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<document id="FB2D10AE22C32AB13F68BA8F5605EF78" ID="10.11646/zootaxa.4162.3.1" ID-DOI="10.11646/zootaxa.4162.3.1" ID-GBIF-Dataset="bc54479e-1a51-4fea-861e-757a5a391348" ID-ISSN="1175-5326" ID-Zenodo-Dep="254973" ID-ZooBank="DDBC36A0-5305-45FD-9C79-170C53B72DA9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1473665070933" checkinUser="plazi" docAuthor="Zvonareva, Sofya &amp; Kantor, Yuri" docDate="2016" docId="F0542475BB45FFDEFF28FDC76D95FB1E" docLanguage="en" docName="zootaxa.4162.3.1.pdf" docOrigin="Zootaxa 4162 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Littoraria scabra Linnaeus 1758" docType="treatment" docVersion="18" lastPageNumber="420" masterDocId="0C6D5C0DBB56FFCDFFBFFF8F6938FFCC" masterDocTitle="Cheсklist of gastropod molluscs in mangroves of Khanh Hoa province, Vietnam" masterLastPageNumber="437" masterPageNumber="401" pageNumber="420" updateTime="1698384908448" updateUser="plazi">
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<mods:title id="798D3FBC3916CB8C68AE0009EF5718DF">Cheсklist of gastropod molluscs in mangroves of Khanh Hoa province, Vietnam</mods:title>
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<mods:namePart id="9D9E67C049347ED3B50051B3F791D82E">Zvonareva, Sofya</mods:namePart>
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<mods:namePart id="81CCB750F66672562326E09408634F43">Kantor, Yuri</mods:namePart>
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<mods:date id="9A8B8EA218C96D2649C268F2034107E1">2016</mods:date>
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<treatment id="F0542475BB45FFDEFF28FDC76D95FB1E" ID-DOI="http://doi.org/10.5281/zenodo.6085779" ID-GBIF-Taxon="124483941" ID-Zenodo-Dep="6085779" LSID="urn:lsid:plazi:treatment:F0542475BB45FFDEFF28FDC76D95FB1E" httpUri="http://treatment.plazi.org/id/F0542475BB45FFDEFF28FDC76D95FB1E" lastPageNumber="420" pageId="19" pageNumber="420">
<subSubSection id="30E7C6E8BB45FFDEFF28FDC76829FD4F" pageId="19" pageNumber="420" type="nomenclature">
<paragraph id="78429563BB45FFDEFF28FDC76B02FDAE" blockId="19.[151,570,584,643]" box="[151,570,584,610]" pageId="19" pageNumber="420">
<heading id="230A220FBB45FFDEFF28FDC76B02FDAE" bold="true" box="[151,570,584,610]" fontSize="11" level="1" pageId="19" pageNumber="420" reason="1">
<taxonomicName id="BFFDEEE0BB45FFDEFF28FDC76B02FDAE" ID-CoL="3VNKD" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[151,570,584,610]" class="Gastropoda" family="Littorinidae" genus="Littoraria" kingdom="Animalia" order="Littorinimorpha" pageId="19" pageNumber="420" phylum="Mollusca" rank="species" species="scabra">
<emphasis id="4A894971BB45FFDEFF28FDC76B02FDAE" bold="true" box="[151,570,584,610]" pageId="19" pageNumber="420">
<emphasis id="4A894971BB45FFDEFF28FDC7685BFDAE" bold="true" box="[151,355,584,610]" italics="true" pageId="19" pageNumber="420">Littoraria scabra</emphasis>
(
