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118 lines
14 KiB
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<mods:title id="15166D6983EE65C4333A2E8D56726133">Re-appraisal of thylacocephalans (Euarthropoda, Thylacocephala) from the Jurassic La Voulte-sur-Rhône Lagerstätte</mods:title>
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<mods:namePart id="219876E1F378D8BE70C2A0E51FAD5598">Laville, Thomas</mods:namePart>
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<mods:affiliation id="53591487BD297163F55085ACC176BDB9">Muséum national d’Histoire naturelle, Centre de Recherche en Paléontologie - Paris (CR 2 P, UMR 7207), Sorbonne Université-MNHN-CNRS, 57 rue Cuvier, 75005 Paris, France. & Biogéosciences, UMR 6282 CNRS, Université de Bourgogne, 6 boulevard Gabriel, 2100 Dijon, France.</mods:affiliation>
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<mods:nameIdentifier id="9DF3C018610E79BE3E681730C38DD9E3" type="email">thomas.laville2@mnhn.fr</mods:nameIdentifier>
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<mods:namePart id="F043937C08FBE5E2A824C35378CED558">Forel, Marie-Béatrice</mods:namePart>
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<mods:affiliation id="354D2511EA082598110E47134F03762F">Muséum national d’Histoire naturelle, Centre de Recherche en Paléontologie - Paris (CR 2 P, UMR 7207), Sorbonne Université-MNHN-CNRS, 57 rue Cuvier, 75005 Paris, France.</mods:affiliation>
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<mods:nameIdentifier id="AD7487F10321F6BAF3A19D47900DD1B7" type="email">marie-beatrice.forel@mnhn.fr</mods:nameIdentifier>
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<mods:namePart id="52BC1F63A16656892D1BE77375B9B969">Charbonnier, Sylvain</mods:namePart>
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<mods:affiliation id="E6B6BDC05AF3808CC36D3D45EDBD0D43">Muséum national d’Histoire naturelle, Centre de Recherche en Paléontologie - Paris (CR 2 P, UMR 7207), Sorbonne Université-MNHN-CNRS, 57 rue Cuvier, 75005 Paris, France.</mods:affiliation>
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<mods:nameIdentifier id="C23ADBE5594573FA156D9973F685449B" type="email">sylvain.charbonnier@mnhn.fr</mods:nameIdentifier>
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<mods:title id="C69037A25488F9DCBCFD3335DD8D9A2D">European Journal of Taxonomy</mods:title>
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<mods:part id="4738AF9595555C3CC2B118281AF5ED7F">
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<mods:date id="7B0F82BC594771E3920B5AB475B8C8A7">2023</mods:date>
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<mods:number id="44E1E9081A914C13C8049A96FC7ACCB2">2023-10-05</mods:number>
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<mods:number id="E8E74BA8FE5BB1494C955753084F3E8B">898</mods:number>
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<mods:url id="613F643E2829283768555EB6F6C3CD97">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2295/9941</mods:url>
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<mods:identifier id="361935E9B1D1097A9A3640BB9595E04F" type="DOI">10.5852/ejt.2023.898.2295</mods:identifier>
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<mods:identifier id="E39D726DCD522B73A382CE355835F961" type="ISSN">2118-9773</mods:identifier>
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<treatment id="C12F87D2997FFFDB2EC9FCDEFA6BF359" ID-DOI="http://doi.org/10.5281/zenodo.10018158" ID-Zenodo-Dep="10018158" LSID="urn:lsid:plazi:treatment:C12F87D2997FFFDB2EC9FCDEFA6BF359" httpUri="http://treatment.plazi.org/id/C12F87D2997FFFDB2EC9FCDEFA6BF359" lastPageNumber="10" pageId="9" pageNumber="10">
