treatments-xml/data/17/72/12/1772122E7811FFD1FF013A44AB0DEFAB.xml

<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik &amp; Sierwald, Petra" docDate="2009" docId="1772122E7811FFD1FF013A44AB0DEFAB" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Colossobolus Wesener &amp; Enghoff &amp; Sierwald 2009" docType="key" docVersion="10" lastPageNumber="24" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="23" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. &amp; Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. &amp; B 54 D 3623309</mods:affiliation>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
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<mods:namePart>Sierwald, Petra</mods:namePart>
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<mods:date>2009</mods:date>
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<mods:number>2009-09-04</mods:number>
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Key to the
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[270,420,1328,1355]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[270,420,1328,1355]" italics="true" pageId="22" pageNumber="23">Colossobolus</emphasis>
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species:
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1 Telopodite of posterior gonopod basally with a finger-shaped coxite branch (
<figureCitation box="[253,345,1434,1460]" captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="22" pageNumber="23">Figs 9C</figureCitation>
,
<figureCitation box="[357,407,1434,1460]" captionStart="Figure 14" captionStartId="26.[140,212,1050,1072]" captionTargetBox="[140,1108,165,1030]" captionTargetId="figure@26.[140,1108,165,1031]" captionTargetPageId="26" captionText="Figure 14. Colossobolus oblongopedus sp. n., male holotype, right posterior gonopod. A anterior view B posterior view. Cx = coxite; Cx-br = coxite branch; ma = membranous area; S-Gr = spermatic groove; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch." figureDoi="http://doi.org/10.5281/zenodo.3820802" httpUri="https://zenodo.org/record/3820802/files/figure.png" pageId="22" pageNumber="23">14A</figureCitation>
,
<figureCitation box="[419,465,1434,1461]" captionStart="Figure 15" captionStartId="28.[140,212,1494,1516]" captionTargetBox="[140,1108,165,1409]" captionTargetId="figure@28.[140,1108,528,1439]" captionTargetPageId="28" captionText="Figure 15. Colossobolus giganteus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. Apo = apodeme; Co = collum; Cx = coxite; Cx-br = coxite branch; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; St = sternite; ST = stipites; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820804" httpUri="https://zenodo.org/record/3820804/files/figure.png" pageId="22" pageNumber="23">15F</figureCitation>
)...................................................................................
<emphasis bold="true" box="[1092,1107,1433,1459]" pageId="22" pageNumber="23">2</emphasis>
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– Telopodite of posterior gonopod lacking coxite branch (
<figureCitation box="[876,975,1469,1495]" captionStart="Figure 16" captionStartId="30.[140,212,1329,1351]" captionTargetBox="[140,1108,165,1309]" captionTargetId="figure@30.[140,1108,165,989]" captionTargetPageId="30" captionText="Figure 16. Colossobolus minor sp. n., male holotype. A legs 1–7, ventral view B telson C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. av = anal valves; Cx = coxite; Gr = groove on anterior gonopods; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820806" httpUri="https://zenodo.org/record/3820806/files/figure.png" pageId="22" pageNumber="23">Figs 16F</figureCitation>
,
<figureCitation box="[987,1037,1469,1495]" captionStart="Figure 19" captionStartId="33.[140,212,694,716]" captionTargetBox="[140,1108,165,674]" captionTargetId="figure@33.[140,1108,165,675]" captionTargetPageId="33" captionText="Figure 19. Colossobolus litoralis sp. n., male holotype, posterior gonopods. A anterior view B posterior view. Cx = coxite; ma = membranous area; S = sternite; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch." figureDoi="http://doi.org/10.5281/zenodo.3820812" httpUri="https://zenodo.org/record/3820812/files/figure.png" pageId="22" pageNumber="23">19A</figureCitation>
, 20G,
<figureCitation box="[246,293,1504,1530]" captionStart="Figure 21" captionStartId="35.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1506]" captionTargetId="figure@35.[320,926,653,1506]" captionTargetPageId="35" captionText="Figure 21. Colossobolus pseudoaculeatus sp. n., male holotype. A head, lateral B telson C anterior gonopods, posterior view D anterior gonopods, anterior view E right posterior gonopod, posterior view F right posterior gonopod, anterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820814" httpUri="https://zenodo.org/record/3820814/files/figure.png" pageId="22" pageNumber="23">21E</figureCitation>
) ..........................................................................................................
<emphasis bold="true" box="[1092,1107,1504,1530]" pageId="22" pageNumber="23">4</emphasis>
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2 Lateral branch of posterior gonopod telopodite curved towards coxite branch, almost forming a circle with latter (
<figureCitation box="[650,771,1575,1601]" captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="22" pageNumber="23">Figs 9H, I</figureCitation>
). Coxite of anterior gonopod in anterior view with a short, wide, well-rounded process (
<figureCitation box="[903,993,1610,1636]" captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="22" pageNumber="23">Fig. 9D</figureCitation>
)............. ......................................................................................
<emphasis bold="true" box="[880,1108,1644,1671]" pageId="22" pageNumber="23">
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[880,1035,1644,1671]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="semicyclus" status="sp. n.">
<emphasis bold="true" box="[880,1035,1644,1671]" italics="true" pageId="22" pageNumber="23">C. semicyclus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1043,1108,1645,1671]" pageId="22" pageNumber="23" rank="species">sp. n.</taxonomicNameLabel>
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– Lateral branch of posterior gonopod telopodite straight (
<figureCitation box="[849,947,167,193]" captionStart="Figure 14" captionStartId="26.[140,212,1050,1072]" captionTargetBox="[140,1108,165,1030]" captionTargetId="figure@26.[140,1108,165,1031]" captionTargetPageId="26" captionText="Figure 14. Colossobolus oblongopedus sp. n., male holotype, right posterior gonopod. A anterior view B posterior view. Cx = coxite; Cx-br = coxite branch; ma = membranous area; S-Gr = spermatic groove; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch." figureDoi="http://doi.org/10.5281/zenodo.3820802" httpUri="https://zenodo.org/record/3820802/files/figure.png" pageId="23" pageNumber="24">Fig. 14A</figureCitation>
). Coxite of anterior gonopod in anterior view with a short pointed process (
<figureCitation box="[905,1001,202,228]" captionStart="Figure 13" captionStartId="21.[140,211,1116,1138]" captionTargetBox="[140,1108,165,1092]" captionTargetId="figure@21.[140,1108,165,1092]" captionTargetPageId="21" captionText="Figure 13. Colossobolus oblongopedus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view. Co = collum; Cx = coxite; Gr = groove on anterior gonopod; IL = incisura lateralis; LL = lamella lingualis; Md = mandible; ms = marginal setae; sf = setiferous foveolae; St = sternite; ST = stipites;T = telopodite;Tp = telopodite process.Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820800" httpUri="https://zenodo.org/record/3820800/files/figure.png" pageId="23" pageNumber="24">Fig. 13C</figureCitation>
)...........
<emphasis bold="true" box="[1092,1107,201,227]" pageId="23" pageNumber="24">3</emphasis>
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<paragraph blockId="23.[140,1108,167,1531]" pageId="23" pageNumber="24">
<keyLead pageId="23" pageNumber="24">
3 Tarsi whitish (
<figureCitation box="[404,505,237,263]" captionStart="Figure 13" captionStartId="21.[140,211,1116,1138]" captionTargetBox="[140,1108,165,1092]" captionTargetId="figure@21.[140,1108,165,1092]" captionTargetPageId="21" captionText="Figure 13. Colossobolus oblongopedus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view. Co = collum; Cx = coxite; Gr = groove on anterior gonopod; IL = incisura lateralis; LL = lamella lingualis; Md = mandible; ms = marginal setae; sf = setiferous foveolae; St = sternite; ST = stipites;T = telopodite;Tp = telopodite process.Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820800" httpUri="https://zenodo.org/record/3820800/files/figure.png" pageId="23" pageNumber="24">Fig. 13B</figureCitation>
). Collum completely red (
<figureCitation box="[798,900,237,263]" captionStart="Figure 13" captionStartId="21.[140,211,1116,1138]" captionTargetBox="[140,1108,165,1092]" captionTargetId="figure@21.[140,1108,165,1092]" captionTargetPageId="21" captionText="Figure 13. Colossobolus oblongopedus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view. Co = collum; Cx = coxite; Gr = groove on anterior gonopod; IL = incisura lateralis; LL = lamella lingualis; Md = mandible; ms = marginal setae; sf = setiferous foveolae; St = sternite; ST = stipites;T = telopodite;Tp = telopodite process.Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820800" httpUri="https://zenodo.org/record/3820800/files/figure.png" pageId="23" pageNumber="24">Fig. 13A</figureCitation>
). Coxite branch of posterior gonopod short, half as long as lateral branch. Lateral branch irregularly curved, becoming apically thicker (
<figureCitation box="[693,828,307,334]" captionStart="Figure 14" captionStartId="26.[140,212,1050,1072]" captionTargetBox="[140,1108,165,1030]" captionTargetId="figure@26.[140,1108,165,1031]" captionTargetPageId="26" captionText="Figure 14. Colossobolus oblongopedus sp. n., male holotype, right posterior gonopod. A anterior view B posterior view. Cx = coxite; Cx-br = coxite branch; ma = membranous area; S-Gr = spermatic groove; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch." figureDoi="http://doi.org/10.5281/zenodo.3820802" httpUri="https://zenodo.org/record/3820802/files/figure.png" pageId="23" pageNumber="24">Figs 14A, B</figureCitation>
). Retrorse process absent from mesal margin of telopodite process of anterior gonopod (
<figureCitation box="[950,1055,343,369]" captionStart="Figure 13" captionStartId="21.[140,211,1116,1138]" captionTargetBox="[140,1108,165,1092]" captionTargetId="figure@21.[140,1108,165,1092]" captionTargetPageId="21" captionText="Figure 13. Colossobolus oblongopedus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view. Co = collum; Cx = coxite; Gr = groove on anterior gonopod; IL = incisura lateralis; LL = lamella lingualis; Md = mandible; ms = marginal setae; sf = setiferous foveolae; St = sternite; ST = stipites;T = telopodite;Tp = telopodite process.Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820800" httpUri="https://zenodo.org/record/3820800/files/figure.png" pageId="23" pageNumber="24">Fig. 13D</figureCitation>
)..... ................................................................................
<emphasis bold="true" box="[842,1108,377,404]" pageId="23" pageNumber="24">
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[842,1035,377,404]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="oblongopedus" status="sp. n.">
<emphasis bold="true" box="[842,1035,377,404]" italics="true" pageId="23" pageNumber="24">C. oblongopedus</emphasis>
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<taxonomicNameLabel box="[1043,1108,378,404]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
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<paragraph blockId="23.[140,1108,167,1531]" pageId="23" pageNumber="24">
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– Legs completely red (
<figureCitation box="[495,600,413,440]" captionStart="Figure 15" captionStartId="28.[140,212,1494,1516]" captionTargetBox="[140,1108,165,1409]" captionTargetId="figure@28.[140,1108,528,1439]" captionTargetPageId="28" captionText="Figure 15. Colossobolus giganteus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. Apo = apodeme; Co = collum; Cx = coxite; Cx-br = coxite branch; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; St = sternite; ST = stipites; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820804" httpUri="https://zenodo.org/record/3820804/files/figure.png" pageId="23" pageNumber="24">Fig. 15B</figureCitation>
). Collum at margins red, median part dark (
<figureCitation box="[254,364,448,475]" captionStart="Figure 15" captionStartId="28.[140,212,1494,1516]" captionTargetBox="[140,1108,165,1409]" captionTargetId="figure@28.[140,1108,528,1439]" captionTargetPageId="28" captionText="Figure 15. Colossobolus giganteus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. Apo = apodeme; Co = collum; Cx = coxite; Cx-br = coxite branch; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; St = sternite; ST = stipites; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820804" httpUri="https://zenodo.org/record/3820804/files/figure.png" pageId="23" pageNumber="24">Fig. 15A</figureCitation>
). Coxite branch of posterior gonopod long, as long as lateral branch. Lateral branch regularly curved (
<figureCitation box="[725,829,483,510]" captionStart="Figure 15" captionStartId="28.[140,212,1494,1516]" captionTargetBox="[140,1108,165,1409]" captionTargetId="figure@28.[140,1108,528,1439]" captionTargetPageId="28" captionText="Figure 15. Colossobolus giganteus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. Apo = apodeme; Co = collum; Cx = coxite; Cx-br = coxite branch; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; St = sternite; ST = stipites; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820804" httpUri="https://zenodo.org/record/3820804/files/figure.png" pageId="23" pageNumber="24">Fig. 15F</figureCitation>
). Mesal margin of telopodite process of anterior gonopod protruding into a very short, but pointed retrorse process (
<figureCitation box="[434,539,554,581]" captionStart="Figure 15" captionStartId="28.[140,212,1494,1516]" captionTargetBox="[140,1108,165,1409]" captionTargetId="figure@28.[140,1108,528,1439]" captionTargetPageId="28" captionText="Figure 15. Colossobolus giganteus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. Apo = apodeme; Co = collum; Cx = coxite; Cx-br = coxite branch; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; St = sternite; ST = stipites; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820804" httpUri="https://zenodo.org/record/3820804/files/figure.png" pageId="23" pageNumber="24">Fig. 15D</figureCitation>
) ..............................................
<emphasis bold="true" box="[891,1108,554,580]" pageId="23" pageNumber="24">
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[891,1036,554,580]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="giganteus" status="sp. n.">
<emphasis bold="true" box="[891,1036,554,580]" italics="true" pageId="23" pageNumber="24">C. giganteus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1043,1108,554,580]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<paragraph blockId="23.[140,1108,167,1531]" pageId="23" pageNumber="24">
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4 Telopodites of posterior gonopods oriented face-to-face to one another. Main branch of telopodite overlapping juxtaposed counterpart (
<figureCitation box="[961,1094,624,651]" captionStart="Figure 16" captionStartId="30.[140,212,1329,1351]" captionTargetBox="[140,1108,165,1309]" captionTargetId="figure@30.[140,1108,165,989]" captionTargetPageId="30" captionText="Figure 16. Colossobolus minor sp. n., male holotype. A legs 1–7, ventral view B telson C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. av = anal valves; Cx = coxite; Gr = groove on anterior gonopods; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820806" httpUri="https://zenodo.org/record/3820806/files/figure.png" pageId="23" pageNumber="24">Figs 16E, F</figureCitation>
). Small,
<quantity box="[323,465,659,686]" metricMagnitude="-2" metricUnit="m" metricValue="9.25" metricValueMax="10.0" metricValueMin="8.5" pageId="23" pageNumber="24" unit="mm" value="92.5" valueMax="100.0" valueMin="85.0">85–100 mm</quantity>
long. Colour of body rings dark brownish (
<figureCitation box="[960,1059,659,686]" captionStart="Figure 16" captionStartId="30.[140,212,1329,1351]" captionTargetBox="[140,1108,165,1309]" captionTargetId="figure@30.[140,1108,165,989]" captionTargetPageId="30" captionText="Figure 16. Colossobolus minor sp. n., male holotype. A legs 1–7, ventral view B telson C anterior gonopods, anterior view D anterior gonopods, posterior view E left posterior gonopod, anterior view F left posterior gonopod, posterior view. av = anal valves; Cx = coxite; Gr = groove on anterior gonopods; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820806" httpUri="https://zenodo.org/record/3820806/files/figure.png" pageId="23" pageNumber="24">Fig. 16B</figureCitation>
).... ............................................................................................
