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<mods:title id="660A2955346730754D8212CE4B9F9BFA">Redefinition And Generic Revision Of The North American Vaejovid Scorpion Subfamily Syntropinae Kraepelin, 1905, With Descriptions Of Six New Genera</mods:title>
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<taxonomicName id="58E52ED0FFC62E6508B79D489714FBDF" ID-CoL="7W89" authority="Soleglad and Fet, 2008" authorityName="Soleglad and Fet" authorityYear="2008" box="[194,588,1041,1064]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC62E6508B79D48946CFBD0" box="[194,308,1041,1064]" italics="true" pageId="53" pageNumber="53">Thorellius</emphasis>
<bibRefCitation id="FB7428A2FFC62E6509499D489714FBDF" author="Soleglad, M. E. &amp; V. Fet" box="[316,588,1041,1063]" pageId="53" pageNumber="53" pagination="1 - 115" refId="ref39584" refString="Soleglad, M. E., and V. Fet. 2008. Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae). Euscorpius 71: 1 - 115." type="journal article" year="2008">Soleglad and Fet, 2008</bibRefCitation>
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<subSubSection id="D7FF06D8FFC62E6508F29D6997DEFBBD" box="[135,646,1072,1094]" pageId="53" pageNumber="53" type="description">
<paragraph id="9F5A5553FFC62E6508F29D6997DEFBBD" blockId="53.[135,646,1041,1094]" box="[135,646,1072,1094]" pageId="53" pageNumber="53">
<figureCitation id="07DE49D6FFC62E6508F29D699451FBBD" box="[135,265,1072,1094]" captionStart="Fig" captionStartId="4.[116,155,1349,1368]" captionTargetBox="[93,1193,211,1319]" captionTargetId="figure-85@4.[93,1193,211,1319]" captionTargetPageId="4" captionText="Fig. 1. Representative species in the North American vaejovid scorpion subfamily Syntropinae Kraepelin, 1905. A. Chihuahuanus bilineatus (Pocock, 1898), comb. nov., ♀; B. Kochius bruneus (Williams, 1970), 8; C. Mesomexovis punctatus (Karsch, 1879), comb. nov., 8; D. Syntropis macrura Kraepelin, 1900, ♀; E. Thorellius sp., 8; F. Vizcaino viscainensis (Williams, 1970), comb. nov., 8. Photos in B, D and E courtesy of R. Mercurio." figureDoi="http://doi.org/10.5281/zenodo.4611869" httpUri="https://zenodo.org/record/4611869/files/figure.png" pageId="53" pageNumber="53">Figures 1E</figureCitation>
, 
<figureCitation id="07DE49D6FFC62E65096D9D699470FBBE" box="[280,296,1072,1094]" captionStart="Fig" captionStartId="9.[150,188,1091,1110]" captionTargetBox="[130,1231,211,1061]" captionTargetId="figure-240@9.[128,1231,211,1061]" captionTargetPageId="9" captionText="Fig. 5. Map of southwestern North America, plotting known locality records for two genera in the vaejovid scorpion subfamily Syntropinae Kraepelin, 1905, based on data collected in the present study: Kochius Soleglad and Fet, 2008 (circles); Thorellius Soleglad and Fet, 2008 (squares)." figureDoi="http://doi.org/10.5281/zenodo.4611881" httpUri="https://zenodo.org/record/4611881/files/figure.png" pageId="53" pageNumber="53">5</figureCitation>
, 
<figureCitation id="07DE49D6FFC62E6509439D69941EFBBD" box="[310,326,1072,1093]" captionStart="Fig" captionStartId="13.[139,178,1308,1328]" captionTargetBox="[341,1397,490,1278]" captionTargetId="figure-0@13.[341,1397,490,1278]" captionTargetPageId="13" captionText="Fig. 7. Most parsimonious tree (length: 10894.593, CI: 0.190, RI: 0.628, fit: 244.08), obtained by simultaneous phylogenetic analysis of 250 qualitative and quantitative morphological characters and 4221 aligned DNA nucleotides from three mitochondrial and two nuclear gene markers, for 18 genera and 61 species in the North American vaejovid scorpion subfamily Syntropinae Kraepelin, 1905. Analyses were conducted with an implied weighting regime (k 518) that maximized average support (González-Santillán and Prendini, in press). Synapomorphies are optimized with accelerated transformation. Black bars indicate uniquely derived apomorphic states, white bars indicate parallel derivations of apomorphic states, numbers above indicate characters, and numbers below indicate states (appendix 1)." figureDoi="http://doi.org/10.5281/zenodo.4611887" httpUri="https://zenodo.org/record/4611887/files/figure.png" pageId="53" pageNumber="53">7</figureCitation>
, 
<figureCitation id="07DE49D6FFC62E6509219D6994F4FBBD" box="[340,428,1072,1093]" captionStart="Fig" captionStartId="32.[116,155,1645,1664]" captionTargetBox="[187,1073,219,1600]" captionTargetId="figure-1@32.[187,1098,211,1615]" captionTargetPageId="32" captionText="Fig. 17. Syntropinae Kraepelin, 1905, dextral pedipalp chela, dorsal (A, C) and retrolateral (B, D) aspects, illustrating carinae and distribution of trichobothria. A, B. Chihuahuanus crassimanus (Pocock, 1898), comb. nov., 8 (AMNH). C, D. Thorellius intrepidus (Thorell, 1876), 8 (AMNH). Scale bars 5 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4611919" httpUri="https://zenodo.org/record/4611919/files/figure.png" pageId="53" pageNumber="53">17C, D</figureCitation>
, 
<figureCitation id="07DE49D6FFC62E6509CF9D6994B2FBBD" box="[442,490,1072,1093]" captionStart="Fig" captionStartId="37.[150,188,1460,1479]" captionTargetBox="[139,1213,218,1430]" captionTargetId="figure-75@37.[128,1228,211,1430]" captionTargetPageId="37" captionText="Fig. 20. Syntropinae Kraepelin, 1905, dextral pedipalp chela movable finger, dorsal aspect, illustrating dentition. A. Balsateres cisnerosi (Ponce-Saavedra and Sissom, 2004), comb. nov., 8 (IBUNAM). B. Thorellius intrepidus (Thorell, 1876), 8 (AMNH). C. Vizcaino viscainensis (Williams, 1970), comb. nov., 8 (AMNH). Scale bars 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4611927" httpUri="https://zenodo.org/record/4611927/files/figure.png" pageId="53" pageNumber="53">20B</figureCitation>
, 
<figureCitation id="07DE49D6FFC62E65098D9D699770FBBD" box="[504,552,1072,1093]" captionStart="Fig" captionStartId="40.[116,155,1317,1336]" captionTargetBox="[93,1182,217,1279]" captionTargetId="figure-140@40.[93,1193,211,1287]" captionTargetPageId="40" captionText="Fig. 21. Syntropinae Kraepelin, 1905, dextral leg III, basitarsus and telotarsus, ventral aspect, illustrating macrosetae and spinules. A. Chihuahuanus globosus (Borelli, 1915), comb. nov., 8 (AMNH). B. Thorellius intrepidus (Thorell, 1876), 8 (AMNH). C. Vizcaino viscainensis (Williams, 1970), comb. nov., 8 (AMNH). Scale bars 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4611931" httpUri="https://zenodo.org/record/4611931/files/figure.png" pageId="53" pageNumber="53">21B</figureCitation>
; 
<tableCitation id="D26760E8FFC62E650A439D6997DEFBBD" box="[566,646,1072,1094]" captionStart="TABLE 1" captionStartId="6.[590,672,214,233]" captionTargetBox="[93,1183,531,993]" captionTargetPageId="6" captionText="TABLE 1 Currently recognized genera in the North American vaejovid scorpion subfamily Syntropinae Kraepelin, 1905, with countries, states and geographical regions from which they have been recorded Abbreviations: Mex. (Mexico): Ags (Aguascalientes), B.C. (Baja California), B.C.S. (Baja California Sur), Chis (Chiapas), Chih. (Chihuahua), Coah. (Coahuila), Col. (Colima), Dgo (Durango), Gto (Guanajuato), Gro (Guerrero), Hgo (Hidalgo), Jal. (Jalisco), Edomex. (Estado de México), Mich. (Michoacán), Mor. (Morelos), Nay. (Nayarit), N.L. (Nuevo León), Oax. (Oaxaca), Pue. (Puebla), Qro (Querétaro), Sin. (Sinaloa), Son. (Sonora), Tamps (Tamaulipas), Tlax. (Tlaxcala), Ver. (Veracruz), Zac. (Zacatecas); U.S.A.: AZ (Arizona), CA (California), NM (New Mexico), NV (Nevada), OR (Oregon), TX (Texas), UT (Utah); Balsas Depression (BD), Baja California Peninsula (BCP), Chihuahuan Desert (CD), Great Basin Desert (GBD), Mojave Desert (MD), Pacific Coast (PC), Sonoran Desert (SD), Trans-Mexican Volcanic Belt (TMVB)." httpUri="http://table.plazi.org/id/CB9A05DBFFF52E560A3B998F95B5FE08" pageId="53" pageNumber="53" tableUuid="CB9A05DBFFF52E560A3B998F95B5FE08">table 1</tableCitation>
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<taxonomicName id="58E52ED0FFC62E6508F59D0794B0FB88" authority="Thorell, 1876" authorityName="Thorell" authorityYear="1876" box="[128,488,1116,1136]" class="Arachnida" family="Vaejovidae" genus="Vejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="species" species="intrepidus">
<emphasis id="AD918941FFC62E6508F59D079467FB88" box="[128,319,1116,1136]" italics="true" pageId="53" pageNumber="53">Vejovis intrepidus</emphasis>
<bibRefCitation id="FB7428A2FFC62E6509249D0494B0FB88" author="Thorell, T." box="[337,488,1117,1136]" pageId="53" pageNumber="53" pagination="1 - 15" refId="ref39938" refString="Thorell, T. 1876. On the classification of scorpions. Annals and Magazine of Natural History 4: 1 - 15." type="journal article" year="1876">Thorell, 1876</bibRefCitation>
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[
<emphasis id="AD918941FFC62E650A749D05974DFB89" box="[513,533,1116,1137]" italics="true" pageId="53" pageNumber="53">=</emphasis>
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<emphasis id="AD918941FFC62E650A5D9D0595A5FB73" italics="true" pageId="53" pageNumber="53">Thorellius intrepidus</emphasis>
(
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)
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species, by original designation.
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<taxonomicName id="58E52ED0FFC62E6508F59DF69592FB39" authorityName="Thorell" authorityYear="1876" box="[128,202,1199,1217]" class="Arachnida" family="Vaejovidae" genus="Vejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC62E6508F59DF69592FB39" box="[128,202,1199,1217]" italics="true" pageId="53" pageNumber="53">Vejovis</emphasis>
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second section (part): 
<treatmentCitation id="1E447342FFC62E6509A19DF6959AFB25" author="Hoffmann, C. C." page="134" pageId="53" pageNumber="53" year="1931">
<bibRefCitation id="FB7428A2FFC62E6509A19DF6959AFB25" author="Hoffmann, C. C." pageId="53" pageNumber="53" pagination="291 - 408" refId="ref37778" refString="Hoffmann, C. C. 1931. Monografias para la entomologia medica de Mexico. Monografia Num. 2, Los escorpiones de Mexico. Primera parte: Diplocentridae, Chactidae, Vaejovidae. Anales del Instituto de Biologia Universidad Nacional Autonoma de Mexico 2: 291 - 408." type="journal article" year="1931">Hoffmann, 1931: 134</bibRefCitation>
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, 139.
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<treatmentCitationGroup id="BFF5727DFFC62E6508F59DBF9482FAEB" pageId="53" pageNumber="53">
<taxonomicName id="58E52ED0FFC62E6508F59DBF958EFB00" ID-CoL="85MN" authorityName="C. L. Koch" authorityYear="1836" box="[128,214,1254,1272]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC62E6508F59DBF958EFB00" box="[128,214,1254,1272]" italics="true" pageId="53" pageNumber="53">Vaejovis</emphasis>
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second section (part): 
<treatmentCitation id="1E447342FFC62E65098E9DBC948EFAEB" author="Francke, O. F. &amp; E. Gonzalez-Santillan" page="590" pageId="53" pageNumber="53" year="2007">
<bibRefCitation id="FB7428A2FFC62E65098E9DBC948EFAEB" author="Francke, O. F. &amp; E. Gonzalez-Santillan" pageId="53" pageNumber="53" pagination="586 - 591" refId="ref37034" refString="Francke, O. F., and E. Gonzalez-Santillan. 2007. A new species belonging to the Vaejovis punctipalpi group (Scorpiones, Vaejovidae) from southern Mexico. Journal of Arachnology 34: 586 - 591." type="journal article" year="2007">Francke and González-Santillán, 2007: 590</bibRefCitation>
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.
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</paragraph>
<paragraph id="9F5A5553FFC62E6508F59C44971AF9FF" blockId="53.[128,655,1116,1653]" pageId="53" pageNumber="53">
<treatmentCitationGroup id="BFF5727DFFC62E6508F59C44971AF9FF" pageId="53" pageNumber="53">
<taxonomicName id="58E52ED0FFC62E6508F59C449466FAD7" authorityName="Thorell" authorityYear="1876" box="[128,318,1307,1327]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="species" species="intrepidus">
<emphasis id="AD918941FFC62E6508F59C449466FAD7" box="[128,318,1307,1327]" italics="true" pageId="53" pageNumber="53">Vaejovis intrepidus</emphasis>
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group (part): 
<treatmentCitation id="1E447342FFC62E6509A59C4597D1FAD7" author="Sissom, W. D." box="[464,649,1308,1327]" page="180" pageId="53" pageNumber="53" year="1989">
<bibRefCitation id="FB7428A2FFC62E6509A59C4597D1FAD7" author="Sissom, W. D." box="[464,649,1308,1327]" pageId="53" pageNumber="53" pagination="179 - 187" refId="ref39026" refString="Sissom, W. D. 1989. Redescription of Vaejovis occidentalis Hoffmann with a revised diagnosis for Vaejovis subcristatus Pocock (Scorpiones: Vaejovidae). Revue Arachnologique 8: 179 - 187." type="journal article" year="1989">Sissom, 1989: 180</bibRefCitation>
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; 
<treatmentCitation id="1E447342FFC62E6508EF9C6E95AAFAB2" author="Sissom, W. D." box="[154,242,1335,1354]" page="24" pageId="53" pageNumber="53" year="1991">
<bibRefCitation id="FB7428A2FFC62E6508EF9C6E95AAFAB2" author="Sissom, W. D." box="[154,242,1335,1354]" pageId="53" pageNumber="53" pagination="4 - 28" refId="ref39100" refString="Sissom, W. D. 1991. Systematic studies on the nitidulus group of the genus Vaejovis, with descriptions of seven new species (Scorpiones, Vaejovidae). Journal of Arachnology 19: 4 - 28." type="journal article" year="1991">1991: 24</bibRefCitation>
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, 26; 
<treatmentCitation id="1E447342FFC62E6509579C6E94AFFAB2" author="Stockwell, S. A." box="[290,503,1335,1354]" page="409" pageId="53" pageNumber="53" year="1992">
<bibRefCitation id="FB7428A2FFC62E6509579C6E94AFFAB2" author="Stockwell, S. A." box="[290,503,1335,1354]" pageId="53" pageNumber="53" pagination="407 - 422" refId="ref39903" refString="Stockwell, S. A. 1992. Systematic observations on North American Scorpionida with a key and checklist of the families and genera. Journal of Medical Entomology 29: 407 - 422." type="journal article" year="1992">Stockwell, 1992: 409</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="1E447342FFC62E650A749C6E95EDFA9D" author="Sissom, W. D." page="68" pageId="53" pageNumber="53" year="1993">
<bibRefCitation id="FB7428A2FFC62E650A749C6E95EDFA9D" author="Sissom, W. D." pageId="53" pageNumber="53" pagination="64 - 68" refId="ref39140" refString="Sissom, W. D. 1993. A new species of Vaejovis (Scorpiones, Vaejovidae) from Western Arizona, with supplemental notes on the male of Vaejovis spicatus Haradon. Journal of Arachnology 21: 64 - 68." type="journal article" year="1993">Sissom, 1993: 68</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="1E447342FFC62E6508B09C0B9776FA9D" author="Lourenco, W. R. &amp; W. D. Sissom" box="[197,558,1362,1381]" page="135" pageId="53" pageNumber="53" year="2000">
<bibRefCitation id="FB7428A2FFC62E6508B09C0B9776FA9D" author="Lourenco, W. R. &amp; W. D. Sissom" box="[197,558,1362,1381]" pageId="53" pageNumber="53" pagination="115 - 135" refId="ref38119" refString="Lourenco, W. R., and W. D. Sissom. 2000. Scorpiones. In B. Llorente, E. Gonzalez, and N. Papavero (editors), Biodiversidad, taxonomia y biogeografia de artropodos de Mexico: hacia una sintesis de su conocimiento: 115 - 135. UNAM, Mexico." type="book chapter" year="2000">Lourenço and Sissom, 2000: 135</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="1E447342FFC62E650A4A9C0B945BFA78" author="Sissom, W. D." page="537" pageId="53" pageNumber="53" year="2000">
<bibRefCitation id="FB7428A2FFC62E650A4A9C0B945BFA78" author="Sissom, W. D." pageId="53" pageNumber="53" pagination="503 - 553" refId="ref39182" refString="Sissom, W. D. 2000. Family Vaejovidae. In V. Fet, W. D. Sissom, G. Lowe, and M. E. Braunwalder. Catalog of the scorpions of the world (1758 - 1998): 503 - 553. New York Entomological Society: New York." type="book chapter" year="2000">Sissom, 2000: 537</bibRefCitation>
</treatmentCitation>
, 538, 551; 
<treatmentCitation id="1E447342FFC62E6509089C3495B0FA63" author="Armas, L. F. de &amp; E. Martin-Frias" page="8" pageId="53" pageNumber="53" year="2001">
<bibRefCitation id="FB7428A2FFC62E6509089C3495B0FA63" author="Armas, L. F. de &amp; E. Martin-Frias" pageId="53" pageNumber="53" pagination="8 - 16" refId="ref36020" refString="Armas, L. F. de., and E. Martin-Frias. 2001. Dos nuevos Vaejovis (Scorpiones: Vaejovidae) de Guerrero y Nayarit, Mexico. Solenodon 1: 8 - 16." type="journal article" year="2001">Armas and Martín-Frías, 2001: 8</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="1E447342FFC62E6508839CD1948BFA63" author="Hendrixson, B. E." box="[246,467,1416,1435]" page="47" pageId="53" pageNumber="53" year="2001">
<bibRefCitation id="FB7428A2FFC62E6508839CD1948BFA63" author="Hendrixson, B. E." box="[246,467,1416,1435]" pageId="53" pageNumber="53" pagination="47 - 55" refId="ref37712" refString="Hendrixson, B. E. 2001. A new species of Vaejovis (Scorpiones, Vaejovidae) from Sonora, Mexico. Journal of Arachnology 29: 47 - 55." type="journal article" year="2001">Hendrixson 2001: 47</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="1E447342FFC62E6509959CD1947FFA4E" author="Gonzalez-Santillan, E." page="30" pageId="53" pageNumber="53" year="2004">
<bibRefCitation id="FB7428A2FFC62E6509959CD1947FFA4E" author="Gonzalez-Santillan, E." pageId="53" pageNumber="53" pagination="25 - 35" refId="ref37379" refString="Gonzalez-Santillan, E. 2004. Diversidad, taxonomia y habitat de alacranes. In A. N. Garcia, and R. Ayala (editors), Artropodos de Chamela: 25 - 35. Instituto de Biologia, UNAM, Mexico." type="book chapter" year="2004">González-Santillán, 2004: 30</bibRefCitation>
</treatmentCitation>
, 31; 
<treatmentCitation id="1E447342FFC62E6509289CFA9452FA29" author="Ponce-Saavedra, J. &amp; W. D. Sissom" page="539" pageId="53" pageNumber="53" year="2004">
<bibRefCitation id="FB7428A2FFC62E6509289CFA9452FA29" author="Ponce-Saavedra, J. &amp; W. D. Sissom" pageId="53" pageNumber="53" pagination="539 - 544" refId="ref38517" refString="Ponce-Saavedra, J., and W. D. Sissom. 2004. A new species of the genus Vaejovis (Scorpiones, Vaejovidae) endemic to the Balsas basin of Michoacan, Mexico. Journal of Arachnology 32: 539 - 544." type="journal article" year="2004">Ponce-Saavedra and Sissom, 2004: 539</bibRefCitation>
</treatmentCitation>
, 541; 
<treatmentCitation id="1E447342FFC62E65092C9CE797D1FA29" author="Graham, M. R. &amp; V. Fet" box="[345,649,1470,1489]" page="7" pageId="53" pageNumber="53" year="2006">
<bibRefCitation id="FB7428A2FFC62E65092C9CE797D1FA29" author="Graham, M. R. &amp; V. Fet" box="[345,649,1470,1489]" pageId="53" pageNumber="53" pagination="1 - 19" refId="ref37461" refString="Graham, M. R., and V. Fet. 2006. Serrula in retrospect: a historical look at scorpion literature (Scorpiones: Orthosterni). Euscorpius 48: 1 - 19." type="journal article" year="2006">Graham and Fet, 2006: 7</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="1E447342FFC62E6508EF9C80943FFA14" author="McWest, K. J." box="[154,359,1497,1516]" page="66" pageId="53" pageNumber="53" year="2009">
<bibRefCitation id="FB7428A2FFC62E6508EF9C80943FFA14" author="McWest, K. J." box="[154,359,1497,1516]" pageId="53" pageNumber="53" pagination="1 - 16" refId="ref38179" refString="McWest, K. J. 2009. Tarsal spinules and setae of vaejovid scorpions (Scorpiones: Vaejovidae). Zootaxa 2001: 1 - 16." type="journal article" year="2009">McWest, 2009: 66</bibRefCitation>
</treatmentCitation>
, 69, 70, 100–102, 
