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<document id="56F8708EA0C3DB0FB6C302D25492E1CA" ID-DOI="10.5281/zenodo.5176593" ID-ISSN="0374-1036" ID-Zenodo-Dep="5176593" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628601849081" checkinUser="carolina" docAuthor="Ježek, Jan &amp; Harten, Antonius van" docDate="2005" docId="9331B306FFBFFFC68EB0FA8858AAFE61" docLanguage="en" docName="ActaEntMusNatPra.45.199-220.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 45" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Psychana rujumensis Ježek &amp; Harten 2005, sp. nov." docType="treatment" docVersion="3" lastPageNumber="211" masterDocId="6F08CB7EFFB7FFCA8F6BFFA15865FF96" masterDocTitle="Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen" masterLastPageNumber="220" masterPageNumber="199" pageNumber="207" updateTime="1699047277773" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="90D497F49AAE8D3608E68D9B07810DB1">Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen</mods:title>
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<mods:name id="30E39C500BD233DBCDB0F73FBFCA7E5B" type="personal">
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<mods:namePart id="124DD49E52C49497A6E02167CD5C134F">Ježek, Jan</mods:namePart>
<mods:affiliation id="841BA905F3C7B826563FBA9FD77B99AE">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: entomologie @ volny. cz</mods:affiliation>
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<mods:namePart id="29834EA8681FFA484DBF9F1CA984E59C">Harten, Antonius van</mods:namePart>
<mods:affiliation id="44FED97BB520C1BC3C4F7C48DC76CB60">UAE Insect Project, P. O. Box 63799, Sharjah, United Arab Emirates; e-mail: vanharten @ zmail. pt</mods:affiliation>
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<mods:title id="D15E0A18F84B24526C1274E54AB09F52">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="597F15E3BA4BCFB12F95C65679CABD0B">
<mods:date id="61290B73E250438790B26D209B6A2243">2005</mods:date>
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<mods:identifier id="339880FF9A7ACCE60FD4079BFAACEB01" type="DOI">10.5281/zenodo.5176593</mods:identifier>
<mods:identifier id="52F181EF1310BE9F23DF1D5510AB533A" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="F50F5874579B6055CE4501D30A768176" type="Zenodo-Dep">5176593</mods:identifier>
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<treatment id="9331B306FFBFFFC68EB0FA8858AAFE61" LSID="urn:lsid:plazi:treatment:9331B306FFBFFFC68EB0FA8858AAFE61" httpUri="http://treatment.plazi.org/id/9331B306FFBFFFC68EB0FA8858AAFE61" lastPageId="12" lastPageNumber="211" pageId="8" pageNumber="207">
<subSubSection id="5382519BFFBFFFC28EB0FA885B46FAD7" box="[475,803,1321,1345]" pageId="8" pageNumber="207" type="nomenclature">
<paragraph id="1B270210FFBFFFC28EB0FA885B46FAD7" blockId="8.[475,803,1321,1345]" box="[475,803,1321,1345]" pageId="8" pageNumber="207">
<heading id="406FB57CFFBFFFC28EB0FA885B46FAD7" bold="true" box="[475,803,1321,1345]" centered="true" fontSize="8" level="2" pageId="8" pageNumber="207" reason="3">
<emphasis id="29ECDE02FFBFFFC28EB0FA885B46FAD7" bold="true" box="[475,803,1321,1345]" pageId="8" pageNumber="207">
<taxonomicName id="DC987993FFBFFFC28EB0FA885AA3FAD7" ID-CoL="8DK2Y" authority="Ježek &amp; Harten, 2005" authorityName="Ježek &amp; Harten" authorityYear="2005" box="[475,710,1321,1345]" class="Insecta" family="Psychodidae" genus="Psychana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="207" phylum="Arthropoda" rank="species" species="rujumensis" status="sp. nov.">
<emphasis id="29ECDE02FFBFFFC28EB0FA885AA3FAD7" bold="true" box="[475,710,1321,1345]" italics="true" pageId="8" pageNumber="207">Psychana rujumensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="32DF6379FFBFFFC28DA6FA885B46FAD7" box="[717,803,1321,1345]" pageId="8" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5382519BFFBFFFC28D2CFAE85AD2FACA" box="[583,695,1353,1372]" pageId="8" pageNumber="207" type="description">
<paragraph id="1B270210FFBFFFC28D2CFAE85AD2FACA" blockId="8.[583,695,1353,1372]" box="[583,695,1353,1372]" pageId="8" pageNumber="207">
(
<figureCitation id="83A31E95FFBFFFC28D26FAE85AD4FACA" box="[589,689,1353,1372]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="9.[113,153,1410,1429]" captionStartId-1="11.[113,152,1439,1458]" captionTargetBox-0="[113,1088,220,1363]" captionTargetBox-1="[121,1084,217,1400]" captionTargetId-0="figure-83@9.[113,1103,205,1363]" captionTargetId-1="figure-18@11.[113,1084,203,1412]" captionTargetPageId-0="9" captionTargetPageId-1="11" captionText-0="Figs. 35-43. Psychana rujumensis sp. nov., J. 35 head; 36 facets; 37 basal antennomeres; 38 apical antennomeres; 39 wing; 40 dorsal view of fore claw; 41 lateral view of aedeagal compex; 42 lateral view of gonocoxite and gonostyle; 43 dorsal view of epandrium and surstyli. Scales = 0.1 mm (Figs. 35-38, 40-43); 1 mm (Fig. 39)." captionText-1="Figs. 44-53. Psychana rujumensis sp. nov., J (44-48) and ♀ (49-53). 44 dorsal view of terminal lobe of labium; 45 maxilla and maxillary palpus; 46 lateral view of thoracic sclerites; 47 dorsal view of aedeagal complex and gonopods; 48 lateral view of epandrium and surstylus; 49 subgenital plate; 50 dorsal view of genital chamber; 51 same, anterior view; 52 same, lateral view; 53 lateral view of cercus. Scales = 0.1 mm (Figs. 44-45, 47-53); 0.5 mm (Fig. 46)." figureDoi-0="http://doi.org/10.5281/zenodo.5176601" figureDoi-1="http://doi.org/10.5281/zenodo.5176603" httpUri-0="https://zenodo.org/record/5176601/files/figure.png" httpUri-1="https://zenodo.org/record/5176603/files/figure.png" pageId="8" pageNumber="207">Figs. 35-53</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5382519BFFBFFFC08FFCFADF5B45FE8E" lastPageId="10" lastPageNumber="209" pageId="8" pageNumber="207" type="materials_examined">
<paragraph id="1B270210FFBFFFC28FFCFADF5C02F9A0" blockId="8.[151,1127,1403,1591]" pageId="8" pageNumber="207">
<emphasis id="29ECDE02FFBFFFC28FFCFADF5958FA00" bold="true" box="[151,317,1406,1430]" pageId="8" pageNumber="207">Type material.</emphasis>
<materialsCitation id="ABF0084DFFBFFFC28E2FFADE5892FA40" collectingDate="2000-10-16" collectingDateMax="2001-01-15" collectingDateMin="2000-10-16" collectionCode="NMPC" collectorName="A. van Harten &amp; Slide &amp; Cat. No. &amp; Inv. No." country="Yemen" latitude="15.483334" location="Ar Rujum" longLatPrecision="1284" longitude="43.683334" pageId="8" pageNumber="207" specimenCount="1" typeStatus="holotype">
<typeStatus id="C423BCB2FFBFFFC28E2FFADE59CEFA03" box="[324,427,1407,1430]" pageId="8" pageNumber="207" type="holotype">HOLOTYPE</typeStatus>
: J,
<emphasis id="29ECDE02FFBFFFC28EB0FADF5A58FA00" bold="true" box="[475,573,1406,1430]" pageId="8" pageNumber="207">
<collectingCountry id="638F4280FFBFFFC28EB0FADF5A58FA00" box="[475,573,1406,1430]" name="Yemen" pageId="8" pageNumber="207">YEMEN</collectingCountry>
</emphasis>
:
<location id="1E4754CBFFBFFFC28D20FADE5ADCFA00" LSID="urn:lsid:plazi:treatment:9331B306FFBFFFC68EB0FA8858AAFE61:1E4754CBFFBFFFC28D20FADE5ADCFA00" box="[587,697,1406,1431]" country="Yemen" latitude="15.483334" longLatPrecision="1284" longitude="43.683334" name="Ar Rujum" pageId="8" pageNumber="207">Ar Rujum</location>
,
<geoCoordinate id="7EAC64D7FFBFFFC28DAFFADF5B78FA00" box="[708,797,1405,1430]" degrees="15" direction="north" minutes="29" orientation="latitude" pageId="8" pageNumber="207" precision="925" value="15.483334">15°29N</geoCoordinate>
<geoCoordinate id="7EAC64D7FFBFFFC28C4EFADF5B18FA00" box="[805,893,1405,1430]" degrees="43" direction="east" minutes="41" orientation="longitude" pageId="8" pageNumber="207" precision="925" value="43.683334">43°41E</geoCoordinate>
,
<date id="6F2624D0FFBFFFC28CE2FADF5B90FA00" box="[905,1013,1406,1430]" pageId="8" pageNumber="207" value="2000-10-16">
<collectingDate id="7F62DD38FFBFFFC28CE2FADF5B90FA00" box="[905,1013,1406,1430]" pageId="8" pageNumber="207" value="2000-10-16">16.x.2000</collectingDate>
</date>
-
<date id="6F2624D0FFBFFFC28C97FADF5C05FA00" box="[1020,1120,1406,1430]" pageId="8" pageNumber="207" value="2001-01-15">
<collectingDate id="7F62DD38FFBFFFC28C97FADF5C05FA00" box="[1020,1120,1406,1430]" pageId="8" pageNumber="207" value="2001-01-15">15.i.2001</collectingDate>
</date>
,
<collectingMethod id="C2D97A07FFBFFFC28FFCFA3F597AFA20" box="[151,287,1438,1462]" pageId="8" pageNumber="207">Malaise trap</collectingMethod>
,
<collectorName id="B66D67C6FFBFFFC28E43FA3E59DAFA20" box="[296,447,1439,1463]" pageId="8" pageNumber="207">A. van Harten</collectorName>
leg.
<collectorName id="B66D67C6FFBFFFC28E9EFA3F5A4BFA20" box="[501,558,1438,1462]" pageId="8" pageNumber="207">Slide</collectorName>
, dissected specimen,
<collectorName id="B66D67C6FFBFFFC28C7FFA3F5B0AFA20" box="[788,879,1438,1462]" pageId="8" pageNumber="207">Cat. No.</collectorName>
34264,
<collectorName id="B66D67C6FFBFFFC28CADFA3E5C7BFA20" box="[966,1054,1439,1462]" pageId="8" pageNumber="207">Inv. No.</collectorName>
14053 (
<collectionCode id="7D899AD5FFBFFFC28FF4FA1F5895FA40" box="[159,240,1470,1494]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="8" pageNumber="207">NMPC</collectionCode>
)
</materialsCitation>
.
<typeStatus id="C423BCB2FFBFFFC28E6BFA1E5909FA43" box="[256,364,1471,1494]" pageId="8" pageNumber="207" type="paratype">
<collectionCode id="7D899AD5FFBFFFC28E6BFA1E5909FA43" box="[256,364,1471,1494]" pageId="8" pageNumber="207">PARATYPES</collectionCode>
</typeStatus>
: 3 JJ
<specimenCount id="0D9EC999FFBFFFC28EDCFA1F5983FA41" box="[439,486,1467,1495]" pageId="8" pageNumber="207" type="female">6 ♀♀</specimenCount>
(slides,
<collectionCode id="7D899AD5FFBFFFC28D2DFA1F5AF3FA40" box="[582,662,1470,1494]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="8" pageNumber="207">NMPC</collectionCode>
), the same locality, collector and trap.
<collectionCode id="7D899AD5FFBFFFC28B4CFA1E58A0FA62" pageId="8" pageNumber="207">ALLO- TYPE</collectionCode>
:
<specimenCount id="0D9EC999FFBFFFC28FBFFA7A5886FA61" box="[212,227,1499,1527]" pageId="8" pageNumber="207" type="female"></specimenCount>
,
<date id="6F2624D0FFBFFFC28F9FFA7F5905FA60" box="[244,352,1502,1526]" pageId="8" pageNumber="207" value="2000-10-16">16.x.2000</date>
-
<date id="6F2624D0FFBFFFC28E0CFA7F59AEFA60" box="[359,459,1502,1526]" pageId="8" pageNumber="207" value="2001-01-15">15.i.2001</date>
, Cat. No. 34265, Inv. No. 14054.
<collectionCode id="7D899AD5FFBFFFC28C5CFA7F5B1FFA62" box="[823,890,1502,1526]" pageId="8" pageNumber="207">OTHER</collectionCode>
<typeStatus id="C423BCB2FFBFFFC28CEAFA455B82FA63" box="[897,999,1508,1525]" pageId="8" pageNumber="207" type="paratype">
<collectionCode id="7D899AD5FFBFFFC28CEAFA455B82FA63" box="[897,999,1508,1525]" pageId="8" pageNumber="207">PARATYPES</collectionCode>
</typeStatus>
: JJ:, same date, Cat. No. 34266-34267, Inv. No. 14055-14056;
<date id="6F2624D0FFBFFFC28DD5FA5F5B32F980" box="[702,855,1534,1558]" pageId="8" pageNumber="207" value="2001-01-16" valueMax="2001-04-09" valueMin="2001-01-16">16.i.-9.iv.2001</date>
, Cat. No. 34268, Inv. No. 14057;
<specimenCount id="0D9EC999FFBFFFC28F85F9BA596EF9A1" box="[238,267,1563,1591]" pageId="8" pageNumber="207" type="female">♀♀</specimenCount>
,
<date id="6F2624D0FFBFFFC28E4FF9BF59F6F9A0" box="[292,403,1566,1590]" pageId="8" pageNumber="207" value="2000-10-16">16.x.2000</date>
-
<date id="6F2624D0FFBFFFC28EF0F9BF5A67F9A0" box="[411,514,1566,1590]" pageId="8" pageNumber="207" value="2001-01-15">15.i.2001</date>
, Cat. No. 34269-34271, Inv. No. 14058-14060; 16.i.-
</paragraph>
<caption id="4FE75298FFBEFFC38F1AFA2358A6FA73" ID-DOI="http://doi.org/10.5281/zenodo.5176601" ID-Zenodo-Dep="5176601" httpUri="https://zenodo.org/record/5176601/files/figure.png" pageId="9" pageNumber="208" startId="9.[113,153,1410,1429]" targetBox="[113,1088,220,1363]" targetPageId="9">
<paragraph id="1B270210FFBEFFC38F1AFA2358A6FA73" blockId="9.[113,1089,1406,1509]" pageId="9" pageNumber="208">
Figs. 35-43.
<taxonomicName id="DC987993FFBEFFC38F8EFA2359FAFA03" authority="Ježek &amp; Harten, 2005" authorityName="Ježek &amp; Harten" authorityYear="2005" box="[229,415,1410,1429]" class="Insecta" family="Psychodidae" genus="Psychana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="208" phylum="Arthropoda" rank="species" species="rujumensis" status="sp. nov.">
<emphasis id="29ECDE02FFBEFFC38F8EFA2359FAFA03" box="[229,415,1410,1429]" italics="true" pageId="9" pageNumber="208">Psychana rujumensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="32DF6379FFBEFFC38ECCFA225995FA00" box="[423,496,1411,1430]" pageId="9" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
, J. 35 head; 36 facets; 37 basal antennomeres; 38 apical antennomeres; 39 wing; 40 dorsal view of fore claw; 41 lateral view of aedeagal compex; 42 lateral view of gonocoxite and gonostyle; 43 dorsal view of epandrium and surstyli. Scales = 0.1 mm (Figs. 35-38, 40-43); 1 mm (Fig. 39).
</paragraph>
</caption>
<paragraph id="1B270210FFBDFFC08FFCFF7E5B45FE8E" blockId="10.[151,1127,223,280]" pageId="10" pageNumber="209">
9.iv.2001, Cat. No. 34272, Inv. No. 14061;
<date id="6F2624D0FFBDFFC08D0CFF7E5B64FF61" box="[615,769,223,247]" pageId="10" pageNumber="209" value="2001-04-09" valueMax="2001-06-05" valueMin="2001-04-09">9.iv.-5.vi.2001</date>
, Cat. No. 34273, Inv. No. 14062. Figures based mainly on the
<typeStatus id="C423BCB2FFBDFFC08EA2FF5E5A4CFE81" box="[457,553,255,279]" pageId="10" pageNumber="209" type="holotype">holotype</typeStatus>
, partially on
<typeStatus id="C423BCB2FFBDFFC08DD8FEA15B79FE8E" box="[691,796,256,280]" pageId="10" pageNumber="209" type="paratype">paratypes</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="5382519BFFBDFFC08FFCFE8F5A27FAF0" pageId="10" pageNumber="209" type="description">
<paragraph id="1B270210FFBDFFC08FFCFE8F5BBCFB50" blockId="10.[151,1128,302,1510]" pageId="10" pageNumber="209">
<emphasis id="29ECDE02FFBDFFC08FFCFE8F5944FED0" bold="true" box="[151,289,302,326]" pageId="10" pageNumber="209">Description.</emphasis>
Male. Eyes separated, C-shaped (
<figureCitation id="83A31E95FFBDFFC08DFEFE8F5A84FED0" box="[661,737,302,326]" captionStart="Figs" captionStartId="9.[113,153,1410,1429]" captionTargetBox="[113,1088,220,1363]" captionTargetId="figure-83@9.[113,1103,205,1363]" captionTargetPageId="9" captionText="Figs. 35-43. Psychana rujumensis sp. nov., J. 35 head; 36 facets; 37 basal antennomeres; 38 apical antennomeres; 39 wing; 40 dorsal view of fore claw; 41 lateral view of aedeagal compex; 42 lateral view of gonocoxite and gonostyle; 43 dorsal view of epandrium and surstyli. Scales = 0.1 mm (Figs. 35-38, 40-43); 1 mm (Fig. 39)." figureDoi="http://doi.org/10.5281/zenodo.5176601" httpUri="https://zenodo.org/record/5176601/files/figure.png" pageId="10" pageNumber="209">Fig 35</figureCitation>
). Minimum distance between upper part of eyes hardly as wide as diameter of facet (
<figureCitation id="83A31E95FFBDFFC08DFDFEEF5A80FEF0" box="[662,741,334,358]" captionStart="Figs" captionStartId="9.[113,153,1410,1429]" captionTargetBox="[113,1088,220,1363]" captionTargetId="figure-83@9.[113,1103,205,1363]" captionTargetPageId="9" captionText="Figs. 35-43. Psychana rujumensis sp. nov., J. 35 head; 36 facets; 37 basal antennomeres; 38 apical antennomeres; 39 wing; 40 dorsal view of fore claw; 41 lateral view of aedeagal compex; 42 lateral view of gonocoxite and gonostyle; 43 dorsal view of epandrium and surstyli. Scales = 0.1 mm (Figs. 35-38, 40-43); 1 mm (Fig. 39)." figureDoi="http://doi.org/10.5281/zenodo.5176601" httpUri="https://zenodo.org/record/5176601/files/figure.png" pageId="10" pageNumber="209">Fig. 36</figureCitation>
). Eye bridge formed by four rows of facets. Frontoclypeus with large central three-lobed patch of insertions of hairs; medial stripe longest, not connected with vertex pits. Antennae 15-segmented (
