275 lines
24 KiB
XML
275 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.129.1202" ID-GBIF-Dataset="4b6a2d34-2ba2-4d8a-b7d5-fcb7963af269" ID-PMC="PMC3175144" ID-Pensoft-Pub="1313-2970-129-1" ID-PubMed="21998552" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-129-1" ModsDocOrigin="ZooKeys 129" ModsDocTitle="A new genus Squamophis of Asteroschematidae (Echinodermata, Ophiuroidea, Euryalida) from Australia" checkinTime="1451249912237" checkinUser="pensoft" docAuthor="Okanishi, Masanori, O'Hara, Timothy D. & Fujita, Toshihiko" docDate="2011" docId="93FDEA6A2532DF480863E4C6615BF4EB" docLanguage="en" docName="ZooKeys 129: 1-15" docOrigin="ZooKeys 129" docSource="http://dx.doi.org/10.3897/zookeys.129.1202" docTitle="Squamophis albozosteres Okanishi, O'Hara & Fujita, 2011, sp. n." docType="treatment" docVersion="4" lastPageNumber="9" masterDocId="120DFFDFD813FFD7FFA4C4322241FFE4" masterDocTitle="A new genus Squamophis of Asteroschematidae (Echinodermata, Ophiuroidea, Euryalida) from Australia" masterLastPageNumber="15" masterPageNumber="1" pageNumber="5" updateTime="1668152085513" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new genus Squamophis of Asteroschematidae (Echinodermata, Ophiuroidea, Euryalida) from Australia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Okanishi, Masanori</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>O'Hara, Timothy D.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fujita, Toshihiko</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>129</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>15</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.129.1202</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.129.1202</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-129-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152030702" LSID="urn:lsid:zoobank.org:act:9D85F117-BE04-4BF6-A03A-68636966B737" httpUri="http://treatment.plazi.org/id/93FDEA6A2532DF480863E4C6615BF4EB" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5">
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<subSubSection pageId="4" pageNumber="5" type="nomenclature">
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<paragraph pageId="4" pageNumber="5">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:9D85F117-BE04-4BF6-A03A-68636966B737" class="Ophiuroidea" family="Euryalidae" genus="Squamophis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamophis albozosteres" order="Euryalida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="species" species="albozosteres">
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<pageBreakToken pageId="4" pageNumber="5" start="start">Squamophis</pageBreakToken>
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albozosteres
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</taxonomicName>
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<taxonomicNameLabel pageId="4" pageNumber="5">sp. n.</taxonomicNameLabel>
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Figs 1B37
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="type materials">
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<paragraph pageId="4" pageNumber="5">Type materials.</paragraph>
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<paragraph pageId="4" pageNumber="5">
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MV F 162657, holotype, stn SS05/2007 116, off Broome, northwestern Australia,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-16.7515">16°45.09'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="121.041336">121°02.48'E</geoCoordinate>
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-
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-16.739334">16°44.36'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="121.03533">121°02.12'E</geoCoordinate>
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, 100-108 m, rocky bottom, 23.3 °C, 30 Jun 2007, epibenthic sled. MV F162658, two paratypes, stn SS05/2007 188, off Ashmore Reef, northwestern Australia,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-12.440333">12°26.42'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="123.6005">123°36.03'E</geoCoordinate>
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-
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-12.443">12°26.58'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="123.605835">123°36.35'E</geoCoordinate>
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, 95-96 m, rocky bottom, 24,8 °C, 7 Jul 2007, benthic dredge (Fig. 2).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="diagnosis">
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<paragraph pageId="4" pageNumber="5">Diagnosis.</paragraph>
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<paragraph pageId="4" pageNumber="5">Epidermal ossicles conspicuous white, slightly domed and round plate-shaped, irregularly placed on aboral side of disc and forming two transverse bands on aboral and lateral sides of each arm segment. Rest of the aboral surface uniformly brown except light purple near the tips of arms and finally without color at the tip.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="description of holotype">
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<paragraph pageId="4" pageNumber="5">Description of holotype.</paragraph>
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<paragraph pageId="4" pageNumber="5">MV F162657: disc diameter 3.4 mm, arm length approximately 50 mm (Fig. 3).</paragraph>
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<paragraph pageId="4" pageNumber="5">
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Disc. Disc five-lobed with slightly notched interradial edges: lacking evidence of fission (Figs 3, 4A). Aboral surface almost flat, but radial shields and their surrounds slightly tumid, covered by white, slightly domed and round plate-shaped epidermal ossicles and brown, flat and polygonal plate-shaped epidermal ossicles (Fig. 4
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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). Epidermal ossicles covered by a thin skin. White epidermal ossicles forming transverse rows at the aboral disc (Fig. 4A), almost uniform in size on aboral disc, 70-120
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<normalizedToken originalValue="µm">µm</normalizedToken>
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long, approximately 100
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<normalizedToken originalValue="µm">µm</normalizedToken>
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thick. Brown epidermal ossicles obscured by skin and cannot observed externally (Fig. 4
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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); relatively large near the periphery, 150-250
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<normalizedToken originalValue="µm">µm</normalizedToken>
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long, approximately 50
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<normalizedToken originalValue="µm">µm</normalizedToken>
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thick, and relatively small at the disc center, 100-150
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<normalizedToken originalValue="µm">µm</normalizedToken>
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long, approximately 50
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<normalizedToken originalValue="µm">µm</normalizedToken>
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thick. Radial shields completely covered by epidermal ossicles, oblong, approximately 1.2 mm long and 0.6 mm wide, not reaching the center of the disc.
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</paragraph>
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<caption pageId="4" pageNumber="5">
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<paragraph pageId="4" pageNumber="5">
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Figure 2. Collected sites of
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<taxonomicName class="Ophiuroidea" family="Euryalidae" genus="Squamophis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamophis albozosteres" order="Euryalida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="species" species="albozosteres">Squamophis albozosteres</taxonomicName>
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sp. n. Northern solid circle is for Ashmore Reef and southern one is off Broome.
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</paragraph>
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</caption>
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<caption pageId="4" pageNumber="5">
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<paragraph pageId="4" pageNumber="5">
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Figure 3.
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<taxonomicName class="Ophiuroidea" family="Euryalidae" genus="Squamophis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamophis albozosteres" order="Euryalida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="species" species="albozosteres">Squamophis albozosteres</taxonomicName>
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sp. n., holotype (MV F162657). A aboral view B oral view. Arrows indicate the arm of another ophiuroid gripped by
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<taxonomicName class="Ophiuroidea" family="Euryalidae" genus="Squamophis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamophis albozosteres" order="Euryalida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="species" species="albozosteres">Squamophis albozosteres</taxonomicName>
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.
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</paragraph>
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</caption>
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<paragraph pageId="5" pageNumber="6">
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<pageBreakToken pageId="5" pageNumber="6" start="start">Oral</pageBreakToken>
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surface of disc entirely covered by only white, flat and polygonal plate-shaped epidermal ossicles, 50-100
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<normalizedToken originalValue="µm">µm</normalizedToken>
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long and approximately 50
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<normalizedToken originalValue="µm">µm</normalizedToken>
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thick (Fig. 4
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<normalizedToken originalValue="D–G">D-G</normalizedToken>
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). Four to five triangular teeth forming a vertical row on dental plate. Domed granule-shaped oral papillae lying on either side of jaw (Fig. 4F).
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</paragraph>
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<paragraph pageId="5" pageNumber="6">Lateral interradial surface of disc nearly vertical, covered by epidermal ossicles similar to those on oral surface (Fig. 4H). Two genital slits (0.6 mm long and 0.3 mm wide) present in each interradius. No distinct ossicles suggesting existence of madreporites or oral plates observed on any oral interradius, and only epidermal ossicles covered these surfaces (Fig. 4H).</paragraph>
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<paragraph pageId="6" pageNumber="7">
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<pageBreakToken pageId="6" pageNumber="7" start="start">Arms</pageBreakToken>
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. Arms simple, five in number, no abrupt change in width near the arm base (Fig. 3). The basal portion of the arm 1.4 mm wide and 1.5 mm high, with an arched aboral surface and flattened oral surface. Arms tapering gradually toward the arm tip (Figs 3, 5A, D, G).
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</paragraph>
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<caption pageId="6" pageNumber="7">
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<paragraph pageId="6" pageNumber="7">
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Figure 4.
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<taxonomicName class="Ophiuroidea" family="Euryalidae" genus="Squamophis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamophis albozosteres" order="Euryalida" pageId="6" pageNumber="7" phylum="Echinodermata" rank="species" species="albozosteres">Squamophis albozosteres</taxonomicName>
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sp. n., holotype (MV F162657). A aboral disc and basal portion of the arms B periphery of the disc and basal portion of the arm C aboral central disc D oral disc E oral central disc F jaws G oral periphery of the disc H lateral interradius of the disc. Abbreviations: GS - genital slit; O - oral papillae; T - teeth; Ten - Tentacles.
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</paragraph>
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</caption>
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<caption pageId="6" pageNumber="7">
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<paragraph pageId="6" pageNumber="7">
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Figure 5.
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<taxonomicName class="Ophiuroidea" family="Euryalidae" genus="Squamophis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamophis albozosteres" order="Euryalida" pageId="6" pageNumber="7" phylum="Echinodermata" rank="species" species="albozosteres">Squamophis albozosteres</taxonomicName>
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sp. n., holotype (MV F162657). A, B aboral basal portion of the arm C oral basal portion of the arm D aboral middle portion of the arm, each arrow head indicates a row of white ossicles E oral middle portion of the arm F lateral middle portion of the arm, each arrow head indicates a row of white ossicles G oral distal portion of the arm H lateral distal portion of the arm. Abbreviations: - AS arm spine; LAP - lateral arm plate; Ten - tentacles.
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</paragraph>
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</caption>
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<paragraph pageId="6" pageNumber="7">
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The aboral and lateral surface of the base of arms completely covered by white epidermal ossicles, 150-300
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<normalizedToken originalValue="µm">µm</normalizedToken>
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long, approximately 60
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<normalizedToken originalValue="µm">µm</normalizedToken>
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thick, and brown epidermal ossicles, 100-300
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<normalizedToken originalValue="µm">µm</normalizedToken>
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long, approximately 50
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<normalizedToken originalValue="µm">µm</normalizedToken>
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thick (Fig. 5A, B), similar to those on aboral periphery of disc. Epidermal ossicles on basal portion of arms covered by thin skin. Brown epidermal ossicles obscured by skin, similar to those on aboral surface of disc. Oral surface of the base of arm covered by white epidermal ossicles, 50-100
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<normalizedToken originalValue="µm">µm</normalizedToken>
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long, approximately 50
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<normalizedToken originalValue="µm">µm</normalizedToken>
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thick, similar to those on oral surface of disc (Fig. 5C). From basal to middle portion of the arms, the size of plate-shaped epidermal ossicles decreasing on both the aboral and lateral surfaces (Fig. 5D, F), the white domed ones to 100-200
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<normalizedToken originalValue="µm">µm</normalizedToken>
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and the brown polygonal ones to 100-150
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<normalizedToken originalValue="µm">µm</normalizedToken>
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. Brown polygonal ones on oral surface decreasing to 50
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(Fig. 5E). The distal sections of arms covered by scattered granule-shaped epidermal ossicles of 30
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, finally disappearing near arm tip (Fig. 5G, H).
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</paragraph>
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<paragraph pageId="6" pageNumber="7">First to third tentacle pores lacking arm spines; 4th and more distal pores with one arm spine. Arm spines on basal one-third of arm ovoid, minute, approximately one-third to one-half the length of corresponding arm segment (Fig. 5C). Arm spines in middle one-third of arm the same length as corresponding arm segment, bearing fine thorns at their apex (Fig. 5E, F). Arm spines on distal one-third of arm hook-shaped with conspicuous lateral secondary teeth along inner edge (Fig. 5G, H). Length of hook-shaped arm spines gradually decreasing to two-thirds of the corresponding arm segment on distal third of arm, and number of secondary teeth decreasing from two to one. All tentacles pores lacking a sheath around the cylindrical, narrow tube feet (Figs 4E, 5G).</paragraph>
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<paragraph pageId="6" pageNumber="7">Lateral and ventral arm plates completely concealed by epidermal ossicles over basal to middle portion of arms, but lateral arm plates visible in distal portion of arms (Fig. 5G).</paragraph>
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<paragraph pageId="6" pageNumber="7">Color.Aboral surface of disc brown, with white spots highlighting the domed epidermal ossicles. Pigmentation on aboral distal portion of arms lighter and appearing purple, finally disappearing at the tip (Fig. 5H). Oral side entirely white (Fig. 3).</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="ossicle morphology of one paratype">
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<paragraph pageId="6" pageNumber="7">Ossicle morphology of one paratype.</paragraph>
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<paragraph pageId="6" pageNumber="7">MVF162658: Disc diameter 5.3 mm, arm length at least 200 mm.</paragraph>
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<paragraph pageId="6" pageNumber="7">
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Flat and polygonal plate-shaped epidermal ossicles at aboral periphery of disc, approximately 236
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<normalizedToken originalValue="µm">µm</normalizedToken>
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long and 43
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<normalizedToken originalValue="µm">µm</normalizedToken>
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thick (Fig. 6A, B), the white, round and domed plate-shaped epidermal ossicles approximately 136
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<normalizedToken originalValue="µm">µm</normalizedToken>
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long and 40
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<normalizedToken originalValue="µm">µm</normalizedToken>
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thick (Fig. 6C, D). On aboral surface at base of arm, domed ossicles slightly oblong, approximately 226
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<normalizedToken originalValue="µm">µm</normalizedToken>
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long and 34
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<normalizedToken originalValue="µm">µm</normalizedToken>
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thick (Fig. 6E, F), whereas the other ossicles flat and round, granule-shaped, 64
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<normalizedToken originalValue="µm">µm</normalizedToken>
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long and 20
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<normalizedToken originalValue="µm">µm</normalizedToken>
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thick (Fig. 6G, H).
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</paragraph>
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<caption pageId="6" pageNumber="7">
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<paragraph pageId="6" pageNumber="7">
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Figure 6.
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<normalizedToken originalValue="A–K">A-K</normalizedToken>
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<taxonomicName class="Ophiuroidea" family="Euryalidae" genus="Squamophis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamophis albozosteres" order="Euryalida" pageId="6" pageNumber="7" phylum="Echinodermata" rank="species" species="albozosteres">Squamophis albozosteres</taxonomicName>
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sp. n., paratype(MV F162658). SEM photographs of internal ossicles. A, B polygonal plate-shaped epidermal ossicles at the aboral periphery of the disc, external (A) and lateral (B) views C, D domed plate-shaped epidermal ossicles at the periphery of the disc, external (C) and lateral (D) views E, F domed plate-shaped epidermal ossicles on the aboral middle portion of the arm, external (E) and lateral (F) views G, H granule-shaped epidermal ossicles on the oral middle portion of the arm, external (G) and lateral (H) views
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<normalizedToken originalValue="I–K">I-K</normalizedToken>
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arm spines from basal (I), middle (J) and distal (K) portion of the arm. Arrows indicate the orientation (B, D, F, H): ext - external side; int - internal side.
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</paragraph>
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</caption>
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<paragraph pageId="6" pageNumber="7">The radial shields flat and oblong, single-layered, approximately 1.15 mm in length and 0.57 mm in width (Fig. 1B).</paragraph>
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<paragraph lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
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<pageBreakToken pageId="7" pageNumber="8" start="start">Arm</pageBreakToken>
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spines on basal one-third of arm ovoid (Fig. 6I), in middle cylindrical, bearing fine thorns at tip (Fig. 6J), and distally, they hook shaped with conspicuous secondary teeth along inner edge (Fig. 6K). Number of secondary teeth decreasing gradually to one along distal quarter of arm
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<pageBreakToken pageId="8" pageNumber="9" start="start">.</pageBreakToken>
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</paragraph>
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<paragraph pageId="8" pageNumber="9">
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Each lateral arm plate associated with one arm spine and has separate muscle and nerve openings (
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<bibRefCitation author="Martynov, A" journalOrPublisher="Zootaxa" pageId="8" pageNumber="9" pagination="1 - 154" title="Reassessment of the classification of the Ophiuroidea (Echinodermata), based on morphological characters. I. General character evaluation and delineation of the families Ophiomyxidae and Ophiacanthidae." volume="2697" year="2010">Martynov 2010</bibRefCitation>
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) (Fig. 7A). Oral side of each arm vertebra with a longitudinal groove along midline, no oral bridge (
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<bibRefCitation pageId="8" pageNumber="9">Okanishi et al. in press</bibRefCitation>
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) formed to surround the radial water vessel and nerve (Fig. 7B, C).
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</paragraph>
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<caption pageId="8" pageNumber="9">
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<paragraph pageId="8" pageNumber="9">
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Figure 7.