<bibRefCitation id="1C6CE892BB45FFDEFECDFDC76B0AFDAE" author="Linnaeus" box="[370,562,584,610]" pageId="19" pageNumber="420" refString="Linnaeus, C. (1758) Systema Naturae per Regna tria Naturae. Vol. 1. 10 th Edition. Salvius, Stockholm, 823 pp." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="78429563BB45FFDEFF28FDE56829FD4F" blockId="19.[151,570,584,643]" box="[151,273,618,643]" pageId="19" pageNumber="420">
<figureCitation id="E0C689E6BB45FFDEFF28FDE569E3FD4F" box="[151,219,618,643]" captionStart="FIGURE 6. A B" captionStartId="16.[151,250,1832,1854]" captionTargetBox="[151,1436,339,1809]" captionTargetId="figure@16.[151,1436,339,1811]" captionTargetPageId="16" captionText="FIGURE 6. A B. Monetaria moneta (26 mm). C D. Monetaria annulus (25 mm). E F. Sermyla riqueti (10 mm). G H. Sermyla riqueti (7 mm). I J. Littoraria lutea (36 mm). K L. Littoraria lutea (23 mm). M N. Littoraria lutea (29 mm). O P. Littoraria ardouiniana (16 mm). Q R. Littoraria scabra (36 mm). S T. Littoraria carinifera (18 mm). U V. Littoraria pallescens (24 mm). W X. Littoraria pallescens (12 mm). Y Z. Littoraria ardouiniana (14 mm). AA AB. Littoraria intermedia (14 mm)." httpUri="https://zenodo.org/record/254977/files/figure.png" pageId="19" pageNumber="420">Figs 6</figureCitation>
QR
</paragraph>
</subSubSection>
<subSubSection id="30E7C6E8BB45FFDEFF28FD3D6D61FCE8" pageId="19" pageNumber="420" type="reference_group">
<paragraph id="78429563BB45FFDEFF28FD3D68CDFD04" blockId="19.[151,1436,690,805]" box="[151,501,690,712]" pageId="19" pageNumber="420">
<treatmentCitationGroup id="58EDB24DBB45FFDEFF28FD3D68CDFD04" box="[151,501,690,712]" pageId="19" pageNumber="420">
<taxonomicName id="BFFDEEE0BB45FFDEFF28FD3D68C9FD04" authority="Linnaeus, 1758: 770" authorityName="Linnaeus" authorityPageNumber="770" authorityYear="1758" box="[151,497,690,712]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="420" phylum="Mollusca" rank="species" species="scabra">
<emphasis id="4A894971BB45FFDEFF28FD3D6821FD04" box="[151,281,690,712]" italics="true" pageId="19" pageNumber="420">Helix scabra</emphasis>
<treatmentCitation id="F95CB372BB45FFDEFEA0FD3D68C9FD04" ID-DOI="http://doi.org/10.5281/zenodo.3918537" ID-Zenodo-Dep="3918537" LSID="urn:lsid:zoobank.org:act:5AA93348-E6DD-42EA-903A-E553B7E70D3B" author="Linnaeus" box="[287,497,690,712]" httpUri="http://treatment.plazi.org/id/481FFA31260DB5B879FFFDCA1BA1E1BB" page="770" pageId="19" pageNumber="420" year="1758">
<bibRefCitation id="1C6CE892BB45FFDEFEA0FD3D6886FD04" author="Linnaeus" box="[287,446,690,712]" pageId="19" pageNumber="420" refString="Linnaeus, C. (1758) Systema Naturae per Regna tria Naturae. Vol. 1. 10 th Edition. Salvius, Stockholm, 823 pp." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
: 770
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="78429563BB45FFDEFF28FD5D6854FCCA" blockId="19.[151,1436,690,805]" pageId="19" pageNumber="420">
<treatmentCitationGroup id="58EDB24DBB45FFDEFF28FD5D6854FCCA" pageId="19" pageNumber="420">