<figureCitation id="83A31E95FFBDFFC08C31FE2F5BBAFE30" box="[858,991,398,422]" captionStart="Figs" captionStartId="9.[113,153,1410,1429]" captionTargetBox="[113,1088,220,1363]" captionTargetId="figure-83@9.[113,1103,205,1363]" captionTargetPageId="9" captionText="Figs. 35-43. Psychana rujumensis sp. nov., J. 35 head; 36 facets; 37 basal antennomeres; 38 apical antennomeres; 39 wing; 40 dorsal view of fore claw; 41 lateral view of aedeagal compex; 42 lateral view of gonocoxite and gonostyle; 43 dorsal view of epandrium and surstyli. Scales = 0.1 mm (Figs. 35-38, 40-43); 1 mm (Fig. 39)." figureDoi="http://doi.org/10.5281/zenodo.5176601" httpUri="https://zenodo.org/record/5176601/files/figure.png" pageId="10" pageNumber="209">Figs. 37, 38</figureCitation>
); scape very short, wide, hardly globular; flagellomeres pitcher-shaped, symmetrical; last three flagellomeres gradually reduced; antennomere 13 pyriform, antennomere 14 almost globular with very short neck, both segments fused; antennomere 15 minute, conical, entirely separated. Sensory filaments long, paired, with three branches. Length ratio of maxillary palpomeres 1: 1.1: 1: 1.6 (
<figureCitation id="83A31E95FFBDFFC08FF4FD8F588AFDD0" box="[159,239,558,582]" captionStart="Figs" captionStartId="11.[113,152,1439,1458]" captionTargetBox="[121,1084,217,1400]" captionTargetId="figure-18@11.[113,1084,203,1412]" captionTargetPageId="11" captionText="Figs. 44-53. Psychana rujumensis sp. nov., J (44-48) and ♀ (49-53). 44 dorsal view of terminal lobe of labium; 45 maxilla and maxillary palpus; 46 lateral view of thoracic sclerites; 47 dorsal view of aedeagal complex and gonopods; 48 lateral view of epandrium and surstylus; 49 subgenital plate; 50 dorsal view of genital chamber; 51 same, anterior view; 52 same, lateral view; 53 lateral view of cercus. Scales = 0.1 mm (Figs. 44-45, 47-53); 0.5 mm (Fig. 46)." figureDoi="http://doi.org/10.5281/zenodo.5176603" httpUri="https://zenodo.org/record/5176603/files/figure.png" pageId="10" pageNumber="209">Fig. 45</figureCitation>
), palpomere 4 not annulate. Terminal lobe of labium with six digital protuberances (
<figureCitation id="83A31E95FFBDFFC08FF4FDEF5894FDF0" box="[159,241,590,614]" captionStart="Figs" captionStartId="11.[113,152,1439,1458]" captionTargetBox="[121,1084,217,1400]" captionTargetId="figure-18@11.[113,1084,203,1412]" captionTargetPageId="11" captionText="Figs. 44-53. Psychana rujumensis sp. nov., J (44-48) and ♀ (49-53). 44 dorsal view of terminal lobe of labium; 45 maxilla and maxillary palpus; 46 lateral view of thoracic sclerites; 47 dorsal view of aedeagal complex and gonopods; 48 lateral view of epandrium and surstylus; 49 subgenital plate; 50 dorsal view of genital chamber; 51 same, anterior view; 52 same, lateral view; 53 lateral view of cercus. Scales = 0.1 mm (Figs. 44-45, 47-53); 0.5 mm (Fig. 46)." figureDoi="http://doi.org/10.5281/zenodo.5176603" httpUri="https://zenodo.org/record/5176603/files/figure.png" pageId="10" pageNumber="209">Fig. 44</figureCitation>
). Maximum length of cibarium equal to 1.3 times length of epipharynx. Thoracic sclerites as in
<figureCitation id="83A31E95FFBDFFC08E5BFDCF591AFD10" box="[304,383,622,646]" captionStart="Figs" captionStartId="11.[113,152,1439,1458]" captionTargetBox="[121,1084,217,1400]" captionTargetId="figure-18@11.[113,1084,203,1412]" captionTargetPageId="11" captionText="Figs. 44-53. Psychana rujumensis sp. nov., J (44-48) and ♀ (49-53). 44 dorsal view of terminal lobe of labium; 45 maxilla and maxillary palpus; 46 lateral view of thoracic sclerites; 47 dorsal view of aedeagal complex and gonopods; 48 lateral view of epandrium and surstylus; 49 subgenital plate; 50 dorsal view of genital chamber; 51 same, anterior view; 52 same, lateral view; 53 lateral view of cercus. Scales = 0.1 mm (Figs. 44-45, 47-53); 0.5 mm (Fig. 46)." figureDoi="http://doi.org/10.5281/zenodo.5176603" httpUri="https://zenodo.org/record/5176603/files/figure.png" pageId="10" pageNumber="209">Fig. 46</figureCitation>
, meropleural sclerite inconspicuously divided. Wing (
<figureCitation id="83A31E95FFBDFFC08CAAFDCF5C6AFD10" box="[961,1039,622,646]" captionStart="Figs" captionStartId="9.[113,153,1410,1429]" captionTargetBox="[113,1088,220,1363]" captionTargetId="figure-83@9.[113,1103,205,1363]" captionTargetPageId="9" captionText="Figs. 35-43. Psychana rujumensis sp. nov., J. 35 head; 36 facets; 37 basal antennomeres; 38 apical antennomeres; 39 wing; 40 dorsal view of fore claw; 41 lateral view of aedeagal compex; 42 lateral view of gonocoxite and gonostyle; 43 dorsal view of epandrium and surstyli. Scales = 0.1 mm (Figs. 35-38, 40-43); 1 mm (Fig. 39)." figureDoi="http://doi.org/10.5281/zenodo.5176601" httpUri="https://zenodo.org/record/5176601/files/figure.png" pageId="10" pageNumber="209">Fig. 39</figureCitation>
) widely lancet-shaped,
<quantity id="DC60AFF5FFBDFFC08E53FD2F59E9FD30" box="[312,396,654,678]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="10" pageNumber="209" unit="mm" value="1.7">1.7 mm</quantity>
(
<typeStatus id="C423BCB2FFBDFFC08EF7FD2F5999FD30" box="[412,508,654,678]" pageId="10" pageNumber="209" type="holotype">holotype</typeStatus>
) and
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(
<typeStatus id="C423BCB2FFBDFFC08DA2FD2F5B57FD30" box="[713,818,654,678]" pageId="10" pageNumber="209" type="paratype">paratypes</typeStatus>
) long, almost clear, clouded between fore margin and R
<quantity id="DC60AFF5FFBDFFC08ED8FD1E59B0FD50" box="[435,469,686,717]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="10" pageNumber="209" unit="in" value="1.0">1 in</quantity>
small narrow area; following veins or their parts strengthened: Sc (conspicuously), R
<subScript id="871C0055FFBDFFC08EECFD7E59EAFD7B" attach="left" box="[391,399,735,749]" fontSize="6" pageId="10" pageNumber="209">1</subScript>
