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<document ID-DOI="http://dx.doi.org/10.3897/zse.99.99227" ID-Pensoft-Pub="1860-0743-1-273" ID-Pensoft-UUID="F3161813062A5D37A0C4DBE590620DF8" ID-ZooBank="86213715F1FC4838BC478464DC0B9F4B" ModsDocID="1860-0743-99-1-273" checkinTime="1680107811176" checkinUser="pensoft" docAuthor="Villarreal, Osvaldo &amp; Chame-Vazquez, David" docDate="2023" docId="5A9658223F055671A90BD59088A40312" docLanguage="en" docName="ZoosystEvol 99(1): 273-280" docOrigin="Zoosystematics and Evolution 99 (1)" docPubDate="2023-03-29" docSource="http://dx.doi.org/10.3897/zse.99.99227" docTitle="Drymusa huberi Villarreal &amp; Chamé-Vázquez 2023, sp. nov." docType="treatment" docUuid="32F46572-64B6-40C6-94E8-5126B59DA6BC" docUuidSource="ZooBank" docVersion="1" id="F3161813062A5D37A0C4DBE590620DF8" lastPageNumber="273" masterDocId="F3161813062A5D37A0C4DBE590620DF8" masterDocTitle="First record of the false violin spider of the family Drymusidae (Araneae, Synspermiata, Scytodoidea) from Venezuela, with the description of a new species" masterLastPageNumber="280" masterPageNumber="273" pageNumber="273" updateTime="1680107811176" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>First record of the false violin spider of the family Drymusidae (Araneae, Synspermiata, Scytodoidea) from Venezuela, with the description of a new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Villarreal, Osvaldo</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5355-3723</mods:nameIdentifier>
<mods:affiliation>Departamento de Invertebrados - Laboratorio de Aracnologia, Campus de Ensino e Pesquisa do Museu Nacional - Predio Administrativo, Avenida Bartolomeu de Gusmao, 875, RJ, CEP 20941 - 160 Sao Cristovao, Rio de Janeiro, Brazil &amp; Museo del Instituto de Zoologia Agricola, Facultad de Agronomia, Universidad Central de Venezuela, Av 19 de Abril c / c Av Casanova Godoy, Maracay, Aragua, Venezuela</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chame-Vazquez, David</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3433-153X</mods:nameIdentifier>
<mods:affiliation>Laboratorio de Aracnologia y Entomologia, Centro de Investigaciones Biologicas del Noroeste S. C., Instituto Politecnico Nacional 195, Col. Playa Palo de Santa Rita Sur, CP. 23096, La Paz, Baja California Sur, Mexico</mods:affiliation>
<mods:nameIdentifier type="email">chamevazquez@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:titleInfo>
<mods:title>Zoosystematics and Evolution</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-29</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>99</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
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<mods:start>273</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zse.99.99227</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.99.99227</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-0743-1-273</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:32F46572-64B6-40C6-94E8-5126B59DA6BC" httpUri="http://treatment.plazi.org/id/5A9658223F055671A90BD59088A40312" lastPageNumber="273" pageId="0" pageNumber="273">
<subSubSection pageId="0" pageNumber="273" type="nomenclature">
<paragraph pageId="0" pageNumber="273">
<taxonomicName LSID="https://zoobank.org/32F46572-64B6-40C6-94E8-5126B59DA6BC" authority="Villarreal &amp; Chamé-Vázquez, 2023" authorityName="Villarreal &amp; Chamé-Vázquez" authorityYear="2023" class="Arachnida" family="Drymusidae" genus="Drymusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymusa huberi" order="Araneae" pageId="0" pageNumber="273" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov.">Drymusa huberi</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="273">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="273" type="description">
<paragraph pageId="0" pageNumber="273">
<figureCitation captionStart="Figures 112" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">Figs 1-12</figureCitation>
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">, 13-20</figureCitation>
<figureCitation captionStart="Figures 2132" captionStartId="F3" captionText="Figures 21 - 32. Drymusa spp., females. 21. D. simoni, posterior part of opisthosoma, lateral view; 22. D. dinora, opisthosoma, ventral view; 23, 25 - 27. Female genitalia, ventral view (23. D. colligata; 25. D. spectata; 26. D. armasi; 27. D. serrana); 24, 28 - 32. Same, dorsal view (24. D. huberi sp. nov.; 28. D. serrana; 29. D. canhemabae; 30. D. colligata; 31. D. rengan; 32. D. spelunca)." figureDoi="10.3897/zse.99.99227.figures21-32" httpUri="https://binary.pensoft.net/fig/831283" pageId="0" pageNumber="273">, 24</figureCitation>