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<taxonomicName class="Ophiuroidea" family="Euryalidae" genus="Squamophis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamophis albozosteres" order="Euryalida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="albozosteres">Squamophis albozosteres</taxonomicName>
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sp. n., paratype(MV F162658). SEM photographs of internal ossicles. A lateral arm plate from middle portion of the arm B, C vertebrae from middle (B) and distal (C) portion of the arm, oral views. Abbreviations: MO - muscle opening; NO - nerve opening.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="9" type="variation">
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<paragraph pageId="8" pageNumber="9">Variation.</paragraph>
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<paragraph pageId="8" pageNumber="9">
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Although only three specimens have been collected, some morphological variation was observed. The smaller holotype (3.4 mm in disc diameter) has no abrupt reduction in arm thickness, but the basal portion of the arm on the two larger paratypes (5.3 mm and 5.6 mm in disc diameter) are slightly widened. The difference between the three specimens may be due to a difference in their sexual maturity or reproductive state similar to the congener,
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<taxonomicName class="Ophiuroidea" family="Euryalidae" genus="Squamophis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamophis amamiensis" order="Euryalida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="amamiensis">Squamophis amamiensis</taxonomicName>
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(
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<bibRefCitation author="Okanishi, M" journalOrPublisher="Species Diversity" pageId="8" pageNumber="9" pagination="115 - 129" title="A new species of Asteroschema (Echinodermata: Ophiuroidea: Asteroschematidae) from southwestern Japan." volume="14" year="2009">Okanishi and Fujita 2009</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="9" type="distribution">
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<paragraph pageId="8" pageNumber="9">Distribution.</paragraph>
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<paragraph pageId="8" pageNumber="9">North-western Australia; 95-108 m. Type locality: off Broome, 100-108 m (Fig. 2).</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="9" type="etymology">
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<paragraph pageId="8" pageNumber="9">Etymology.</paragraph>
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<paragraph pageId="8" pageNumber="9">
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The specific name is a masculine noun in apposition formed as a compound of Latin words, albus (adjective, meaning
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<normalizedToken originalValue="“whitish”">"whitish"</normalizedToken>
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) and a plural form of zoster (masculine noun, meaning
|
||
<normalizedToken originalValue="“ring”">"ring"</normalizedToken>
|
||
), referring to the rings of white plate-shaped dermal ossicles of arms.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="9" type="remarks">
|
||
<paragraph pageId="8" pageNumber="9">Remarks.</paragraph>
|
||
<paragraph pageId="8" pageNumber="9">
|
||
<taxonomicName class="Ophiuroidea" family="Euryalidae" genus="Squamophis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamophis albozosteres" order="Euryalida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="albozosteres">Squamophis albozosteres</taxonomicName>
|
||
sp. n. and its congener,
|
||
<taxonomicName class="Ophiuroidea" family="Euryalidae" genus="Squamophis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamophis amamiensis" order="Euryalida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="amamiensis">Squamophis amamiensis</taxonomicName>
|
||
, are similar to each other, however, they can be distinguished by the morphology of the epidermal ossicles on the aboral body and by pigmentation.
|
||
<taxonomicName class="Ophiuroidea" family="Euryalidae" genus="Squamophis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamophis albozosteres" order="Euryalida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="albozosteres">Squamophis albozosteres</taxonomicName>
|
||
has conspicuously white, domed and plate-shaped epidermal ossicles on the aboral side of the disc and basal to middle portion of the arms, forming two transverse rows on the lateral and aboral surfaces of each arm segments (Fig. 5F). Whereas
|
||
<taxonomicName class="Ophiuroidea" family="Euryalidae" genus="Squamophis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamophis amamiensis" order="Euryalida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="amamiensis">Squamophis amamiensis</taxonomicName>
|
||
has only uniform coloured, flattened and plate-shaped epidermal ossicles on the corresponding surfaces. The aboral body surface of
|
||
<taxonomicName class="Ophiuroidea" family="Euryalidae" genus="Squamophis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamophis albozosteres" order="Euryalida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="albozosteres">Squamophis albozosteres</taxonomicName>
|
||
is basically brown with white spots and the tips of the arms are light purple, finally with no color, but that of
|
||
<taxonomicName class="Ophiuroidea" family="Euryalidae" genus="Squamophis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamophis amamiensis" order="Euryalida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="amamiensis">Squamophis amamiensis</taxonomicName>
|
||
is uniformly orange or pinkish brown (
|
||
<bibRefCitation author="Okanishi, M" journalOrPublisher="Species Diversity" pageId="8" pageNumber="9" pagination="115 - 129" title="A new species of Asteroschema (Echinodermata: Ophiuroidea: Asteroschematidae) from southwestern Japan." volume="14" year="2009">Okanishi and Fujita 2009</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |