treatments-xml/data/17/72/12/1772122E780CFFCBFF013B62AD95E9F5.xml

<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik &amp; Sierwald, Petra" docDate="2009" docId="1772122E780CFFCBFF013B62AD95E9F5" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Sanguinobolus Wesener &amp; Enghoff &amp; Sierwald 2009, gen. n." docType="treatment" docUuid="C699F031-ACE7-4624-8B68-4C153FAE59E1" docUuidSource="ZooBank" docVersion="10" lastPageNumber="14" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="12" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. &amp; Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. &amp; B 54 D 3623309</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sierwald, Petra</mods:namePart>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-09-04</mods:number>
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<mods:number>19</mods:number>
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<mods:number>19</mods:number>
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<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791380" ID-GBIF-Taxon="163747659" ID-Zenodo-Dep="3791380" LSID="urn:lsid:zoobank.org:act:C699F031-ACE7-4624-8B68-4C153FAE59E1" httpUri="http://treatment.plazi.org/id/1772122E780CFFCBFF013B62AD95E9F5" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
<subSubSection box="[140,516,1046,1073]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph blockId="11.[140,964,1046,1179]" box="[140,516,1046,1073]" pageId="11" pageNumber="12">
<heading box="[140,516,1046,1073]" fontSize="11" level="2" pageId="11" pageNumber="12" reason="3">
<emphasis bold="true" box="[140,516,1046,1073]" pageId="11" pageNumber="12">
<taxonomicName authority="Wesener" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,423,1046,1073]" class="Diplopoda" family="Pachybolidae" genus="Sanguinobolus" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis bold="true" box="[140,315,1046,1073]" inLineHeading="true" italics="true" pageId="11" pageNumber="12" reason="1">Sanguinobolus</emphasis>
Wesener
</taxonomicName>
, 
<taxonomicNameLabel box="[435,516,1047,1073]" pageId="11" pageNumber="12" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act: 
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C699F031- 
<collectingRegion box="[554,616,1082,1109]" country="Indonesia" name="Aceh" pageId="11" pageNumber="12">ACE</collectingRegion>
7-4624-8B68-4C153FAE59E1
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</paragraph>
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<subSubSection box="[140,780,1152,1179]" pageId="11" pageNumber="12" type="type_taxon">
<paragraph blockId="11.[140,964,1046,1179]" box="[140,780,1152,1179]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[140,287,1152,1178]" pageId="11" pageNumber="12">
<emphasis box="[140,198,1152,1178]" inLineHeading="true" pageId="11" pageNumber="12" reason="1">
<typeStatus box="[140,198,1152,1178]" pageId="11" pageNumber="12">Type</typeStatus>
</emphasis>
species
</emphasis>
: 
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[302,574,1152,1178]" class="Diplopoda" family="Pachybolidae" genus="Sanguinobolus" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="maculosus" status="sp. n.">
<emphasis box="[302,574,1152,1178]" italics="true" pageId="11" pageNumber="12">Sanguinobolus maculosus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[581,644,1153,1179]" pageId="11" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
, monotypic
</paragraph>
</subSubSection>
<subSubSection lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="diagnosis">
<paragraph blockId="11.[140,1108,1222,1672]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[140,259,1222,1248]" inLineHeading="true" pageId="11" pageNumber="12" reason="1">Diagnosis</emphasis>
: distinguished by male tarsal pads less prominently developed than in other large-bodied Malagasy genera of 
<taxonomicName authorityName="Cook" authorityYear="1895" box="[513,643,1258,1284]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
(
<figureCitation box="[658,788,1258,1284]" captionStart="Figure 7" captionStartId="14.[140,212,1263,1285]" captionTargetBox="[141,1107,165,1244]" captionTargetId="figure@14.[140,1107,165,1247]" captionTargetPageId="14" captionText="Figure 7. Sanguinobolus maculosus sp. n., male paratype, SEM. A antenna, apical view, arrow indicates area shown in 7B B antennomere 6 with sensilla basiconica C male left midbody leg D detail of tarsus E left mandible, mesal view. 3iT = 3-combed internal teeth; eT = external tooth; LT = lateral tooth; mp = molar plate; pL = pectinate lamella; sb = sensilla basiconica." figureDoi="http://doi.org/10.5281/zenodo.3767292" httpUri="https://zenodo.org/record/3767292/files/figure.png" pageId="11" pageNumber="12">Figs 7C, D</figureCitation>
), and by anterior telopodite process with a wide and apically swollen tip, unlike in any other Malagasy spirobolid genus (
<figureCitation box="[220,344,1328,1355]" captionStart="Figure 6" captionStartId="13.[140,212,440,462]" captionTargetId="figure@13.[140,1108,165,421]" captionTargetPageId="13" captionText="Figure 6. Sanguinobolus maculosus sp. n., male holotype. A left half of anterior gonopod, anterior view B left half, posterior view C left posterior gonopod, anterior view D left posterior gonopod, posterior view. Cx = coxite; Cx-br = coxite branch; Gr = groove on anterior gonopod; St = sternite; T = telopodite;T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820790" httpUri="https://zenodo.org/record/3820790/files/figure.png" pageId="11" pageNumber="12">Figs 6A, B</figureCitation>
). Sperm canal on posterior gonopod discharging into membranous area of telopodite main branch (
<figureCitation box="[505,598,1363,1390]" captionStart="Figure 6" captionStartId="13.[140,212,440,462]" captionTargetId="figure@13.[140,1108,165,421]" captionTargetPageId="13" captionText="Figure 6. Sanguinobolus maculosus sp. n., male holotype. A left half of anterior gonopod, anterior view B left half, posterior view C left posterior gonopod, anterior view D left posterior gonopod, posterior view. Cx = coxite; Cx-br = coxite branch; Gr = groove on anterior gonopod; St = sternite; T = telopodite;T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820790" httpUri="https://zenodo.org/record/3820790/files/figure.png" pageId="11" pageNumber="12">Fig. 6D</figureCitation>
). Shares a large body size (&gt; 
<quantity box="[915,1015,1364,1390]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="12" unit="mm" value="100.0">100 mm</quantity>
) and an aposematic red-black colour pattern with 
<taxonomicName authority="Silvestri, 1897" authorityName="Silvestri" authorityYear="1897" box="[625,972,1399,1425]" class="Diplopoda" family="Pachybolidae" genus="Aphistogoniulus" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis box="[625,795,1399,1425]" italics="true" pageId="11" pageNumber="12">Aphistogoniulus</emphasis>
<bibRefCitation author="Silvestri" box="[805,972,1399,1425]" firstAuthor="Silvestri" pageId="11" pageNumber="12" pagination="651 - 653" refId="ref57740" refString="Silvestri F (1897) Systema Diplopodum. Annali del Museo civico di storia naturale di Genova 38: 651 - 653." type="journal article" year="1897">Silvestri, 1897</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Wesener &amp; Enghoff, 2008" authorityName="Wesener &amp; Enghoff" authorityYear="2008" class="Diplopoda" family="Pachybolidae" genus="Madabolus" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis box="[987,1107,1399,1425]" italics="true" pageId="11" pageNumber="12">Madabolus</emphasis>
<bibRefCitation author="Wesener &amp; Enghoff" box="[140,446,1434,1460]" firstAuthor="Wesener" pageId="11" pageNumber="12" pagination="37 - 53" refId="ref58000" refString="Wesener T, Enghoff H, Wagele J-W (2008) Pachybolini - a tribe of giant Afrotropical millipedes: arguments for monophyly and the description of a new genus from Madagascar (Diplopoda: Spirobolida: Pachybolidae). Invertebrate Systematics 22: 37 - 53. [http: // www. publish. csiro. au /? act = view _ file &amp; file _ id = IS 07008. pdf]" type="journal article" year="2008">Wesener &amp; Enghoff, 2008</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[458,592,1434,1460]" class="Diplopoda" family="Pachybolidae" genus="Corallobolus" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis box="[458,592,1434,1460]" italics="true" pageId="11" pageNumber="12">Corallobolus</emphasis>
</taxonomicName>
and 
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[652,784,1434,1460]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. n">
<emphasis box="[652,784,1434,1460]" italics="true" pageId="11" pageNumber="12">Colossobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[793,866,1434,1460]" pageId="11" pageNumber="12" rank="genus">gen. n</taxonomicNameLabel>
.. Differs from 
<taxonomicName authority="Silvestri, 1897" authorityName="Silvestri" authorityYear="1897" class="Diplopoda" family="Pachybolidae" genus="Aphistogoniulus" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="11" pageNumber="12">Aphistogoniulus</emphasis>
<bibRefCitation author="Silvestri" box="[257,420,1469,1495]" firstAuthor="Silvestri" pageId="11" pageNumber="12" pagination="651 - 653" refId="ref57740" refString="Silvestri F (1897) Systema Diplopodum. Annali del Museo civico di storia naturale di Genova 38: 651 - 653." type="journal article" year="1897">Silvestri, 1897</bibRefCitation>
</taxonomicName>
and 
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[476,608,1469,1495]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[476,608,1469,1495]" italics="true" pageId="11" pageNumber="12">Colossobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[616,693,1470,1496]" pageId="11" pageNumber="12" rank="genus">gen. n.</taxonomicNameLabel>