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<subSubSection id="019C654F997FFFDB2EC9FCDEFD9EF536" box="[417,1171,786,812]" pageId="9" pageNumber="10" type="nomenclature">
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<paragraph id="493936C4997FFFDB2EC9FCDEFD9EF536" blockId="9.[417,1171,786,812]" box="[417,1171,786,812]" pageId="9" pageNumber="10">
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Class
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<taxonomicName id="8E864D47997FFFDB2E80FCDEFD9EF536" authority="Pinna, Arduini, Pesarini & Teruzzi, 1982" authorityName="Pinna, Arduini, Pesarini & Teruzzi" authorityYear="1982" box="[488,1171,786,812]" class="Thylacocephala" kingdom="Animalia" pageId="9" pageNumber="10" rank="class">
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<emphasis id="7BF2EAD6997FFFDB2E80FCDEFBA0F536" bold="true" box="[488,685,786,812]" pageId="9" pageNumber="10">Thylacocephala</emphasis>
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<bibRefCitation id="2D174B35997FFFDB2DDCFCDEFD9EF536" author="Pinna G. & Arduini P. & Pesarini C. & Teruzzi G." box="[692,1171,786,812]" pageId="9" pageNumber="10" pagination="469 - 482" refId="ref28096" refString="Pinna G., Arduini P., Pesarini C. & Teruzzi G. 1982. Thylacocephala: una nuova classe di crostacei fossili. Atti della Societa italiana di Scienze naturali e del Museo civico di Storia naturale di Milano 123 (4): 469 - 482." type="journal article" year="1982">Pinna, Arduini, Pesarini & Teruzzi, 1982</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="019C654F997FFFDB2FD5FC9FFB51F276" pageId="9" pageNumber="10" type="diagnosis">
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<paragraph id="493936C4997FFFDB2FD5FC9FFB55F574" blockId="9.[189,600,851,878]" box="[189,600,851,878]" pageId="9" pageNumber="10">
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<emphasis id="7BF2EAD6997FFFDB2FD5FC9FF8AFF577" bold="true" box="[189,418,851,877]" pageId="9" pageNumber="10">Original diagnosis</emphasis>
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(
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<bibRefCitation id="2D174B35997FFFDB2ED9FC98FB5DF574" author="Schram F. R." box="[433,592,852,878]" pageId="9" pageNumber="10" pagination="1 - 16" refId="ref28898" refString="Schram F. R. 1990. On Mazon Creek Thylacocephala. Proceedings of the San Diego Society of Natural History 3: 1 - 16." type="journal article" year="1990">Schram 1990</bibRefCitation>
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)
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</paragraph>
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<paragraph id="493936C4997FFFDB2FD5FC4EFB51F276" blockId="9.[188,1400,898,1132]" pageId="9" pageNumber="10">
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These are arthropods with a small to large bilobed carapace enclosing the entire body. Compound eyes very well developed as either large, sessile bodies situated in optic notches or as organs that virtually cover the surface of a protrudent, sac-like cephalon. Body appendages are of
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<specimenCount id="5F80FD4D997FFFDB2B15FC04FDFBF5F8" box="[1149,1270,968,994]" pageId="9" pageNumber="10" type="generic" typeStatus="types">two types</specimenCount>
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: anteriorly 3 pairs of subchelate, raptorial limbs, and posteriorly a set of from 8 to 16 pairs of apparently paddle-like limbs. Eight sets of well-developed gills are associated with the subchelate limbs. The posterior trunk is marked with pronounced pleurites, probably developed internally as an endrophragmal skeleton, associated with the posterior limbs.