<taxonomicName id="BFFDEEE0BB45FFDEFF28FD5D6B44FD2B" ID-CoL="3VNKD" authority="Reid 1986: 94" authorityName="Reid" authorityPageNumber="94" authorityYear="1986" box="[151,636,721,743]" class="Gastropoda" family="Littorinidae" genus="Littoraria" kingdom="Animalia" order="Littorinimorpha" pageId="19" pageNumber="420" phylum="Mollusca" rank="species" species="scabra" subGenus="Littorinopsis">
<emphasis id="4A894971BB45FFDEFF28FD5D69C3FD2B" box="[151,251,722,743]" italics="true" pageId="19" pageNumber="420">Littoraria</emphasis>
(
<emphasis id="4A894971BB45FFDEFEB6FD5D68B3FD2B" box="[265,395,722,743]" italics="true" pageId="19" pageNumber="420">Littorinopsis</emphasis>
)
<emphasis id="4A894971BB45FFDEFE25FD5E68E7FD2B" box="[410,479,721,743]" italics="true" pageId="19" pageNumber="420">scabra</emphasis>
:
<treatmentCitation id="F95CB372BB45FFDEFE53FD5E6B44FD2B" author="Reid" box="[492,636,721,743]" page="94" pageId="19" pageNumber="420" year="1986">
<bibRefCitation id="1C6CE892BB45FFDEFE53FD5E6B60FD2B" author="Reid" box="[492,600,721,743]" pageId="19" pageNumber="420" refString="Reid, D. G. (1986) The littorinid molluscs of mangrove forests in the Indo-Pacific region: the genus Littoraria. British Museum (Natural History), London, 227 pp." type="book" year="1986">Reid 1986</bibRefCitation>
: 94
</treatmentCitation>
</taxonomicName>
103, figs 3, 5, 2730 (full synonymy);
<treatmentCitation id="F95CB372BB45FFDEFBB0FD5E6DF5FD2B" author="Okutani" box="[1039,1229,721,743]" page="138" pageId="19" pageNumber="420" year="2000">
<bibRefCitation id="1C6CE892BB45FFDEFBB0FD5E6DA6FD2B" author="Okutani" box="[1039,1182,721,743]" pageId="19" pageNumber="420" refString="Okutani, T. (2000) Marine Mollusks in Japan. Tokai University Press, Tokyo, 1173 pp." type="book" year="2000">Okutani 2000</bibRefCitation>
: 138
</treatmentCitation>
139, pl. 69, fig. 15;
<treatmentCitation id="F95CB372BB45FFDEFF78FD7F6850FCCA" author="Thach" box="[199,360,752,774]" page="53" pageId="19" pageNumber="420" year="2005">
<bibRefCitation id="1C6CE892BB45FFDEFF78FD7F687AFCCA" author="Thach" box="[199,322,752,774]" pageId="19" pageNumber="420" refString="Thach, N. N. (2005) Shells of Vietnam. Conchbooks, Hackenheim, 338 pp." type="book" year="2005">Thach 2005</bibRefCitation>
: 53
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="78429563BB45FFDEFF28FC9F6D61FCE8" blockId="19.[151,1436,690,805]" box="[151,1113,782,805]" pageId="19" pageNumber="420">
<treatmentCitationGroup id="58EDB24DBB45FFDEFF28FC9F6D61FCE8" box="[151,1113,782,805]" pageId="19" pageNumber="420">
<taxonomicName id="BFFDEEE0BB45FFDEFF28FC9F6B40FCE8" ID-CoL="3VNKD" authority="Lozouet &amp; Plaziat 2008: 116" authorityName="Lozouet &amp; Plaziat" authorityPageNumber="116" authorityYear="2008" box="[151,632,782,805]" class="Gastropoda" family="Littorinidae" genus="Littoraria" kingdom="Animalia" order="Littorinimorpha" pageId="19" pageNumber="420" phylum="Mollusca" rank="species" species="scabra">
<emphasis id="4A894971BB45FFDEFF28FC9F687EFCE8" box="[151,326,782,805]" italics="true" pageId="19" pageNumber="420">Littoraria scabra</emphasis>