, R
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, M
<subScript id="871C0055FFBDFFC08E8BFD7E599CFD7B" attach="left" box="[480,505,735,749]" fontSize="6" pageId="10" pageNumber="209">1+2</subScript>
, M
<subScript id="871C0055FFBDFFC08D4BFD7E5A4DFD7B" attach="left" box="[544,552,735,749]" fontSize="6" pageId="10" pageNumber="209">4</subScript>
and Cu; basal costal nodes well visible, Sc uninterrupted; M
<subScript id="871C0055FFBDFFC08E69FD5E596FFC9B" attach="left" box="[258,266,767,781]" fontSize="6" pageId="10" pageNumber="209">3</subScript>
and Cu hardly with connection to M
<subScript id="871C0055FFBDFFC08DFDFD5E5AFBFC9B" attach="left" box="[662,670,767,781]" fontSize="6" pageId="10" pageNumber="209">4</subScript>
; R
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extends in apex of wing. Medial wing angle 79° (BCD); wing indices: AB: AC: AD = 1.4: 2.1: 1.6; BC: CD: BD = 1: 1.1: 1.4; maximum length of wing equal to 2.2 times its maximum width. Maximum length of haltere equal to 2.8 times its maximum width. Ratios of lengths of femora, tibiae and first tarsomeres: P 1 = 2.8: 2.6: 1; P 2 = 2.9: 3.2: 1.2; P 3 = 3.3: 3.8: 1.3. Fore claws as in
<figureCitation id="83A31E95FFBDFFC08CA4FCCF5C45FC10" box="[975,1056,878,902]" captionStart="Figs" captionStartId="9.[113,153,1410,1429]" captionTargetBox="[113,1088,220,1363]" captionTargetId="figure-83@9.[113,1103,205,1363]" captionTargetPageId="9" captionText="Figs. 35-43. Psychana rujumensis sp. nov., J. 35 head; 36 facets; 37 basal antennomeres; 38 apical antennomeres; 39 wing; 40 dorsal view of fore claw; 41 lateral view of aedeagal compex; 42 lateral view of gonocoxite and gonostyle; 43 dorsal view of epandrium and surstyli. Scales = 0.1 mm (Figs. 35-38, 40-43); 1 mm (Fig. 39)." figureDoi="http://doi.org/10.5281/zenodo.5176601" httpUri="https://zenodo.org/record/5176601/files/figure.png" pageId="10" pageNumber="209">Fig. 40</figureCitation>
. Basal apodeme (
<figureCitation id="83A31E95FFBDFFC08E6FFC2F593CFC30" box="[260,345,910,934]" captionStart="Figs" captionStartId="3.[113,152,1081,1100]" captionTargetBox="[113,1088,220,1050]" captionTargetId="figure-170@3.[113,1088,220,1050]" captionTargetPageId="3" captionText="Figs. 1-9. Copropsychoda bulbosa sp. nov., J (1-7) and ♀ (8-9). 1 head; 2 basal antennomeres; 3 dorsal view of terminal lobe of labium; 4 dorsal view of fore claw; 5 dorsal view of aedeagal complex and gonopods; 6 lateral view of gonocoxite and gonostyle; 7 lateral view of epandrium and surstylus; 8 anterior view of genital chamber; 9 lateral view of subgenital plate and cercus. Scales = 0.1 mm (Figs. 1-3, 5-9); 0.05 mm (Fig. 4)." figureDoi="http://doi.org/10.5281/zenodo.5176595" httpUri="https://zenodo.org/record/5176595/files/figure.png" pageId="10" pageNumber="209">Figs. 41</figureCitation>
,
<figureCitation id="83A31E95FFBDFFC08E0DFC2F59E7FC30" box="[358,386,910,934]" captionStart="Figs" captionStartId="11.[113,152,1439,1458]" captionTargetBox="[121,1084,217,1400]" captionTargetId="figure-18@11.[113,1084,203,1412]" captionTargetPageId="11" captionText="Figs. 44-53. Psychana rujumensis sp. nov., J (44-48) and ♀ (49-53). 44 dorsal view of terminal lobe of labium; 45 maxilla and maxillary palpus; 46 lateral view of thoracic sclerites; 47 dorsal view of aedeagal complex and gonopods; 48 lateral view of epandrium and surstylus; 49 subgenital plate; 50 dorsal view of genital chamber; 51 same, anterior view; 52 same, lateral view; 53 lateral view of cercus. Scales = 0.1 mm (Figs. 44-45, 47-53); 0.5 mm (Fig. 46)." figureDoi="http://doi.org/10.5281/zenodo.5176603" httpUri="https://zenodo.org/record/5176603/files/figure.png" pageId="10" pageNumber="209">47</figureCitation>
) of male genitalia bent, narrow from dorsal view, widened from lateral one, pointed proximally; distal part of basal apodeme forked in two caudal arms. Copulatory organ with smooth surface outside, phallomeres of characteristic shape, sclerotized in some parts of lobes. Gonocoxite (
<figureCitation id="83A31E95FFBDFFC08EA0FC4F5A42FB90" box="[459,551,1006,1030]" captionStart="Figs" captionStartId="9.[113,153,1410,1429]" captionTargetBox="[113,1088,220,1363]" captionTargetId="figure-83@9.[113,1103,205,1363]" captionTargetPageId="9" captionText="Figs. 35-43. Psychana rujumensis sp. nov., J. 35 head; 36 facets; 37 basal antennomeres; 38 apical antennomeres; 39 wing; 40 dorsal view of fore claw; 41 lateral view of aedeagal compex; 42 lateral view of gonocoxite and gonostyle; 43 dorsal view of epandrium and surstyli. Scales = 0.1 mm (Figs. 35-38, 40-43); 1 mm (Fig. 39)." figureDoi="http://doi.org/10.5281/zenodo.5176601" httpUri="https://zenodo.org/record/5176601/files/figure.png" pageId="10" pageNumber="209">Figs. 42</figureCitation>
,
<figureCitation id="83A31E95FFBDFFC08D5EFC4F5A34FB90" box="[565,593,1006,1030]" captionStart="Figs" captionStartId="11.[113,152,1439,1458]" captionTargetBox="[121,1084,217,1400]" captionTargetId="figure-18@11.[113,1084,203,1412]" captionTargetPageId="11" captionText="Figs. 44-53. Psychana rujumensis sp. nov., J (44-48) and ♀ (49-53). 44 dorsal view of terminal lobe of labium; 45 maxilla and maxillary palpus; 46 lateral view of thoracic sclerites; 47 dorsal view of aedeagal complex and gonopods; 48 lateral view of epandrium and surstylus; 49 subgenital plate; 50 dorsal view of genital chamber; 51 same, anterior view; 52 same, lateral view; 53 lateral view of cercus. Scales = 0.1 mm (Figs. 44-45, 47-53); 0.5 mm (Fig. 46)." figureDoi="http://doi.org/10.5281/zenodo.5176603" httpUri="https://zenodo.org/record/5176603/files/figure.png" pageId="10" pageNumber="209">47</figureCitation>