by the absence of a mesal branch on the posterior gonopod telopodite (
<figureCitation box="[540,633,1504,1531]" captionStart="Figure 6" captionStartId="13.[140,212,440,462]" captionTargetId="figure@13.[140,1108,165,421]" captionTargetPageId="13" captionText="Figure 6. Sanguinobolus maculosus sp. n., male holotype. A left half of anterior gonopod, anterior view B left half, posterior view C left posterior gonopod, anterior view D left posterior gonopod, posterior view. Cx = coxite; Cx-br = coxite branch; Gr = groove on anterior gonopod; St = sternite; T = telopodite;T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.3820790" httpUri="https://zenodo.org/record/3820790/files/figure.png" pageId="11" pageNumber="12">Fig. 6C</figureCitation>
). The sperm canal in 
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[890,1047,1504,1530]" class="Diplopoda" family="Pachybolidae" genus="Sanguinobolus" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis box="[890,1047,1504,1530]" italics="true" pageId="11" pageNumber="12">Sanguinobolus</emphasis>
</taxonomicName>
runs through the lateral branch, in 
<taxonomicName authorityName="Silvestri" authorityYear="1897" box="[499,669,1539,1565]" class="Diplopoda" family="Pachybolidae" genus="Aphistogoniulus" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis box="[499,669,1539,1565]" italics="true" pageId="11" pageNumber="12">Aphistogoniulus</emphasis>
</taxonomicName>
and 
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[734,866,1539,1565]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[734,866,1539,1565]" italics="true" pageId="11" pageNumber="12">Colossobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[877,958,1540,1566]" pageId="11" pageNumber="12" rank="genus">gen. n.</taxonomicNameLabel>
through the mesal branch. Differs from 
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[446,578,1575,1601]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[446,578,1575,1601]" italics="true" pageId="11" pageNumber="12">Colossobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[584,660,1575,1601]" pageId="11" pageNumber="12" rank="genus">gen. n.</taxonomicNameLabel>
in the absence of a central membranous area basally to the main branch of the posterior gonopod telopodite. 
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[950,1107,1610,1636]" class="Diplopoda" family="Pachybolidae" genus="Sanguinobolus" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis box="[950,1107,1610,1636]" italics="true" pageId="11" pageNumber="12">Sanguinobolus</emphasis>
</taxonomicName>
resembles 
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[258,390,1645,1671]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[258,390,1645,1671]" italics="true" pageId="11" pageNumber="12">Colossobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[400,480,1646,1672]" pageId="11" pageNumber="12" rank="genus">gen. n.</taxonomicNameLabel>
in the presence of a short and wide mesal process on
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767290" ID-Zenodo-Dep="3767290" box="[140,970,1654,1676]" httpUri="https://zenodo.org/record/3767290/files/figure.png" pageId="12" pageNumber="13" startId="12.[140,212,1654,1676]" targetBox="[151,1090,171,1630]" targetPageId="12">
<paragraph blockId="12.[140,970,1654,1676]" box="[140,970,1654,1676]" pageId="12" pageNumber="13">
<emphasis bold="true" box="[140,238,1654,1676]" pageId="12" pageNumber="13">Figure 5.</emphasis>
Distribution map for the genera 
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[546,656,1654,1675]" class="Diplopoda" family="Pachybolidae" genus="Corallobolus" kingdom="Animalia" order="Spirobolida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[546,656,1654,1675]" italics="true" pageId="12" pageNumber="13">Corallobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[662,726,1655,1676]" pageId="12" pageNumber="13" rank="genus">gen. n.</taxonomicNameLabel>
and 
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[772,900,1654,1675]" class="Diplopoda" family="Pachybolidae" genus="Sanguinobolus" kingdom="Animalia" order="Spirobolida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[772,900,1654,1675]" italics="true" pageId="12" pageNumber="13">Sanguinobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[906,970,1655,1676]" pageId="12" pageNumber="13" rank="genus">gen. n.</taxonomicNameLabel>
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3820790" ID-Zenodo-Dep="3820790" httpUri="https://zenodo.org/record/3820790/files/figure.png" pageId="13" pageNumber="14" targetBox="[140,1108,165,420]" targetPageId="13">
<paragraph pageId="13" pageNumber="14">
<emphasis bold="true" box="[140,237,440,462]" pageId="13" pageNumber="14">Figure 6.</emphasis>
<emphasis box="[242,463,440,461]" italics="true" pageId="13" pageNumber="14">Sanguinobolus maculosus</emphasis>
sp. n., male holotype. 