<emphasis bold="true" box="[928,1108,695,721]" pageId="23" pageNumber="24">
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[928,1035,695,721]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="minor" status="sp. n.">
<emphasis bold="true" box="[928,1035,695,721]" italics="true" pageId="23" pageNumber="24">C. minor</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1043,1108,695,721]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph blockId="23.[140,1108,167,1531]" pageId="23" pageNumber="24">
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– Telopodites of posterior gonopods oriented parallel to one another. Telopodite main branches not overlapping (
<figureCitation box="[692,830,765,792]" captionStart="Figure 19" captionStartId="33.[140,212,694,716]" captionTargetBox="[140,1108,165,674]" captionTargetId="figure@33.[140,1108,165,675]" captionTargetPageId="33" captionText="Figure 19. Colossobolus litoralis sp. n., male holotype, posterior gonopods. A anterior view B posterior view. Cx = coxite; ma = membranous area; S = sternite; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch." figureDoi="http://doi.org/10.5281/zenodo.3820812" httpUri="https://zenodo.org/record/3820812/files/figure.png" pageId="23" pageNumber="24">Figs 19A, B</figureCitation>
). Body large,&gt;
<quantity box="[1003,1104,765,792]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="23" pageNumber="24" unit="mm" value="120.0">120 mm</quantity>
. Colour of body rings always partly red.......................................................
<emphasis bold="true" box="[1092,1107,800,826]" pageId="23" pageNumber="24">5</emphasis>
</keyLead>
</paragraph>
</keyStep>
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<paragraph blockId="23.[140,1108,167,1531]" pageId="23" pageNumber="24">
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5 Mesal margin of telopodite process of anterior gonopod widely projecting, almost protruding up to lateral margin (
<figureCitation box="[674,772,871,897]" captionStart="Figure 18" captionStartId="32.[140,212,1558,1580]" captionTargetBox="[179,1067,165,1538]" captionTargetId="figure@32.[179,1068,165,1541]" captionTargetPageId="32" captionText="Figure 18. Colossobolus litoralis sp. n., male holotype.A head, lateral B telson C legs 1–7, ventral view D coxae of midbody leg, posterior view E anterior gonopods, anterior view F anterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; LL = lamella lingualis; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale;T = telopodite; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820810" httpUri="https://zenodo.org/record/3820810/files/figure.png" pageId="23" pageNumber="24">Fig. 18F</figureCitation>
). Coxite process extraordinary long, but wide and well-rounded, longer than sternite, but far shorter than process of telopodite (
<figureCitation box="[432,528,941,967]" captionStart="Figure 18" captionStartId="32.[140,212,1558,1580]" captionTargetBox="[179,1067,165,1538]" captionTargetId="figure@32.[179,1068,165,1541]" captionTargetPageId="32" captionText="Figure 18. Colossobolus litoralis sp. n., male holotype.A head, lateral B telson C legs 1–7, ventral view D coxae of midbody leg, posterior view E anterior gonopods, anterior view F anterior gonopods, posterior view. av = anal valves; Co = collum; Cx = coxite; LL = lamella lingualis; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale;T = telopodite; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820810" httpUri="https://zenodo.org/record/3820810/files/figure.png" pageId="23" pageNumber="24">Fig. 18E</figureCitation>
). Telopodite of posterior gonopod not U-shaped. Lateral branch weakly developed, very slender (
<figureCitation box="[726,825,976,1002]" captionStart="Figure 19" captionStartId="33.[140,212,694,716]" captionTargetBox="[140,1108,165,674]" captionTargetId="figure@33.[140,1108,165,675]" captionTargetPageId="33" captionText="Figure 19. Colossobolus litoralis sp. n., male holotype, posterior gonopods. A anterior view B posterior view. Cx = coxite; ma = membranous area; S = sternite; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch." figureDoi="http://doi.org/10.5281/zenodo.3820812" httpUri="https://zenodo.org/record/3820812/files/figure.png" pageId="23" pageNumber="24">Fig. 19A</figureCitation>
). Telopodite main branch separated into two parts, lateral part with large and long membranes, mesal part short and sclerotized. Telopodite lateral branch and both parts of main branch orientated parallel to one another (
<figureCitation box="[621,751,1082,1108]" captionStart="Figure 19" captionStartId="33.[140,212,694,716]" captionTargetBox="[140,1108,165,674]" captionTargetId="figure@33.[140,1108,165,675]" captionTargetPageId="33" captionText="Figure 19. Colossobolus litoralis sp. n., male holotype, posterior gonopods. A anterior view B posterior view. Cx = coxite; ma = membranous area; S = sternite; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch." figureDoi="http://doi.org/10.5281/zenodo.3820812" httpUri="https://zenodo.org/record/3820812/files/figure.png" pageId="23" pageNumber="24">Figs 19A, B</figureCitation>
) ......................
<emphasis bold="true" box="[917,1107,1081,1108]" pageId="23" pageNumber="24">
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[917,1039,1081,1108]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="litoralis" status="sp. n.">
<emphasis bold="true" box="[917,1039,1081,1108]" italics="true" pageId="23" pageNumber="24">C. litoralis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1045,1107,1082,1108]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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– Mesal margin of telopodite process of anterior gonopod sharp, but not projecting (
<figureCitation box="[296,394,1152,1178]" captionStart="Figure 20" captionStartId="33.[140,212,1494,1516]" captionTargetBox="[140,1108,834,1475]" captionTargetPageId="33" captionText="Figure 20. Colossobolus aculeatus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C telson D anterior gonopods, anterior view E right posterior gonopod, anterior view F anterior gonopods, posterior view G right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="23" pageNumber="24">Fig. 20F</figureCitation>
). Coxite process present, but short and well-rounded (
<figureCitation box="[995,1096,1152,1178]" captionStart="Figure 20" captionStartId="33.[140,212,1494,1516]" captionTargetBox="[140,1108,834,1475]" captionTargetPageId="33" captionText="Figure 20. Colossobolus aculeatus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C telson D anterior gonopods, anterior view E right posterior gonopod, anterior view F anterior gonopods, posterior view G right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="23" pageNumber="24">Fig. 20A</figureCitation>
). Telopodite of posterior gonopod U-shaped (
<figureCitation box="[720,817,1187,1214]" captionStart="Figure 20" captionStartId="33.[140,212,1494,1516]" captionTargetBox="[140,1108,834,1475]" captionTargetPageId="33" captionText="Figure 20. Colossobolus aculeatus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C telson D anterior gonopods, anterior view E right posterior gonopod, anterior view F anterior gonopods, posterior view G right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="23" pageNumber="24">Fig. 20E</figureCitation>
). Lateral and main branches apically of similar width. Telopodite main branch not divided into two parts, branch basally sclerotized, apically membranous (
<figureCitation box="[779,912,1258,1284]" captionStart="Figure 20" captionStartId="33.[140,212,1494,1516]" captionTargetBox="[140,1108,834,1475]" captionTargetPageId="33" captionText="Figure 20. Colossobolus aculeatus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C telson D anterior gonopods, anterior view E right posterior gonopod, anterior view F anterior gonopods, posterior view G right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="23" pageNumber="24">Figs 20E, G</figureCitation>
)........................