<tableCitation id="D26760E8FFC62E650A359C8097D0FA14" box="[576,648,1497,1516]" captionStart="TABLE 1" captionStartId="6.[590,672,214,233]" captionTargetBox="[93,1183,531,993]" captionTargetPageId="6" captionText="TABLE 1 Currently recognized genera in the North American vaejovid scorpion subfamily Syntropinae Kraepelin, 1905, with countries, states and geographical regions from which they have been recorded Abbreviations: Mex. (Mexico): Ags (Aguascalientes), B.C. (Baja California), B.C.S. (Baja California Sur), Chis (Chiapas), Chih. (Chihuahua), Coah. (Coahuila), Col. (Colima), Dgo (Durango), Gto (Guanajuato), Gro (Guerrero), Hgo (Hidalgo), Jal. (Jalisco), Edomex. (Estado de México), Mich. (Michoacán), Mor. (Morelos), Nay. (Nayarit), N.L. (Nuevo León), Oax. (Oaxaca), Pue. (Puebla), Qro (Querétaro), Sin. (Sinaloa), Son. (Sonora), Tamps (Tamaulipas), Tlax. (Tlaxcala), Ver. (Veracruz), Zac. (Zacatecas); U.S.A.: AZ (Arizona), CA (California), NM (New Mexico), NV (Nevada), OR (Oregon), TX (Texas), UT (Utah); Balsas Depression (BD), Baja California Peninsula (BCP), Chihuahuan Desert (CD), Great Basin Desert (GBD), Mojave Desert (MD), Pacific Coast (PC), Sonoran Desert (SD), Trans-Mexican Volcanic Belt (TMVB)." httpUri="http://table.plazi.org/id/CB9A05DBFFF52E560A3B998F95B5FE08" pageId="53" pageNumber="53" tableUuid="CB9A05DBFFF52E560A3B998F95B5FE08">table 1</tableCitation>
; 
<treatmentCitation id="1E447342FFC62E6508EF9CAD9766F9FF" author="Santibanez-Lopez, C. &amp; W. D. Sissom" box="[154,574,1524,1543]" page="52" pageId="53" pageNumber="53" year="2010">
<bibRefCitation id="FB7428A2FFC62E6508EF9CAD9766F9FF" author="Santibanez-Lopez, C. &amp; W. D. Sissom" box="[154,574,1524,1543]" pageId="53" pageNumber="53" pagination="49 - 58" refId="ref38931" refString="Santibanez-Lopez, C., and W. D. Sissom. 2010. A new species of the Vaejovis eusthenura group in Oaxaca, Mexico (Scorpiones: Vaejovidae). Zootaxa 2493: 49 - 58." type="journal article" year="2010">Santibáñez-López and Sissom, 2010: 52</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9F5A5553FFC62E6508F59F49975CF9C6" blockId="53.[128,655,1116,1653]" pageId="53" pageNumber="53">
<treatmentCitationGroup id="BFF5727DFFC62E6508F59F49975CF9C6" pageId="53" pageNumber="53">
<taxonomicName id="58E52ED0FFC62E6508F59F499413F9DB" authority="Fet et al., 2006" box="[128,331,1551,1571]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="species" species="punctipalpi">
<emphasis id="AD918941FFC62E6508F59F499413F9DB" box="[128,331,1551,1571]" italics="true" pageId="53" pageNumber="53">Vaejovis punctipalpi</emphasis>
</taxonomicName>
group (part): 
<bibRefCitation id="FB7428A2FFC62E6509959F4997D3F9DB" author="Fet, V., M. E. &amp; M. S. Brewer" box="[480,651,1552,1571]" pageId="53" pageNumber="53" pagination="1 - 19" refId="ref36799" refString="Fet, V., M. E. Soleglad, and M. S. Brewer. 2006 a. Laterobasal aculear serrations (LAS) in scorpion family Vaejovidae (Scorpiones: Chactoidea). Euscorpius 45: 1 - 19." type="journal article" year="2006">Fet et al., 2006a</bibRefCitation>
: 
<tableCitation id="D26760E8FFC62E6508EF9F7295BAF9C6" box="[154,226,1579,1598]" captionStart="TABLE 1" captionStartId="6.[590,672,214,233]" captionTargetBox="[93,1183,531,993]" captionTargetPageId="6" captionText="TABLE 1 Currently recognized genera in the North American vaejovid scorpion subfamily Syntropinae Kraepelin, 1905, with countries, states and geographical regions from which they have been recorded Abbreviations: Mex. (Mexico): Ags (Aguascalientes), B.C. (Baja California), B.C.S. (Baja California Sur), Chis (Chiapas), Chih. (Chihuahua), Coah. (Coahuila), Col. (Colima), Dgo (Durango), Gto (Guanajuato), Gro (Guerrero), Hgo (Hidalgo), Jal. (Jalisco), Edomex. (Estado de México), Mich. (Michoacán), Mor. (Morelos), Nay. (Nayarit), N.L. (Nuevo León), Oax. (Oaxaca), Pue. (Puebla), Qro (Querétaro), Sin. (Sinaloa), Son. (Sonora), Tamps (Tamaulipas), Tlax. (Tlaxcala), Ver. (Veracruz), Zac. (Zacatecas); U.S.A.: AZ (Arizona), CA (California), NM (New Mexico), NV (Nevada), OR (Oregon), TX (Texas), UT (Utah); Balsas Depression (BD), Baja California Peninsula (BCP), Chihuahuan Desert (CD), Great Basin Desert (GBD), Mojave Desert (MD), Pacific Coast (PC), Sonoran Desert (SD), Trans-Mexican Volcanic Belt (TMVB)." httpUri="http://table.plazi.org/id/CB9A05DBFFF52E560A3B998F95B5FE08" pageId="53" pageNumber="53" tableUuid="CB9A05DBFFF52E560A3B998F95B5FE08">table 1</tableCitation>
; 
<treatmentCitation id="1E447342FFC62E6508859F729758F9C6" author="Soleglad, M. E. &amp; V. Fet" box="[240,512,1579,1598]" page="6" pageId="53" pageNumber="53" year="2006">
<bibRefCitation id="FB7428A2FFC62E6508859F729758F9C6" author="Soleglad, M. E. &amp; V. Fet" box="[240,512,1579,1598]" pageId="53" pageNumber="53" pagination="1 - 32" refId="ref39541" refString="Soleglad, M. E., and V. Fet. 2006. Contributions to scorpion systematics. II. Stahnkeini, a new tribe in scorpion family Vaejovidae (Scorpiones: Chactoidea). Euscorpius 40: 1 - 32." type="journal article" year="2006">Soleglad and Fet, 2006: 6</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9F5A5553FFC62E6508F59F1F9756F98C" blockId="53.[128,655,1116,1653]" pageId="53" pageNumber="53">
<treatmentCitationGroup id="BFF5727DFFC62E6508F59F1F9756F98C" pageId="53" pageNumber="53">
<treatmentCitation id="1E447342FFC62E6508F59F1F94A8F9A1" author="Soleglad, M. E. &amp; V. Fet" box="[128,496,1606,1626]" page="1" pageId="53" pageNumber="53" year="2008">
<taxonomicName id="58E52ED0FFC62E6508F59F1F94A8F9A1" ID-CoL="7W89" authority="Soleglad and Fet, 2008: 1" authorityName="Soleglad and Fet" authorityPageNumber="1" authorityYear="2008" box="[128,496,1606,1626]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC62E6508F59F1F95BEF9A2" box="[128,230,1606,1626]" italics="true" pageId="53" pageNumber="53">Thorellius</emphasis>
<bibRefCitation id="FB7428A2FFC62E65089E9F1F94A8F9A1" author="Soleglad, M. E. &amp; V. Fet" box="[235,496,1606,1626]" pageId="53" pageNumber="53" pagination="1 - 115" refId="ref39584" refString="Soleglad, M. E., and V. Fet. 2008. Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae). Euscorpius 71: 1 - 115." type="journal article" year="2008">Soleglad and Fet, 2008: 1</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 95, 102 (part); 
<treatmentCitation id="1E447342FFC62E6508EF9F3B949BF98C" author="Ayrey, R. F. &amp; M. E. Soleglad" box="[154,451,1633,1653]" page="1" pageId="53" pageNumber="53" year="2011">
<bibRefCitation id="FB7428A2FFC62E6508EF9F3B949BF98C" author="Ayrey, R. F. &amp; M. E. Soleglad" box="[154,451,1633,1653]" pageId="53" pageNumber="53" pagination="1 - 15" refId="ref36088" refString="Ayrey, R. F., and M. E. Soleglad. 2011. A new species of Vaejovis from Prescott, Arizona (Scorpiones: Vaejovidae). Euscorpius 114: 1 - 15." type="journal article" year="2011">Ayrey and Soleglad, 2011: 1</bibRefCitation>