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</paragraph>
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</subSubSection>
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<subSubSection id="019C654F997FFFDB2FD5FB5FFA6BF359" pageId="9" pageNumber="10" type="discussion">
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<paragraph id="493936C4997FFFDB2FD5FB5FF822F2B7" blockId="9.[189,303,1171,1197]" box="[189,303,1171,1197]" pageId="9" pageNumber="10">
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<heading id="127181A8997FFFDB2FD5FB5FF822F2B7" bold="true" box="[189,303,1171,1197]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
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<emphasis id="7BF2EAD6997FFFDB2FD5FB5FF822F2B7" bold="true" box="[189,303,1171,1197]" pageId="9" pageNumber="10">Remarks</emphasis>
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</heading>
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</paragraph>
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<paragraph id="493936C4997FFFDB2FD5FB0DFA6BF359" blockId="9.[189,1399,1217,1347]" pageId="9" pageNumber="10">
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We decide not to follow the attempt of classification proposed by
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<bibRefCitation id="2D174B35997FFFDB2CA1FB0DFD74F2C1" author="Schram F. R." box="[969,1145,1217,1243]" pageId="9" pageNumber="10" pagination="340 - 363" refId="ref28925" refString="Schram F. R. 2014. Family level classification within Thylacocephala, with comments on their evolution and possible relationships. Crustaceana 87 (3): 340 - 363. https: // doi. org / 10.1163 / 15685403 - 00003289" type="journal article" year="2014">Schram (2014)</bibRefCitation>
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as it has been shown to present many issues (e.g., polyphyly of Concavicarida
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<bibRefCitation id="2D174B35997FFFDB2C1AFB2FFD7AF2E7" author="Briggs D. E. G. & Rolfe W. D. I." box="[882,1143,1251,1277]" pageId="9" pageNumber="10" pagination="249 - 276" refId="ref24663" refString="Briggs D. E. G. & Rolfe W. D. I. 1983. New Concavicarida (new order:? Crustacea) from the Upper Devonian of Gogo, Western Australia, and the palaeoecology and affinities of the group. Special Papers in Palaeontology 30: 249 - 276." type="journal article" year="1983">Briggs & Rolfe, 1983</bibRefCitation>
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and Conchyliocarida
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<bibRefCitation id="2D174B35997FFFDB2FD7FACAF87FF33A" author="Secretan S. & Riou B." box="[191,370,1286,1312]" pageId="9" pageNumber="10" pagination="59 - 97" refId="ref29121" refString="Secretan S. & Riou B. 1983. Un groupe enigmatique de Crustaces: ses representants du Callovien de la Voulte-sur-Rhone (France). Annales de Paleontologie 69 (4): 59 - 97." type="journal article" year="1983">Secrétan, 1983</bibRefCitation>
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, problems with the definition of autapomorphies for many taxa; see
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<bibRefCitation id="2D174B35997FFFDB2BCCFACBFC7AF33A" author="Hegna T. A. & Vega F. J. & Gonzalez-Rodriguez K. A." box="[1188,1399,1286,1313]" pageId="9" pageNumber="10" pagination="606 - 616" refId="ref26392" refString="Hegna T. A., Vega F. J. & Gonzalez-Rodriguez K. A. 2014. First Mesozoic Thylacocephalans (Arthropoda,? Crustacea; Cretaceous) in the Western Hemisphere: new discoveries from the Muhi Quarry Lagerstatte. Journal of Paleontology 88 (3): 606 - 616. https: // doi. org / 10.1666 / 13 - 131" type="journal article" year="2014">
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Hegna
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<emphasis id="7BF2EAD6997FFFDB2B9FFACBFC3EF33A" box="[1271,1331,1286,1312]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
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2014
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</bibRefCitation>
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and
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<bibRefCitation id="2D174B35997FFFDB2F98FAE5F8D8F359" author="Laville T. & Haug C. & Haug J. T. & Forel M. - B. & Charbonnier S." box="[240,469,1321,1347]" pageId="9" pageNumber="10" pagination="289 - 320" refId="ref27343" refString="Laville T., Haug C., Haug J. T., Forel M. - B. & Charbonnier S. 2021 a. Morphology and anatomy of the Late Jurassic Mayrocaris bucculata (Eucrustacea?, Thylacocephala) with comments on the tagmosis of Thylacocephala. Journal of Systematic Palaeontology 19 (4): 289 - 320. https: // doi. org / 10.1080 / 14772019.2021.1910584" type="journal article" year="2021">
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Laville
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<emphasis id="7BF2EAD6997FFFDB2E21FAE5F889F359" box="[329,388,1321,1347]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
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2021a
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</bibRefCitation>
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,
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<bibRefCitation id="2D174B35997FFFDB2E89FAE5FB26F359" author="Laville T. & Haug J. T. & Haug C." box="[481,555,1321,1347]" pageId="9" pageNumber="10" pagination="295 - 310" refId="ref27416" refString="Laville T., Haug J. T. & Haug C. 2021 b. New species of Thylacocephala, Eodollocaris keithflinti n. gen., n. sp., from the Mazon Creek Lagerstatte, Illinois, United States (c. 307 Ma) and redescription of other Mazon Creek thylacocephalans. Geodiversitas 43 (10): 295 - 310. https: // doi. org / 10.5252 / geodiversitas 2021 v 43 a 10" type="journal article" year="2021">2021b</bibRefCitation>
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for detailed explanations).
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</paragraph>
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</subSubSection>
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</treatment>
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