<emphasis bold="true" box="[1092,1107,1257,1283]" pageId="23" pageNumber="24">6</emphasis>
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6 Preanal process apically sharp-edged, slightly shorter than anal valves (
<figureCitation captionStart="Figure 20" captionStartId="33.[140,212,1494,1516]" captionTargetBox="[140,1108,834,1475]" captionTargetPageId="33" captionText="Figure 20. Colossobolus aculeatus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C telson D anterior gonopods, anterior view E right posterior gonopod, anterior view F anterior gonopods, posterior view G right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="23" pageNumber="24">Fig. 20C</figureCitation>
). Main branch of posterior gonopod telopodite of equal width all along (
<figureCitation box="[254,355,1363,1390]" captionStart="Figure 20" captionStartId="33.[140,212,1494,1516]" captionTargetBox="[140,1108,834,1475]" captionTargetPageId="33" captionText="Figure 20. Colossobolus aculeatus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C telson D anterior gonopods, anterior view E right posterior gonopod, anterior view F anterior gonopods, posterior view G right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="23" pageNumber="24">Fig. 20E</figureCitation>
), longer than lateral branch (
<figureCitation box="[683,787,1363,1390]" captionStart="Figure 20" captionStartId="33.[140,212,1494,1516]" captionTargetBox="[140,1108,834,1475]" captionTargetPageId="33" captionText="Figure 20. Colossobolus aculeatus sp. n., male holotype. A head, lateral B legs 1–7, ventral view C telson D anterior gonopods, anterior view E right posterior gonopod, anterior view F anterior gonopods, posterior view G right posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; IL = incisura lateralis; LL = lamella lingualis; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3767296" httpUri="https://zenodo.org/record/3767296/files/figure.png" pageId="23" pageNumber="24">Fig. 20G</figureCitation>
) ............
<emphasis bold="true" box="[890,1108,1363,1390]" pageId="23" pageNumber="24">
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[890,1035,1363,1390]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="aculeatus" status="sp. n.">
<emphasis bold="true" box="[890,1035,1363,1390]" italics="true" pageId="23" pageNumber="24">C. aculeatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1043,1108,1363,1389]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
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– Preanal process apically rounded, shorter than anal valves (
<figureCitation box="[920,1024,1399,1425]" captionStart="Figure 21" captionStartId="35.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1506]" captionTargetId="figure@35.[320,926,653,1506]" captionTargetPageId="35" captionText="Figure 21. Colossobolus pseudoaculeatus sp. n., male holotype. A head, lateral B telson C anterior gonopods, posterior view D anterior gonopods, anterior view E right posterior gonopod, posterior view F right posterior gonopod, anterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820814" httpUri="https://zenodo.org/record/3820814/files/figure.png" pageId="23" pageNumber="24">Fig. 21B</figureCitation>
). Main branch of posterior gonopod telopodite basally very slender, but with a wide, swollen tip (
<figureCitation box="[384,485,1469,1495]" captionStart="Figure 21" captionStartId="35.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1506]" captionTargetId="figure@35.[320,926,653,1506]" captionTargetPageId="35" captionText="Figure 21. Colossobolus pseudoaculeatus sp. n., male holotype. A head, lateral B telson C anterior gonopods, posterior view D anterior gonopods, anterior view E right posterior gonopod, posterior view F right posterior gonopod, anterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820814" httpUri="https://zenodo.org/record/3820814/files/figure.png" pageId="23" pageNumber="24">Fig. 21E</figureCitation>
), shorter than lateral branch (
<figureCitation box="[821,919,1469,1495]" captionStart="Figure 21" captionStartId="35.[140,212,1526,1548]" captionTargetBox="[140,1108,165,1506]" captionTargetId="figure@35.[320,926,653,1506]" captionTargetPageId="35" captionText="Figure 21. Colossobolus pseudoaculeatus sp. n., male holotype. A head, lateral B telson C anterior gonopods, posterior view D anterior gonopods, anterior view E right posterior gonopod, posterior view F right posterior gonopod, anterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820814" httpUri="https://zenodo.org/record/3820814/files/figure.png" pageId="23" pageNumber="24">Fig. 21F</figureCitation>
) ....................... ............................................................................
<emphasis bold="true" box="[809,1108,1504,1531]" pageId="23" pageNumber="24">
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[809,1036,1504,1531]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="pseudoaculeatus" status="sp. n.">
<emphasis bold="true" box="[809,1036,1504,1531]" italics="true" pageId="23" pageNumber="24">C. pseudoaculeatus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1043,1108,1504,1530]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>