<figureCitation captionStart="Figures 4953" captionStartId="F5" captionText="Figures 49 - 53. 49, 50. Drymusa huberi sp. nov. live specimens from La Guaquira, Yaracuy State, Venezuela. 49. Male, paratype; 50. Female, holotype; 51. Habitat, tropical dry forest in La Guaquira, Yaracuy State; 52. Leaf litter on the forest floor, microhabitat of the species; 53. Known distribution of all species of the genus Drymusa. Photo credit: O. Villarreal: 49, 50, 51; D. Romero: 52." figureDoi="10.3897/zse.99.99227.figures49-53" httpUri="https://binary.pensoft.net/fig/831285" pageId="0" pageNumber="273">, 49</figureCitation>
<figureCitation captionStart="Figures 4953" captionStartId="F5" captionText="Figures 49 - 53. 49, 50. Drymusa huberi sp. nov. live specimens from La Guaquira, Yaracuy State, Venezuela. 49. Male, paratype; 50. Female, holotype; 51. Habitat, tropical dry forest in La Guaquira, Yaracuy State; 52. Leaf litter on the forest floor, microhabitat of the species; 53. Known distribution of all species of the genus Drymusa. Photo credit: O. Villarreal: 49, 50, 51; D. Romero: 52." figureDoi="10.3897/zse.99.99227.figures49-53" httpUri="https://binary.pensoft.net/fig/831285" pageId="0" pageNumber="273">, 50</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="273" type="type data">
<paragraph pageId="0" pageNumber="273">Type data.</paragraph>
<paragraph pageId="0" pageNumber="273">
Venezuela • ♀
<emphasis bold="true" italics="true" pageId="0" pageNumber="273">holotype</emphasis>
: MIZA0105853; Yaracuy, La
<normalizedToken originalValue="Guáquira">Guaquira</normalizedToken>
;
<geoCoordinate degrees="10.2951" direction="north" orientation="latitude" precision="5" value="10.2951">10.2951°N</geoCoordinate>
,
<geoCoordinate degrees="68.6535" direction="west" orientation="longitude" precision="5" value="-68.6535">68.6535°W</geoCoordinate>
; 120 m a.s.l.; 16 Feb 2020; O. Villarreal, B.A. Huber, Q. Arias C. leg.; forest along stream • Paratype: 1 ♂ same data as the holotype.
</paragraph>
<caption doi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273" start="Figures 112" startId="F1">
<paragraph pageId="0" pageNumber="273">
<emphasis bold="true" pageId="0" pageNumber="273">Figures 1-12.</emphasis>
<taxonomicName authorityName="Villarreal &amp; Chamé-Vázquez" authorityYear="2023" class="Arachnida" family="Drymusidae" genus="Drymusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymusa huberi" order="Araneae" pageId="0" pageNumber="273" phylum="Arthropoda" rank="species" species="huberi">
<emphasis italics="true" pageId="0" pageNumber="273">Drymusa huberi</emphasis>
</taxonomicName>
sp. nov., female holotype (
<emphasis bold="true" pageId="0" pageNumber="273">1-4, 9, 11</emphasis>
) and male paratype (
<emphasis bold="true" pageId="0" pageNumber="273">5-8, 10, 12</emphasis>
).
<emphasis bold="true" pageId="0" pageNumber="273">1, 5.</emphasis>
Prosoma, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="273">3, 7.</emphasis>
Same, ventral view;
<emphasis bold="true" pageId="0" pageNumber="273">2, 6.</emphasis>
Opisthosoma, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="273">4, 8, 11</emphasis>
Same, ventral view (
<emphasis bold="true" pageId="0" pageNumber="273">11</emphasis>
depilated);
<emphasis bold="true" pageId="0" pageNumber="273">9, 10.</emphasis>
Prosoma, frontal view;
<emphasis bold="true" pageId="0" pageNumber="273">12.</emphasis>
Same, lateral view. Scale bars: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="273">1-12</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273" start="Figures 1320" startId="F2">
<paragraph pageId="0" pageNumber="273">
<emphasis bold="true" pageId="0" pageNumber="273">Figures 13-20.</emphasis>
<taxonomicName authorityName="Villarreal &amp; Chamé-Vázquez" authorityYear="2023" class="Arachnida" family="Drymusidae" genus="Drymusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymusa huberi" order="Araneae" pageId="0" pageNumber="273" phylum="Arthropoda" rank="species" species="huberi">
<emphasis italics="true" pageId="0" pageNumber="273">Drymusa huberi</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="273">13.</emphasis>
Female genitalia, ventral view;
<emphasis bold="true" pageId="0" pageNumber="273">14.</emphasis>
Same (cleared), dorsal view;
<emphasis bold="true" pageId="0" pageNumber="273">15.</emphasis>
Male palp, prolateral view;
<emphasis bold="true" pageId="0" pageNumber="273">16, 20.</emphasis>
Same, ventral view (
<emphasis bold="true" pageId="0" pageNumber="273">20</emphasis>