) long, cylindrical in lateral view and protruding laterally in dorsal view. Gonostylus somewhat shorter than gonocoxite, with acuminate apex, slightly arched. Epandrium (
<figureCitation id="83A31E95FFBDFFC08EA4FB8F5A4EFBD0" box="[463,555,1070,1094]" captionStart="Figs" captionStartId="9.[113,153,1410,1429]" captionTargetBox="[113,1088,220,1363]" captionTargetId="figure-83@9.[113,1103,205,1363]" captionTargetPageId="9" captionText="Figs. 35-43. Psychana rujumensis sp. nov., J. 35 head; 36 facets; 37 basal antennomeres; 38 apical antennomeres; 39 wing; 40 dorsal view of fore claw; 41 lateral view of aedeagal compex; 42 lateral view of gonocoxite and gonostyle; 43 dorsal view of epandrium and surstyli. Scales = 0.1 mm (Figs. 35-38, 40-43); 1 mm (Fig. 39)." figureDoi="http://doi.org/10.5281/zenodo.5176601" httpUri="https://zenodo.org/record/5176601/files/figure.png" pageId="10" pageNumber="209">Figs. 43</figureCitation>
,
<figureCitation id="83A31E95FFBDFFC08D52FB8F5A30FBD0" box="[569,597,1070,1094]" captionStart="Figs" captionStartId="11.[113,152,1439,1458]" captionTargetBox="[121,1084,217,1400]" captionTargetId="figure-18@11.[113,1084,203,1412]" captionTargetPageId="11" captionText="Figs. 44-53. Psychana rujumensis sp. nov., J (44-48) and ♀ (49-53). 44 dorsal view of terminal lobe of labium; 45 maxilla and maxillary palpus; 46 lateral view of thoracic sclerites; 47 dorsal view of aedeagal complex and gonopods; 48 lateral view of epandrium and surstylus; 49 subgenital plate; 50 dorsal view of genital chamber; 51 same, anterior view; 52 same, lateral view; 53 lateral view of cercus. Scales = 0.1 mm (Figs. 44-45, 47-53); 0.5 mm (Fig. 46)." figureDoi="http://doi.org/10.5281/zenodo.5176603" httpUri="https://zenodo.org/record/5176603/files/figure.png" pageId="10" pageNumber="209">48</figureCitation>
) with two large patches of hair areas connected caudally, inside with two conspicuous spine-shaped protuberances, sclerotized remainders of tergite and sternite 10 missing. Basal aperture developed. Hypandrium (
<figureCitation id="83A31E95FFBDFFC08CE0FBCF5B85FB10" box="[907,992,1134,1158]" captionStart="Figs" captionStartId="3.[113,152,1081,1100]" captionTargetBox="[113,1088,220,1050]" captionTargetId="figure-170@3.[113,1088,220,1050]" captionTargetPageId="3" captionText="Figs. 1-9. Copropsychoda bulbosa sp. nov., J (1-7) and ♀ (8-9). 1 head; 2 basal antennomeres; 3 dorsal view of terminal lobe of labium; 4 dorsal view of fore claw; 5 dorsal view of aedeagal complex and gonopods; 6 lateral view of gonocoxite and gonostyle; 7 lateral view of epandrium and surstylus; 8 anterior view of genital chamber; 9 lateral view of subgenital plate and cercus. Scales = 0.1 mm (Figs. 1-3, 5-9); 0.05 mm (Fig. 4)." figureDoi="http://doi.org/10.5281/zenodo.5176595" httpUri="https://zenodo.org/record/5176595/files/figure.png" pageId="10" pageNumber="209">Figs. 41</figureCitation>
,
<figureCitation id="83A31E95FFBDFFC08C86FBCF5C6CFB10" box="[1005,1033,1134,1158]" captionStart="Figs" captionStartId="11.[113,152,1439,1458]" captionTargetBox="[121,1084,217,1400]" captionTargetId="figure-18@11.[113,1084,203,1412]" captionTargetPageId="11" captionText="Figs. 44-53. Psychana rujumensis sp. nov., J (44-48) and ♀ (49-53). 44 dorsal view of terminal lobe of labium; 45 maxilla and maxillary palpus; 46 lateral view of thoracic sclerites; 47 dorsal view of aedeagal complex and gonopods; 48 lateral view of epandrium and surstylus; 49 subgenital plate; 50 dorsal view of genital chamber; 51 same, anterior view; 52 same, lateral view; 53 lateral view of cercus. Scales = 0.1 mm (Figs. 44-45, 47-53); 0.5 mm (Fig. 46)." figureDoi="http://doi.org/10.5281/zenodo.5176603" httpUri="https://zenodo.org/record/5176603/files/figure.png" pageId="10" pageNumber="209">47</figureCitation>
) narrow, bare, slightly widened in middle. Epiproct and hypoproct small (
<figureCitation id="83A31E95FFBDFFC08C5EFB2F5BE9FB30" box="[821,908,1166,1190]" captionStart="Figs" captionStartId="9.[113,153,1410,1429]" captionTargetBox="[113,1088,220,1363]" captionTargetId="figure-83@9.[113,1103,205,1363]" captionTargetPageId="9" captionText="Figs. 35-43. Psychana rujumensis sp. nov., J. 35 head; 36 facets; 37 basal antennomeres; 38 apical antennomeres; 39 wing; 40 dorsal view of fore claw; 41 lateral view of aedeagal compex; 42 lateral view of gonocoxite and gonostyle; 43 dorsal view of epandrium and surstyli. Scales = 0.1 mm (Figs. 35-38, 40-43); 1 mm (Fig. 39)." figureDoi="http://doi.org/10.5281/zenodo.5176601" httpUri="https://zenodo.org/record/5176601/files/figure.png" pageId="10" pageNumber="209">Figs. 43</figureCitation>
,
<figureCitation id="83A31E95FFBDFFC08CFEFB2F5BD7FB30" box="[917,946,1166,1190]" captionStart="Figs" captionStartId="11.[113,152,1439,1458]" captionTargetBox="[121,1084,217,1400]" captionTargetId="figure-18@11.[113,1084,203,1412]" captionTargetPageId="11" captionText="Figs. 44-53. Psychana rujumensis sp. nov., J (44-48) and ♀ (49-53). 44 dorsal view of terminal lobe of labium; 45 maxilla and maxillary palpus; 46 lateral view of thoracic sclerites; 47 dorsal view of aedeagal complex and gonopods; 48 lateral view of epandrium and surstylus; 49 subgenital plate; 50 dorsal view of genital chamber; 51 same, anterior view; 52 same, lateral view; 53 lateral view of cercus. Scales = 0.1 mm (Figs. 44-45, 47-53); 0.5 mm (Fig. 46)." figureDoi="http://doi.org/10.5281/zenodo.5176603" httpUri="https://zenodo.org/record/5176603/files/figure.png" pageId="10" pageNumber="209">48</figureCitation>
), hairy. Surstylus long, C-shaped, twice as long as epandrium, subapically with one retinaculum.
