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<document id="F73732FC279F16FCCDFC806264AFEFF5" ID-DOI="10.5281/zenodo.196078" ID-GBIF-Dataset="3afa748f-2467-40ae-83ec-00984bfe7362" ID-ISSN="1175-5326" ID-Zenodo-Dep="196078" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460238428527" checkinUser="plazi" docAuthor="Fjellberg, Arne" docDate="2010" docId="504F2D6CFFD5FF8497BEF5CEFB431F21" docLanguage="en" docName="zt02513p049.pdf" docOrigin="Zootaxa 2513" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Desoria olympica Fjellberg, 2010, sp. n." docType="treatment" docVersion="14" lastPageNumber="40" masterDocId="AC765514FFD9FF899729F019FFC71D06" masterDocTitle="Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A." masterLastPageNumber="49" masterPageNumber="27" pageNumber="39" updateTime="1698239971832" updateUser="plazi">
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<mods:title id="4F8A94910D930F25258D298641A8BE1B">Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A.</mods:title>
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<mods:namePart id="76963858BFDD06B7DB90A9B6124CBE89">Fjellberg, Arne</mods:namePart>
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<mods:date id="5B25CE15A8923371F1F5D02E510A63B4">2010</mods:date>
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<mods:identifier id="CA33618DBD75839DA3B6AA1C5F8CA646" type="DOI">10.5281/zenodo.196078</mods:identifier>
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<treatment id="504F2D6CFFD5FF8497BEF5CEFB431F21" ID-DOI="http://doi.org/10.5281/zenodo.5670731" ID-GBIF-Taxon="119389108" ID-Zenodo-Dep="5670731" LSID="urn:lsid:plazi:treatment:504F2D6CFFD5FF8497BEF5CEFB431F21" httpUri="http://treatment.plazi.org/id/504F2D6CFFD5FF8497BEF5CEFB431F21" lastPageId="13" lastPageNumber="40" pageId="12" pageNumber="39">
<subSubSection id="90FCCFF1FFD5FF8597BEF5CEFEF31B12" pageId="12" pageNumber="39" type="nomenclature">
<paragraph id="D8599C7AFFD5FF8597BEF5CEFE7418F7" blockId="12.[151,435,1495,1556]" box="[151,435,1495,1521]" pageId="12" pageNumber="39">
<heading id="83112B16FFD5FF8597BEF5CEFE7418F7" bold="true" box="[151,435,1495,1521]" fontSize="11" level="1" pageId="12" pageNumber="39" reason="1">
<emphasis id="EA924068FFD5FF8597BEF5CEFE7418F7" bold="true" box="[151,435,1495,1521]" pageId="12" pageNumber="39">
<taxonomicName id="1FE6E7F9FFD5FF8597BEF5CEFEAD18F7" box="[151,362,1495,1521]" class="Entognatha" family="Isotomidae" genus="Desoria" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="39" phylum="Arthropoda" rank="species" species="olympica" status="sp. nov.">
<emphasis id="EA924068FFD5FF8597BEF5CEFEAD18F7" bold="true" box="[151,362,1495,1521]" italics="true" pageId="12" pageNumber="39">Desoria olympica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F1A1FD13FFD5FF859658F5CEFE7418F7" box="[369,435,1495,1521]" pageId="12" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D8599C7AFFD5FF8597BEF5E3FEF31B12" blockId="12.[151,435,1495,1556]" box="[151,308,1530,1556]" pageId="12" pageNumber="39">
(
<figureCitation id="40DD80FFFFD5FF8597B6F5E3FEEA1B12" box="[159,301,1530,1556]" captionStart="FIGURES 57 77" captionStartId="14.[151,270,1589,1613]" captionTargetBox="[159,1433,207,1545]" captionTargetId="figure@14.[151,1436,197,1568]" captionTargetPageId="14" captionText="FIGURES 57 77. (57 63) Desoria cryophila sp. n., (64 70) Desoria rosea sp. nov., (71 76) Desoria olympica sp. n., (77) Desoria capra sp. n., (57) sensillary chaetotaxy, positions of macrochaetae indicated, (58) eye-field and PAO, (59) left mucro, (60) sensilla on ventral side of right ant. 1, (61) ditto, right ant. 3, dorsal, (62) ditto, right ant. 2, lateral, (63) setae along midline on inner side of tib. 1, (64) eye-field and PAO, (65) sensilla on right ant. 2, lateral. es: erect sensilla, (66) ditto, ant. 3, (67) setae in right basomedian field of labium, the additional 5 th. seta encircled, (68) right mucro, lateral, (69) ditto, dorsal, with the secondary inner tooth formed by the truncated inner lamella marked, (70) dorsal setae in basal half of dens, (71) sensillary chaetotaxy, positions of macrochaetae indicated, (72) setae along midline on inner side of tib. 1, (73) left mucro and apical part of dens, (74) ventral side of manubrium, (75) eye-field and PAO, (76) labrum and frontoclypeal field, (77) right maxilla with details of capitular teeth (mt: membranous tooth)." httpUri="https://zenodo.org/record/196081/files/figure.png" pageId="12" pageNumber="39">Figs. 7176</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="90FCCFF1FFD5FF8597BEF65CFE281BAA" pageId="12" pageNumber="39" type="materials_examined">
<paragraph id="D8599C7AFFD5FF8597BEF65CFE281BAA" blockId="12.[151,1437,1605,2007]" pageId="12" pageNumber="39">
<emphasis id="EA924068FFD5FF8597BEF65CFE971B59" bold="true" box="[151,336,1605,1631]" pageId="12" pageNumber="39">
<typeStatus id="075D22D8FFD5FF8597BEF65CFF1F1B59" box="[151,216,1605,1631]" pageId="12" pageNumber="39">Type</typeStatus>
material
</emphasis>
(all in slides).
<typeStatus id="075D22D8FFD5FF859539F65CFD441B59" box="[528,643,1605,1631]" pageId="12" pageNumber="39" type="holotype">Holotype</typeStatus>
(male) and 10
<typeStatus id="075D22D8FFD5FF85946AF65CFC7D1B59" box="[835,954,1605,1631]" pageId="12" pageNumber="39" type="paratype">paratypes</typeStatus>
(in 4 slides) from &quot;
<collectingCountry id="A0F1DCEAFFD5FF859399F65CFB331B59" box="[1200,1268,1605,1631]" name="United States of America" pageId="12" pageNumber="39">USA</collectingCountry>
, Washington, Olympic National Park, Hurricane Ridge
<geoCoordinate id="BDD2FABDFFD5FF8595ACF674FD231B81" box="[645,740,1642,1671]" direction="north" orientation="latitude" pageId="12" pageNumber="39" precision="925" value="47.983334">47o59'N</geoCoordinate>
,
<geoCoordinate id="BDD2FABDFFD5FF8595D8F674FCA31B81" box="[753,868,1642,1671]" direction="west" orientation="longitude" pageId="12" pageNumber="39" precision="925" value="-123.51667">123o31'W</geoCoordinate>
,
<date id="AC58BABAFFD5FF859458F674FC3C1B81" box="[881,1019,1645,1671]" pageId="12" pageNumber="39" value="1983-09-10">10.IX.1983</date>
. Wet moss at snow edge,
<quantity id="1F1E319FFFD5FF859205F674FA551B81" box="[1324,1426,1645,1671]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="39" unit="m" value="1800.0">1,800 m</quantity>
., A. Fjellberg 422/83&quot; (
<collectionCode id="BEF704BFFFD5FF8596B1F68BFE251BAA" LSID="urn:lsid:biocol.org:col:34797" box="[408,482,1682,1708]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="12" pageNumber="39">INHS</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="90FCCFF1FFD5FF8497EDF6A1FB991C24" lastPageId="13" lastPageNumber="40" pageId="12" pageNumber="39" type="description">
<paragraph id="D8599C7AFFD5FF8497EDF6A1FB991C24" blockId="12.[151,1437,1605,2007]" lastBlockId="13.[151,1437,152,551]" lastPageId="13" lastPageNumber="40" pageId="12" pageNumber="39">
<emphasis id="EA924068FFD5FF8597EDF6A1FE901BD4" bold="true" box="[196,343,1720,1746]" pageId="12" pageNumber="39">Description</emphasis>
. Size up to 1.0 mm. Colour white, eye-spots black. Body shape as in two previous species, abd. 56 weakly separated. Characters of ocelli, labrum, labial and maxillary palps, maxillae and mandibles as in previous two species. PAO roundish, the size of a ocellus (
<figureCitation id="40DD80FFFFD5FF859457F71BFC1E1A1A" box="[894,985,1794,1820]" captionStart="FIGURES 57 77" captionStartId="14.[151,270,1589,1613]" captionTargetBox="[159,1433,207,1545]" captionTargetId="figure@14.[151,1436,197,1568]" captionTargetPageId="14" captionText="FIGURES 57 77. (57 63) Desoria cryophila sp. n., (64 70) Desoria rosea sp. nov., (71 76) Desoria olympica sp. n., (77) Desoria capra sp. n., (57) sensillary chaetotaxy, positions of macrochaetae indicated, (58) eye-field and PAO, (59) left mucro, (60) sensilla on ventral side of right ant. 1, (61) ditto, right ant. 3, dorsal, (62) ditto, right ant. 2, lateral, (63) setae along midline on inner side of tib. 1, (64) eye-field and PAO, (65) sensilla on right ant. 2, lateral. es: erect sensilla, (66) ditto, ant. 3, (67) setae in right basomedian field of labium, the additional 5 th. seta encircled, (68) right mucro, lateral, (69) ditto, dorsal, with the secondary inner tooth formed by the truncated inner lamella marked, (70) dorsal setae in basal half of dens, (71) sensillary chaetotaxy, positions of macrochaetae indicated, (72) setae along midline on inner side of tib. 1, (73) left mucro and apical part of dens, (74) ventral side of manubrium, (75) eye-field and PAO, (76) labrum and frontoclypeal field, (77) right maxilla with details of capitular teeth (mt: membranous tooth)." httpUri="https://zenodo.org/record/196081/files/figure.png" pageId="12" pageNumber="39">Fig. 75</figureCitation>
). Frontoclypeal field with three setae (
<figureCitation id="40DD80FFFFD5FF859789F731FF3D1A44" box="[160,250,1832,1858]" captionStart="FIGURES 57 77" captionStartId="14.[151,270,1589,1613]" captionTargetBox="[159,1433,207,1545]" captionTargetId="figure@14.[151,1436,197,1568]" captionTargetPageId="14" captionText="FIGURES 57 77. (57 63) Desoria cryophila sp. n., (64 70) Desoria rosea sp. nov., (71 76) Desoria olympica sp. n., (77) Desoria capra sp. n., (57) sensillary chaetotaxy, positions of macrochaetae indicated, (58) eye-field and PAO, (59) left mucro, (60) sensilla on ventral side of right ant. 1, (61) ditto, right ant. 3, dorsal, (62) ditto, right ant. 2, lateral, (63) setae along midline on inner side of tib. 1, (64) eye-field and PAO, (65) sensilla on right ant. 2, lateral. es: erect sensilla, (66) ditto, ant. 3, (67) setae in right basomedian field of labium, the additional 5 th. seta encircled, (68) right mucro, lateral, (69) ditto, dorsal, with the secondary inner tooth formed by the truncated inner lamella marked, (70) dorsal setae in basal half of dens, (71) sensillary chaetotaxy, positions of macrochaetae indicated, (72) setae along midline on inner side of tib. 1, (73) left mucro and apical part of dens, (74) ventral side of manubrium, (75) eye-field and PAO, (76) labrum and frontoclypeal field, (77) right maxilla with details of capitular teeth (mt: membranous tooth)." httpUri="https://zenodo.org/record/196081/files/figure.png" pageId="12" pageNumber="39">Fig. 76</figureCitation>
). Ant. 1 with only two ventroapical short sensilla in addition to a few setaceous ones, ant. 23 with only the standard set of sensilla. Ventromedian field of labium with 4 setae. Head with 3+3(4) postlabial setae. Body integument smooth; setal cover sparse and uniform. Macrochaetae distinct, smooth. Median macrochaetae on abd. 5 about 2.7 times as long as inner edge of last claw. Number of ordinary sensilla on tergites as 22/11135 (
<figureCitation id="40DD80FFFFD5FF8596BBF7A4FE2C1AD1" box="[402,491,1981,2007]" captionStart="FIGURES 57 77" captionStartId="14.[151,270,1589,1613]" captionTargetBox="[159,1433,207,1545]" captionTargetId="figure@14.[151,1436,197,1568]" captionTargetPageId="14" captionText="FIGURES 57 77. (57 63) Desoria cryophila sp. n., (64 70) Desoria rosea sp. nov., (71 76) Desoria olympica sp. n., (77) Desoria capra sp. n., (57) sensillary chaetotaxy, positions of macrochaetae indicated, (58) eye-field and PAO, (59) left mucro, (60) sensilla on ventral side of right ant. 1, (61) ditto, right ant. 3, dorsal, (62) ditto, right ant. 2, lateral, (63) setae along midline on inner side of tib. 1, (64) eye-field and PAO, (65) sensilla on right ant. 2, lateral. es: erect sensilla, (66) ditto, ant. 3, (67) setae in right basomedian field of labium, the additional 5 th. seta encircled, (68) right mucro, lateral, (69) ditto, dorsal, with the secondary inner tooth formed by the truncated inner lamella marked, (70) dorsal setae in basal half of dens, (71) sensillary chaetotaxy, positions of macrochaetae indicated, (72) setae along midline on inner side of tib. 1, (73) left mucro and apical part of dens, (74) ventral side of manubrium, (75) eye-field and PAO, (76) labrum and frontoclypeal field, (77) right maxilla with details of capitular teeth (mt: membranous tooth)." httpUri="https://zenodo.org/record/196081/files/figure.png" pageId="12" pageNumber="39">Fig. 71</figureCitation>
). Microsensilla as 11/001. Posterior sensilla on th. 2abd.
<date id="AC58BABAFFD5FF8593B7F7A4FB111AD1" box="[1182,1238,1981,2007]" pageId="12" pageNumber="39">4 set</date>
in the p-row; no ordinary sensillum laterally to the spine-like ms on abd. 3. Ventral tube with only 1+1 frontal setae; retinaculum with only two setae (
<specimenCount id="CEE057F3FFD4FF849535F0A4FD7C1DD1" box="[540,699,189,215]" pageId="13" pageNumber="40" type="generic">10 specimens</specimenCount>
examined); ventral central field of manubrium with fewer than 20 setae (
<figureCitation id="40DD80FFFFD4FF84962CF0FBFE9B1DFA" box="[261,348,226,252]" captionStart="FIGURES 57 77" captionStartId="14.[151,270,1589,1613]" captionTargetBox="[159,1433,207,1545]" captionTargetId="figure@14.[151,1436,197,1568]" captionTargetPageId="14" captionText="FIGURES 57 77. (57 63) Desoria cryophila sp. n., (64 70) Desoria rosea sp. nov., (71 76) Desoria olympica sp. n., (77) Desoria capra sp. n., (57) sensillary chaetotaxy, positions of macrochaetae indicated, (58) eye-field and PAO, (59) left mucro, (60) sensilla on ventral side of right ant. 1, (61) ditto, right ant. 3, dorsal, (62) ditto, right ant. 2, lateral, (63) setae along midline on inner side of tib. 1, (64) eye-field and PAO, (65) sensilla on right ant. 2, lateral. es: erect sensilla, (66) ditto, ant. 3, (67) setae in right basomedian field of labium, the additional 5 th. seta encircled, (68) right mucro, lateral, (69) ditto, dorsal, with the secondary inner tooth formed by the truncated inner lamella marked, (70) dorsal setae in basal half of dens, (71) sensillary chaetotaxy, positions of macrochaetae indicated, (72) setae along midline on inner side of tib. 1, (73) left mucro and apical part of dens, (74) ventral side of manubrium, (75) eye-field and PAO, (76) labrum and frontoclypeal field, (77) right maxilla with details of capitular teeth (mt: membranous tooth)." httpUri="https://zenodo.org/record/196081/files/figure.png" pageId="13" pageNumber="40">Fig. 74</figureCitation>
); dens with 7 dorsal setae; mucro 5-toothed (
<figureCitation id="40DD80FFFFD4FF84944EF0FBFC781DFA" box="[871,959,226,252]" captionStart="FIGURES 57 77" captionStartId="14.[151,270,1589,1613]" captionTargetBox="[159,1433,207,1545]" captionTargetId="figure@14.[151,1436,197,1568]" captionTargetPageId="14" captionText="FIGURES 57 77. (57 63) Desoria cryophila sp. n., (64 70) Desoria rosea sp. nov., (71 76) Desoria olympica sp. n., (77) Desoria capra sp. n., (57) sensillary chaetotaxy, positions of macrochaetae indicated, (58) eye-field and PAO, (59) left mucro, (60) sensilla on ventral side of right ant. 1, (61) ditto, right ant. 3, dorsal, (62) ditto, right ant. 2, lateral, (63) setae along midline on inner side of tib. 1, (64) eye-field and PAO, (65) sensilla on right ant. 2, lateral. es: erect sensilla, (66) ditto, ant. 3, (67) setae in right basomedian field of labium, the additional 5 th. seta encircled, (68) right mucro, lateral, (69) ditto, dorsal, with the secondary inner tooth formed by the truncated inner lamella marked, (70) dorsal setae in basal half of dens, (71) sensillary chaetotaxy, positions of macrochaetae indicated, (72) setae along midline on inner side of tib. 1, (73) left mucro and apical part of dens, (74) ventral side of manubrium, (75) eye-field and PAO, (76) labrum and frontoclypeal field, (77) right maxilla with details of capitular teeth (mt: membranous tooth)." httpUri="https://zenodo.org/record/196081/files/figure.png" pageId="13" pageNumber="40">Fig. 73</figureCitation>
). Inner side of tib. 1-2 with three pairs of setae along median line (
<figureCitation id="40DD80FFFFD4FF849692F111FDD51C24" box="[443,530,264,290]" captionStart="FIGURES 57 77" captionStartId="14.[151,270,1589,1613]" captionTargetBox="[159,1433,207,1545]" captionTargetId="figure@14.[151,1436,197,1568]" captionTargetPageId="14" captionText="FIGURES 57 77. (57 63) Desoria cryophila sp. n., (64 70) Desoria rosea sp. nov., (71 76) Desoria olympica sp. n., (77) Desoria capra sp. n., (57) sensillary chaetotaxy, positions of macrochaetae indicated, (58) eye-field and PAO, (59) left mucro, (60) sensilla on ventral side of right ant. 1, (61) ditto, right ant. 3, dorsal, (62) ditto, right ant. 2, lateral, (63) setae along midline on inner side of tib. 1, (64) eye-field and PAO, (65) sensilla on right ant. 2, lateral. es: erect sensilla, (66) ditto, ant. 3, (67) setae in right basomedian field of labium, the additional 5 th. seta encircled, (68) right mucro, lateral, (69) ditto, dorsal, with the secondary inner tooth formed by the truncated inner lamella marked, (70) dorsal setae in basal half of dens, (71) sensillary chaetotaxy, positions of macrochaetae indicated, (72) setae along midline on inner side of tib. 1, (73) left mucro and apical part of dens, (74) ventral side of manubrium, (75) eye-field and PAO, (76) labrum and frontoclypeal field, (77) right maxilla with details of capitular teeth (mt: membranous tooth)." httpUri="https://zenodo.org/record/196081/files/figure.png" pageId="13" pageNumber="40">Fig. 72</figureCitation>
). Males present, reproductive specimens not seen.
</paragraph>
</subSubSection>
<subSubSection id="90FCCFF1FFD4FF8497EDF134FC3A1C41" box="[196,1021,301,327]" pageId="13" pageNumber="40" type="etymology">
<paragraph id="D8599C7AFFD4FF8497EDF134FC3A1C41" blockId="13.[151,1437,152,551]" box="[196,1021,301,327]" pageId="13" pageNumber="40">
<emphasis id="EA924068FFD4FF8497EDF134FE961C41" bold="true" box="[196,337,301,327]" pageId="13" pageNumber="40">Etymology.</emphasis>
Named after the
<typeStatus id="075D22D8FFD4FF849535F134FD881C41" box="[540,591,301,327]" pageId="13" pageNumber="40">type</typeStatus>
locality, the Olympic National Park.
</paragraph>
</subSubSection>
<subSubSection id="90FCCFF1FFD4FF8497EDF14BFBF21CDA" pageId="13" pageNumber="40" type="discussion">
<paragraph id="D8599C7AFFD4FF8497EDF14BFBF21CDA" blockId="13.[151,1437,152,551]" pageId="13" pageNumber="40">
<emphasis id="EA924068FFD4FF8497EDF14BFE8F1C6A" bold="true" box="[196,328,338,364]" pageId="13" pageNumber="40">Discussion</emphasis>
. The character which immediately identifies
<taxonomicName id="1FE6E7F9FFD4FF849475F14BFC0F1C6D" box="[860,968,338,363]" class="Entognatha" family="Isotomidae" genus="Desoria" kingdom="Animalia" order="Collembola" pageId="13" pageNumber="40" phylum="Arthropoda" rank="species" species="olympica">
<emphasis id="EA924068FFD4FF849475F14BFC0F1C6D" box="[860,968,338,363]" italics="true" pageId="13" pageNumber="40">olympica</emphasis>
</taxonomicName>
is the reduced number of tergal sensilla (22/11135), which is probably the lowest number observed in any
<taxonomicName id="1FE6E7F9FFD4FF849486F161FBC91C97" box="[943,1038,376,401]" class="Entognatha" family="Isotomidae" genus="Desoria" kingdom="Animalia" order="Collembola" pageId="13" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="EA924068FFD4FF849486F161FBC91C97" box="[943,1038,376,401]" italics="true" pageId="13" pageNumber="40">Desoria</emphasis>
</taxonomicName>
. Unlike the two previous species the posterior sensilla on th. 2abd. 4 are set in the p-row. It also differs by the reduced number of antennal sensilla and the low number of setae on ventral tube and retinaculum (
<tableCitation id="9564A9C1FFD4FF8494E5F1DBFBE31CDA" box="[972,1060,450,476]" captionStart="TABLE 1" captionStartId="1.[151,242,151,175]" captionTargetBox="[151,1437,234,769]" captionTargetPageId="1" captionText="TABLE 1. Morphological characteristics of adult specimens of Myopia (M.), Desoria (D.) and Gnathisotoma (G.) species from the Northwestern Rocky Mountains." httpUri="http://table.plazi.org/id/8C99CCF2FFD8FF8897BEF08EFD6D1DC9" pageId="13" pageNumber="40" tableUuid="8C99CCF2FFD8FF8897BEF08EFD6D1DC9">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="90FCCFF1FFD4FF8497EDF1FEFB431F21" pageId="13" pageNumber="40" type="distribution">
<paragraph id="D8599C7AFFD4FF8497EDF1FEFB431F21" blockId="13.[151,1437,152,551]" pageId="13" pageNumber="40">
<emphasis id="EA924068FFD4FF8497EDF1FEFDC51F07" bold="true" box="[196,514,487,513]" pageId="13" pageNumber="40">Distribution and ecology.</emphasis>
This distinct species is only known from the
<typeStatus id="075D22D8FFD4FF84930CF1F1FB9F1F04" box="[1061,1112,488,514]" pageId="13" pageNumber="40">type</typeStatus>
series which appeared in a wet snow edge community in the coastal mountains above Port Angeles, Washington.
</paragraph>
</subSubSection>
</treatment>
</document>