9">
|
||
Adambulacral plates quadrate in shape. Furrow spines,
|
||
<quantity id="4CD989D1DD71FFA4FAA0FAA7FCB28819" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.286" metricValueMin="1.524" pageId="8" pageNumber="9" unit="in" value="7.5" valueMax="9.0" valueMin="6.0">6–9 in</quantity>
|
||
straight to weakly convex series. Spines blunt, quadrate in cross-section. Adjacent to furrow spines, a large paddle-shaped pedicellaria with rectangular valves and smooth edges. Remainder of adambulacral plate covered by a single series of four subambulacral spines, pointed but thick and quadrate in cross-section and relatively low, thick granules with pointed tips similar to peripheral granules on actinal plates. Oral plates with approximately 15 furrow spines, similar to those on the adambulacral plates, blunt and quadrate in cross-section. A larger spine, triangular in cross-section, pointed into mouth. Remainder of oral plate with an irregular number of short blunt spines, 2–4 per plate, and large rectangular pedicellariae, 1–4, identical to those on the adambulacral plates.
|
||
</paragraph>
|
||
<paragraph id="8B9E2434DD71FFA4FCF0F912FC2A8A0F" blockId="8.[808,1442,1776,1902]" pageId="8" pageNumber="9">
|
||
<emphasis id="B955F826DD71FFA4FCF0F912FC558A62" box="[809,986,1776,1795]" italics="true" pageId="8" pageNumber="9">Material examined</emphasis>
|
||
.
|
||
<materialsCitation id="3B492E69DD71FFA4FC29F912FC2A8A0F" collectingDate="2022-12-03" collectionCode="WAM, CSIRO, RV, R" elevation="2013" latitude="-20.802267" location="Gascoyne Marine Preserve" longLatPrecision="1" longitude="111.61544" pageId="8" pageNumber="9" specimenCode="Z110039" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
|
||
<emphasis id="B955F826DD71FFA4FC29F912FBC38A62" bold="true" box="[1008,1100,1776,1795]" pageId="8" pageNumber="9">
|
||
<typeStatus id="549A9A96DD71FFA4FC29F912FBC78A62" box="[1008,1096,1776,1795]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="ED30BCF1DD71FFA4FB85F912FB078A62" box="[1116,1160,1776,1795]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="8" pageNumber="9" type="Museum">WAM</collectionCode>
|
||
<specimenCode id="DB878C4FDD71FFA4FB7DF912FB698A62" box="[1188,1254,1776,1795]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="8" pageNumber="9" type="Museum">Z110039</specimenCode>
|
||
(
|
||
<collectionCode id="ED30BCF1DD71FFA4FADFF912FAB28A62" box="[1286,1341,1776,1795]" country="Australia" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="8" pageNumber="9" type="Museum">CSIRO</collectionCode>
|
||
barcode 10051582),
|
||
<location id="8EFE72EFDD71FFA4FC4BF8E9FB048A7F" LSID="urn:lsid:plazi:treatment:03889522DD7CFFA4FCE8FA01FC2A8A0F:8EFE72EFDD71FFA4FC4BF8E9FB048A7F" box="[914,1163,1802,1822]" latitude="-20.802267" longLatPrecision="1" longitude="111.61544" name="Gascoyne Marine Preserve" pageId="8" pageNumber="9" stateProvince="Western Australia">Gascoyne Marine Preserve</location>
|
||
, circa
|
||
<locationDeviation id="9A2C3D16DD71FFA4FB13F8E9FCD48A59" location="Low Point" pageId="8" pageNumber="9">
|
||
<quantity id="4CD989D1DD71FFA4FB13F8E9FA9F8A7C" box="[1226,1296,1802,1822]" metricMagnitude="5" metricUnit="m" metricValue="2.73" pageId="8" pageNumber="9" unit="km" value="273.0">273 km</quantity>
|
||
WNW of Low Point
|
||
</locationDeviation>
|
||
,
|
||
<collectingRegion id="49E5EAD6DD71FFA4FCB8F8C4FB8B8A59" box="[865,1028,1829,1849]" country="Australia" name="Western Australia" pageId="8" pageNumber="9">Western Australia</collectingRegion>
|
||
.
|
||
<geoCoordinate id="EE1542F3DD71FFA4FBD3F8C7FB2B8A59" box="[1034,1188,1829,1848]" degrees="20" direction="south" minutes="48" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="08.1612" value="-20.802267">20º 48'08.1612'' S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1542F3DD71FFA4FB72F8C7FADC8A58" box="[1195,1363,1829,1849]" degrees="111" direction="east" minutes="36" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="55.5984" value="111.61544">111º 36'55.5984'' E</geoCoordinate>
|
||
,
|
||
<quantity id="4CD989D1DD71FFA4FA82F8C7FA128A58" box="[1371,1437,1829,1849]" metricMagnitude="3" metricUnit="m" metricValue="2.013" pageId="8" pageNumber="9" unit="m" value="2013.0">
|
||
<elevation id="000CC307DD71FFA4FA82F8C7FA128A58" box="[1371,1437,1829,1849]" metricMagnitude="3" metricUnit="m" metricValue="2.013" pageId="8" pageNumber="9" unit="m" value="2013.0">2013 m</elevation>
|
||
</quantity>
|
||
, coll.
|
||
<collectionCode id="ED30BCF1DD71FFA4FC8FF8A2FCFB8A32" box="[854,884,1856,1875]" country="Italy" name="Collection of Leptospira Strains" pageId="8" pageNumber="9">RV</collectionCode>
|
||
<taxonomicName id="4C215FB7DD71FFA4FCA5F8A3FB388A32" authority=", Gascoyne Expedition" authorityName="Gascoyne Expedition" box="[892,1207,1856,1875]" class="Actinopterygii" family="Nemichthyidae" genus="Investigator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
|
||
<emphasis id="B955F826DD71FFA4FCA5F8A3FC638A32" box="[892,1004,1857,1875]" italics="true" pageId="8" pageNumber="9">Investigator,</emphasis>
|
||
Gascoyne Expedition
|
||
</taxonomicName>
|
||
,
|
||
<date id="FF9F02F4DD71FFA4FB18F8A2FAA08A32" box="[1217,1327,1856,1875]" pageId="8" pageNumber="9" value="2022-12-03">
|
||
<collectingDate id="EFDBFB1CDD71FFA4FB18F8A2FAA08A32" box="[1217,1327,1856,1875]" pageId="8" pageNumber="9" value="2022-12-03">3 Dec 2022</collectingDate>
|
||
</date>
|
||
. 1 wet spec,
|
||
<collectionCode id="ED30BCF1DD71FFA4FCF0F8B9FCB78A0F" box="[809,824,1883,1902]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="9">R</collectionCode>
|
||
=11.6, r=3.4.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |