treatments-xml/data/FD/3F/87/FD3F87E6FFC9FFB94BE12E47FA8D1C3B.xml
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<document id="E6271237427DD5E3234CBD604B486455" ID-DOI="10.11646/zootaxa.5406.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10618162" ID-ZooBank="DA9C74DC-0A99-42F8-BE57-8797A3964BDF" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1707132132110" checkinUser="plazi" docAuthor="Sassi, Davide" docDate="2024" docId="FD3F87E6FFC9FFB94BE12E47FA8D1C3B" docLanguage="en" docName="zootaxa.5406.2.1.pdf" docOrigin="Zootaxa 5406 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5406.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Griburius tredecimpunctatus Suffr. D. Sassi 2024, comb. nov." docType="treatment" docVersion="3" lastPageNumber="226" masterDocId="0106FF9EFFDFFFA04B762B6CFFC41B3E" masterDocTitle="Revision of the Griburius posticatus (Suffrian, 1866) species group (Coleoptera: Chrysomelidae: Cryptocephalinae)" masterLastPageNumber="237" masterPageNumber="201" pageNumber="223" updateTime="1707162799537" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title id="77A97828A9D57FA6EC74E40F2BD0E01F">Revision of the Griburius posticatus (Suffrian, 1866) species group (Coleoptera: Chrysomelidae: Cryptocephalinae)</mods:title>
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<treatment id="FD3F87E6FFC9FFB94BE12E47FA8D1C3B" LSID="urn:lsid:plazi:treatment:FD3F87E6FFC9FFB94BE12E47FA8D1C3B" httpUri="http://treatment.plazi.org/id/FD3F87E6FFC9FFB94BE12E47FA8D1C3B" lastPageId="25" lastPageNumber="226" pageId="22" pageNumber="223">
<subSubSection id="3D8C657BFFC9FFB64BE12E47FC8B1E7B" box="[151,847,1323,1350]" pageId="22" pageNumber="223" type="nomenclature">
<paragraph id="752936F0FFC9FFB64BE12E47FC8B1E7B" blockId="22.[151,847,1323,1385]" box="[151,847,1323,1350]" pageId="22" pageNumber="223">
<heading id="2E61819CFFC9FFB64BE12E47FC8B1E7B" bold="true" box="[151,847,1323,1350]" fontSize="11" level="1" pageId="22" pageNumber="223" reason="1">
<emphasis id="47E2EAE2FFC9FFB64BE12E47FC8B1E7B" bold="true" box="[151,847,1323,1350]" pageId="22" pageNumber="223">
<taxonomicName id="B2964D73FFC9FFB64BE12E47FD051E7B" authority="(Suffrian, 1866)" authorityName="Suffr. D. Sassi" authorityYear="2024" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[151,705,1323,1350]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="tredecimpunctatus" status="comb. nov.">
<emphasis id="47E2EAE2FFC9FFB64BE12E47FE311E78" bold="true" box="[151,501,1323,1350]" italics="true" pageId="22" pageNumber="223">Griburius tredecimpunctatus</emphasis>
(
<bibRefCitation id="11074B01FFC9FFB649722E47FD7D1E7B" author="Suffrian, E." box="[516,697,1323,1349]" pageId="22" pageNumber="223" pagination="1 - 483" refId="ref27961" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1 - 483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="5CD15799FFC9FFB649BF2E47FC8B1E7B" box="[713,847,1323,1349]" pageId="22" pageNumber="223" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3D8C657BFFC9FFB64BE12E23FE9B1E57" box="[151,351,1359,1385]" pageId="22" pageNumber="223" type="description">
<paragraph id="752936F0FFC9FFB64BE12E23FE9B1E57" blockId="22.[151,847,1323,1385]" box="[151,351,1359,1385]" pageId="22" pageNumber="223">(Figs 8; 11e; 13b)</paragraph>
</subSubSection>
<subSubSection id="3D8C657BFFC9FFB64BE12EF8FD4B1D36" pageId="22" pageNumber="223" type="reference_group">
<paragraph id="752936F0FFC9FFB64BE12EF8FDAE1E95" blockId="22.[151,1437,1427,1544]" box="[151,618,1427,1452]" pageId="22" pageNumber="223">
<treatmentCitationGroup id="558611DEFFC9FFB64BE12EF8FDAE1E95" box="[151,618,1427,1452]" pageId="22" pageNumber="223">
<emphasis id="47E2EAE2FFC9FFB64BE12EF8FE5C1E95" box="[151,408,1428,1452]" italics="true" pageId="22" pageNumber="223">
<taxonomicName id="B2964D73FFC9FFB64BE12EF8FECB1E92" baseAuthorityName="Sassi" baseAuthorityYear="2018" box="[151,271,1428,1452]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">Metallactus</taxonomicName>
13 punctatus
</emphasis>
<treatmentCitation id="F43710E1FFC9FFB64AE82EF8FDA21E95" author="Suffrian, E." box="[414,614,1427,1452]" page="325" pageId="22" pageNumber="223" year="1866">
<bibRefCitation id="11074B01FFC9FFB64AE82EF8FDA21E95" author="Suffrian, E." box="[414,614,1427,1452]" pageId="22" pageNumber="223" pagination="1 - 483" refId="ref27961" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1 - 483." type="journal article" year="1866">Suffrian, 1866: 325</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="752936F0FFC9FFB64BE12EDEFB5D1EF4" blockId="22.[151,1437,1427,1544]" box="[151,1177,1458,1482]" pageId="22" pageNumber="223">
<treatmentCitationGroup id="558611DEFFC9FFB64BE12EDEFB5D1EF4" box="[151,1177,1458,1482]" pageId="22" pageNumber="223">
<treatmentCitation id="F43710E1FFC9FFB64BE12EDEFD661EF4" author="Clavareau, C. H." box="[151,674,1458,1482]" page="94" pageId="22" pageNumber="223" year="1913">
<taxonomicName id="B2964D73FFC9FFB64BE12EDEFD661EF4" authority="Clavareau, 1913: 94" authorityName="Clavareau" authorityPageNumber="94" authorityYear="1913" box="[151,674,1458,1482]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="tredecimpunctatus">
<emphasis id="47E2EAE2FFC9FFB64BE12EDEFE0B1EF4" box="[151,463,1458,1482]" italics="true" pageId="22" pageNumber="223">Metallactus tredecimpunctatus</emphasis>
<bibRefCitation id="11074B01FFC9FFB64AA02EDEFD661EF4" author="Clavareau, C. H." box="[470,674,1458,1482]" pageId="22" pageNumber="223" pagination="1 - 278" refId="ref26948" refString="Clavareau, C. H. (1913) Pars 53: Chrysomelidae: Megascelinae, Megalopodinae, Clytrinae, Cryptocephalinae, Chlamydinae, Lamprosominae. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. W. Junk, Berlin, pp. 1 - 278." type="book chapter" year="1913">Clavareau, 1913: 94</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(catalogue);
<treatmentCitation id="F43710E1FFC9FFB6485E2EDEFBDE1EF4" author="Blackwelder, R. E." box="[808,1050,1458,1482]" page="641" pageId="22" pageNumber="223" year="1946">
<bibRefCitation id="11074B01FFC9FFB6485E2EDEFBDE1EF4" author="Blackwelder, R. E." box="[808,1050,1458,1482]" pageId="22" pageNumber="223" pagination="551 - 763" refId="ref26838" refString="Blackwelder, R. E. (1946) Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America. Part 4. Bulletin of the United States National Museum, 185, 551 - 763. https: // doi. org / 10.5479 / si. 03629236.185.4" type="journal article" year="1946">Blackwelder, 1946: 641</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph id="752936F0FFC9FFB64BE12EBEFD4B1D36" blockId="22.[151,1437,1427,1544]" pageId="22" pageNumber="223">
<treatmentCitationGroup id="558611DEFFC9FFB64BE12EBEFBDA1ED7" box="[151,1054,1489,1513]" pageId="22" pageNumber="223">
<emphasis id="47E2EAE2FFC9FFB64BE12EBEFF6C1ED7" box="[151,168,1490,1513]" italics="true" pageId="22" pageNumber="223">=</emphasis>
<treatmentCitation id="F43710E1FFC9FFB64BDA2EBDFDB71ED7" author="Suffrian, E." box="[172,627,1489,1513]" page="95" pageId="22" pageNumber="223" year="1866">
<taxonomicName id="B2964D73FFC9FFB64BDA2EBDFDB71ED7" authority="Suffrian, 1866: 95" authorityName="Suffrian" authorityPageNumber="95" authorityYear="1866" box="[172,627,1489,1513]" class="Insecta" family="Chrysomelidae" genus="Scolochrus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="incomparabilis">
<emphasis id="47E2EAE2FFC9FFB64BDA2EBDFE7D1ED7" box="[172,441,1489,1513]" italics="true" pageId="22" pageNumber="223">Scolochrus incomparabilis</emphasis>
<bibRefCitation id="11074B01FFC9FFB64ACB2EBDFDB71ED7" author="Suffrian, E." box="[445,627,1489,1513]" pageId="22" pageNumber="223" pagination="1 - 483" refId="ref27961" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1 - 483." type="journal article" year="1866">Suffrian, 1866: 95</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="F43710E1FFC9FFB6490D2EBDFC8A1ED7" author="Burmeister, H." box="[635,846,1489,1513]" page="65" pageId="22" pageNumber="223" year="1877">
<bibRefCitation id="11074B01FFC9FFB6490D2EBDFC8A1ED7" author="Burmeister, H." box="[635,846,1489,1513]" pageId="22" pageNumber="223" pagination="52 - 71" refId="ref26928" refString="Burmeister, H. (1877) Phytophaga Argentina. Stettiner Entomologische Zeitung, 38, 52 - 71." type="journal article" year="1877">Burmeister, 1877: 65</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="F43710E1FFC9FFB648212EBDFBDA1ED7" author="Clavareau, C. H." box="[855,1054,1489,1513]" page="90" pageId="22" pageNumber="223" year="1913">
<bibRefCitation id="11074B01FFC9FFB648212EBDFBDA1ED7" author="Clavareau, C. H." box="[855,1054,1489,1513]" pageId="22" pageNumber="223" pagination="1 - 278" refId="ref26948" refString="Clavareau, C. H. (1913) Pars 53: Chrysomelidae: Megascelinae, Megalopodinae, Clytrinae, Cryptocephalinae, Chlamydinae, Lamprosominae. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. W. Junk, Berlin, pp. 1 - 278." type="book chapter" year="1913">Clavareau, 1913: 90</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
<treatmentCitationGroup id="558611DEFFC9FFB64F542EBDFD4B1D36" pageId="22" pageNumber="223">
[
<taxonomicName id="B2964D73FFC9FFB64F5C2EBDFB4A1ED7" ID-CoL="4R3T" authorityName="Haldeman" authorityYear="1849" box="[1066,1166,1489,1513]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="47E2EAE2FFC9FFB64F5C2EBDFB4A1ED7" box="[1066,1166,1489,1513]" italics="true" pageId="22" pageNumber="223">Griburius</emphasis>
</taxonomicName>
] (catalogue);
<treatmentCitation id="F43710E1FFC9FFB64E612EBDFEE91D36" author="Blackwelder, R. E." page="640" pageId="22" pageNumber="223" year="1946">
<bibRefCitation id="11074B01FFC9FFB64E612EBDFEE91D36" author="Blackwelder, R. E." pageId="22" pageNumber="223" pagination="551 - 763" refId="ref26838" refString="Blackwelder, R. E. (1946) Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America. Part 4. Bulletin of the United States National Museum, 185, 551 - 763. https: // doi. org / 10.5479 / si. 03629236.185.4" type="journal article" year="1946">Blackwelder, 1946: 640</bibRefCitation>
</treatmentCitation>
[
<taxonomicName id="B2964D73FFC9FFB64A4A2E9CFE641D36" ID-CoL="4R3T" authorityName="Haldeman" authorityYear="1849" box="[316,416,1520,1544]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="47E2EAE2FFC9FFB64A4A2E9CFE641D36" box="[316,416,1520,1544]" italics="true" pageId="22" pageNumber="223">Griburius</emphasis>
</taxonomicName>
] (catalogue).
<emphasis id="47E2EAE2FFC9FFB6495B2E83FD4B1D36" bold="true" box="[557,655,1519,1544]" pageId="22" pageNumber="223">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="3D8C657BFFC9FFB74BE12D5AFCC21A7E" lastPageId="23" lastPageNumber="224" pageId="22" pageNumber="223" type="materials_examined">
<paragraph id="752936F0FFC9FFB74BE12D5AFCAF1BEB" blockId="22.[151,1437,1590,2012]" lastBlockId="23.[151,1437,151,2023]" lastPageId="23" lastPageNumber="224" pageId="22" pageNumber="223">
<emphasis id="47E2EAE2FFC9FFB64BE12D5AFF241D6E" bold="true" box="[151,224,1590,1616]" italics="true" pageId="22" pageNumber="223">Types.</emphasis>
In the original description (
<bibRefCitation id="11074B01FFC9FFB649442D5AFD1F1D6E" author="Suffrian, E." box="[562,731,1590,1616]" pageId="22" pageNumber="223" pagination="1 - 483" refId="ref27961" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1 - 483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
) the author did not report the number of the specimens of
<taxonomicName id="B2964D73FFC9FFB64BE12D36FE1F1D4A" authorityName="Suffr. D. Sassi" authorityYear="2024" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[151,475,1626,1652]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="tredecimpunctatus">
<emphasis id="47E2EAE2FFC9FFB64BE12D36FE1F1D4A" box="[151,475,1626,1652]" italics="true" pageId="22" pageNumber="223">Griburius tredecimpunctatus</emphasis>
</taxonomicName>
available for the study, but he pointed out that they all were males. It was possible to trace down
<specimenCount id="6390FD79FFC9FFB64A6B2D12FE721DA6" box="[285,438,1662,1688]" count="4" pageId="22" pageNumber="223" type="generic" typeStatus="syntype">four syntypes</specimenCount>
in BMNH (2), in NMV (1) and in MNHUB (1). The typification was made as follows, in order to stabilize the epithet.
<typeStatus id="AA2D8852FFC9FFB649672DCEFD481D82" box="[529,652,1698,1724]" pageId="22" pageNumber="223" type="lectotype">LECTOTYPE</typeStatus>
(by present designation): (MNHUB)
<specimenCount id="6390FD79FFC9FFB64F332DCEFBAD1D82" box="[1093,1129,1698,1724]" count="1" pageId="22" pageNumber="223" type="male">1♂</specimenCount>
, pinned // “23983” [white label, printed] // “13 punctatus Suffr. San Joam [sic] del Rey Sellow” [blue label, handwritten] // “
<taxonomicName id="B2964D73FFC9FFB64E582DAAFDE11C3A" authority="(Suffrian, 1866)" authorityName="Suffr. D. Sassi" authorityYear="2024" baseAuthorityName="Suffrian" baseAuthorityYear="1866" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="tredecimpunctatus">
<emphasis id="47E2EAE2FFC9FFB64E582DAAFEA11C3A" italics="true" pageId="22" pageNumber="223">Griburius tredecimpunctatus</emphasis>
(
<bibRefCitation id="11074B01FFC9FFB64A002D86FDD91C3A" author="Suffrian, E." box="[374,541,1770,1796]" pageId="22" pageNumber="223" pagination="1 - 483" refId="ref27961" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1 - 483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
)
</taxonomicName>
(
<emphasis id="47E2EAE2FFC9FFB6494E2D86FC9C1C3A" box="[568,856,1770,1796]" italics="true" pageId="22" pageNumber="223">
<taxonomicName id="B2964D73FFC9FFB6494E2D86FD781C3A" baseAuthorityName="Sassi" baseAuthorityYear="2018" box="[568,700,1770,1796]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">Metallactus</taxonomicName>
13 punctatus
</emphasis>
)
<typeStatus id="AA2D8852FFC9FFB6481C2D86FC361C3A" box="[874,1010,1770,1796]" pageId="22" pageNumber="223" type="lectotype">LECTOTYPUS</typeStatus>
D. Sassi des.” [red label, printed] //. The label information and the presence of a single
<typeStatus id="AA2D8852FFC9FFB649A92C62FCF21C16" box="[735,822,1806,1832]" pageId="22" pageNumber="223" type="syntype">syntype</typeStatus>
for the
<typeStatus id="AA2D8852FFC9FFB648E22C62FC3F1C16" box="[916,1019,1806,1832]" pageId="22" pageNumber="223" type="lectotype">lectotype</typeStatus>
match the registration data from the old catalogue of the MNHUB (“23983. 13-punctatus Suffr.*. 1. St. João d. R. Sello.”.