cleared);
<emphasis bold="true" pageId="0" pageNumber="273">17.</emphasis>
Detail of vulva (cleared), dorsal view;
<emphasis bold="true" pageId="0" pageNumber="273">18.</emphasis>
Male palpal bulb, apical view;
<emphasis bold="true" pageId="0" pageNumber="273">19.</emphasis>
Same, ventral view. Scale bars: 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="273">13-16, 18-20</emphasis>
). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="273" type="etymology">
<paragraph pageId="0" pageNumber="273">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="273">The specific epithet is a name in apposition to honour the arachnologist Bernhard A. Huber in recognition of his prolific and excellent contributions to the taxonomy and systematics of the pholcid spiders of Venezuela and the world. Bernhard was also one of the collectors of the type specimens.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="273" type="diagnosis">
<paragraph pageId="0" pageNumber="273">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="273">
The vulva of
<taxonomicName lsidName="D. huberi" pageId="0" pageNumber="273" rank="species" species="huberi">
<emphasis italics="true" pageId="0" pageNumber="273">D. huberi</emphasis>
</taxonomicName>
sp. nov. resembles those of
<taxonomicName lsidName="D. canhemabae" pageId="0" pageNumber="273" rank="species" species="canhemabae">
<emphasis italics="true" pageId="0" pageNumber="273">D. canhemabae</emphasis>
</taxonomicName>
Brescovit, Bonaldo &amp; Rheims, 2004 and
<taxonomicName lsidName="D. colligata" pageId="0" pageNumber="273" rank="species" species="colligata">
<emphasis italics="true" pageId="0" pageNumber="273">D. colligata</emphasis>
</taxonomicName>
Bonaldo, Rheims &amp; Brescovit, 2006 by the unbranched, long spermathecae (Figs
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">14</figureCitation>
,
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">17</figureCitation>
). They differ by the spermathecae with a long stalk that is not coiled and the distal part globular (Figs
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">14</figureCitation>
,
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">17</figureCitation>
), whereas
<taxonomicName lsidName="D. canhemabae" pageId="0" pageNumber="273" rank="species" species="canhemabae">
<emphasis italics="true" pageId="0" pageNumber="273">D. canhemabae</emphasis>
</taxonomicName>
has coiled spermathecae (see
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.697.1.1" author="Brescovit, AD" journalOrPublisher="Zootaxa" pageId="0" pageNumber="273" pagination="1 - 5" refId="B3" refString="Brescovit, AD, Bonaldo, AB, Rheims, CA, 2004. A new species of Drymusa Simon, 1891 (Araneae, Drymusidae) from Brazil. Zootaxa 697 (1): 1 - 5, DOI: https://doi.org/10.11646/zootaxa.697.1.1" title="A new species of Drymusa Simon, 1891 (Araneae, Drymusidae) from Brazil." url="https://doi.org/10.11646/zootaxa.697.1.1" volume="697" year="2004">Brescovit et al. (2004)</bibRefCitation>
: fig. 5; Fig.
<figureCitation captionStart="Figures 2132" captionStartId="F3" captionText="Figures 21 - 32. Drymusa spp., females. 21. D. simoni, posterior part of opisthosoma, lateral view; 22. D. dinora, opisthosoma, ventral view; 23, 25 - 27. Female genitalia, ventral view (23. D. colligata; 25. D. spectata; 26. D. armasi; 27. D. serrana); 24, 28 - 32. Same, dorsal view (24. D. huberi sp. nov.; 28. D. serrana; 29. D. canhemabae; 30. D. colligata; 31. D. rengan; 32. D. spelunca)." figureDoi="10.3897/zse.99.99227.figures21-32" httpUri="https://binary.pensoft.net/fig/831283" pageId="0" pageNumber="273">29</figureCitation>
) and
<taxonomicName lsidName="D. colligata" pageId="0" pageNumber="273" rank="species" species="colligata">
<emphasis italics="true" pageId="0" pageNumber="273">D. colligata</emphasis>
</taxonomicName>
has shorter spermatheca, with its distal part swollen (not globular) (see
<bibRefCitation DOI="https://doi.org/10.1590/S0101-81752006000200020" author="Bonaldo, AB" journalOrPublisher="Revista Brasileira de Zoologia" pageId="0" pageNumber="273" pagination="455 - 459" refId="B2" refString="Bonaldo, AB, Rheims, CA, Brescovit, AD, 2006. Four new species of Drymusa Simon (Araneae, Drymusidae) from Brazilian Oriental Amazonia. Revista Brasileira de Zoologia 23 (2): 455 - 459, DOI: https://doi.org/10.1590/S0101-81752006000200020" title="Four new species of Drymusa Simon (Araneae, Drymusidae) from Brazilian Oriental Amazonia." url="https://doi.org/10.1590/S0101-81752006000200020" volume="23" year="2006">Bonaldo et al. (2006)</bibRefCitation>
: fig. 6; Fig.
<figureCitation captionStart="Figures 2132" captionStartId="F3" captionText="Figures 21 - 32. Drymusa spp., females. 21. D. simoni, posterior part of opisthosoma, lateral view; 22. D. dinora, opisthosoma, ventral view; 23, 25 - 27. Female genitalia, ventral view (23. D. colligata; 25. D. spectata; 26. D. armasi; 27. D. serrana); 24, 28 - 32. Same, dorsal view (24. D. huberi sp. nov.; 28. D. serrana; 29. D. canhemabae; 30. D. colligata; 31. D. rengan; 32. D. spelunca)." figureDoi="10.3897/zse.99.99227.figures21-32" httpUri="https://binary.pensoft.net/fig/831283" pageId="0" pageNumber="273">30</figureCitation>
). Females of
<taxonomicName lsidName="D. huberi" pageId="0" pageNumber="273" rank="species" species="huberi">
<emphasis italics="true" pageId="0" pageNumber="273">D. huberi</emphasis>
</taxonomicName>
sp. nov. also resemble those of
<taxonomicName lsidName="D. dinora" pageId="0" pageNumber="273" rank="species" species="dinora">
<emphasis italics="true" pageId="0" pageNumber="273">D. dinora</emphasis>
</taxonomicName>
Valerio, 1971 by the long and broad epigastric plate, but
<taxonomicName lsidName="D. huberi" pageId="0" pageNumber="273" rank="species" species="huberi">
<emphasis italics="true" pageId="0" pageNumber="273">D. huberi</emphasis>
</taxonomicName>
sp. nov. has a median hood on the post-epigastric plate, absent in
<taxonomicName lsidName="D. dinora" pageId="0" pageNumber="273" rank="species" species="dinora">
<emphasis italics="true" pageId="0" pageNumber="273">D. dinora</emphasis>
</taxonomicName>
(see
<bibRefCitation DOI="https://doi.org/10.2307/3493570" author="Valerio, CE" journalOrPublisher="The Florida Entomologist" pageId="0" pageNumber="273" pagination="193 - 200" refId="B10" refString="Valerio, CE, 1971. The spider genus Drymusa in the New World (Araneae: Scytodidae). The Florida Entomologist 54 (2): 193 - 200, DOI: https://doi.org/10.2307/3493570" title="The spider genus Drymusa in the New World (Araneae: Scytodidae)." url="https://doi.org/10.2307/3493570" volume="54" year="1971">Valerio (1971)</bibRefCitation>
: figs 8, 9; Fig.
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">20</figureCitation>
),
<taxonomicName lsidName="D. canhemabae" pageId="0" pageNumber="273" rank="species" species="canhemabae">
<emphasis italics="true" pageId="0" pageNumber="273">D. canhemabae</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="D. colligata" pageId="0" pageNumber="273" rank="species" species="colligata">
<emphasis italics="true" pageId="0" pageNumber="273">D. colligata</emphasis>
</taxonomicName>
. Males of
<taxonomicName lsidName="D. huberi" pageId="0" pageNumber="273" rank="species" species="huberi">
<emphasis italics="true" pageId="0" pageNumber="273">D. huberi</emphasis>
</taxonomicName>
sp. nov. can be easily distinguished from all congeneric species by having the embolus shorter than the bulb length (Figs
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">15</figureCitation>
,
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">16</figureCitation>
,
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">18</figureCitation>
,
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="273" type="description">
<paragraph pageId="0" pageNumber="273">Description.</paragraph>
<paragraph pageId="0" pageNumber="273">
<emphasis bold="true" pageId="0" pageNumber="273">
Female (Holotype).
<emphasis italics="true" pageId="0" pageNumber="273">Habitus</emphasis>
.
</emphasis>
Carapace pear-shaped, pars cephalica elongated, with many long, procumbent setae (Figs
<figureCitation captionStart="Figures 112" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">1</figureCitation>
,
<figureCitation captionStart="Figures 112" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">9</figureCitation>
). Opisthosoma elongated, with posterior two thirds attenuated and covered sparsely with long, brown setae (Fig.
<figureCitation captionStart="Figures 112" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">2</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="273">
<emphasis italics="true" pageId="0" pageNumber="273">Colouration</emphasis>
.
</emphasis>
Carapace yellowish, margins on pars thoracica tinged with purple-brown and a median broad V-shaped pattern; pars cephalica reticulated with purple-brown maculations (Fig.
<figureCitation captionStart="Figures 112" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">1</figureCitation>
). Chelicerae orange, with brownish maculations. Endites and labium purple-brown, the latter darker than the former. Sternum purple-brown, except on lateral margins, which are yellowish, with one darker spot on the posterior margin (arrow in Fig.
<figureCitation captionStart="Figures 112" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">3</figureCitation>
). Palp with femur and patella mostly yellowish, tinged with purple-brown on lateral sides; tibia and tarsus mostly purple-brown.
<emphasis bold="true" italics="true" pageId="0" pageNumber="273">Legs</emphasis>
: femora mostly purple-brown, with one basal and one distal yellowish ring; patellae mostly purple-brown; tibiae mostly yellowish, with one basal, thin purple-brown ring and one distal purple-brown ring; metatarsi and tarsi yellowish. Dorsum of prosoma grey-white with three W-shaped dark brown bands, posterior third mostly dark brown (Fig.
<figureCitation captionStart="Figures 112" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">2</figureCitation>
). Venter purple-brown, with pale-yellowish spots on the epigastric area (Fig.
<figureCitation captionStart="Figures 112" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">4</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="273">
<emphasis italics="true" pageId="0" pageNumber="273">Female genitalia</emphasis>
.
</emphasis>
Epigastric plate rectangular, large and about half the opisthosoma length (Figs
<figureCitation captionStart="Figures 112" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">4</figureCitation>
,
<figureCitation captionStart="Figures 112" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">11</figureCitation>
); posterior corners are more sclerotised than the plate (arrow in Fig.
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">13</figureCitation>
). Post-epigastric plate crescent-shaped, with a small median hood. Vulva with the distal part of spermatheca globular, on long, thin stalks, arising on lateral corners of the epigastric plate (Figs
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">14</figureCitation>
,
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">17</figureCitation>
). Spermatheca with gland ductules well-spaced on the distal half (Fig.
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">17</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="273">
<emphasis italics="true" pageId="0" pageNumber="273">Measurements</emphasis>
.
</emphasis>
Carapace 1.32 long, 1.02 wide; eye diameters: PME 0.10, ALE 0.09, PLE 0.06; labium 0.30 long, 0.30 wide; sternum 0.69 long, 0.65 wide. Legs: I 10.8 (2.82, 0.36, 2.97, 3.06, 0.87), II 8.76 (2.46, 0.34, 2.49, 2.65, 0.82), III 6.87 (1.95, 0.39, 1.74, 2.04, 0.75), IV 8.59 (2.35, 0.33, 2.43, 2.61, 0.87). Leg formula: 1243.
</paragraph>
<caption doi="10.3897/zse.99.99227.figures21-32" httpUri="https://binary.pensoft.net/fig/831283" pageId="0" pageNumber="273" start="Figures 2132" startId="F3">
<paragraph pageId="0" pageNumber="273">
<emphasis bold="true" pageId="0" pageNumber="273">Figures 21-32.</emphasis>
<taxonomicName class="Arachnida" family="Drymusidae" genus="Drymusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymusa" order="Araneae" pageId="0" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="273">Drymusa</emphasis>
</taxonomicName>
spp., females.
<emphasis bold="true" pageId="0" pageNumber="273">21.</emphasis>
<taxonomicName lsidName="D. simoni" pageId="0" pageNumber="273" rank="species" species="simoni">
<emphasis italics="true" pageId="0" pageNumber="273">D. simoni</emphasis>
</taxonomicName>
, posterior part of opisthosoma, lateral view;
<emphasis bold="true" pageId="0" pageNumber="273">22.</emphasis>
<taxonomicName lsidName="D. dinora" pageId="0" pageNumber="273" rank="species" species="dinora">
<emphasis italics="true" pageId="0" pageNumber="273">D. dinora</emphasis>
</taxonomicName>
, opisthosoma, ventral view;
<emphasis bold="true" pageId="0" pageNumber="273">23, 25-27.</emphasis>
Female genitalia, ventral view (
<emphasis bold="true" pageId="0" pageNumber="273">23.</emphasis>
<taxonomicName lsidName="D. colligata" pageId="0" pageNumber="273" rank="species" species="colligata">
<emphasis italics="true" pageId="0" pageNumber="273">D. colligata</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="273">25.</emphasis>
<taxonomicName lsidName="D. spectata" pageId="0" pageNumber="273" rank="species" species="spectata">
<emphasis italics="true" pageId="0" pageNumber="273">D. spectata</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="273">26.</emphasis>
<taxonomicName lsidName="D. armasi" pageId="0" pageNumber="273" rank="species" species="armasi">
<emphasis italics="true" pageId="0" pageNumber="273">D. armasi</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="273">27.</emphasis>
<taxonomicName lsidName="D. serrana" pageId="0" pageNumber="273" rank="species" species="serrana">
<emphasis italics="true" pageId="0" pageNumber="273">D. serrana</emphasis>
</taxonomicName>
);
<emphasis bold="true" pageId="0" pageNumber="273">24, 28-32.</emphasis>
Same, dorsal view (
<emphasis bold="true" pageId="0" pageNumber="273">24.</emphasis>
<taxonomicName lsidName="D. huberi" pageId="0" pageNumber="273" rank="species" species="huberi">
<emphasis italics="true" pageId="0" pageNumber="273">D. huberi</emphasis>
</taxonomicName>
sp. nov.;
<emphasis bold="true" pageId="0" pageNumber="273">28.</emphasis>
<taxonomicName lsidName="D. serrana" pageId="0" pageNumber="273" rank="species" species="serrana">
<emphasis italics="true" pageId="0" pageNumber="273">D. serrana</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="273">29.</emphasis>
<taxonomicName lsidName="D. canhemabae" pageId="0" pageNumber="273" rank="species" species="canhemabae">
<emphasis italics="true" pageId="0" pageNumber="273">D. canhemabae</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="273">30.</emphasis>
<taxonomicName lsidName="D. colligata" pageId="0" pageNumber="273" rank="species" species="colligata">
<emphasis italics="true" pageId="0" pageNumber="273">D. colligata</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="273">31.</emphasis>
<taxonomicName lsidName="D. rengan" pageId="0" pageNumber="273" rank="species" species="rengan">
<emphasis italics="true" pageId="0" pageNumber="273">D. rengan</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="273">32.</emphasis>
<taxonomicName lsidName="D. spelunca" pageId="0" pageNumber="273" rank="species" species="spelunca">
<emphasis italics="true" pageId="0" pageNumber="273">D. spelunca</emphasis>
</taxonomicName>
).
</paragraph>
</caption>
<caption doi="10.3897/zse.99.99227.figures33-48" httpUri="https://binary.pensoft.net/fig/831284" pageId="0" pageNumber="273" start="Figures 3348" startId="F4">
<paragraph pageId="0" pageNumber="273">
<emphasis bold="true" pageId="0" pageNumber="273">Figures 33-48.</emphasis>
<taxonomicName class="Arachnida" family="Drymusidae" genus="Drymusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymusa" order="Araneae" pageId="0" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="273">Drymusa</emphasis>
</taxonomicName>
spp., male palp (
<emphasis bold="true" pageId="0" pageNumber="273">33, 34.</emphasis>
<taxonomicName lsidName="D. rengan" pageId="0" pageNumber="273" rank="species" species="rengan">
<emphasis italics="true" pageId="0" pageNumber="273">D. rengan</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="273">35, 36.</emphasis>
<taxonomicName lsidName="D. serrana" pageId="0" pageNumber="273" rank="species" species="serrana">
<emphasis italics="true" pageId="0" pageNumber="273">D. serrana</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="273">37, 38.</emphasis>
<taxonomicName lsidName="D. spelunca" pageId="0" pageNumber="273" rank="species" species="spelunca">
<emphasis italics="true" pageId="0" pageNumber="273">D. spelunca</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="273">39.</emphasis>
<taxonomicName lsidName="D. dinora" pageId="0" pageNumber="273" rank="species" species="dinora">
<emphasis italics="true" pageId="0" pageNumber="273">D. dinora</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="273">40.</emphasis>
<taxonomicName lsidName="D. armasi" pageId="0" pageNumber="273" rank="species" species="armasi">
<emphasis italics="true" pageId="0" pageNumber="273">D. armasi</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="273">41.</emphasis>
<taxonomicName lsidName="D. spectata" pageId="0" pageNumber="273" rank="species" species="spectata">
<emphasis italics="true" pageId="0" pageNumber="273">D. spectata</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="273">42.</emphasis>
<taxonomicName lsidName="D. tobyi" pageId="0" pageNumber="273" rank="species" species="tobyi">
<emphasis italics="true" pageId="0" pageNumber="273">D. tobyi</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="273">43.</emphasis>
<taxonomicName lsidName="D. canhemabae" pageId="0" pageNumber="273" rank="species" species="canhemabae">
<emphasis italics="true" pageId="0" pageNumber="273">D. canhemabae</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="273">44, 45.</emphasis>
<taxonomicName lsidName="D. colligata" pageId="0" pageNumber="273" rank="species" species="colligata">
<emphasis italics="true" pageId="0" pageNumber="273">D. colligata</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="273">46, 47.</emphasis>
<taxonomicName lsidName="D. philomatica" pageId="0" pageNumber="273" rank="species" species="philomatica">
<emphasis italics="true" pageId="0" pageNumber="273">D. philomatica</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="273">48.</emphasis>
<taxonomicName lsidName="D. simoni" pageId="0" pageNumber="273" rank="species" species="simoni">
<emphasis italics="true" pageId="0" pageNumber="273">D. simoni</emphasis>
</taxonomicName>
).
<emphasis bold="true" pageId="0" pageNumber="273">33, 35, 37, 40, 41, 44, 46, 48.</emphasis>
Palp, prolateral view;
<emphasis bold="true" pageId="0" pageNumber="273">34, 36, 38, 39, 42, 43, 45, 47.</emphasis>
Palp, retrolateral view.
</paragraph>
</caption>
<caption doi="10.3897/zse.99.99227.figures49-53" httpUri="https://binary.pensoft.net/fig/831285" pageId="0" pageNumber="273" start="Figures 4953" startId="F5">
<paragraph pageId="0" pageNumber="273">
<emphasis bold="true" pageId="0" pageNumber="273">Figures 49-53.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="273">49, 50.</emphasis>
<taxonomicName authorityName="Villarreal &amp; Chamé-Vázquez" authorityYear="2023" class="Arachnida" family="Drymusidae" genus="Drymusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymusa huberi" order="Araneae" pageId="0" pageNumber="273" phylum="Arthropoda" rank="species" species="huberi">
<emphasis italics="true" pageId="0" pageNumber="273">Drymusa huberi</emphasis>
</taxonomicName>
sp. nov. live specimens from La
<normalizedToken originalValue="Guáquira">Guaquira</normalizedToken>
, Yaracuy State, Venezuela.
<emphasis bold="true" pageId="0" pageNumber="273">49.</emphasis>
Male, paratype;
<emphasis bold="true" pageId="0" pageNumber="273">50.</emphasis>
Female, holotype;
<emphasis bold="true" pageId="0" pageNumber="273">51.</emphasis>
Habitat, tropical dry forest in La
<normalizedToken originalValue="Guáquira">Guaquira</normalizedToken>
, Yaracuy State;
<emphasis bold="true" pageId="0" pageNumber="273">52.</emphasis>
Leaf litter on the forest floor, microhabitat of the species;
<emphasis bold="true" pageId="0" pageNumber="273">53.</emphasis>
Known distribution of all species of the genus
<taxonomicName class="Arachnida" family="Drymusidae" genus="Drymusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymusa" order="Araneae" pageId="0" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="273">Drymusa</emphasis>
</taxonomicName>
. Photo credit: O. Villarreal:
<emphasis bold="true" pageId="0" pageNumber="273">49, 50, 51</emphasis>
; D. Romero:
<emphasis bold="true" pageId="0" pageNumber="273">52</emphasis>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="273">
<emphasis bold="true" pageId="0" pageNumber="273">Male (paratype).</emphasis>
Habitus as the female (Figs
<figureCitation captionStart="Figures 112" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">5-8</figureCitation>
,
<figureCitation captionStart="Figures 112" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">10</figureCitation>
,
<figureCitation captionStart="Figures 112" captionStartId="F1" captionText="Figures 1 - 12. Drymusa huberi sp. nov., female holotype (1 - 4, 9, 11) and male paratype (5 - 8, 10, 12). 1, 5. Prosoma, dorsal view; 3, 7. Same, ventral view; 2, 6. Opisthosoma, dorsal view; 4, 8, 11 Same, ventral view (11 depilated); 9, 10. Prosoma, frontal view; 12. Same, lateral view. Scale bars: 0.5 mm (1 - 12)." figureDoi="10.3897/zse.99.99227.figures1-12" httpUri="https://binary.pensoft.net/fig/831281" pageId="0" pageNumber="273">12</figureCitation>
), except pars cephalic with few short setae, mostly on clypeal area, opisthosoma with posterior third attenuated.
<emphasis bold="true" pageId="0" pageNumber="273">
<emphasis italics="true" pageId="0" pageNumber="273">Colouration</emphasis>
.
</emphasis>
As the female, but the purple-brown median pattern and lateral margins are narrower; the sternum maculation is light purple; palp yellowish with scattered purple-brown maculations, tarsus darker than other segments. Dorsum of opisthosoma with transversal bands narrower, elongated, fused with the next one; venter yellowish, except posterior third, which is light purple.
<emphasis bold="true" pageId="0" pageNumber="273">
<emphasis italics="true" pageId="0" pageNumber="273">Palp</emphasis>
.
</emphasis>
Sub-cylindrical femur longer than tibia; tibia slightly more thickened than femur and slightly swollen at mid-length (Figs
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">15</figureCitation>
,
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">16</figureCitation>
,
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">20</figureCitation>
). Bulb subspherical, slightly longer than wide (Figs
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">15</figureCitation>
,
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">16</figureCitation>
,
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">18</figureCitation>
). Embolus spiniform, curved, directed ventral-retrolaterally and relatively short, about half the length of bulb (Figs
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">18</figureCitation>
,
<figureCitation captionStart="Figures 1320" captionStartId="F2" captionText="Figures 13 - 20. Drymusa huberi sp. nov. 13. Female genitalia, ventral view; 14. Same (cleared), dorsal view; 15. Male palp, prolateral view; 16, 20. Same, ventral view (20 cleared); 17. Detail of vulva (cleared), dorsal view; 18. Male palpal bulb, apical view; 19. Same, ventral view. Scale bars: 0.2 mm (13 - 16, 18 - 20). Abbreviations: E-embolus, ep-epigastric plate, F-fundus, Fm-femur, Ho-hood of the post-epigastric plate, pp-post-epigastric plate, Pt-patella, S-spermathecae, Sd-Sperm duct, Ss-Spermatheca stalk, Ta-tarsus, Ti-tibia." figureDoi="10.3897/zse.99.99227.figures13-20" httpUri="https://binary.pensoft.net/fig/831282" pageId="0" pageNumber="273">19</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="273">
<emphasis italics="true" pageId="0" pageNumber="273">Measurements</emphasis>
.
</emphasis>
Carapace 1.26 long, 1.05 wide; eye diameters: PME 0.12, ALE 0.09, PLE 0.07; labium 0.24 long, 0.27 wide; sternum 0.72 long, 0.72 wide. Legs: I missing, II 10.62 (2.97, 0.39, 3.09, 3.24, 0.93), III 7.77 (2.13, 0.36, 2.04, 2.40, 0.84), IV 10.20 (2.82, 0.36, 2.82, 3.09, 0.93). Leg formula: (1)243.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="273" type="distribution">
<paragraph pageId="0" pageNumber="273">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="273">
Only known from the type locality. See Fig.
<figureCitation captionStart="Figures 4953" captionStartId="F5" captionText="Figures 49 - 53. 49, 50. Drymusa huberi sp. nov. live specimens from La Guaquira, Yaracuy State, Venezuela. 49. Male, paratype; 50. Female, holotype; 51. Habitat, tropical dry forest in La Guaquira, Yaracuy State; 52. Leaf litter on the forest floor, microhabitat of the species; 53. Known distribution of all species of the genus Drymusa. Photo credit: O. Villarreal: 49, 50, 51; D. Romero: 52." figureDoi="10.3897/zse.99.99227.figures49-53" httpUri="https://binary.pensoft.net/fig/831285" pageId="0" pageNumber="273">53</figureCitation>
for the known distribution of all
<taxonomicName class="Arachnida" family="Drymusidae" genus="Drymusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymusa" order="Araneae" pageId="0" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="273">Drymusa</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="273" type="natural history">
<paragraph pageId="0" pageNumber="273">Natural history.</paragraph>
<paragraph pageId="0" pageNumber="273">
Both specimens were found in the leaf litter in a tropical dry forest along a streambed (Figs
<figureCitation captionStart="Figures 4953" captionStartId="F5" captionText="Figures 49 - 53. 49, 50. Drymusa huberi sp. nov. live specimens from La Guaquira, Yaracuy State, Venezuela. 49. Male, paratype; 50. Female, holotype; 51. Habitat, tropical dry forest in La Guaquira, Yaracuy State; 52. Leaf litter on the forest floor, microhabitat of the species; 53. Known distribution of all species of the genus Drymusa. Photo credit: O. Villarreal: 49, 50, 51; D. Romero: 52." figureDoi="10.3897/zse.99.99227.figures49-53" httpUri="https://binary.pensoft.net/fig/831285" pageId="0" pageNumber="273">51</figureCitation>
,
<figureCitation captionStart="Figures 4953" captionStartId="F5" captionText="Figures 49 - 53. 49, 50. Drymusa huberi sp. nov. live specimens from La Guaquira, Yaracuy State, Venezuela. 49. Male, paratype; 50. Female, holotype; 51. Habitat, tropical dry forest in La Guaquira, Yaracuy State; 52. Leaf litter on the forest floor, microhabitat of the species; 53. Known distribution of all species of the genus Drymusa. Photo credit: O. Villarreal: 49, 50, 51; D. Romero: 52." figureDoi="10.3897/zse.99.99227.figures49-53" httpUri="https://binary.pensoft.net/fig/831285" pageId="0" pageNumber="273">52</figureCitation>
); this species shares the microhabitat with
<taxonomicName class="Arachnida" family="Ochyroceratidae" genus="Ochyrocera" kingdom="Animalia" lsidName="Ochyrocera" order="Araneae" pageId="0" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="273">Ochyrocera</emphasis>
</taxonomicName>
sp. (
<taxonomicName authorityName="Fage" authorityYear="1912" class="Arachnida" family="Ochyroceratidae" kingdom="Animalia" lsidName="" order="Araneae" pageId="0" pageNumber="273" phylum="Arthropoda" rank="family">Ochyroceratidae</taxonomicName>
). For live specimens of
<taxonomicName lsidName="D. huberi" pageId="0" pageNumber="273" rank="species" species="huberi">
<emphasis italics="true" pageId="0" pageNumber="273">D. huberi</emphasis>
</taxonomicName>
sp. nov., see Figs
<figureCitation captionStart="Figures 4953" captionStartId="F5" captionText="Figures 49 - 53. 49, 50. Drymusa huberi sp. nov. live specimens from La Guaquira, Yaracuy State, Venezuela. 49. Male, paratype; 50. Female, holotype; 51. Habitat, tropical dry forest in La Guaquira, Yaracuy State; 52. Leaf litter on the forest floor, microhabitat of the species; 53. Known distribution of all species of the genus Drymusa. Photo credit: O. Villarreal: 49, 50, 51; D. Romero: 52." figureDoi="10.3897/zse.99.99227.figures49-53" httpUri="https://binary.pensoft.net/fig/831285" pageId="0" pageNumber="273">49</figureCitation>
,
<figureCitation captionStart="Figures 4953" captionStartId="F5" captionText="Figures 49 - 53. 49, 50. Drymusa huberi sp. nov. live specimens from La Guaquira, Yaracuy State, Venezuela. 49. Male, paratype; 50. Female, holotype; 51. Habitat, tropical dry forest in La Guaquira, Yaracuy State; 52. Leaf litter on the forest floor, microhabitat of the species; 53. Known distribution of all species of the genus Drymusa. Photo credit: O. Villarreal: 49, 50, 51; D. Romero: 52." figureDoi="10.3897/zse.99.99227.figures49-53" httpUri="https://binary.pensoft.net/fig/831285" pageId="0" pageNumber="273">50</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>