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<document id="5E69E8284ACC965C07C72BF7F2A41EF3" ID-CLB-Dataset="6318" ID-DOI="10.11646/zootaxa.4970.2.1" ID-GBIF-Dataset="89d3c795-198b-4162-8903-9c6cf3f1e665" ID-ISSN="1175-5326" ID-Zenodo-Dep="4761544" ID-ZooBank="6DBA0C02-9E52-4FD0-8B48-8C7B405713A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620984897660" checkinUser="plazi" docAuthor="Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M. &amp; Brescovit, Antonio Domingos" docDate="2021" docId="052487CFFFC1E52946C6B2C68F8DFA64" docLanguage="en" docName="zootaxa.4970.2.1.pdf" docOrigin="Zootaxa 4970 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Arthrosolaenomeris Schubart 1943" docType="treatment" docVersion="7" lastPageNumber="212" masterDocId="F91DFFB7FFCAE5224651B2518D76FFF7" masterDocTitle="Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae)" masterLastPageNumber="256" masterPageNumber="201" pageNumber="212" updateTime="1698939836712" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2EC01596E6D0F79EED349708DFD14517">Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae)</mods:title>
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<mods:namePart id="671C96770DFBA9A547AE4EDF929AF712">Bouzan, Rodrigo Salvador</mods:namePart>
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<mods:namePart id="B85F0C30DAC5C7D9DA88D6D346AB2BAB">Iniesta, Luiz Felipe M.</mods:namePart>
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<mods:affiliation id="788F0D4A47C1708757E77E4AEFD182B2">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brasil. &amp; luiz-moretti @ hotmail. com; https: // orcid. org / 0000 - 0002 - 0529 - 4162</mods:affiliation>
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<taxonomicName id="4A8D4D5AFFC1E52946C6B2C68F3BFF46" ID-CoL="7NR8T" authority="Schubart, 1943" authorityName="Schubart" authorityYear="1943" box="[151,589,151,178]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="genus">
<emphasis id="BFF9EACBFFC1E52946C6B2C68F3BFF46" bold="true" box="[151,589,151,178]" pageId="11" pageNumber="212">
<emphasis id="BFF9EACBFFC1E52946C6B2C68CFEFF45" bold="true" box="[151,392,151,178]" italics="true" pageId="11" pageNumber="212">Arthrosolaenomeris</emphasis>
<bibRefCitation id="E91C4B28FFC1E52947C1B2C68F3BFF46" author="Schubart, O." box="[400,589,151,177]" pageId="11" pageNumber="212" pagination="127 - 164" refId="ref33096" refString="Schubart, O. (1943) Especies novas das familias Strongylosomidae e Leptodesmidae da ordem Proterospermophora do interior dos Estados de Sao Paulo e Mato Grosso. Papeis Avulsos de Zoologia, 3, 127 - 164." type="journal article" year="1943">Schubart, 1943</bibRefCitation>
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<treatmentCitation id="0C2C10C8FFC1E52946C6B28D8F42FF03" author="Schubart, O." box="[151,564,220,244]" page="143" pageId="11" pageNumber="212" year="1943">
<taxonomicName id="4A8D4D5AFFC1E52946C6B28D8F42FF03" ID-CoL="7NR8T" authority="Schubart, 1943: 143" authorityName="Schubart" authorityPageNumber="143" authorityYear="1943" box="[151,564,220,244]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="genus">
<emphasis id="BFF9EACBFFC1E52946C6B28D8C17FF03" box="[151,353,220,244]" italics="true" pageId="11" pageNumber="212">Arthrosolaenomeris</emphasis>
<bibRefCitation id="E91C4B28FFC1E5294737B28D8F42FF03" author="Schubart, O." box="[358,564,220,244]" pageId="11" pageNumber="212" pagination="127 - 164" refId="ref33096" refString="Schubart, O. (1943) Especies novas das familias Strongylosomidae e Leptodesmidae da ordem Proterospermophora do interior dos Estados de Sao Paulo e Mato Grosso. Papeis Avulsos de Zoologia, 3, 127 - 164." type="journal article" year="1943">Schubart, 1943: 143</bibRefCitation>
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.
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<typeStatus id="5236887BFFC1E529446DB28D8F18FF03" box="[572,622,220,244]" pageId="11" pageNumber="212">Type</typeStatus>
species:
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<emphasis id="BFF9EACBFFC1E5294499B28D8E16FF03" box="[712,864,220,244]" italics="true" pageId="11" pageNumber="212">A. chapadensis</emphasis>
<bibRefCitation id="E91C4B28FFC1E5294537B28D8976FF03" author="Schubart, O." box="[870,1024,220,244]" pageId="11" pageNumber="212" pagination="127 - 164" refId="ref33096" refString="Schubart, O. (1943) Especies novas das familias Strongylosomidae e Leptodesmidae da ordem Proterospermophora do interior dos Estados de Sao Paulo e Mato Grosso. Papeis Avulsos de Zoologia, 3, 127 - 164." type="journal article" year="1943">Schubart, 1943</bibRefCitation>
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, by original designation;
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<bibRefCitation id="E91C4B28FFC1E52942ACB28D8D86FEE5" author="Schubart, O." pageId="11" pageNumber="212" pagination="203 - 252" refId="ref33204" refString="Schubart, O. (1958) Sobre alguns Diplopoda de Mato Grosso e Goias, Brasil e a familia Spirostreptidae. Arquivos do Museu Nacional, 46, 203 - 252." type="journal article" year="1958">Schubart, 1958: 205</bibRefCitation>
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;
<treatmentCitation id="0C2C10C8FFC1E52946AAB2AB8CBCFEE5" author="Schubart, O." box="[251,458,250,274]" page="453" pageId="11" pageNumber="212" year="1960">
<bibRefCitation id="E91C4B28FFC1E52946AAB2AB8CBCFEE5" author="Schubart, O." box="[251,458,250,274]" pageId="11" pageNumber="212" pagination="453 - 464" refId="ref33237" refString="Schubart, O. (1960) Leptodesmidae Brasileiras. VIII. Novas especies do Estado de Sao Paulo (Diplopoda, Proterospermophora). Revista Brasileira de Biologia, 20 (4), 453 - 464." type="journal article" year="1960">Schubart, 1960: 453</bibRefCitation>
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;
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<bibRefCitation id="E91C4B28FFC1E5294784B2AB8FFCFEE5" author="Jeekel, C. A. W." box="[469,650,250,274]" pageId="11" pageNumber="212" pagination="1 - 412" refId="ref32646" refString="Jeekel, C. A. W. (1971) Nomenclator generum et familiarum Diplopodorum: A list of the genus and family-group names in the Class Diplopoda from the 10 th edition of Linnaeus, 1758, to the end of 1957. Monografieen van de Nederlandse Entomologische Vereniging, 5, 1 - 412." type="journal article" year="1971">Jeekel, 1971: 249</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0C2C10C8FFC1E52944C5B2AB8E13FEE5" author="Hoffman, R. L." box="[660,869,250,274]" page="177" pageId="11" pageNumber="212" year="1976">
<bibRefCitation id="E91C4B28FFC1E52944C5B2AB8E13FEE5" author="Hoffman, R. L." box="[660,869,250,274]" pageId="11" pageNumber="212" pagination="171 - 183" refId="ref32248" refString="Hoffman, R. L. (1976) Chelodesmid studies IX. A synopsis of the new Brazilian tribe Arthrosolaenomeridini (Diplopoda; Polydesmida). Papeis Avulsos de Zoologia, 30, 171 - 183." type="journal article" year="1976">Hoffman, 1976: 177</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0C2C10C8FFC1E529453EB2AB8936FEE5" author="Hoffman, R. L." box="[879,1088,250,274]" page="151" pageId="11" pageNumber="212" year="1980">
<bibRefCitation id="E91C4B28FFC1E529453EB2AB8936FEE5" author="Hoffman, R. L." box="[879,1088,250,274]" pageId="11" pageNumber="212" refId="ref32397" refString="Hoffman, R. L. (1980) Classification of the Diplopoda. Museum d'histoire naturelle, Geneve, 237 pp." type="book" year="1980">Hoffman, 1980: 151</bibRefCitation>
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.
</paragraph>
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<paragraph id="8D3236D9FFC1E52946C6B3118F0EFDF8" blockId="11.[151,1437,320,1427]" pageId="11" pageNumber="212">
<emphasis id="BFF9EACBFFC1E52946C6B3118C7CFEAD" bold="true" box="[151,266,320,346]" pageId="11" pageNumber="212">Diagnosis</emphasis>
. Males of
<taxonomicName id="4A8D4D5AFFC1E529472AB3118F2CFEAD" authorityName="Schubart" authorityYear="1943" box="[379,602,320,346]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="genus">
<emphasis id="BFF9EACBFFC1E529472AB3118F2CFEAD" box="[379,602,320,346]" italics="true" pageId="11" pageNumber="212">Arthrosolaenomeris</emphasis>
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differ from those of other
<taxonomicName id="4A8D4D5AFFC1E5294524B3118902FEAD" authorityName="Hoffman" authorityYear="1976" box="[885,1140,320,346]" class="Diplopoda" family="Chelodesmidae" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="tribe" tribe="Arthrosolaenomeridini">Arthrosolaenomeridini</taxonomicName>
genera by the combination of the following characters: rectangular ventral projections in the post-gonopodal sternites (
<figureCitation id="15B62A5CFFC1E52942CEB3358874FE88" box="[1183,1282,356,383]" captionStart="FIGURE 30" captionStartId="47.[151,250,1703,1727]" captionTargetBox="[161,1426,181,1679]" captionTargetId="figure-20@47.[161,1426,181,1679]" captionTargetPageId="47" captionText="FIGURE 30. Sternites, ventral projections. A) Arthrosolaenomeris pantanalensis Schubart, 1943 (IBSP 530), 6th body ring; B) Dioplosternus salvatrix Hoffman, 2005 (IBSP 1084), 6th body ring; C) Arthrosolaenomeris saci n. sp. (IBSP 1148), 7th body ring; D) Folcloredesmus thomasi n. sp. (CZUFMT 833), 9th body ring; E) A. saci n. sp., 9th body ring; F) Abiliodesmus planaltensis (Schubart, 1960) comb. nov. (IBSP 3570), 9th body ring. Scale bars: AC, 1 mm.; D, 0,5 mm.; E, 0,3 mm.; F, 0,3 mm." figureDoi="http://doi.org/10.5281/zenodo.4761651" httpUri="https://zenodo.org/record/4761651/files/figure.png" pageId="11" pageNumber="212">Fig. 30E</figureCitation>
) (acuminated in
<taxonomicName id="4A8D4D5AFFC1E52946E5B3D88C43FE55" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[180,309,393,419]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="curupira" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E52946E5B3D88C43FE55" box="[180,309,393,419]" italics="true" pageId="11" pageNumber="212">A. curupira</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E529476AB3D88C0CFE54" bold="true" box="[315,378,393,419]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E529476AB3D88C0CFE54" box="[315,378,393,419]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4A8D4D5AFFC1E52947E1B3D88F50FE55" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[432,550,393,419]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="caipora" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E52947E1B3D88F50FE55" box="[432,550,393,419]" italics="true" pageId="11" pageNumber="212">A. caipora</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E529447CB3D88F1DFE54" bold="true" box="[557,619,393,419]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E529447CB3D88F1DFE54" box="[557,619,393,419]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
); posterior edge of the gonopod aperture with a small dentiform process; presence of a secondary process in medial portion in the prefemoral process of the gonopod (
<figureCitation id="15B62A5CFFC1E5294292B3FD884DFE30" box="[1219,1339,428,455]" captionStart="FIGURE 4" captionStartId="13.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@13.[161,1426,181,1680]" captionTargetPageId="13" captionText="FIGURE 4. Arthrosolaenomeris chapadensis Schubart, 1943 (holotype, MZSP 1092), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761555" httpUri="https://zenodo.org/record/4761555/files/figure.png" pageId="11" pageNumber="212">Fig. 4DF</figureCitation>
) (absent in
<taxonomicName id="4A8D4D5AFFC1E52946E6B3808C7EFE1D" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[183,264,465,491]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="iara" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E52946E6B3808C7EFE1D" box="[183,264,465,491]" italics="true" pageId="11" pageNumber="212">A. iara</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E5294743B3808C25FE1C" bold="true" box="[274,339,465,491]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E5294743B3808C25FE1C" box="[274,339,465,491]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
); presence of a rounded extension of the gonocoxae, in oral view (
<figureCitation id="15B62A5CFFC1E5294234B38189A7FE1C" box="[1125,1233,464,491]" captionStart="FIGURE 32" captionStartId="49.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@49.[161,1426,181,1680]" captionTargetPageId="49" captionText="FIGURE 32. Gonocoxae.A) Arthrosolaenomeris pantanalensis Schubart, 1943 (IBSP 530); B) Gangugia tapirapensis Schubart, 1947 (MNRJ 12008); C) Folcloredesmus thomasi n. sp. (CZUFMT 833); D) A. pantanalensis Schubart, 1943 (IBSP 530); E) F. thomasi n. sp. (CZUFMT 833); F) Abiliodesmus mapinguari n. sp. (CZUFMT 836). Scale bars: AB, DF) 1 mm.; C) 0,2 mm. Abbreviations: Cx, gonocoxae; RE, rounded extension; SP, spiniform process." figureDoi="http://doi.org/10.5281/zenodo.4761655" httpUri="https://zenodo.org/record/4761655/files/figure.png" pageId="11" pageNumber="212">Fig. 32D</figureCitation>
); and apex of the solenomere abruptly descending (
<figureCitation id="15B62A5CFFC1E5294440B3A58F1AFDF8" box="[529,620,500,527]" captionStart="FIGURE 4" captionStartId="13.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@13.[161,1426,181,1680]" captionTargetPageId="13" captionText="FIGURE 4. Arthrosolaenomeris chapadensis Schubart, 1943 (holotype, MZSP 1092), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761555" httpUri="https://zenodo.org/record/4761555/files/figure.png" pageId="11" pageNumber="212">Fig. 4D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C5976552FFC1E5294696B0498C4FFAD0" pageId="11" pageNumber="212" type="description">
<paragraph id="8D3236D9FFC1E5294696B04988F0FC14" blockId="11.[151,1437,320,1427]" pageId="11" pageNumber="212">
<emphasis id="BFF9EACBFFC1E5294696B0498C2FFDC5" bold="true" box="[199,345,536,562]" pageId="11" pageNumber="212">Description.</emphasis>
<emphasis id="BFF9EACBFFC1E529470EB0498F4FFDC5" box="[351,569,536,562]" italics="true" pageId="11" pageNumber="212">General characters</emphasis>
: Body length between
<quantity id="4A759B3CFFC1E5294568B0488EDDFDC4" box="[825,939,537,563]" metricMagnitude="-2" metricUnit="m" metricValue="4.778" pageId="11" pageNumber="212" unit="mm" value="47.78">47.78 mm</quantity>
(
<taxonomicName id="4A8D4D5AFFC1E52945EBB0488959FDC5" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[954,1071,537,563]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="caipora" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E52945EBB0488959FDC5" box="[954,1071,537,563]" italics="true" pageId="11" pageNumber="212">A. caipora</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E5294264B0488905FDC4" bold="true" box="[1077,1139,537,563]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E5294264B0488905FDC4" box="[1077,1139,537,563]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
) and
<quantity id="4A759B3CFFC1E52942E3B0498855FDC4" box="[1202,1315,536,563]" metricMagnitude="-2" metricUnit="m" metricValue="6.942" pageId="11" pageNumber="212" unit="mm" value="69.42">69.42 mm</quantity>
(
<taxonomicName id="4A8D4D5AFFC1E5294363B04888F6FDC5" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[1330,1408,537,563]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="iara" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E5294363B04888F6FDC5" box="[1330,1408,537,563]" italics="true" pageId="11" pageNumber="212">A. iara</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E52943D7B0488DCEFDA0" bold="true" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E52943D7B0488DCEFDA0" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
). Coloration (specimens long preserved in 70% ethanol) variable between species, ranging from ocher to reddish brown, paranota tip and posterior border of the metaterga slightly whitish (some specimens of
<taxonomicName id="4A8D4D5AFFC1E52942F2B030882AFD8D" authorityName="Schubart" authorityYear="1943" box="[1187,1372,608,635]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="pantanalensis">
<emphasis id="BFF9EACBFFC1E52942F2B030882AFD8D" box="[1187,1372,608,635]" italics="true" pageId="11" pageNumber="212">A. pantanalensis</emphasis>
</taxonomicName>
show a purple coloration for the whole body). Gnathochilarium: lingual plate covered by several setae; mentum centrally covered by setae, with smooth edges and stipes covered by thick setae (sparsely covered in
<taxonomicName id="4A8D4D5AFFC1E52942DAB0F8887DFD35" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[1163,1291,681,707]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="curupira" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E52942DAB0F8887DFD35" box="[1163,1291,681,707]" italics="true" pageId="11" pageNumber="212">A. curupira</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E5294340B0F88839FD34" bold="true" box="[1297,1359,681,707]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E5294340B0F88839FD34" box="[1297,1359,681,707]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4A8D4D5AFFC1E52943D5B0F88D98FD11" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="caipora" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E52943D5B0F88D98FD11" italics="true" pageId="11" pageNumber="212">A. caipora</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E52946A5B09C8C44FD10" bold="true" box="[244,306,717,743]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E52946A5B09C8C44FD10" box="[244,306,717,743]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
). Body rings: cuticle slightly rough; alignment of paranota in posterior view curved ventrally (straight in
<taxonomicName id="4A8D4D5AFFC1E52946E5B0A08C5FFCFD" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[180,297,753,779]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="caipora" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E52946E5B0A08C5FFCFD" box="[180,297,753,779]" italics="true" pageId="11" pageNumber="212">A. caipora</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E5294761B0A08C18FCFC" bold="true" box="[304,366,753,779]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E5294761B0A08C18FCFC" box="[304,366,753,779]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
). Sternite of the fourth segment with a pair of projections, sternite of the fifth segment with two pairs of projections covered with setae. Sternite of sixth body ring with two pairs of ventral projections; seventh body ring with a pair of ventral projections (absent in
<taxonomicName id="4A8D4D5AFFC1E52944A6B1688ED7FCA5" authorityName="Schubart" authorityYear="1943" box="[759,929,824,851]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="chapadensis">
<emphasis id="BFF9EACBFFC1E52944A6B1688ED7FCA5" box="[759,929,824,851]" italics="true" pageId="11" pageNumber="212">A. chapadensis</emphasis>
</taxonomicName>
) and post-gonopodal sternites with two pairs of rectangular ventral projections (acuminate in
<taxonomicName id="4A8D4D5AFFC1E52944E6B10C8E4FFC81" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[695,825,861,887]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="curupira" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E52944E6B10C8E4FFC81" box="[695,825,861,887]" italics="true" pageId="11" pageNumber="212">A. curupira</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E5294511B10C8EF6FC80" bold="true" box="[832,896,861,887]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E5294511B10C8EF6FC80" box="[832,896,861,887]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4A8D4D5AFFC1E52945E9B10C8958FC81" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[952,1070,861,887]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="caipora" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E52945E9B10C8958FC81" box="[952,1070,861,887]" italics="true" pageId="11" pageNumber="212">A. caipora</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E5294267B10C8909FC81" bold="true" box="[1078,1151,860,887]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E5294267B10C8900FC81" box="[1078,1142,860,887]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
)
</emphasis>
. Paranota form: round to slightly rectangular (
<figureCitation id="15B62A5CFFC1E529472EB1D18CADFC6C" box="[383,475,896,923]" captionStart="FIGURE 4" captionStartId="13.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@13.[161,1426,181,1680]" captionTargetPageId="13" captionText="FIGURE 4. Arthrosolaenomeris chapadensis Schubart, 1943 (holotype, MZSP 1092), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761555" httpUri="https://zenodo.org/record/4761555/files/figure.png" pageId="11" pageNumber="212">Figs 4B</figureCitation>
,
<figureCitation id="15B62A5CFFC1E52947B6B1D08F7DFC6C" box="[487,523,897,923]" captionStart="FIGURE 8" captionStartId="18.[151,250,1939,1963]" captionTargetBox="[186,1401,475,1915]" captionTargetId="figure-73@18.[186,1401,475,1915]" captionTargetPageId="18" captionText="FIGURE 8. Arthrosolaenomeris caipora n. sp. (holotype, IBSP 1569), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 0,5mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761569" httpUri="https://zenodo.org/record/4761569/files/figure.png" pageId="11" pageNumber="212">8B</figureCitation>
). Legs: podomeres with several thick ventral setae (
<figureCitation id="15B62A5CFFC1E5294203B1D189CEFC6C" box="[1106,1208,896,923]" captionStart="FIGURE 31" captionStartId="48.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-24@48.[161,1426,181,1680]" captionTargetPageId="48" captionText="FIGURE 31. Legs. A) Arthrosolaenomeris pantanalensis Schubart, 1943 (IBSP 530); B) Gangugia boto n. sp. (holotype, IBSP 7539); CD) Abiliodesmus planaltensis (Schubart, 1960) comb. nov. (IBSP 3570); EF) Gangugia tapirapensis Schubart, 1947 (MNRJ 12008). Scale bars: A) 2 mm.; B) 1 mm.; C) 1 mm.; D) 1 mm.; E) 500 µm.; F)1 mm. Abbreviations: co, coxae; f, femur; prf, prefemur; pst, postfemur; t, tarsus; tb, tibia; tc; tarsal claw." figureDoi="http://doi.org/10.5281/zenodo.4761653" httpUri="https://zenodo.org/record/4761653/files/figure.png" pageId="11" pageNumber="212">Fig. 31A</figureCitation>
) and granules in the tibia and tarsus (
<figureCitation id="15B62A5CFFC1E529471EB1F58CC6FC48" box="[335,432,932,959]" captionStart="FIGURE 31" captionStartId="48.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-24@48.[161,1426,181,1680]" captionTargetPageId="48" captionText="FIGURE 31. Legs. A) Arthrosolaenomeris pantanalensis Schubart, 1943 (IBSP 530); B) Gangugia boto n. sp. (holotype, IBSP 7539); CD) Abiliodesmus planaltensis (Schubart, 1960) comb. nov. (IBSP 3570); EF) Gangugia tapirapensis Schubart, 1947 (MNRJ 12008). Scale bars: A) 2 mm.; B) 1 mm.; C) 1 mm.; D) 1 mm.; E) 500 µm.; F)1 mm. Abbreviations: co, coxae; f, femur; prf, prefemur; pst, postfemur; t, tarsus; tb, tibia; tc; tarsal claw." figureDoi="http://doi.org/10.5281/zenodo.4761653" httpUri="https://zenodo.org/record/4761653/files/figure.png" pageId="11" pageNumber="212">Fig. 31F</figureCitation>
) (except in
<taxonomicName id="4A8D4D5AFFC1E5294464B1F48FC0FC49" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[565,694,933,959]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="curupira" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E5294464B1F48FC0FC49" box="[565,694,933,959]" italics="true" pageId="11" pageNumber="212">A. curupira</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E52944ECB1F48F8DFC48" bold="true" box="[701,763,933,959]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E52944ECB1F48F8DFC48" box="[701,763,933,959]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4A8D4D5AFFC1E5294560B1F48ED1FC49" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[817,935,933,959]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="caipora" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E5294560B1F48ED1FC49" box="[817,935,933,959]" italics="true" pageId="11" pageNumber="212">A. caipora</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E52945FFB1F48E9BFC48" bold="true" box="[942,1005,933,959]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E52945FFB1F48E9BFC48" box="[942,1005,933,959]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
), dorsal lobe of prefemur conspicuous of the anterior legs (
<figureCitation id="15B62A5CFFC1E5294728B1998CA9FC14" box="[377,479,968,995]" captionStart="FIGURE 31" captionStartId="48.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-24@48.[161,1426,181,1680]" captionTargetPageId="48" captionText="FIGURE 31. Legs. A) Arthrosolaenomeris pantanalensis Schubart, 1943 (IBSP 530); B) Gangugia boto n. sp. (holotype, IBSP 7539); CD) Abiliodesmus planaltensis (Schubart, 1960) comb. nov. (IBSP 3570); EF) Gangugia tapirapensis Schubart, 1947 (MNRJ 12008). Scale bars: A) 2 mm.; B) 1 mm.; C) 1 mm.; D) 1 mm.; E) 500 µm.; F)1 mm. Abbreviations: co, coxae; f, femur; prf, prefemur; pst, postfemur; t, tarsus; tb, tibia; tc; tarsal claw." figureDoi="http://doi.org/10.5281/zenodo.4761653" httpUri="https://zenodo.org/record/4761653/files/figure.png" pageId="11" pageNumber="212">Fig. 31A</figureCitation>
) and a ventroapical projection in the prefemur of the post-gonopod legs (
<figureCitation id="15B62A5CFFC1E5294345B199880FFC14" box="[1300,1401,968,995]" captionStart="FIGURE 31" captionStartId="48.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-24@48.[161,1426,181,1680]" captionTargetPageId="48" captionText="FIGURE 31. Legs. A) Arthrosolaenomeris pantanalensis Schubart, 1943 (IBSP 530); B) Gangugia boto n. sp. (holotype, IBSP 7539); CD) Abiliodesmus planaltensis (Schubart, 1960) comb. nov. (IBSP 3570); EF) Gangugia tapirapensis Schubart, 1947 (MNRJ 12008). Scale bars: A) 2 mm.; B) 1 mm.; C) 1 mm.; D) 1 mm.; E) 500 µm.; F)1 mm. Abbreviations: co, coxae; f, femur; prf, prefemur; pst, postfemur; t, tarsus; tb, tibia; tc; tarsal claw." figureDoi="http://doi.org/10.5281/zenodo.4761653" httpUri="https://zenodo.org/record/4761653/files/figure.png" pageId="11" pageNumber="212">Fig. 31E</figureCitation>
).
</paragraph>
<paragraph id="8D3236D9FFC1E5294696B1BD8F3CFB28" blockId="11.[151,1437,320,1427]" pageId="11" pageNumber="212">
<emphasis id="BFF9EACBFFC1E5294696B1BD8CF5FBF1" box="[199,387,1004,1030]" italics="true" pageId="11" pageNumber="212">Male characters</emphasis>
: Leg pair of the third body ring with the coxae presenting the rectangular genital papilla. Gonopod aperture on seventh body ring with the posterior margin excavated and with a small process. Gonopods: gonocoxae equivalent to about half the length of the telopodite, cylindrical, without a spiniform process. Prefemoral process (
<figureCitation id="15B62A5CFFC1E52946ADB6098C0AFB84" box="[252,380,1112,1139]" captionStart="FIGURE 4" captionStartId="13.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@13.[161,1426,181,1680]" captionTargetPageId="13" captionText="FIGURE 4. Arthrosolaenomeris chapadensis Schubart, 1943 (holotype, MZSP 1092), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761555" httpUri="https://zenodo.org/record/4761555/files/figure.png" pageId="11" pageNumber="212">Figs 4DF</figureCitation>
): elongate with a dorso-basal process and a secondary process in its medial portion (long in
<taxonomicName id="4A8D4D5AFFC1E52946C6B62C8C34FB61" authorityName="Schubart" authorityYear="1943" box="[151,322,1148,1175]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="chapadensis">
<emphasis id="BFF9EACBFFC1E52946C6B62C8C34FB61" box="[151,322,1148,1175]" italics="true" pageId="11" pageNumber="212">A. chapadensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4A8D4D5AFFC1E5294700B62C8F7BFB61" authorityName="Schubart" authorityYear="1943" box="[337,525,1148,1175]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="pantanalensis">
<emphasis id="BFF9EACBFFC1E5294700B62C8F7BFB61" box="[337,525,1148,1175]" italics="true" pageId="11" pageNumber="212">A. pantanalensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4A8D4D5AFFC1E529444DB62C8F1FFB61" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[540,617,1149,1175]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="saci" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E529444DB62C8F1FFB61" box="[540,617,1149,1175]" italics="true" pageId="11" pageNumber="212">A. saci</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E5294423B62C8FC4FB60" bold="true" box="[626,690,1149,1175]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E5294423B62C8FC4FB60" box="[626,690,1149,1175]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4A8D4D5AFFC1E5294490B62C8E67FB61" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[705,785,1149,1175]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="iara" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E5294490B62C8E67FB61" box="[705,785,1149,1175]" italics="true" pageId="11" pageNumber="212">A. iara</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E5294548B62C8E2FFB60" bold="true" box="[793,857,1149,1175]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E5294548B62C8E2FFB60" box="[793,857,1149,1175]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and short in
<taxonomicName id="4A8D4D5AFFC1E52945A0B62C8905FB61" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[1009,1139,1149,1175]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="curupira" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E52945A0B62C8905FB61" box="[1009,1139,1149,1175]" italics="true" pageId="11" pageNumber="212">A. curupira</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E529422DB62C89CAFB60" bold="true" box="[1148,1212,1149,1175]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E529422DB62C89CAFB60" box="[1148,1212,1149,1175]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4A8D4D5AFFC1E529429DB62C8835FB61" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[1228,1347,1149,1175]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="caipora" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E529429DB62C8835FB61" box="[1228,1347,1149,1175]" italics="true" pageId="11" pageNumber="212">A. caipora</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E529431AB62C88FAFB60" bold="true" box="[1355,1420,1149,1175]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E529431AB62C88FAFB60" box="[1355,1420,1149,1175]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
). Cingulum in an apical position (
<figureCitation id="15B62A5CFFC1E5294455B6F18F2EFB4C" box="[516,600,1184,1211]" captionStart="FIGURE 4" captionStartId="13.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@13.[161,1426,181,1680]" captionTargetPageId="13" captionText="FIGURE 4. Arthrosolaenomeris chapadensis Schubart, 1943 (holotype, MZSP 1092), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761555" httpUri="https://zenodo.org/record/4761555/files/figure.png" pageId="11" pageNumber="212">Fig. 4F</figureCitation>
) (middle position in
<taxonomicName id="4A8D4D5AFFC1E5294517B6F08ECBFB4D" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[838,957,1185,1211]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="caipora" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E5294517B6F08ECBFB4D" box="[838,957,1185,1211]" italics="true" pageId="11" pageNumber="212">A. caipora</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E5294594B6F08973FB4C" bold="true" box="[965,1029,1185,1211]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E5294594B6F08973FB4C" box="[965,1029,1185,1211]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
). Solenomere long and thin with its apex abruptly descending (
<figureCitation id="15B62A5CFFC1E5294794B6958F48FB28" box="[453,574,1220,1247]" captionStart="FIGURE 4" captionStartId="13.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@13.[161,1426,181,1680]" captionTargetPageId="13" captionText="FIGURE 4. Arthrosolaenomeris chapadensis Schubart, 1943 (holotype, MZSP 1092), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761555" httpUri="https://zenodo.org/record/4761555/files/figure.png" pageId="11" pageNumber="212">Fig. 4DF</figureCitation>
).
</paragraph>
<paragraph id="8D3236D9FFC1E5294696B6B98C4FFAD0" blockId="11.[151,1437,320,1427]" pageId="11" pageNumber="212">
<emphasis id="BFF9EACBFFC1E5294696B6B98CEFFAF5" box="[199,409,1256,1282]" italics="true" pageId="11" pageNumber="212">Female characters</emphasis>
: Vulvae: oval-shaped, having equal proportions in lateral view, with an operculum advanced (
<figureCitation id="15B62A5CFFC1E52946CEB75D8C5BFAD0" box="[159,301,1292,1319]" captionStart="FIGURE 10" captionStartId="21.[151,250,1798,1822]" captionTargetBox="[314,1273,181,1773]" captionTargetId="figure-20@21.[314,1273,181,1773]" captionTargetPageId="21" captionText="FIGURE 10. Left vulva. Arthrosolaenomeris chapadensis Schubart, 1943 (IBSP 1049): (A), ventral view; (B), lateral view. Arthrosolaenomeris pantanalensis Schubart, 1943 (CZUFMT 847): (C), ventral view; (D), lateral view. Arthrosolaenomeris saci n. sp. (paratype, IBSP 2577): (E), ventral view; (F), lateral view.Arthrosolaenomeris iara n. sp. (paratype, IBSP 7537). (G), ventral view; (H), lateral view. Scale bars: 0,5 mm. Abbreviations: EV, external valve; IV, internal valve; Op, operculum." figureDoi="http://doi.org/10.5281/zenodo.4761577" httpUri="https://zenodo.org/record/4761577/files/figure.png" pageId="11" pageNumber="212">Figs 10AB</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C5976552FFC1E5294696B7618F8DFA64" pageId="11" pageNumber="212" type="distribution">
<paragraph id="8D3236D9FFC1E5294696B7618910FABC" blockId="11.[151,1437,320,1427]" box="[199,1126,1328,1355]" pageId="11" pageNumber="212">
<emphasis id="BFF9EACBFFC1E5294696B7618C17FABD" bold="true" box="[199,353,1328,1354]" pageId="11" pageNumber="212">Distribution.</emphasis>
States of
<collectingRegion id="4F49F83BFFC1E5294781B7608F12FABC" box="[464,612,1329,1355]" country="Brazil" name="Mato Grosso" pageId="11" pageNumber="212">Mato Grosso</collectingRegion>
,
<collectingRegion id="4F49F83BFFC1E529443EB7608E26FABD" box="[623,848,1328,1355]" country="Brazil" name="Mato Grosso do Sul" pageId="11" pageNumber="212">Mato Grosso do Sul</collectingRegion>