<typeStatus id="AA2D8852FFC9FFB64F1D2C5EFAEF1C72" box="[1131,1323,1842,1868]" pageId="22" pageNumber="223" type="paralectotype">PARALECTOTYPES</typeStatus>
: (BMNH)
<specimenCount id="6390FD79FFC9FFB64BE12C3AFF781C4E" box="[151,188,1878,1904]" count="1" pageId="22" pageNumber="223" type="male">1♂</specimenCount>
, pinned // “
<taxonomicName id="B2964D73FFC9FFB64A4D2C3AFE041C4E" baseAuthorityName="Sassi" baseAuthorityYear="2018" box="[315,448,1878,1904]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="237" phylum="Arthropoda" rank="genus">Metallactus</taxonomicName>
13punctatus Suffr
<collectingCountry id="0D817660FFC9FFB649E02C3AFD1F1C4E" box="[662,731,1878,1904]" name="Brazil" pageId="22" pageNumber="223">Brasil</collectingCountry>
” [white label, printed] // “Type Suffrian Coll. Deyrolle” [white label, printed] //; (BMNH)
<specimenCount id="6390FD79FFC9FFB64ABB2C16FE361CAA" box="[461,498,1914,1940]" count="1" pageId="22" pageNumber="223" type="male">1♂</specimenCount>
, pinned // “a D. Suffrian determinat” [White label, printed] // “Met. 13 punctatus” [White label, handwritten] // “67-56” [White label, printed] // “123” [Blue label, handwritten] // “
<typeStatus id="AA2D8852FFC9FFB64FA22CF2FAF61C86" box="[1236,1330,1950,1976]" pageId="22" pageNumber="223" type="syntype">SYNTYPE</typeStatus>
” [White-blue label, printed] //; (NMV)
<specimenCount id="6390FD79FFC9FFB64A9C2CAEFDCA1CE2" box="[490,526,1986,2012]" count="1" pageId="22" pageNumber="223" type="male">1♂</specimenCount>
, pinned // “N. c. M.” [blue label, handwritten] // “Alte Sammlg.
<collectingCountry id="0D817660FFC9FFB64FAB2CAEFA811CE2" box="[1245,1349,1986,2012]" name="Brazil" pageId="22" pageNumber="223">Brasilien</collectingCountry>
” [white label, printed] //. All
<typeStatus id="AA2D8852FFC8FFB74A082BFBFDE41B8F" box="[382,544,151,177]" pageId="23" pageNumber="224" type="paralectotype">paralectotypes</typeStatus>
were labelled: // “
<taxonomicName id="B2964D73FFC8FFB7499D2BFBFB231B8F" authority="(Suffrian, 1866)" authorityName="Suffr. D. Sassi" authorityYear="2024" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[747,1255,151,177]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="tredecimpunctatus">
<emphasis id="47E2EAE2FFC8FFB7499D2BFBFBEA1B8F" box="[747,1070,151,177]" italics="true" pageId="23" pageNumber="224">Griburius tredecimpunctatus</emphasis>
(
<bibRefCitation id="11074B01FFC8FFB74F4D2BFBFB1A1B8F" author="Suffrian, E." box="[1083,1246,151,177]" pageId="23" pageNumber="224" pagination="1 - 483" refId="ref27961" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1 - 483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
)
</taxonomicName>
(
<emphasis id="47E2EAE2FFC8FFB74F802BFBFEC01BEA" italics="true" pageId="23" pageNumber="224">
<taxonomicName id="B2964D73FFC8FFB74F802BFBFABE1B8F" baseAuthorityName="Sassi" baseAuthorityYear="2018" box="[1270,1402,151,177]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">Metallactus</taxonomicName>
13 punctatus
</emphasis>
)
<typeStatus id="AA2D8852FFC8FFB74A622BD7FE161BEB" box="[276,466,187,213]" pageId="23" pageNumber="224" type="paralectotype">PARALECTOTYPUS</typeStatus>
D. Sassi des.” [red label, printed] //.
</paragraph>
<paragraph id="752936F0FFC8FFB74BB12BB3FCC21A7E" blockId="23.[151,1437,151,2023]" pageId="23" pageNumber="224">
The species was described as belonging to the genus
<taxonomicName id="B2964D73FFC8FFB748522BB3FC6C1BC7" baseAuthorityName="Sassi" baseAuthorityYear="2018" box="[804,936,223,249]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="47E2EAE2FFC8FFB748522BB3FC6C1BC7" box="[804,936,223,249]" italics="true" pageId="23" pageNumber="224">Metallactus</emphasis>
</taxonomicName>
, but it clearly belongs to
<taxonomicName id="B2964D73FFC8FFB74FBB2BB3FAFF1BC7" authorityName="Haldeman" authorityYear="1849" box="[1229,1339,223,249]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="47E2EAE2FFC8FFB74FBB2BB3FAFF1BC7" box="[1229,1339,223,249]" italics="true" pageId="23" pageNumber="224">Griburius</emphasis>
</taxonomicName>
(at least until the latter genus is redefined and circumscribed through a new, more up-to-date diagnosis based on a careful examination of all the species currently attributed to it).
</paragraph>
</subSubSection>
<subSubSection id="3D8C657BFFC8FFB74BB12A25FB421EEC" pageId="23" pageNumber="224" type="diagnosis">
<paragraph id="752936F0FFC8FFB74BB12A25FCBD1F16" blockId="23.[151,1437,151,2023]" pageId="23" pageNumber="224">
Regarding
<taxonomicName id="B2964D73FFC8FFB74A332A25FDD71A5D" authorityName="Suffrian" authorityYear="1866" box="[325,531,329,355]" class="Insecta" family="Chrysomelidae" genus="Scolochrus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="incomparabilis">
<emphasis id="47E2EAE2FFC8FFB74A332A25FDD71A5D" box="[325,531,329,355]" italics="true" pageId="23" pageNumber="224">G. incomparabilis</emphasis>
</taxonomicName>
, in the original description (
<bibRefCitation id="11074B01FFC8FFB7482D2A25FBC51A5D" author="Suffrian, E." box="[859,1025,329,355]" pageId="23" pageNumber="224" pagination="1 - 483" refId="ref27961" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1 - 483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
) the author again did not report the number of the available specimens, but he pointed out that they all were females. It was possible to trace down
<specimenCount id="6390FD79FFC8FFB74E182A00FF3D1A95" count="4" pageId="23" pageNumber="224" type="generic" typeStatus="syntype">four syntypes</specimenCount>
in BMNH (2) and in NMV (2). The typification was made as follows, in order to stabilize the epithet.
<typeStatus id="AA2D8852FFC8FFB74BE12AD8FED61AF0" box="[151,274,436,462]" pageId="23" pageNumber="224" type="lectotype">LECTOTYPE</typeStatus>
(by present designation): (BMNH)
<specimenCount id="6390FD79FFC8FFB749D52AD8FD021AF0" box="[675,710,436,462]" count="1" pageId="23" pageNumber="224" type="female">1♀</specimenCount>
, pinned // “
<taxonomicName id="B2964D73FFC8FFB748312ADFFB771AF3" authority="Suffr" authorityName="Suffrian" authorityYear="1866" box="[839,1203,435,461]" class="Insecta" family="Chrysomelidae" genus="Scolochrus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="incomparabilis">Scolochrus incomparabilis Suffr</taxonomicName>
<collectingCountry id="0D817660FFC8FFB74FCC2ADFFB3B1AF3" box="[1210,1279,435,461]" name="Brazil" pageId="23" pageNumber="224">Brasil</collectingCountry>
” [white label, handwritten] //“Type Suffr.coll Deyrolle” [white label, handwritten] //“Baly Coll.”[white label, printed] //“
<taxonomicName id="B2964D73FFC8FFB74E582ABBFDC1192A" authority="(Suffrian, 1866)" baseAuthorityName="Suffrian" baseAuthorityYear="1866" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="incomparabilis">
<emphasis id="47E2EAE2FFC8FFB74E582ABBFE85192A" italics="true" pageId="23" pageNumber="224">Griburius incomparabilis</emphasis>
(
<bibRefCitation id="11074B01FFC8FFB74A222A96FE39192A" author="Suffrian, E." box="[340,509,506,532]" pageId="23" pageNumber="224" pagination="1 - 483" refId="ref27961" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1 - 483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
)
</taxonomicName>
(
<taxonomicName id="B2964D73FFC8FFB7496C2A96FC89192A" authorityName="Suffrian" authorityYear="1866" box="[538,845,506,532]" class="Insecta" family="Chrysomelidae" genus="Scolochrus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="incomparabilis">
<emphasis id="47E2EAE2FFC8FFB7496C2A96FC89192A" box="[538,845,506,532]" italics="true" pageId="23" pageNumber="224">Scolochrus incomparabilis</emphasis>
</taxonomicName>
)
<typeStatus id="AA2D8852FFC8FFB748172A97FC2D192B" box="[865,1001,507,533]" pageId="23" pageNumber="224" type="lectotype">LECTOTYPUS</typeStatus>
D. Sassi des.” [red label, printed] // “
<taxonomicName id="B2964D73FFC8FFB74BD22972FD481906" authority="Suffr. D. Sassi" authorityName="Suffr. D. Sassi" authorityYear="2024" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[164,652,542,568]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="tredecimpunctatus">
<emphasis id="47E2EAE2FFC8FFB74BD22972FE221906" box="[164,486,542,568]" italics="true" pageId="23" pageNumber="224">Griburius tredecimpunctatus</emphasis>
Suffr. D. Sassi
</taxonomicName>
det. 2023” [white label, printed] //.
<typeStatus id="AA2D8852FFC8FFB74F612972FB151906" box="[1047,1233,542,568]" pageId="23" pageNumber="224" type="paralectotype">PARALECTOTYPES</typeStatus>
(3): (BMNH)
<specimenCount id="6390FD79FFC8FFB74E042972FA511906" box="[1394,1429,542,568]" count="1" pageId="23" pageNumber="224" type="female">1♀</specimenCount>
, pinned // “
<collectingCountry id="0D817660FFC8FFB74A7A292DFE951965" box="[268,337,577,603]" name="Brazil" pageId="23" pageNumber="224">Brasil</collectingCountry>
… [unreadable word]” [white label, handwritten] //
<taxonomicName id="B2964D73FFC8FFB748D4292DFB911965" authorityName="Suffrian" authorityYear="1866" box="[930,1109,577,603]" class="Insecta" family="Chrysomelidae" genus="Scolochrus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="incomparabilis">“incomparabilis</taxonomicName>
m” [blue label, handwritten] // “8” [blue label, handwritten] // “67-56” [white label, printed] // “TYPE” [white label, printed] // “E. Coll. Chevr.
<superScript id="82E39BB8FFC8FFB74BE129E4FF5819A8" attach="none" box="[151,156,648,662]" fontSize="6" pageId="23" pageNumber="224">t</superScript>
” [white label, printed] //; (NMV)
<specimenCount id="6390FD79FFC8FFB7495029E4FD8E199C" box="[550,586,648,674]" count="2" pageId="23" pageNumber="224" type="female">2♀</specimenCount>
, pinned // “
<collectingRegion id="B752F812FFC8FFB749A629E4FCBE199C" box="[720,890,648,674]" country="Brazil" name="Rio de Janeiro" pageId="23" pageNumber="224">Rio de Janeiro</collectingRegion>
” [white label, printed] //. All
<typeStatus id="AA2D8852FFC8FFB74FB929E4FAB5199C" box="[1231,1393,648,674]" pageId="23" pageNumber="224" type="paralectotype">paralectotypes</typeStatus>
are labelled: // “
<taxonomicName id="B2964D73FFC8FFB74A6929C0FD3219F8" authority="(Suffrian, 1866)" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[287,758,684,710]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="incomparabilis">
<emphasis id="47E2EAE2FFC8FFB74A6929C0FDF919F8" box="[287,573,684,710]" italics="true" pageId="23" pageNumber="224">Griburius incomparabilis</emphasis>
(
<bibRefCitation id="11074B01FFC8FFB7493C29C0FD2919F8" author="Suffrian, E." box="[586,749,684,710]" pageId="23" pageNumber="224" pagination="1 - 483" refId="ref27961" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1 - 483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
)
</taxonomicName>
(
<taxonomicName id="B2964D73FFC8FFB7487329C0FBF519F8" authorityName="Suffrian" authorityYear="1866" box="[773,1073,684,710]" class="Insecta" family="Chrysomelidae" genus="Scolochrus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="incomparabilis">
<emphasis id="47E2EAE2FFC8FFB7487329C0FBF519F8" box="[773,1073,684,710]" italics="true" pageId="23" pageNumber="224">Scolochrus incomparabilis</emphasis>
</taxonomicName>
)
<typeStatus id="AA2D8852FFC8FFB74F3629C0FB3A19F8" box="[1088,1278,684,710]" pageId="23" pageNumber="224" type="paralectotype">PARALECTOTYPUS</typeStatus>
D. Sassi des.” [red label, printed] // “
<taxonomicName id="B2964D73FFC8FFB74AE629A3FCBE19D7" authority="Suffr. D. Sassi" authorityName="Suffr. D. Sassi" authorityYear="2024" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[400,890,719,746]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="tredecimpunctatus">
<emphasis id="47E2EAE2FFC8FFB74AE629A3FD1719D7" box="[400,723,719,745]" italics="true" pageId="23" pageNumber="224">Griburius tredecimpunctatus</emphasis>
Suffr. D. Sassi
</taxonomicName>
det. 2023” [white label, printed] //. Furthermore, MNHUB holds
<specimenCount id="6390FD79FFC8FFB74A26299FFDC31833" box="[336,519,755,781]" count="3" pageId="23" pageNumber="224" type="generic">three specimens</specimenCount>
which most likely belong to the type series, in my opinion, but whose data do not correspond satisfactorily with what Suffrian pointed out in the original description. In fact, Suffrian reported specimens from the Berlin Museum collected by “Sello” (Friedrich Sellow) in Collo del Sacramento, while the specimens housed there, labelled with the catalog number “24030”, bear collection data as “
<collectingCountry id="0D817660FFC8FFB74FEF2831FB251849" box="[1177,1249,861,887]" name="Brazil" pageId="23" pageNumber="224">Brasil</collectingCountry>
, v. Olf.” In other cases, for example for the
<typeStatus id="AA2D8852FFC8FFB74ABA28EDFDEA18A5" box="[460,558,897,923]" pageId="23" pageNumber="224" type="syntype">syntypes</typeStatus>
kept in the Vienna Museum, the non-correspondence can be explained with arguments that I consider valid enough (see for example
<bibRefCitation id="11074B01FFC8FFB7486128C8FC5F1880" author="Sassi, D." box="[791,923,932,958]" pageId="23" pageNumber="224" pagination="201 - 245" refId="ref27561" refString="Sassi, D. (2019) Revision of the Metallactus hamifer species group (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa, 4657 (2), 201 - 245. https: // doi. org / 10.11646 / zootaxa. 4657.2.1" type="journal article" year="2019">Sassi, 2019</bibRefCitation>
), but in the case of MNHUB, the coincidence between the information attached to the type series of the different species described by Suffrian and what he wrote in the original descriptions, is usually very good. Therefore, it is considered consequent to exclude the Berlin material from the designation of the
<typeStatus id="AA2D8852FFC8FFB749462F62FD161F16" box="[560,722,1038,1064]" pageId="23" pageNumber="224" type="paralectotype">paralectotypes</typeStatus>
of the species.
</paragraph>
<paragraph id="752936F0FFC8FFB74BB12F5EFB421EEC" blockId="23.[151,1437,151,2023]" pageId="23" pageNumber="224">
Despite the notable difference in dorsal coloration pattern, a careful evaluation of the specimens origins leads to the conclusion that the two entities described as
<taxonomicName id="B2964D73FFC8FFB749932F39FBC31F51" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[741,1031,1109,1135]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="incomparabilis">
<emphasis id="47E2EAE2FFC8FFB749932F39FBC31F51" box="[741,1031,1109,1135]" italics="true" pageId="23" pageNumber="224">Griburius incomparabilis</emphasis>
</taxonomicName>
and
<taxonomicName id="B2964D73FFC8FFB74F372F39FA581F51" authorityName="Clavareau" authorityYear="1913" box="[1089,1436,1109,1135]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="tredecimpunctatus">
<emphasis id="47E2EAE2FFC8FFB74F372F39FA581F51" box="[1089,1436,1109,1135]" italics="true" pageId="23" pageNumber="224">Metallactus tredecimpunctatus</emphasis>
</taxonomicName>
are actually the female and the male of the same species, respectively. As the male genitalia are more informative, at least in light of the current knowledge of Cryptocephalinae in general, it is considered appropriate to give priority to the name originally attributed to the males, even though such name appears further down in the same work (
<bibRefCitation id="11074B01FFC8FFB74B962F8FFE4D1FC3" author="Suffrian, E." box="[224,393,1251,1277]" pageId="23" pageNumber="224" pagination="1 - 483" refId="ref27961" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1 - 483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
). Since both names were published simultaneously, the precedence between them cannot be objectively determined, as outlined in Article 24 of the International Code of Zoological Nomenclature (
<bibRefCitation id="11074B01FFC8FFB74E242E6BFF171E7A" author="ICZN &amp; International Commission on Zoological Nomenclature" pageId="23" pageNumber="224" refId="ref27251" refString="ICZN [International Commission on Zoological Nomenclature] (1999) International code of zoological nomenclature. Fourth Edition. International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN, 1999</bibRefCitation>
). Therefore, I consider my decision justified by the same article (24.2.2. Determination of precedence of names or acts by the First Reviser. If two or more names, different or identical, and based on the same or different
<typeStatus id="AA2D8852FFC8FFB74E2C2E22FA5C1E56" box="[1370,1432,1358,1384]" pageId="23" pageNumber="224">types</typeStatus>
, or two or more nomenclatural acts, are published on the same date in the
<emphasis id="47E2EAE2FFC8FFB748AE2E1EFBD61EB5" box="[984,1042,1394,1419]" italics="true" pageId="23" pageNumber="224">same</emphasis>
or different works, the precedence of the names or acts is fixed by the First Reviser). Consequently, the following synonymy is proposed:
<taxonomicName id="B2964D73FFC8FFB74E582EF8FDE41EEC" authority="(Suffrian, 1866)" authorityName="Suffr. D. Sassi" authorityYear="2024" baseAuthorityName="Suffrian" baseAuthorityYear="1866" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="tredecimpunctatus">
<emphasis id="47E2EAE2FFC8FFB74E582EF8FEA11EEC" italics="true" pageId="23" pageNumber="224">Griburius tredecimpunctatus</emphasis>
(
<bibRefCitation id="11074B01FFC8FFB74A052ED4FDDC1EEC" author="Suffrian, E." box="[371,536,1464,1490]" pageId="23" pageNumber="224" pagination="1 - 483" refId="ref27961" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1 - 483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
)
</taxonomicName>
=
<taxonomicName id="B2964D73FFC8FFB749482ED4FBDC1EEC" authority="(Suffrian, 1866)" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[574,1048,1464,1490]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="incomparabilis" status="syn. nov.">
<emphasis id="47E2EAE2FFC8FFB749482ED4FC991EEC" box="[574,861,1464,1490]" italics="true" pageId="23" pageNumber="224">Griburius incomparabilis</emphasis>
(
<bibRefCitation id="11074B01FFC8FFB7481D2ED4FBCB1EEC" author="Suffrian, E." box="[875,1039,1464,1490]" pageId="23" pageNumber="224" pagination="1 - 483" refId="ref27961" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1 - 483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
)
</taxonomicName>
<emphasis id="47E2EAE2FFC8FFB74F692ED4FB421EEC" bold="true" box="[1055,1158,1464,1490]" pageId="23" pageNumber="224">
<taxonomicNameLabel id="5CD15799FFC8FFB74F692ED4FB421EEC" box="[1055,1158,1464,1490]" pageId="23" pageNumber="224">syn. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="3D8C657BFFC8FFB74BB12EB0FCCF1C0B" pageId="23" pageNumber="224" type="materials_examined">
<paragraph id="752936F0FFC8FFB74BB12EB0FAB41ECB" blockId="23.[151,1437,151,2023]" box="[199,1392,1499,1526]" pageId="23" pageNumber="224">
<emphasis id="47E2EAE2FFC8FFB74BB12EB0FD921ECB" box="[199,598,1499,1526]" italics="true" pageId="23" pageNumber="224">
<emphasis id="47E2EAE2FFC8FFB74BB12EB0FE9B1EC8" bold="true" box="[199,351,1499,1526]" italics="true" pageId="23" pageNumber="224">
<typeStatus id="AA2D8852FFC8FFB74BB12EB0FF3A1EC8" box="[199,254,1500,1526]" pageId="23" pageNumber="224">Type</typeStatus>
locality.
</emphasis>
<taxonomicName id="B2964D73FFC8FFB74A102EB0FD921ECB" authorityName="Suffr. D. Sassi" authorityYear="2024" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[358,598,1499,1526]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="tredecimpunctatus">G. tredecimpunctatus</taxonomicName>
</emphasis>
: São João del Rei (
<collectingRegion id="B752F812FFC8FFB748592EB7FC0E1ECB" box="[815,970,1499,1525]" country="Brazil" name="Minas Gerais" pageId="23" pageNumber="224">Minas Gerais</collectingRegion>
,
<collectingCountry id="0D817660FFC8FFB748A32EB7FBDA1ECB" box="[981,1054,1499,1525]" name="Brazil" pageId="23" pageNumber="224">Brazil</collectingCountry>
).
<taxonomicName id="B2964D73FFC8FFB74F462EB0FB391ECB" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[1072,1277,1499,1526]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="incomparabilis">
<emphasis id="47E2EAE2FFC8FFB74F462EB0FB391ECB" box="[1072,1277,1499,1526]" italics="true" pageId="23" pageNumber="224">G. incomparabilis</emphasis>
</taxonomicName>
: “
<collectingCountry id="0D817660FFC8FFB74E602EB7FAA51ECB" box="[1302,1377,1499,1525]" name="Brazil" pageId="23" pageNumber="224">Brazil</collectingCountry>
”.
</paragraph>
<paragraph id="752936F0FFC8FFB74BB12E92FCF91D5E" blockId="23.[151,1437,151,2023]" pageId="23" pageNumber="224">
<emphasis id="47E2EAE2FFC8FFB74BB12E92FDE01D27" bold="true" box="[199,548,1534,1561]" italics="true" pageId="23" pageNumber="224">Additional material examined</emphasis>
.
<materialsCitation id="C5FE3CADFFC8FFB749432E93FA951D27" box="[565,1361,1535,1561]" collectingDate="2021-12-07" collectionCode="GBIF" collectorName="Domingos Martins" country="Brazil" location="Espirito Santo" pageId="23" pageNumber="224" specimenCount="1" stateProvince="Espirito Santo">
<emphasis id="47E2EAE2FFC8FFB749432E93FD641D27" bold="true" box="[565,672,1535,1561]" pageId="23" pageNumber="224">
<collectingCountry id="0D817660FFC8FFB749432E93FD641D27" box="[565,672,1535,1561]" name="Brazil" pageId="23" pageNumber="224">BRAZIL</collectingCountry>
</emphasis>
:
<emphasis id="47E2EAE2FFC8FFB749C42E93FCA41D27" bold="true" box="[690,864,1535,1561]" pageId="23" pageNumber="224">
<collectingRegion id="B752F812FFC8FFB749C42E93FCA41D27" box="[690,864,1535,1561]" country="Brazil" name="Espirito Santo" pageId="23" pageNumber="224">Espirito Santo</collectingRegion>
</emphasis>
:
<collectorName id="D8635326FFC8FFB748042E93FB8E1D27" box="[882,1098,1535,1561]" pageId="23" pageNumber="224">Domingos Martins</collectorName>
,
<date id="01281030FFC8FFB74F2E2E93FB161D27" box="[1112,1234,1535,1561]" pageId="23" pageNumber="224" value="2021-12-07">
<collectingDate id="116CE9D8FFC8FFB74F2E2E93FB161D27" box="[1112,1234,1535,1561]" pageId="23" pageNumber="224" value="2021-12-07">7.XII.2021</collectingDate>
</date>
(1,
<collectionCode id="1387AE35FFC8FFB74E722E93FA8D1D27" box="[1284,1353,1535,1561]" pageId="23" pageNumber="224">GBIF</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C5FE3CADFFC8FFB74E292E93FE431D03" collectingDate="2021-12-07" collectionCode="HNHMB" collectorName="Domingos Martins" country="Brazil" isTailEnumeration="true" location="Espirito Santo" pageId="23" pageNumber="224" specimenCount="1" stateProvince="Esp. Santo">
<collectingRegion id="B752F812FFC8FFB74E1C2E93FF1D1D03" inferredBySuffix="true" pageId="23" pageNumber="224">Esp. Santo</collectingRegion>
” (3,
<collectionCode id="1387AE35FFC8FFB74A642D4FFEBB1D03" box="[274,383,1571,1597]" pageId="23" pageNumber="224">HNHMB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C5FE3CADFFC8FFB74AE52D4EFCC51D03" box="[403,769,1570,1597]" collectionCode="USNMNH" country="Brazil" location="Espirito Santo" pageId="23" pageNumber="224" specimenCount="1" stateProvince="Espirito Santo">
<collectingRegion id="B752F812FFC8FFB74AEB2D4EFD861D03" box="[413,578,1570,1597]" inferredBySuffix="true" pageId="23" pageNumber="224">Espirito Santo</collectingRegion>
” (1,
<collectionCode id="1387AE35FFC8FFB7490C2D4FFD3D1D03" box="[634,761,1571,1597]" pageId="23" pageNumber="224">USNMNH</collectionCode>
)
</materialsCitation>
. “Brasil” (1,
<collectionCode id="1387AE35FFC8FFB748E02D4FFBC71D03" box="[918,1027,1571,1597]" pageId="23" pageNumber="224">MNHUB</collectionCode>
); “Bresil” (1,
<collectionCode id="1387AE35FFC8FFB74FD72D4FFB381D03" box="[1185,1276,1571,1597]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="224">MNHN</collectionCode>
);
<materialsCitation id="C5FE3CADFFC8FFB74E792D4EFDDA1D5E" collectionCode="USNMNH" collectorName="F. C. Bowditch" country="Brazil" pageId="23" pageNumber="224" specimenCount="1" stateProvince="Espirito Santo">
“Brazil, Gift of
<collectorName id="D8635326FFC8FFB74BC02D2AFEA51D5E" box="[182,353,1606,1632]" pageId="23" pageNumber="224">F. C. Bowditch</collectorName>
” (1,
<collectionCode id="1387AE35FFC8FFB74AE12D2AFDD21D5E" box="[407,534,1606,1632]" pageId="23" pageNumber="224">USNMNH</collectionCode>
)
</materialsCitation>
. “Brasilia” (1,
<collectionCode id="1387AE35FFC8FFB749B22D2AFCF51D5E" box="[708,817,1606,1632]" pageId="23" pageNumber="224">MNHUB</collectionCode>
).
</paragraph>
<paragraph id="752936F0FFC8FFB74BB12D05FA461DF4" blockId="23.[151,1437,151,2023]" pageId="23" pageNumber="224">
<emphasis id="47E2EAE2FFC8FFB74BB12D05FDF91DBA" bold="true" box="[199,573,1641,1668]" italics="true" pageId="23" pageNumber="224">Additional data from literature.</emphasis>
<bibRefCitation id="11074B01FFC8FFB7493E2D05FD3F1DBD" author="Suffrian, E." box="[584,763,1641,1667]" pageId="23" pageNumber="224" pagination="1 - 483" refId="ref27961" refString="Suffrian, E. (1866) Zur Kenntnis der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1 - 483." type="journal article" year="1866">Suffrian (1866)</bibRefCitation>
, as stated above, reported “Collo del Sagramento” for
<taxonomicName id="B2964D73FFC8FFB74EF72D06FE851D99" baseAuthorityName="Suffrian" baseAuthorityYear="1866" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="incomparabilis">
<emphasis id="47E2EAE2FFC8FFB74EF72D06FE851D99" italics="true" pageId="23" pageNumber="224">G. incomparabilis</emphasis>
</taxonomicName>
as Brazilian collection locality for part of the
<typeStatus id="AA2D8852FFC8FFB748272DE1FC731D99" box="[849,951,1677,1703]" pageId="23" pageNumber="224" type="syntype">syntypes</typeStatus>
. As previously discussed elsewhere (
<bibRefCitation id="11074B01FFC8FFB74E2C2DE1FF171DF4" author="Sassi, D." pageId="23" pageNumber="224" pagination="201 - 245" refId="ref27561" refString="Sassi, D. (2019) Revision of the Metallactus hamifer species group (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa, 4657 (2), 201 - 245. https: // doi. org / 10.11646 / zootaxa. 4657.2.1" type="journal article" year="2019">Sassi, 2019</bibRefCitation>
), I am inclined to deem that the toponym must be intended as
<collectingRegion id="B752F812FFC8FFB748FD2DDCFB5C1DF4" box="[907,1176,1712,1738]" country="Uruguay" name="Colonia" pageId="23" pageNumber="224">Colonia del Sacramento</collectingRegion>
(
<collectingRegion id="B752F812FFC8FFB74FD12DDCFAC01DF4" box="[1191,1284,1712,1738]" country="Uruguay" name="Colonia" pageId="23" pageNumber="224">Colonia</collectingRegion>
,
<collectingCountry id="0D817660FFC8FFB74E792DDCFAB21DF4" box="[1295,1398,1712,1738]" name="Uruguay" pageId="23" pageNumber="224">Uruguay</collectingCountry>
).
</paragraph>
<paragraph id="752936F0FFC8FFB74BB12DB8FCCF1C0B" blockId="23.[151,1437,151,2023]" pageId="23" pageNumber="224">
Blackwelder (1877) described two new species from
<collectingCountry id="0D817660FFC8FFB7485A2DB8FC5F1DD0" box="[812,923,1748,1774]" name="Argentina" pageId="23" pageNumber="224">Argentina</collectingCountry>
hinting at
<taxonomicName id="B2964D73FFC8FFB74F6A2DB8FB2F1DD0" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[1052,1259,1748,1774]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="incomparabilis">
<emphasis id="47E2EAE2FFC8FFB74F6A2DB8FB2F1DD0" box="[1052,1259,1748,1774]" italics="true" pageId="23" pageNumber="224">G. incomparabilis</emphasis>
</taxonomicName>
as the possible closest taxon. As a matter of fact, these new species (
<taxonomicName id="B2964D73FFC8FFB7498B2D94FC681C2F" box="[765,940,1784,1810]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="octoguttatus">
<emphasis id="47E2EAE2FFC8FFB7498B2D94FC681C2F" box="[765,940,1784,1810]" italics="true" pageId="23" pageNumber="224">G. octoguttatus</emphasis>
</taxonomicName>
and
<taxonomicName id="B2964D73FFC8FFB748902D94FBBC1C2F" box="[998,1144,1783,1810]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="persimilis">
<emphasis id="47E2EAE2FFC8FFB748902D94FBBC1C2F" box="[998,1144,1783,1810]" italics="true" pageId="23" pageNumber="224">G. persimilis</emphasis>
</taxonomicName>
) are clearly different and turned out to belong to genus
<taxonomicName id="B2964D73FFC8FFB74A902C77FCC21C0B" authority="(Sassi, 2019)" baseAuthorityName="Sassi" baseAuthorityYear="2019" box="[486,774,1819,1845]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="47E2EAE2FFC8FFB74A902C77FDAE1C0B" box="[486,618,1819,1845]" italics="true" pageId="23" pageNumber="224">Metallactus</emphasis>
(
<bibRefCitation id="11074B01FFC8FFB7490F2C77FD3B1C0B" author="Sassi, D." box="[633,767,1819,1845]" pageId="23" pageNumber="224" pagination="201 - 245" refId="ref27561" refString="Sassi, D. (2019) Revision of the Metallactus hamifer species group (Coleoptera: Chrysomelidae: Cryptocephalinae). Zootaxa, 4657 (2), 201 - 245. https: // doi. org / 10.11646 / zootaxa. 4657.2.1" type="journal article" year="2019">Sassi, 2019</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="3D8C657BFFC8FFB74BB12C52FCBD1C67" box="[199,889,1854,1881]" pageId="23" pageNumber="224" type="distribution">
<paragraph id="752936F0FFC8FFB74BB12C52FCBD1C67" blockId="23.[151,1437,151,2023]" box="[199,889,1854,1881]" pageId="23" pageNumber="224">
<emphasis id="47E2EAE2FFC8FFB74BB12C52FE9E1C67" bold="true" box="[199,346,1854,1881]" italics="true" pageId="23" pageNumber="224">Distribution.</emphasis>
<collectingCountry id="0D817660FFC8FFB74A172C52FE611C66" box="[353,421,1854,1880]" name="Brazil" pageId="23" pageNumber="224">Brazil</collectingCountry>
(
<collectingRegion id="B752F812FFC8FFB74AC22C52FD9D1C66" box="[436,601,1854,1880]" country="Brazil" name="Espirito Santo" pageId="23" pageNumber="224">Espirito Santo</collectingRegion>
,
<collectingRegion id="B752F812FFC8FFB749152C52FD391C66" box="[611,765,1854,1880]" country="Brazil" name="Minas Gerais" pageId="23" pageNumber="224">Minas Gerais</collectingRegion>
),
<collectingCountry id="0D817660FFC8FFB748672C53FCB11C67" box="[785,885,1855,1881]" name="Uruguay" pageId="23" pageNumber="224">Uruguay</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="3D8C657BFFC8FFB74BB12C0DFE6B1CD9" pageId="23" pageNumber="224" type="diagnosis">
<paragraph id="752936F0FFC8FFB74BB12C0DFE6B1CD9" blockId="23.[151,1437,151,2023]" pageId="23" pageNumber="224">
<emphasis id="47E2EAE2FFC8FFB74BB12C0DFEFE1C45" bold="true" box="[199,314,1889,1915]" italics="true" pageId="23" pageNumber="224">Diagnosis</emphasis>
. The robust silhouette, with a highly transverse pronotum and, above all, the distinctive dorsal coloration featuring seven black spots on the pronotum arranged in two rows, the wide dark elytral spot in females, and, in males, the characteristically elongated small black elytral spots, make this species easily distinguishable from others in the group.
</paragraph>
</subSubSection>
<caption id="21E96678FFC7FFB84BE12C19FC821CC4" ID-DOI="http://doi.org/10.5281/zenodo.10618187" ID-Zenodo-Dep="10618187" httpUri="https://zenodo.org/record/10618187/files/figure.png" pageId="24" pageNumber="225" startId="24.[151,234,1909,1933]" targetBox="[219,1369,202,1879]" targetPageId="24" targetType="plate">
<paragraph id="752936F0FFC7FFB84BE12C19FC821CC4" blockId="24.[151,1437,1909,2042]" pageId="24" pageNumber="225">
<emphasis id="47E2EAE2FFC7FFB84BE12C19FEC71CB3" bold="true" box="[151,259,1909,1933]" pageId="24" pageNumber="225">PLATE 8.</emphasis>
<emphasis id="47E2EAE2FFC7FFB84A7C2C1AFE2F1CB3" box="[266,491,1909,1933]" italics="true" pageId="24" pageNumber="225">
<taxonomicName id="B2964D73FFC7FFB84A7C2C1AFE2C1CB3" authorityName="Suffr. D. Sassi" authorityYear="2024" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[266,488,1909,1933]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="species" species="tredecimpunctatus">G. tredecimpunctatus</taxonomicName>
.
</emphasis>
ab: habitus, dorsal and lateral view (LT); c: frontal view, male (LT); d: frontal view, female (LT of
<taxonomicName id="B2964D73FFC7FFB84B972CF6FE5F1C8F" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[225,411,1945,1969]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="species" species="incomparabilis">
<emphasis id="47E2EAE2FFC7FFB84B972CF6FE5F1C8F" box="[225,411,1945,1969]" italics="true" pageId="24" pageNumber="225">G. incomparabilis</emphasis>
</taxonomicName>
); eh: dorsal variations (d: male, “Brasilia”; f: female, LT of
<taxonomicName id="B2964D73FFC7FFB84F6B2CF6FB121C8F" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[1053,1238,1945,1969]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="species" species="incomparabilis">
<emphasis id="47E2EAE2FFC7FFB84F6B2CF6FB121C8F" box="[1053,1238,1945,1969]" italics="true" pageId="24" pageNumber="225">G. incomparabilis</emphasis>
</taxonomicName>
; g: female, PLT of
<taxonomicName id="B2964D73FFC7FFB84BE12CD2FE941CEB" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[151,336,1981,2005]" class="Insecta" family="Chrysomelidae" genus="Griburius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="species" species="incomparabilis">
<emphasis id="47E2EAE2FFC7FFB84BE12CD2FE941CEB" box="[151,336,1981,2005]" italics="true" pageId="24" pageNumber="225">G. incomparabilis</emphasis>
</taxonomicName>
; h: female, Espirito Santo, Brazil); kn: median lobe of aedeagus (“Brasilia”) in lateral (k), ventral (m) and dorsal (n) view [1.0 mm]. Square brackets: length of scale segment.