:
<treatmentCitation id="F95CB372BB45FFDEFEECFC806B40FCE8" author="Lozouet" box="[339,632,782,804]" page="116" pageId="19" pageNumber="420" year="2008">
<bibRefCitation id="1C6CE892BB45FFDEFEECFC806B71FCE8" author="Lozouet" box="[339,585,782,804]" pageId="19" pageNumber="420" refString="Lozouet, P. &amp; Plaziat, J. - C. (2008) Mangrove environments and mollusks. Abatan River, Bohol and Panglao Islands, Central Philippines. ConchBooks, Hackenheim, 166 pp." type="book" year="2008">Lozouet &amp; Plaziat 2008</bibRefCitation>
: 116
</treatmentCitation>
</taxonomicName>
117, pl. 22, figs 12;
<treatmentCitation id="F95CB372BB45FFDEFCE4FC806D6DFCE8" author="Zvonareva" box="[859,1109,782,805]" page="8" pageId="19" pageNumber="420" year="2015">
<bibRefCitation id="1C6CE892BB45FFDEFCE4FC806D05FCE8" author="Zvonareva" box="[859,1085,782,805]" pageId="19" pageNumber="420" refString="Zvonareva, S., Kantor Yu., Xinzheng, L. &amp; Britayev, T. (2015) Long-term monitoring of Gastropoda (Mollusca) fauna in planted mangroves in central Vietnam. Zoological studies, 54, 1 - 39." type="journal article" year="2015">
Zvonareva
<emphasis id="4A894971BB45FFDEFC72FC9F6AC2FCE8" box="[973,1018,782,805]" italics="true" pageId="19" pageNumber="420">et al</emphasis>
. 2015
</bibRefCitation>
: 8
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="30E7C6E8BB45FFDEFF28FCDE68F6FC42" pageId="19" pageNumber="420" type="materials_examined">
<paragraph id="78429563BB45FFDEFF28FCDE6D41FCA6" blockId="19.[151,1436,849,1234]" box="[151,1145,849,874]" pageId="19" pageNumber="420">
<materialsCitation id="C8959F3EBB45FFDEFF28FCDE6D41FCA6" ID-GBIF-Occurrence="1306554424" box="[151,1145,849,874]" county="Dam Bay" location="Material" municipality="Nha Phu Bay" pageId="19" pageNumber="420" specimenCount="15">
<emphasis id="4A894971BB45FFDEFF28FCDE68B8FCA6" bold="true" box="[151,384,849,874]" pageId="19" pageNumber="420">
<location id="7D22C3B8BB45FFDEFF28FCDE69C7FCA6" LSID="urn:lsid:plazi:treatment:F0542475BB45FFDEFF28FDC76D95FB1E:7D22C3B8BB45FFDEFF28FCDE69C7FCA6" box="[151,255,849,874]" county="Dam Bay" municipality="Nha Phu Bay" name="Material" pageId="19" pageNumber="420">Material</location>
examined.
</emphasis>
<collectingMunicipality id="98260F19BB45FFDEFE38FCDE6B25FCA6" box="[391,541,849,874]" pageId="19" pageNumber="420">Nha Phu Bay</collectingMunicipality>
,
<collectingCounty id="9123EDEFBB45FFDEFD96FCDD6BACFCA6" box="[553,660,850,874]" pageId="19" pageNumber="420">Dam Bay</collectingCounty>
, mangrove plantation, about
<specimenCount id="6EFB5EEABB45FFDEFC66FCDE6D4DFCA6" box="[985,1141,849,874]" pageId="19" pageNumber="420" type="generic">15 specimens</specimenCount>
.
</materialsCitation>
</paragraph>
<paragraph id="78429563BB45FFDEFF78FCFA68F6FC42" blockId="19.[151,1436,849,1234]" box="[199,462,885,910]" pageId="19" pageNumber="420">
<emphasis id="4A894971BB45FFDEFF78FCFA686FFC42" bold="true" box="[199,343,885,910]" pageId="19" pageNumber="420">Abundance.</emphasis>
Common.