</paragraph>
<paragraph id="1B270210FFBDFFC08FDDFB6F5A27FAF0" blockId="10.[151,1128,302,1510]" pageId="10" pageNumber="209">
Female. Wing 1.6-2.0 mm long. Subgenital plate (
<figureCitation id="83A31E95FFBDFFC08DBBFB6F5B7AFB70" box="[720,799,1230,1254]" captionStart="Figs" captionStartId="11.[113,152,1439,1458]" captionTargetBox="[121,1084,217,1400]" captionTargetId="figure-18@11.[113,1084,203,1412]" captionTargetPageId="11" captionText="Figs. 44-53. Psychana rujumensis sp. nov., J (44-48) and ♀ (49-53). 44 dorsal view of terminal lobe of labium; 45 maxilla and maxillary palpus; 46 lateral view of thoracic sclerites; 47 dorsal view of aedeagal complex and gonopods; 48 lateral view of epandrium and surstylus; 49 subgenital plate; 50 dorsal view of genital chamber; 51 same, anterior view; 52 same, lateral view; 53 lateral view of cercus. Scales = 0.1 mm (Figs. 44-45, 47-53); 0.5 mm (Fig. 46)." figureDoi="http://doi.org/10.5281/zenodo.5176603" httpUri="https://zenodo.org/record/5176603/files/figure.png" pageId="10" pageNumber="209">Fig. 49</figureCitation>
) of characteristic shape, hairy, with large bilobed sclerotized basis; two small caudal lobuli separated by medial cleft. Sensory digit developed. Mesh-like structures of complicated sclerotized forms in the area of genital chamber not developed (
<figureCitation id="83A31E95FFBDFFC08E9BFA8F5A2FFAD0" box="[496,586,1326,1350]" captionStart="Figs" captionStartId="11.[113,152,1439,1458]" captionTargetBox="[121,1084,217,1400]" captionTargetId="figure-18@11.[113,1084,203,1412]" captionTargetPageId="11" captionText="Figs. 44-53. Psychana rujumensis sp. nov., J (44-48) and ♀ (49-53). 44 dorsal view of terminal lobe of labium; 45 maxilla and maxillary palpus; 46 lateral view of thoracic sclerites; 47 dorsal view of aedeagal complex and gonopods; 48 lateral view of epandrium and surstylus; 49 subgenital plate; 50 dorsal view of genital chamber; 51 same, anterior view; 52 same, lateral view; 53 lateral view of cercus. Scales = 0.1 mm (Figs. 44-45, 47-53); 0.5 mm (Fig. 46)." figureDoi="http://doi.org/10.5281/zenodo.5176603" httpUri="https://zenodo.org/record/5176603/files/figure.png" pageId="10" pageNumber="209">Figs. 50</figureCitation>
,
<figureCitation id="83A31E95FFBDFFC08D3DFA8F5A15FAD0" box="[598,624,1326,1350]" captionStart="Figs" captionStartId="3.[113,152,1081,1100]" captionTargetBox="[113,1088,220,1050]" captionTargetId="figure-170@3.[113,1088,220,1050]" captionTargetPageId="3" captionText="Figs. 1-9. Copropsychoda bulbosa sp. nov., J (1-7) and ♀ (8-9). 1 head; 2 basal antennomeres; 3 dorsal view of terminal lobe of labium; 4 dorsal view of fore claw; 5 dorsal view of aedeagal complex and gonopods; 6 lateral view of gonocoxite and gonostyle; 7 lateral view of epandrium and surstylus; 8 anterior view of genital chamber; 9 lateral view of subgenital plate and cercus. Scales = 0.1 mm (Figs. 1-3, 5-9); 0.05 mm (Fig. 4)." figureDoi="http://doi.org/10.5281/zenodo.5176595" httpUri="https://zenodo.org/record/5176595/files/figure.png" pageId="10" pageNumber="209">51</figureCitation>
,
<figureCitation id="83A31E95FFBDFFC08D14FA8F5AF8FAD0" box="[639,669,1326,1350]" captionStart="Figs" captionStartId="11.[113,152,1439,1458]" captionTargetBox="[121,1084,217,1400]" captionTargetId="figure-18@11.[113,1084,203,1412]" captionTargetPageId="11" captionText="Figs. 44-53. Psychana rujumensis sp. nov., J (44-48) and ♀ (49-53). 44 dorsal view of terminal lobe of labium; 45 maxilla and maxillary palpus; 46 lateral view of thoracic sclerites; 47 dorsal view of aedeagal complex and gonopods; 48 lateral view of epandrium and surstylus; 49 subgenital plate; 50 dorsal view of genital chamber; 51 same, anterior view; 52 same, lateral view; 53 lateral view of cercus. Scales = 0.1 mm (Figs. 44-45, 47-53); 0.5 mm (Fig. 46)." figureDoi="http://doi.org/10.5281/zenodo.5176603" httpUri="https://zenodo.org/record/5176603/files/figure.png" pageId="10" pageNumber="209">52</figureCitation>
). Cercus (
<figureCitation id="83A31E95FFBDFFC08C61FA8F5B3DFAD0" box="[778,856,1326,1350]" captionStart="Figs" captionStartId="11.[113,152,1439,1458]" captionTargetBox="[121,1084,217,1400]" captionTargetId="figure-18@11.[113,1084,203,1412]" captionTargetPageId="11" captionText="Figs. 44-53. Psychana rujumensis sp. nov., J (44-48) and ♀ (49-53). 44 dorsal view of terminal lobe of labium; 45 maxilla and maxillary palpus; 46 lateral view of thoracic sclerites; 47 dorsal view of aedeagal complex and gonopods; 48 lateral view of epandrium and surstylus; 49 subgenital plate; 50 dorsal view of genital chamber; 51 same, anterior view; 52 same, lateral view; 53 lateral view of cercus. Scales = 0.1 mm (Figs. 44-45, 47-53); 0.5 mm (Fig. 46)." figureDoi="http://doi.org/10.5281/zenodo.5176603" httpUri="https://zenodo.org/record/5176603/files/figure.png" pageId="10" pageNumber="209">Fig. 53</figureCitation>
) slightly bent, more then three times as long as its basal diameter.