</treatmentCitation>
(part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D7FF06D8FFC62E6508E99FD596E8FB36" pageId="53" pageNumber="53" type="diagnosis">
<paragraph id="9F5A5553FFC62E6508E99FD59127FDF7" blockId="53.[128,654,1676,1759]" lastBlockId="53.[701,1228,213,1759]" pageId="53" pageNumber="53">
DIAGNOSIS: 
<taxonomicName id="58E52ED0FFC62E6509409FD494FFF95C" authorityName="Soleglad and Fet" authorityYear="2008" box="[309,423,1677,1700]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC62E6509409FD494FFF95C" box="[309,423,1677,1700]" italics="true" pageId="53" pageNumber="53">Thorellius</emphasis>
</taxonomicName>
may be separated from other genera of 
<taxonomicName id="58E52ED0FFC62E6509EB9FF29773F939" authorityName="KRAEPELIN" authorityYear="1905" box="[414,555,1707,1729]" class="Arachnida" family="Vaejovidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="64" phylum="Arthropoda" rank="subFamily" subFamily="Syntropinae">Syntropinae</taxonomicName>
by the presence of nine fully developed retroventral macrosetae on the basitarsus of leg III (fig. 21B). Most of the other genera bear six or seven fully developed retroventral macrosetae, the other retroventral setae being smaller (
<bibRefCitation id="FB7428A2FFC62E650B5298139682FE98" author="McWest, K. J." box="[807,986,330,352]" pageId="53" pageNumber="53" pagination="1 - 16" refId="ref38179" refString="McWest, K. J. 2009. Tarsal spinules and setae of vaejovid scorpions (Scorpiones: Vaejovidae). Zootaxa 2001: 1 - 16." type="journal article" year="2009">McWest, 2009</bibRefCitation>
), whereas 
<taxonomicName id="58E52ED0FFC62E650C2A98139194FEA7" authorityName="Kraepelin" authorityYear="1900" box="[1119,1228,330,351]" class="Arachnida" family="Vaejovidae" genus="Syntropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC62E650C2A98139194FEA7" box="[1119,1228,330,351]" italics="true" pageId="53" pageNumber="53">Syntropis</emphasis>
</taxonomicName>
has more than 10. The pedipalp patellar rlds and rlm carinae are weakly developed and smooth, the dorsal surface of the pedipalp femur finely and sparsely granular, and the ventral surface of the telson vesicle shagreened in all species of 
<taxonomicName id="58E52ED0FFC62E650C7398A09120FDE8" authorityName="Soleglad and Fet" authorityYear="2008" box="[1030,1144,505,528]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC62E650C7398A09120FDE8" box="[1030,1144,505,528]" italics="true" pageId="53" pageNumber="53">Thorellius</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="9F5A5553FFC62E650AAE9B4E96E8FB36" blockId="53.[701,1228,213,1759]" pageId="53" pageNumber="53">
<taxonomicName id="58E52ED0FFC62E650AAE9B4E9615FDD6" authorityName="Soleglad and Fet" authorityYear="2008" box="[731,845,535,558]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC62E650AAE9B4E9615FDD6" box="[731,845,535,558]" italics="true" pageId="53" pageNumber="53">Thorellius</emphasis>
</taxonomicName>
is most closely related to 
<taxonomicName id="58E52ED0FFC62E650CF69B4E97ABFDB2" authority="González-Santillán &amp; Prendini, 2013" authorityName="González-Santillán &amp; Prendini" authorityYear="2013" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="AD918941FFC62E650CF69B4E97ABFDB2" italics="true" pageId="53" pageNumber="53">Balsateres</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="B6A2343AFFC62E650B739B6C962FFDB2" box="[774,887,565,586]" pageId="53" pageNumber="53" rank="genus">gen. nov.</taxonomicNameLabel>
, with which it shares broad pedipalpal and metasomal carinae. However, the carinae of 
<taxonomicName id="58E52ED0FFC62E650B019B3696BEFD7E" authorityName="Soleglad and Fet" authorityYear="2008" box="[884,998,623,646]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC62E650B019B3696BEFD7E" box="[884,998,623,646]" italics="true" pageId="53" pageNumber="53">Thorellius</emphasis>
</taxonomicName>
are moderately to densely granular, instead of smooth as in 
<taxonomicName id="58E52ED0FFC62E650AC89BF0966AFD38" authority="González-Santillán &amp; Prendini, 2013" authorityName="González-Santillán &amp; Prendini" authorityYear="2013" box="[701,818,681,704]" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="AD918941FFC62E650AC89BF0966AFD38" box="[701,818,681,704]" italics="true" pageId="53" pageNumber="53">Balsateres</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="B6A2343AFFC62E650B349BF396F3FD47" box="[833,939,682,703]" pageId="53" pageNumber="53" rank="genus">gen. nov.</taxonomicNameLabel>
Both sexes of 
<taxonomicName id="58E52ED0FFC62E650C279BF0919CFD38" authorityName="Soleglad and Fet" authorityYear="2008" box="[1106,1220,681,704]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC62E650C279BF0919CFD38" box="[1106,1220,681,704]" italics="true" pageId="53" pageNumber="53">Thorellius</emphasis>
</taxonomicName>
, especially the female, exhibit a distinct proximal gap between the fixed and movable fingers, when closed, which is absent in 
<taxonomicName id="58E52ED0FFC62E650AC89A47966AFCCD" authority="González-Santillán &amp; Prendini, 2013" authorityName="González-Santillán &amp; Prendini" authorityYear="2013" box="[701,818,798,821]" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="AD918941FFC62E650AC89A47966AFCCD" box="[701,818,798,821]" italics="true" pageId="53" pageNumber="53">Balsateres</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="B6A2343AFFC62E650B3C9A4696E3FCCC" box="[841,955,799,820]" pageId="53" pageNumber="53" rank="genus">gen. nov.</taxonomicNameLabel>
The two genera also differ in base coloration and infuscation. 
<taxonomicName id="58E52ED0FFC62E650AC89A009677FC88" authorityName="Soleglad and Fet" authorityYear="2008" box="[701,815,857,880]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC62E650AC89A009677FC88" box="[701,815,857,880]" italics="true" pageId="53" pageNumber="53">Thorellius</emphasis>
</taxonomicName>
are dark and reddish in color, with various degrees of infuscation on the carapace and tergites, whereas 
<taxonomicName id="58E52ED0FFC62E650C659ACA91DDFC52" authority="González-Santillán &amp; Prendini, 2013" authorityName="González-Santillán &amp; Prendini" authorityYear="2013" box="[1040,1157,915,938]" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="AD918941FFC62E650C659ACA91DDFC52" box="[1040,1157,915,938]" italics="true" pageId="53" pageNumber="53">Balsateres</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="B6A2343AFFC62E650CEE9ACD97ABFC3F" pageId="53" pageNumber="53" rank="genus">gen. nov.</taxonomicNameLabel>
, is yellow and almost immaculate, except for the ocular tubercle, which is infuscate. 