</paragraph>
</subSubSection>
<subSubSection id="30E7C6E8BB45FFDEFF78FC166ABAFB46" pageId="19" pageNumber="420" type="description">
<paragraph id="78429563BB45FFDEFF78FC166B95FC7E" blockId="19.[151,1436,849,1234]" box="[199,685,921,946]" pageId="19" pageNumber="420">
<emphasis id="4A894971BB45FFDEFF78FC166845FC7E" bold="true" box="[199,381,921,946]" pageId="19" pageNumber="420">Measurements.</emphasis>
Up to
<quantity id="BF053886BB45FFDEFE75FC166B21FC7D" box="[458,537,921,946]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="19" pageNumber="420" unit="mm" value="35.0">35 mm</quantity>
(
<bibRefCitation id="1C6CE892BB45FFDEFD98FC166B99FC7E" author="Reid" box="[551,673,921,946]" pageId="19" pageNumber="420" refString="Reid, D. G. (1986) The littorinid molluscs of mangrove forests in the Indo-Pacific region: the genus Littoraria. British Museum (Natural History), London, 227 pp." type="book" year="1986">Reid 1986</bibRefCitation>
).
</paragraph>
<paragraph id="78429563BB45FFDEFF78FC326A10FC1A" blockId="19.[151,1436,849,1234]" box="[199,808,957,982]" pageId="19" pageNumber="420">
<emphasis id="4A894971BB45FFDEFF78FC3268B6FC1A" bold="true" box="[199,398,957,982]" pageId="19" pageNumber="420">Ecological notes.</emphasis>
Ovoviviparous species (
<bibRefCitation id="1C6CE892BB45FFDEFD1DFC326A23FC1A" author="Reid" box="[674,795,957,982]" pageId="19" pageNumber="420" refString="Reid, D. G. (1986) The littorinid molluscs of mangrove forests in the Indo-Pacific region: the genus Littoraria. British Museum (Natural History), London, 227 pp." type="book" year="1986">Reid 1986</bibRefCitation>
).
</paragraph>
<paragraph id="78429563BB45FFDEFF78FC6D6DA3FB8E" blockId="19.[151,1436,849,1234]" pageId="19" pageNumber="420">
An oceanic species, typically on trunks and roots of
<taxonomicName id="BFFDEEE0BB45FFDEFC96FC6E6A94FC35" box="[809,940,993,1017]" class="Magnoliopsida" family="Rhizophoraceae" genus="Rhizophora" kingdom="Plantae" order="Malpighiales" pageId="19" pageNumber="420" phylum="Tracheophyta" rank="genus">
<emphasis id="4A894971BB45FFDEFC96FC6E6A94FC35" box="[809,940,993,1017]" italics="true" pageId="19" pageNumber="420">Rhizophora</emphasis>
</taxonomicName>
and less frequently on trunks of
<taxonomicName id="BFFDEEE0BB45FFDEFA92FC6D6CA3FC35" box="[1325,1435,994,1017]" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" kingdom="Plantae" order="Lamiales" pageId="19" pageNumber="420" phylum="Tracheophyta" rank="genus">
<emphasis id="4A894971BB45FFDEFA92FC6D6CA3FC35" box="[1325,1435,994,1017]" italics="true" pageId="19" pageNumber="420">Avicennia</emphasis>
</taxonomicName>
near seaward edge of mangrove forest and on isolated trees,
<quantity id="BF053886BB45FFDEFCD0FB8A6AC8FBD1" box="[879,1008,1029,1054]" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="19" pageNumber="420" unit="m" value="1.25" valueMax="2.0" valueMin="0.5">
0.5 to
<quantity id="BF053886BB45FFDEFC7FFB896AC8FBD1" box="[960,1008,1030,1054]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="420" unit="m" value="2.0">2 m</quantity>
</quantity>
above the ground. Rarely found on driftwood on sandy shores where mangroves are absent and on sheltered rocks (
<bibRefCitation id="1C6CE892BB45FFDEFBAAFBA66DB6FB8E" author="Reid" box="[1045,1166,1065,1090]" pageId="19" pageNumber="420" refString="Reid, D. G. (1986) The littorinid molluscs of mangrove forests in the Indo-Pacific region: the genus Littoraria. British Museum (Natural History), London, 227 pp." type="book" year="1986">Reid 1986</bibRefCitation>
).