</paragraph>
</subSubSection>
<subSubSection id="5382519BFFBDFFC68FFCFACF59B1FEE1" lastPageId="12" lastPageNumber="211" pageId="10" pageNumber="209" type="diagnosis">
<paragraph id="1B270210FFBDFFC68FFCFACF59B1FEE1" blockId="10.[151,1128,302,1510]" lastBlockId="12.[151,1127,223,503]" lastPageId="12" lastPageNumber="211" pageId="10" pageNumber="209">
<emphasis id="29ECDE02FFBDFFC08FFCFACF59F4FA10" bold="true" box="[151,401,1390,1414]" pageId="10" pageNumber="209">Differential diagnosis.</emphasis>
The new species is similar to
<taxonomicName id="DC987993FFBDFFC08DB7FACF5C20FA10" authority="Quate, 1965" authorityName="Quate" authorityYear="1965" box="[732,1093,1390,1414]" class="Insecta" family="Psychodidae" genus="Psychana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="209" phylum="Arthropoda" rank="species" species="umbratica">
<emphasis id="29ECDE02FFBDFFC08DB7FACF5BDDFA10" box="[732,952,1390,1414]" italics="true" pageId="10" pageNumber="209">Psychana umbratica</emphasis>
<bibRefCitation id="7F097FE1FFBDFFC08CABFACF5C20FA10" author="QUATE L. W." box="[960,1093,1390,1414]" pageId="10" pageNumber="209" pagination="815 - 902" refId="ref10464" refString="QUATE L. W. 1965: A taxonomic study of Philippine Psychodidae (Diptera). Pacific Insects 7: 815 - 902." type="journal article" year="1965">Quate, 1965</bibRefCitation>
</taxonomicName>
, in the shape of the terminal flagellomere, the form of male gonopode as well as surstylus and the shape of female subgenital plate, but is quite different in the structures of male genitalia.
<taxonomicName id="DC987993FFBDFFC08FFCFA6F591FFA70" authority="Ježek &amp; Harten, 2005" authorityName="Ježek &amp; Harten" authorityYear="2005" box="[151,378,1486,1510]" class="Insecta" family="Psychodidae" genus="Psychana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="209" phylum="Arthropoda" rank="species" species="rujumensis" status="sp. nov.">
<emphasis id="29ECDE02FFBDFFC08FFCFA6F591FFA70" box="[151,378,1486,1510]" italics="true" pageId="10" pageNumber="209">Psychana rujumensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="32DF6379FFBDFFC08EE9FA6E59B0FA70" box="[386,469,1487,1510]" pageId="10" pageNumber="209" rank="species">sp. nov.</taxonomicNameLabel>
is diagnosed by the conspicuously developed basal apodeme and ventral phallomere, short and partially fused dorsal phallomeres, and conspicuously sclerotized sides of the base of female subgenital plate (present as largely widened patches).
<taxonomicName id="DC987993FFBBFFC68FFCFEBE5912FEA1" authorityName="Quate" authorityYear="1965" box="[151,375,287,311]" class="Insecta" family="Psychodidae" genus="Psychana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="211" phylum="Arthropoda" rank="species" species="umbratica">
<emphasis id="29ECDE02FFBBFFC68FFCFEBE5912FEA1" box="[151,375,287,311]" italics="true" pageId="12" pageNumber="211">Psychana umbratica</emphasis>
</taxonomicName>
has a rudimentary basal apodeme of the male genitalia and ventral phallomere, very long and entirely fused dorsal paired phallomeres, and a narrowly sclerotized base of subgenital plate.
</paragraph>
</subSubSection>
<caption id="4FE75298FFBCFFC18F1AFA3E596EF98A" ID-DOI="http://doi.org/10.5281/zenodo.5176603" ID-Zenodo-Dep="5176603" httpUri="https://zenodo.org/record/5176603/files/figure.png" pageId="11" pageNumber="210" startId="11.[113,152,1439,1458]" targetBox="[121,1084,217,1400]" targetPageId="11">
<paragraph id="1B270210FFBCFFC18F1AFA3E596EF98A" blockId="11.[113,1090,1435,1565]" pageId="11" pageNumber="210">
Figs. 44-53.
<taxonomicName id="DC987993FFBCFFC18FB5FA3E59F6FA27" authority="Ježek &amp; Harten, 2005" authorityName="Ježek &amp; Harten" authorityYear="2005" box="[222,403,1439,1458]" class="Insecta" family="Psychodidae" genus="Psychana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="210" phylum="Arthropoda" rank="species" species="rujumensis" status="sp. nov.">
<emphasis id="29ECDE02FFBCFFC18FB5FA3E59F6FA27" box="[222,403,1439,1458]" italics="true" pageId="11" pageNumber="210">Psychana rujumensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="32DF6379FFBCFFC18EF3FA3E59B8FA24" box="[408,477,1439,1458]" pageId="11" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
, J (44-48) and ♀ (49-53). 44 dorsal view of terminal lobe of labium; 45 maxilla and maxillary palpus; 46 lateral view of thoracic sclerites; 47 dorsal view of aedeagal complex and gonopods; 48 lateral view of epandrium and surstylus; 49 subgenital plate; 50 dorsal view of genital chamber; 51 same, anterior view; 52 same, lateral view; 53 lateral view of cercus. Scales = 0.1 mm (Figs. 44-45, 47-53); 0.5 mm (Fig. 46).
</paragraph>
</caption>
<subSubSection id="5382519BFFBBFFC68FFCFEDE5A3CFE01" box="[151,601,383,408]" pageId="12" pageNumber="211" type="etymology">
<paragraph id="1B270210FFBBFFC68FFCFEDE5A3CFE01" blockId="12.[151,1127,223,503]" box="[151,601,383,408]" pageId="12" pageNumber="211">
<emphasis id="29ECDE02FFBBFFC68FFCFEDE5972FE01" bold="true" box="[151,279,383,407]" pageId="12" pageNumber="211">Etymology.</emphasis>
Named after the
<typeStatus id="C423BCB2FFBBFFC68EA4FE215998FE0E" box="[463,509,384,408]" pageId="12" pageNumber="211">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="5382519BFFBBFFC68FFCFE3E58AAFE61" pageId="12" pageNumber="211" type="distribution">
<paragraph id="1B270210FFBBFFC68FFCFE3E59E5FE21" blockId="12.[151,1127,223,503]" box="[151,384,415,439]" pageId="12" pageNumber="211">
<emphasis id="29ECDE02FFBBFFC68FFCFE3E594DFE21" bold="true" box="[151,296,415,439]" pageId="12" pageNumber="211">Distribution.</emphasis>
<collectingCountry id="638F4280FFBBFFC68E45FE01591EFE21" box="[302,379,416,439]" name="Yemen" pageId="12" pageNumber="211">Yemen</collectingCountry>
.
</paragraph>
<paragraph id="1B270210FFBBFFC68FFCFE1E58AAFE61" blockId="12.[151,1127,223,503]" pageId="12" pageNumber="211">
<emphasis id="29ECDE02FFBBFFC68FFCFE1E5967FE41" bold="true" box="[151,258,447,471]" pageId="12" pageNumber="211">Bionomy.</emphasis>
Unknown. The adults were trapped in a Malaise trap at Ar Rujum from October to June.
</paragraph>
</subSubSection>
</treatment>
</document>