<taxonomicName id="58E52ED0FFC62E650B3F9AB297A5FBE7" authority="(Thorell, 1876)" baseAuthorityName="Thorell" baseAuthorityYear="1876" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="species" species="intrepidus">
<emphasis id="AD918941FFC62E650B3F9AB29110FBFA" box="[842,1096,1003,1026]" italics="true" pageId="53" pageNumber="53">Thorellius intrepidus</emphasis>
(
<bibRefCitation id="FB7428A2FFC62E650C1B9AB597ADFBE7" author="Thorell, T." pageId="53" pageNumber="53" pagination="1 - 15" refId="ref39938" refString="Thorell, T. 1876. On the classification of scorpions. Annals and Magazine of Natural History 4: 1 - 15." type="journal article" year="1876">Thorell, 1876</bibRefCitation>
)
</taxonomicName>
shares with 
<taxonomicName id="58E52ED0FFC62E650BE99D519149FBE7" authority="González-Santillán &amp; Prendini, 2013" authorityName="González-Santillán &amp; Prendini" authorityYear="2013" box="[924,1041,1032,1055]" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="AD918941FFC62E650BE99D519149FBE7" box="[924,1041,1032,1055]" italics="true" pageId="53" pageNumber="53">Balsateres</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="B6A2343AFFC62E650C509D5091CFFBE6" box="[1061,1175,1033,1054]" pageId="53" pageNumber="53" rank="genus">gen. nov.</taxonomicNameLabel>
, the presence of double basal prolateral denticles on the movable finger of the pedipalp chela (fig. 20A, B), but is readily separated from the latter by the densely granular carinae and intercarinal surfaces, which are smooth in 
<taxonomicName id="58E52ED0FFC62E650AC89DE1966AFB37" authority="González-Santillán &amp; Prendini, 2013" authorityName="González-Santillán &amp; Prendini" authorityYear="2013" box="[701,818,1208,1231]" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="AD918941FFC62E650AC89DE1966AFB37" box="[701,818,1208,1231]" italics="true" pageId="53" pageNumber="53">Balsateres</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="B6A2343AFFC62E650B369DE096E8FB36" box="[835,944,1209,1230]" pageId="53" pageNumber="53" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="D7FF06D8FFC62E670AAE9D8E972DFEDD" lastPageId="55" lastPageNumber="55" pageId="53" pageNumber="53" type="discussion">
<paragraph id="9F5A5553FFC62E650AAE9D8E910EF9E9" blockId="53.[701,1228,213,1759]" pageId="53" pageNumber="53">
Species of 
<taxonomicName id="58E52ED0FFC62E650B229D8F9691FB15" authorityName="Soleglad and Fet" authorityYear="2008" box="[855,969,1238,1261]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC62E650B229D8F9691FB15" box="[855,969,1238,1261]" italics="true" pageId="53" pageNumber="53">Thorellius</emphasis>
</taxonomicName>
superficially resemble 
<taxonomicName id="58E52ED0FFC62E650AC89DAA964FFAF2" baseAuthorityName="Soleglad and Fet" baseAuthorityYear="2008" box="[701,791,1267,1290]" class="Arachnida" family="Vaejovidae" genus="Kochius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC62E650AC89DAA964FFAF2" box="[701,791,1267,1290]" italics="true" pageId="53" pageNumber="53">Kochius</emphasis>
</taxonomicName>
, but may be distinguished from the latter by the broad, raised pedipalp carinae, each comprising clustered granules (fig. 17C, D). The pedipalp carinae of 
<taxonomicName id="58E52ED0FFC62E650C4E9C1291CDFA9A" baseAuthorityName="Soleglad and Fet" baseAuthorityYear="2008" box="[1083,1173,1355,1378]" class="Arachnida" family="Vaejovidae" genus="Kochius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC62E650C4E9C1291CDFA9A" box="[1083,1173,1355,1378]" italics="true" pageId="53" pageNumber="53">Kochius</emphasis>
</taxonomicName>
are comparatively shallower and the granulation usually moniliform (fig. 18A, B). The two genera also differ in the macrosculpture of the vsm carinae of metasomal segments I–III, which are costate to weakly denticulate in 
<taxonomicName id="58E52ED0FFC62E650AC89CA29677F9EA" authorityName="Soleglad and Fet" authorityYear="2008" box="[701,815,1531,1554]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC62E650AC89CA29677F9EA" box="[701,815,1531,1554]" italics="true" pageId="53" pageNumber="53">Thorellius</emphasis>
</taxonomicName>
but granular in 
<taxonomicName id="58E52ED0FFC62E650B819CA29116F9EA" baseAuthorityName="Soleglad and Fet" baseAuthorityYear="2008" box="[1012,1102,1531,1554]" class="Arachnida" family="Vaejovidae" genus="Kochius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC62E650B819CA29116F9EA" box="[1012,1102,1531,1554]" italics="true" pageId="53" pageNumber="53">Kochius</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="9F5A5553FFC62E650AAE9F409174F927" blockId="53.[701,1228,213,1759]" pageId="53" pageNumber="53">
<taxonomicName id="58E52ED0FFC62E650AAE9F409615F9C8" authorityName="Soleglad and Fet" authorityYear="2008" box="[731,845,1561,1584]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC62E650AAE9F409615F9C8" box="[731,845,1561,1584]" italics="true" pageId="53" pageNumber="53">Thorellius</emphasis>
</taxonomicName>
contains the largest and most robust scorpions in the subfamily; the 
<typeStatus id="405EEBF1FFC62E650CFB9F6E97B7F992" pageId="53" pageNumber="53" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="58E52ED0FFC62E650B5D9F0C9699F992" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[808,961,1619,1642]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="species" species="intrepidus">
<emphasis id="AD918941FFC62E650B5D9F0C9699F992" box="[808,961,1619,1642]" italics="true" pageId="53" pageNumber="53">T. intrepidus</emphasis>
</taxonomicName>
is 
<quantity id="581DF8B6FFC62E650B869F0D9114F992" box="[1011,1100,1620,1642]" metricMagnitude="-2" metricUnit="m" metricValue="9.4" pageId="53" pageNumber="53" unit="mm" value="94.0">94 mm</quantity>
in length (
<bibRefCitation id="FB7428A2FFC62E650AB09F289634F97F" author="Sissom, W. D." box="[709,876,1649,1671]" pageId="53" pageNumber="53" pagination="503 - 553" refId="ref39182" refString="Sissom, W. D. 2000. Family Vaejovidae. In V. Fet, W. D. Sissom, G. Lowe, and M. E. Braunwalder. Catalog of the scorpions of the world (1758 - 1998): 503 - 553. New York Entomological Society: New York." type="book chapter" year="2000">Sissom, 2000</bibRefCitation>
). 
<taxonomicName id="58E52ED0FFC62E650BFC9F2B96ADF97F" authorityName="Kraepelin" authorityYear="1900" box="[905,1013,1650,1671]" class="Arachnida" family="Vaejovidae" genus="Syntropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="53" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC62E650BFC9F2B96ADF97F" box="[905,1013,1650,1671]" italics="true" pageId="53" pageNumber="53">Syntropis</emphasis>
</taxonomicName>
, the only genus with adults similar in total length, is comparatively slender, with narrow, elongated pedipalps, legs, and metasoma.
</paragraph>
<caption id="CB9A05DBFFC52E6608019F3E9468F932" ID-DOI="http://doi.org/10.5281/zenodo.4611949" ID-Zenodo-Dep="4611949" httpUri="https://zenodo.org/record/4611949/files/figure.png" pageId="54" pageNumber="54" startId="54.[116,155,1639,1658]" targetBox="[232,1046,211,1594]" targetPageId="54">
<paragraph id="9F5A5553FFC52E6608019F3E9468F932" blockId="54.[93,1194,1639,1739]" pageId="54" pageNumber="54">
Fig. 27. 
<taxonomicName id="58E52ED0FFC52E6608A99F3E9751F982" authority="Kraepelin, 1905" authorityName="KRAEPELIN" authorityYear="1905" box="[220,521,1639,1658]" class="Arachnida" family="Vaejovidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="64" phylum="Arthropoda" rank="subFamily" subFamily="Syntropinae">
Syntropinae 
<bibRefCitation id="FB7428A2FFC52E6609149F3E9751F982" author="Kraepelin, K." box="[353,521,1639,1658]" pageId="54" pageNumber="54" pagination="321 - 364" refId="ref37966" refString="Kraepelin, K. 1905. Die geographische Verbreitung der Skorpione. Zoologische Jahrbucher, Abtheilung fur Systematik 22: 321 - 364." type="journal article" year="1905">Kraepelin, 1905</bibRefCitation>
</taxonomicName>
, metasomal segments I–V and telson, ventral aspect, illustrating carinae. 