</paragraph>
<paragraph id="78429563BB45FFDEFF78FBC16D49FBA9" blockId="19.[151,1436,849,1234]" box="[199,1137,1101,1126]" pageId="19" pageNumber="420">
In studied sites it was found on trunks and roots of young and old
<taxonomicName id="BFFDEEE0BB45FFDEFC14FBC26D16FBA9" box="[939,1070,1101,1125]" class="Magnoliopsida" family="Rhizophoraceae" genus="Rhizophora" kingdom="Plantae" order="Malpighiales" pageId="19" pageNumber="420" phylum="Tracheophyta" rank="genus">
<emphasis id="4A894971BB45FFDEFC14FBC26D16FBA9" box="[939,1070,1101,1125]" italics="true" pageId="19" pageNumber="420">Rhizophora</emphasis>
</taxonomicName>
trees.
</paragraph>
<paragraph id="78429563BB45FFDEFF78FBFE6ABAFB46" blockId="19.[151,1436,849,1234]" box="[199,898,1137,1162]" pageId="19" pageNumber="420">
<emphasis id="4A894971BB45FFDEFF78FBFE68E2FB46" bold="true" box="[199,474,1137,1162]" pageId="19" pageNumber="420">Relation to mangroves.</emphasis>
Predominantly mangrove-associated.
</paragraph>
</subSubSection>
<subSubSection id="30E7C6E8BB45FFDEFF78FB1A6D95FB1E" pageId="19" pageNumber="420" type="distribution">
<paragraph id="78429563BB45FFDEFF78FB1A6D4AFB62" blockId="19.[151,1436,849,1234]" box="[199,1138,1173,1198]" pageId="19" pageNumber="420">
<emphasis id="4A894971BB45FFDEFF78FB1A68D3FB62" bold="true" box="[199,491,1173,1198]" pageId="19" pageNumber="420">
Distribution in
<collectingCountry id="00EAD5F3BB45FFDEFEC1FB1A68DEFB62" box="[382,486,1173,1198]" name="Vietnam" pageId="19" pageNumber="420">Vietnam</collectingCountry>
.
</emphasis>
From northern-east coast to southern delta (
<bibRefCitation id="1C6CE892BB45FFDEFC65FB1A6D5EFB62" author="Thach" box="[986,1126,1173,1198]" pageId="19" pageNumber="420" refString="Thach, N. N. (2005) Shells of Vietnam. Conchbooks, Hackenheim, 338 pp." type="book" year="2005">Thach 2005</bibRefCitation>
).
</paragraph>
<paragraph id="78429563BB45FFDEFF78FB366D95FB1E" blockId="19.[151,1436,849,1234]" box="[199,1197,1209,1234]" pageId="19" pageNumber="420">
<emphasis id="4A894971BB45FFDEFF78FB3668FBFB1E" bold="true" box="[199,451,1209,1234]" pageId="19" pageNumber="420">General distribution.</emphasis>
Throughout the tropical and subtropical Indo-Pacific (
<bibRefCitation id="1C6CE892BB45FFDEFB99FB366D98FB1E" author="Reid" box="[1062,1184,1209,1234]" pageId="19" pageNumber="420" refString="Reid, D. G. (1986) The littorinid molluscs of mangrove forests in the Indo-Pacific region: the genus Littoraria. British Museum (Natural History), London, 227 pp." type="book" year="1986">Reid 1986</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>