and
<collectingRegion id="4F49F83BFFC1E52945D7B7608E81FABD" box="[902,1015,1328,1355]" country="Brazil" name="Sao Paulo" pageId="11" pageNumber="212">São Paulo</collectingRegion>
(
<figureCitation id="15B62A5CFFC1E5294257B761892EFABC" box="[1030,1112,1328,1355]" captionStart="FIGURE 11" captionStartId="22.[151,250,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-24@22.[151,1436,181,1337]" captionTargetPageId="22" captionText="FIGURE 11. Distribution map of Arthrosolaenomeris Schubart, 1943 species." figureDoi="http://doi.org/10.5281/zenodo.4761582" httpUri="https://zenodo.org/record/4761582/files/figure.png" pageId="11" pageNumber="212">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="8D3236D9FFC1E5294696B7058F8DFA64" blockId="11.[151,1437,320,1427]" pageId="11" pageNumber="212">
<emphasis id="BFF9EACBFFC1E5294696B7058C11FA99" bold="true" box="[199,359,1364,1390]" pageId="11" pageNumber="212">Composition.</emphasis>
<taxonomicName id="4A8D4D5AFFC1E5294721B7058EEAFA98" authority="Schubart, 1943" authorityName="Schubart" authorityYear="1943" box="[368,924,1364,1391]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="chapadensis">
<emphasis id="BFF9EACBFFC1E5294721B7058F95FA99" box="[368,739,1364,1390]" italics="true" pageId="11" pageNumber="212">Arthrosolaenomeris chapadensis</emphasis>
<bibRefCitation id="E91C4B28FFC1E52944BCB7058EEAFA98" author="Schubart, O." box="[749,924,1364,1391]" pageId="11" pageNumber="212" pagination="127 - 164" refId="ref33096" refString="Schubart, O. (1943) Especies novas das familias Strongylosomidae e Leptodesmidae da ordem Proterospermophora do interior dos Estados de Sao Paulo e Mato Grosso. Papeis Avulsos de Zoologia, 3, 127 - 164." type="journal article" year="1943">Schubart, 1943</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4A8D4D5AFFC1E52945F8B7048869FA98" authority="Schubart, 1943" authorityName="Schubart" authorityYear="1943" box="[937,1311,1364,1391]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="pantanalensis">
<emphasis id="BFF9EACBFFC1E52945F8B7048911FA99" box="[937,1127,1364,1391]" italics="true" pageId="11" pageNumber="212">A. pantanalensis</emphasis>
<bibRefCitation id="E91C4B28FFC1E5294221B7058869FA98" author="Schubart, O." box="[1136,1311,1364,1391]" pageId="11" pageNumber="212" pagination="127 - 164" refId="ref33096" refString="Schubart, O. (1943) Especies novas das familias Strongylosomidae e Leptodesmidae da ordem Proterospermophora do interior dos Estados de Sao Paulo e Mato Grosso. Papeis Avulsos de Zoologia, 3, 127 - 164." type="journal article" year="1943">Schubart, 1943</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4A8D4D5AFFC1E529437CB704880AFA99" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[1325,1404,1365,1391]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="saci" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E529437CB704880AFA99" box="[1325,1404,1365,1391]" italics="true" pageId="11" pageNumber="212">A. saci</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E52943D7B7048DCEFA64" bold="true" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E52943D7B7048DCEFA64" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<taxonomicName id="4A8D4D5AFFC1E5294696B7288C3EFA65" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[199,328,1401,1427]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="curupira" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E5294696B7288C3EFA65" box="[199,328,1401,1427]" italics="true" pageId="11" pageNumber="212">A. curupira</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E529471EB7288CF8FA64" bold="true" box="[335,398,1401,1427]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E529471EB7288CF8FA64" box="[335,398,1401,1427]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<taxonomicName id="4A8D4D5AFFC1E52947CCB7288F65FA65" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[413,531,1401,1427]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="caipora" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E52947CCB7288F65FA65" box="[413,531,1401,1427]" italics="true" pageId="11" pageNumber="212">A. caipora</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E529444BB7288F2FFA64" bold="true" box="[538,601,1401,1427]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E529444BB7288F2FFA64" box="[538,601,1401,1427]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<taxonomicName id="4A8D4D5AFFC1E5294439B7288FC0FA65" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[616,694,1401,1427]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="iara" status="sp. nov.">
<emphasis id="BFF9EACBFFC1E5294439B7288FC0FA65" box="[616,694,1401,1427]" italics="true" pageId="11" pageNumber="212">A. iara</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFC1E52944ECB7288F8DFA64" bold="true" box="[701,763,1401,1427]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="A4CA57B0FFC1E52944ECB7288F8DFA64" box="[701,763,1401,1427]" pageId="11" pageNumber="212" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>