<emphasis id="AD918941FFC52E6608C39FDB9596F96D" bold="true" box="[182,206,1666,1685]" pageId="54" pageNumber="54">A.</emphasis>
<taxonomicName id="58E52ED0FFC52E6608A09FD89726F96D" authority="(Williams, 1968)" authorityName="González-Santillán &amp; Prendini" authorityYear="2013" baseAuthorityName="Williams" baseAuthorityYear="1968" box="[213,638,1665,1685]" class="Arachnida" family="Vaejovidae" genus="Chihuahuanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="54" phylum="Arthropoda" rank="species" species="coahuilae" status="comb. nov.">
<emphasis id="AD918941FFC52E6608A09FD89492F96D" box="[213,458,1665,1685]" italics="true" pageId="54" pageNumber="54">Chihuahuanus coahuilae</emphasis>
(
<bibRefCitation id="FB7428A2FFC52E6609AD9FDB972FF96D" author="Williams, S. C." box="[472,631,1666,1685]" pageId="54" pageNumber="54" pagination="1 - 24" refId="ref40171" refString="Williams, S. C. 1968. Scorpions from northern Mexico: five new species of Vejovis from Coahuila, Mexico. Occasional Papers of the California Academy of Sciences 68: 1 - 24." type="journal article" year="1968">Williams, 1968</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="B6A2343AFFC52E660AFC9FDB9659F96D" box="[649,769,1666,1685]" pageId="54" pageNumber="54" rank="species">comb. nov.</taxonomicNameLabel>
, ♂ (AMNH). 
<emphasis id="AD918941FFC52E660BEC9FDB96E8F96D" bold="true" box="[921,944,1666,1685]" pageId="54" pageNumber="54">B.</emphasis>
<taxonomicName id="58E52ED0FFC52E660BC29FDB95A0F948" authority="(Karsch, 1879)" authorityName="González-Santillán &amp; Prendini" authorityYear="2013" baseAuthorityName="Karsch" baseAuthorityYear="1879" class="Arachnida" family="Vaejovidae" genus="Mesomexovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="54" phylum="Arthropoda" rank="species" species="punctatus" status="comb. nov.">
<emphasis id="AD918941FFC52E660BC29FDB91F1F96C" box="[951,1193,1666,1684]" italics="true" pageId="54" pageNumber="54">Mesomexovis punctatus</emphasis>
(Karsch, 1879)
</taxonomicName>
, 
<taxonomicNameLabel id="B6A2343AFFC52E6609769FC59422F948" box="[259,378,1692,1712]" pageId="54" pageNumber="54" rank="species">comb. nov.</taxonomicNameLabel>
, ♂ (AMNH). 
<emphasis id="AD918941FFC52E660A659FC59770F957" bold="true" box="[528,552,1692,1711]" pageId="54" pageNumber="54">C.</emphasis>
<taxonomicName id="58E52ED0FFC52E660A5B9FC496C0F948" authority="(Williams, 1970)" authorityName="González-Santillán &amp; Prendini" authorityYear="2013" baseAuthorityName="Williams" baseAuthorityYear="1970" box="[558,920,1692,1712]" class="Arachnida" family="Vaejovidae" genus="Paravaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="54" phylum="Arthropoda" rank="species" species="diazi" status="comb. nov.">
<emphasis id="AD918941FFC52E660A5B9FC497BDF948" box="[558,741,1692,1712]" italics="true" pageId="54" pageNumber="54">Paravaejovis diazi</emphasis>
(Williams, 1970)
</taxonomicName>
, 
<taxonomicNameLabel id="B6A2343AFFC52E660BD79FC59141F948" box="[930,1049,1692,1712]" pageId="54" pageNumber="54" rank="species">comb. nov.</taxonomicNameLabel>
, ♂ (AMNH). Scale bars 
<emphasis id="AD918941FFC52E6608BA9FEF95BBF933" box="[207,227,1718,1739]" italics="true" pageId="54" pageNumber="54">=</emphasis>
2 mm.
</paragraph>
</caption>
<paragraph id="9F5A5553FFC42E6708E9998A972DFEDD" blockId="55.[128,654,211,1759]" pageId="55" pageNumber="55">
INCLUDED SPECIES: 
<taxonomicName id="58E52ED0FFC42E6709FC998D944DFEF0" authority="(Hoffmann, 1931)" baseAuthorityName="Hoffmann" baseAuthorityYear="1931" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="species" species="atrox">
<emphasis id="AD918941FFC42E6709FC998D9766FF12" box="[393,574,212,235]" italics="true" pageId="55" pageNumber="55">Thorellius atrox</emphasis>
(
<bibRefCitation id="FB7428A2FFC42E670A3E998C9454FEF0" author="Hoffmann, C. C." pageId="55" pageNumber="55" pagination="291 - 408" refId="ref37778" refString="Hoffmann, C. C. 1931. Monografias para la entomologia medica de Mexico. Monografia Num. 2, Los escorpiones de Mexico. Primera parte: Diplocentridae, Chactidae, Vaejovidae. Anales del Instituto de Biologia Universidad Nacional Autonoma de Mexico 2: 291 - 408." type="journal article" year="1931">Hoffmann, 1931</bibRefCitation>
)
</taxonomicName>
; 
<taxonomicName id="58E52ED0FFC42E67095799A8959AFEDD" authority="(Pocock, 1898)" baseAuthorityName="Pocock" baseAuthorityYear="1898" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="species" species="cristimanus">
<emphasis id="AD918941FFC42E67095799A89747FEF0" box="[290,543,241,264]" italics="true" pageId="55" pageNumber="55">Thorellius cristimanus</emphasis>
(Pocock, 1898)
</taxonomicName>
; 
<taxonomicName id="58E52ED0FFC42E6708BA98569729FEDD" authority="(Thorell, 1876)" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[207,625,271,294]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="species" species="intrepidus">
<emphasis id="AD918941FFC42E6708BA985694E0FEDE" box="[207,440,271,294]" italics="true" pageId="55" pageNumber="55">Thorellius intrepidus</emphasis>
(
<bibRefCitation id="FB7428A2FFC42E6709BD98569731FEDD" author="Thorell, T." box="[456,617,271,293]" pageId="55" pageNumber="55" pagination="1 - 15" refId="ref39938" refString="Thorell, T. 1876. On the classification of scorpions. Annals and Magazine of Natural History 4: 1 - 15." type="journal article" year="1876">Thorell, 1876</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="D7FF06D8FFC42E6708E998759476FDB2" pageId="55" pageNumber="55" type="distribution">
<paragraph id="9F5A5553FFC42E6708E998759476FDB2" blockId="55.[128,654,211,1759]" pageId="55" pageNumber="55">
DISTRIBUTION: 
<taxonomicName id="58E52ED0FFC42E67092A98749489FEBC" authorityName="Soleglad and Fet" authorityYear="2008" box="[351,465,301,324]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC42E67092A98749489FEBC" box="[351,465,301,324]" italics="true" pageId="55" pageNumber="55">Thorellius</emphasis>
</taxonomicName>
is endemic to 
<collectingCountry id="E7F215C3FFC42E6708F598129582FE99" box="[128,218,331,353]" name="Mexico" pageId="55" pageNumber="55">Mexico</collectingCountry>
, and recorded from nine states on the mainland: 
<collectingRegion id="5D219BB1FFC42E67088E983194F3FE86" box="[251,427,360,382]" country="Mexico" name="Aguascalientes" pageId="55" pageNumber="55">Aguascalientes</collectingRegion>
, 
<collectingRegion id="5D219BB1FFC42E6709C398319757FE86" box="[438,527,360,382]" country="Mexico" name="Colima" pageId="55" pageNumber="55">Colima</collectingRegion>
, Estado de 
<collectingCountry id="E7F215C3FFC42E6708F598DC9582FE63" box="[128,218,389,411]" name="Mexico" pageId="55" pageNumber="55">México</collectingCountry>
, 
<collectingRegion id="5D219BB1FFC42E67089D98DC942FFE63" box="[232,375,389,411]" country="Mexico" name="Guanajuato" pageId="55" pageNumber="55">Guanajuato</collectingRegion>
, 
<collectingRegion id="5D219BB1FFC42E6709F098DC94ACFE62" box="[389,500,389,410]" country="Mexico" name="Guerrero" pageId="55" pageNumber="55">Guerrero</collectingRegion>
, 
<collectingRegion id="5D219BB1FFC42E670A7698DC970CFE63" box="[515,596,389,411]" country="Mexico" name="Jalisco" pageId="55" pageNumber="55">Jalisco</collectingRegion>
, 
<collectingRegion id="5D219BB1FFC42E670A1798DC95BBFE40" country="Mexico" name="Michoacan de Ocampo" pageId="55" pageNumber="55">Michoacán</collectingRegion>
, 
<collectingRegion id="5D219BB1FFC42E67088598FB9415FE40" box="[240,333,418,440]" country="Mexico" name="Nayarit" pageId="55" pageNumber="55">Nayarit</collectingRegion>
, and 
<collectingRegion id="5D219BB1FFC42E6709FA98FB94BEFE40" box="[399,486,418,440]" country="Mexico" name="Sinaloa" pageId="55" pageNumber="55">Sinaloa</collectingRegion>
(fig. 5). 