</paragraph>
</caption>
<subSubSection id="3D8C657BFFC6FFB94BB12BFBFA8D1C3B" pageId="25" pageNumber="226" type="description">
<paragraph id="752936F0FFC6FFB94BB12BFBFD2E1BEB" blockId="25.[151,1437,150,1797]" pageId="25" pageNumber="226">
<emphasis id="47E2EAE2FFC6FFB94BB12BFBFE761B8F" bold="true" box="[199,434,150,177]" italics="true" pageId="25" pageNumber="226">Description of male.</emphasis>
Habitus in figs 8ab (LT). BL = 6.0
<quantity id="B26E9B15FFC6FFB9482C2BFAFC751B8F" box="[858,945,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="25" pageNumber="226" unit="mm" value="6.5">6.5 mm</quantity>
, BW =
<quantity id="B26E9B15FFC6FFB94F7F2BFBFB561B8F" box="[1033,1170,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.4" metricValueMin="3.9" pageId="25" pageNumber="226" unit="mm" value="4.15" valueMax="4.4" valueMin="3.9">3.94.4 mm</quantity>
, PL =
<quantity id="B26E9B15FFC6FFB94FAB2BFBFAA31B8F" box="[1245,1383,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.1" metricValueMin="1.7" pageId="25" pageNumber="226" unit="mm" value="1.9" valueMax="2.1" valueMin="1.7">1.72.1 mm</quantity>
, PW = 3.64.0 mm. Interocular distance 8.39.2 % of BL.
</paragraph>
<paragraph id="752936F0FFC6FFB94BB12BB3FBD01A93" blockId="25.[151,1437,150,1797]" pageId="25" pageNumber="226">Head totally black. Labrum light yellow. Head surface covered with punctures and short appressed setosity, denser and coarser on frontoclypeal surface, sparser on vertex. Mid-cranial suture short but well impressed between eyes. Frontoclypeal sutures hardly detectable. Upper lobes of eyes broadly separated from each other. Ocular line slender, strictly adhering to ocular rim up to ocular canthus. Ocular canthus large, rounded, with surface hidden by short, appressed, dense setae. Antennae with first five antennomeres yellowish, reminders brownish. Antennomeres 35 bright subcylindrical; 611 dull, more flattened and more diffusedly setose.</paragraph>
<paragraph id="752936F0FFC6FFB94BA62ADBFD2319BB" blockId="25.[151,1437,150,1797]" pageId="25" pageNumber="226">Pronotum yellow with seven black spots arranged in two lines: four spots along anterior margin, inner ones larger; three spots along posterior margin, outer ones larger, nearly rectangular, mid one just in front of scutellum, small, usually diamond- or teardrop-shaped. Pronotal shape elliptical, markedly transverse and particularly flattened. Lateral margins narrow, scarcely visible from above, regularly curved, with maximum width nearly at middle. Surface moderately shiny with shallow, sparse punctures irregularly distributed across whole pronotal surface. Posterolateral impressions shallow, scarcely oblique.</paragraph>
<paragraph id="752936F0FFC6FFB94BB129E3FAA51997" blockId="25.[151,1437,150,1797]" box="[199,1377,655,681]" pageId="25" pageNumber="226">Scutellum black, trapezoidal. Surface shallowly micropunctured and covered with dense whitish setosity.</paragraph>
<paragraph id="752936F0FFC6FFB94BB129DFFA4218BF" blockId="25.[151,1437,150,1797]" pageId="25" pageNumber="226">Elytron yellow with three black spots, two of them lined along basal margin, longitudinally arranged, third one, lengthened as well, on apical clivus. Elytral outline with sides slightly convergent toward apex. Surface noticeably flattened on disc. Lateral margins narrow, almost invisible from above. Scutellar area fairly raised, so that dorsal profile clearly angular in lateral view. Humeral callus moderately prominent, impunctate. Surface rather shiny with distinctly impressed punctures irregularly arranged anteriorly, with remnants of rows on elytral clivus. Intervals flat. Epipleuron slightly convex, smooth, with surface mostly shiny and impunctate but finely rugulose along margin.</paragraph>
<paragraph id="752936F0FFC6FFB94BB128E7FAA5189B" blockId="25.[151,1437,150,1797]" box="[199,1377,907,933]" pageId="25" pageNumber="226">Pygidium totally yellow. Surface matt, covered with shallow punctures and short, appressed, dense setae.</paragraph>
<paragraph id="752936F0FFC6FFB94BB128C3FE5C1F9F" blockId="25.[151,1437,150,1797]" pageId="25" pageNumber="226">Ventral parts of thorax dark brown to black, with lateral part of hypomera yellowish. Abdominal ventrites brownish, with lighter colouring along sides. Hypomera shiny, almost devoid of punctation, covered with sparse, short setosity. Remainder of ventral part of thorax and abdominal surface matt, with dense, short, regularly distributed whitish setae and fine, shallow, dense punctures. Prosternal process large, with sides almost straight between anterior coxae; apex broadly rounded; surface almost flat and closely but shallowly punctured, covered with long, semi-erect setae. Legs totally black. Sexual dimorphism apparent in anterior leg shape, with femora robust, strengthened and tarsi stocky, thickened.</paragraph>
<paragraph id="752936F0FFC6FFB94BB12FC7FB2E1E6B" blockId="25.[151,1437,150,1797]" pageId="25" pageNumber="226">Median depression on fifth abdominal ventrite shallow, closely covered with short, appressed setae, hardly distinguishable from remainder of ventrite surface. Posterior margin of fifth abdominal ventrite straight. Median lobe of aedeagus (Figs 8kn) terminated with large, acute triangular apex, tapered anteriorly in lateral view with apex faintly bent ventrally. Setose depressions narrow, shifted on sides, setose only along dorsal margins. Glabrous depressions well delimited, oval-shaped. Aedeagal ventral surface with long, sharp, median carina.</paragraph>
<paragraph id="752936F0FFC6FFB94BB12E33FEB11EA3" blockId="25.[151,1437,150,1797]" pageId="25" pageNumber="226">
<emphasis id="47E2EAE2FFC6FFB94BB12E33FECB1E47" box="[199,271,1375,1401]" italics="true" pageId="25" pageNumber="226">Femal</emphasis>
e. BL =
<quantity id="B26E9B15FFC6FFB94A042E32FE3A1E47" box="[370,510,1374,1401]" metricMagnitude="-3" metricUnit="m" metricValue="6.449999999999999" metricValueMax="7.1" metricValueMin="5.8" pageId="25" pageNumber="226" unit="mm" value="6.449999999999999" valueMax="7.1" valueMin="5.8">5.87.1 mm</quantity>
, BW =
<quantity id="B26E9B15FFC6FFB9492B2E33FD2C1E47" box="[605,744,1375,1401]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.6" metricValueMin="3.9" pageId="25" pageNumber="226" unit="mm" value="4.25" valueMax="4.6" valueMin="3.9">3.94.6 mm</quantity>
, PL = 1.72.0 mm, PW =
<quantity id="B26E9B15FFC6FFB94F572E33FB6A1E47" box="[1057,1198,1375,1401]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.1" metricValueMin="3.7" pageId="25" pageNumber="226" unit="mm" value="3.9" valueMax="4.1" valueMin="3.7">3.74.1 mm</quantity>
. Interocular distance 11.312.1 % of BL.
</paragraph>
<paragraph id="752936F0FFC6FFB94BB12ECBFE761D6F" blockId="25.[151,1437,150,1797]" pageId="25" pageNumber="226">Females differ in more elliptical outline, in slightly larger interocular distance and in the anterior legs more typically shaped, with tarsi slenderer. Additionally, the most apparent difference is given by the elytral colour pattern, where a large oval black spot occupies the whole surface, leaving free only the lateral, apical and sutural margins. Sometimes, such spot is chestnut on its central part and black only along the margin, giving rise to a threecoloured pattern (Fig. 8f)</paragraph>
<paragraph id="752936F0FFC6FFB94BB12D37FA8D1C3B" blockId="25.[151,1437,150,1797]" pageId="25" pageNumber="226">Fifth abdominal ventrite in females with quite large and shallow pit. Bottom of pit glabrous, matt, impunctate but covered by tiny wrinkles. Vasculum of spermatheca (Fig. 11e) scarcely pigmented with slender proximal lobe slightly bent at base. Distal lobe slender as well, long, tapered towards apex which is pointed and mildly bent downward. Ampulla short, not pigmented. Duct insertion and sperm gland insertion perceptibly distinct. Duct uniform in size, slender, coiled with coils rather loose. Insertion on bursa copulatrix simple, not pigmented.</paragraph>
</subSubSection>
</treatment>
</document>