<emphasis bold="true" box="[671,690,440,462]" pageId="13" pageNumber="14">A</emphasis>
left half of anterior gonopod, anterior view 
<emphasis bold="true" box="[1091,1108,440,462]" pageId="13" pageNumber="14">B</emphasis>
left half, posterior view 
<emphasis bold="true" box="[359,378,472,494]" pageId="13" pageNumber="14">C</emphasis>
left posterior gonopod, anterior view 
<emphasis bold="true" box="[729,748,472,494]" pageId="13" pageNumber="14">D</emphasis>
left posterior gonopod, posterior view. Cx = coxite; Cx-br = coxite branch; Gr = groove on anterior gonopod; St = sternite; T = telopodite; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale.
</paragraph>
</caption>
<paragraph blockId="13.[140,1108,659,932]" pageId="13" pageNumber="14">the coxite of the anterior gonopod which is shorter or only slightly longer than the telopodite and sternite.</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="14" type="distribution">
<paragraph blockId="13.[140,1108,659,932]" pageId="13" pageNumber="14">
<emphasis bold="true" box="[185,487,729,756]" pageId="13" pageNumber="14">
<emphasis box="[185,336,729,755]" inLineHeading="true" pageId="13" pageNumber="14" reason="1">Distribution</emphasis>
and ecology
</emphasis>
: only known from the isolated rainforest in Montagne d’Ambre, north 
<collectingCountry box="[327,460,765,791]" name="Madagascar" pageId="13" pageNumber="14">Madagascar</collectingCountry>
(
<figureCitation box="[477,546,765,792]" captionStart="Figure 5" captionStartId="12.[140,212,1654,1676]" captionTargetBox="[151,1090,171,1630]" captionTargetId="figure@12.[156,1092,165,1635]" captionTargetPageId="12" captionText="Figure 5. Distribution map for the genera Corallobolus gen. n. and Sanguinobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3767290" httpUri="https://zenodo.org/record/3767290/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation>
). Species of the genus are probably active on the soil surface because of their relatively long legs and aposematic colour pattern.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="14" type="etymology">
<paragraph blockId="13.[140,1108,659,932]" pageId="13" pageNumber="14">
<emphasis bold="true" box="[185,315,835,861]" inLineHeading="true" pageId="13" pageNumber="14" reason="1">Etymology</emphasis>
: 
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[334,491,835,861]" class="Diplopoda" family="Pachybolidae" genus="Sanguinobolus" kingdom="Animalia" order="Spirobolida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis box="[334,491,835,861]" italics="true" pageId="13" pageNumber="14">Sanguinobolus</emphasis>
</taxonomicName>
, masculine, is composed of 
<emphasis box="[829,935,836,861]" italics="true" pageId="13" pageNumber="14">sanguinus</emphasis>
, blood, which refers to the blood-red colour pattern of the single constituent species, and - 
<emphasis box="[1030,1085,871,897]" italics="true" pageId="13" pageNumber="14">bolus</emphasis>
a common suffix for genera of the order 
<taxonomicName authorityName="Cook" authorityYear="1895" box="[577,711,906,932]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>