<bibRefCitation id="FB7428A2FFC42E670A3898FB947FFE2E" author="Hoffmann, C. C." pageId="55" pageNumber="55" pagination="291 - 408" refId="ref37778" refString="Hoffmann, C. C. 1931. Monografias para la entomologia medica de Mexico. Monografia Num. 2, Los escorpiones de Mexico. Primera parte: Diplocentridae, Chactidae, Vaejovidae. Anales del Instituto de Biologia Universidad Nacional Autonoma de Mexico 2: 291 - 408." type="journal article" year="1931">Hoffmann’s (1931)</bibRefCitation>
records of 
<taxonomicName id="58E52ED0FFC42E6709C398999711FE2E" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[438,585,447,470]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="species" species="intrepidus">
<emphasis id="AD918941FFC42E6709C398999711FE2E" box="[438,585,447,470]" italics="true" pageId="55" pageNumber="55">T. intrepidus</emphasis>
</taxonomicName>
from 
<collectingRegion id="5D219BB1FFC42E6708F5988495B1FE0A" box="[128,233,477,498]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="55" pageNumber="55">Veracruz</collectingRegion>
(Catemaco) on the eastern coast of 
<collectingCountry id="E7F215C3FFC42E6708F598A3958FFDE8" box="[128,215,506,528]" name="Mexico" pageId="55" pageNumber="55">Mexico</collectingCountry>
are probably erroneous; extensive fieldwork has not yielded new collections from this area.
</paragraph>
</subSubSection>
<subSubSection id="D7FF06D8FFC42E6708E99B08973BFBFB" pageId="55" pageNumber="55" type="biology_ecology">
<paragraph id="9F5A5553FFC42E6708E99B08973BFBFB" blockId="55.[128,654,211,1759]" pageId="55" pageNumber="55">
NATURAL HISTORY: There are relatively few data on the ecology of 
<taxonomicName id="58E52ED0FFC42E6709CC9B299773FD7F" authorityName="Soleglad and Fet" authorityYear="2008" box="[441,555,624,647]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC42E6709CC9B299773FD7F" box="[441,555,624,647]" italics="true" pageId="55" pageNumber="55">Thorellius</emphasis>
</taxonomicName>
. Species of the genus have mostly been collected on the surface at night with UV light detection or by turning stones during the day in subtropical deciduous forest from sea level to 
<quantity id="581DF8B6FFC42E6708D39A5B9459FCE0" box="[166,257,770,792]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="55" pageNumber="55" unit="m" value="1760.0">1760 m</quantity>
altitude. As with 
<taxonomicName id="58E52ED0FFC42E6709999A5A97DEFCE0" authority="González-Santillán &amp; Prendini, 2013" authorityName="González-Santillán &amp; Prendini" authorityYear="2013" box="[492,646,771,792]" class="Arachnida" family="Vaejovidae" genus="Mesomexovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="AD918941FFC42E6709999A5A97DEFCE0" box="[492,646,771,792]" italics="true" pageId="55" pageNumber="55">Mesomexovis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="B6A2343AFFC42E6708F59A7995A9FCCD" box="[128,241,800,821]" pageId="55" pageNumber="55" rank="genus">gen. nov.</taxonomicNameLabel>
, 
<taxonomicName id="58E52ED0FFC42E6709779A46942CFCCE" authorityName="Soleglad and Fet" authorityYear="2008" box="[258,372,799,822]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC42E6709779A46942CFCCE" box="[258,372,799,822]" italics="true" pageId="55" pageNumber="55">Thorellius</emphasis>
</taxonomicName>
species are only active during the rainy season, and estivate during the dry season. Although 
<bibRefCitation id="FB7428A2FFC42E6709C59A039591FC76" author="Gonzalez-Santillan, E." pageId="55" pageNumber="55" pagination="25 - 35" refId="ref37379" refString="Gonzalez-Santillan, E. 2004. Diversidad, taxonomia y habitat de alacranes. In A. N. Garcia, and R. Ayala (editors), Artropodos de Chamela: 25 - 35. Instituto de Biologia, UNAM, Mexico." type="book chapter" year="2004">González-Santillán (2004)</bibRefCitation>
suggested that 
<taxonomicName id="58E52ED0FFC42E6709FA9A21977EFC76" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[399,550,887,910]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="species" species="intrepidus">
<emphasis id="AD918941FFC42E6709FA9A21977EFC76" box="[399,550,887,910]" italics="true" pageId="55" pageNumber="55">T. intrepidus</emphasis>
</taxonomicName>
is pelophilous, the absence of fossorial adaptations suggests otherwise. The habitat and habitus of 
<taxonomicName id="58E52ED0FFC42E6708D19A96944EFC1E" authorityName="Soleglad and Fet" authorityYear="2008" box="[164,278,975,998]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC42E6708D19A96944EFC1E" box="[164,278,975,998]" italics="true" pageId="55" pageNumber="55">Thorellius</emphasis>
</taxonomicName>
are consistent with the lapidicolous ecomorphotype (
<bibRefCitation id="FB7428A2FFC42E6709EF9AB4970FFBFB" author="Prendini, L." box="[410,599,1005,1027]" pageId="55" pageNumber="55" pagination="113 - 139" refId="ref38594" refString="Prendini, L. 2001 a. Substratum specialization and speciation in southern African scorpions: the Effect Hypothesis revisited. In V. Fet, and P. A. Selden (editors), Scorpions 2001. In memoriam Gary A. Polis: 113 - 139. British Arachnological Society: Burnham Beeches, Buckinghamshire, UK." type="book chapter" year="2001">Prendini, 2001a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D7FF06D8FFC42E6708E99D50977CF927" pageId="55" pageNumber="55" type="discussion">
<paragraph id="9F5A5553FFC42E6708E99D50977CF927" blockId="55.[128,654,211,1759]" pageId="55" pageNumber="55">
REMARKS: As redefined here, 
<taxonomicName id="58E52ED0FFC42E670A699D5397D6FBD9" authorityName="Soleglad and Fet" authorityYear="2008" box="[540,654,1034,1057]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC42E670A699D5397D6FBD9" box="[540,654,1034,1057]" italics="true" pageId="55" pageNumber="55">Thorellius</emphasis>
</taxonomicName>
accommodates a subset of species previously assigned to 
<bibRefCitation id="FB7428A2FFC42E67095F9D1C974FFBA4" author="Hoffmann, C. C." box="[298,535,1093,1116]" pageId="55" pageNumber="55" pagination="291 - 408" refId="ref37778" refString="Hoffmann, C. C. 1931. Monografias para la entomologia medica de Mexico. Monografia Num. 2, Los escorpiones de Mexico. Primera parte: Diplocentridae, Chactidae, Vaejovidae. Anales del Instituto de Biologia Universidad Nacional Autonoma de Mexico 2: 291 - 408." type="journal article" year="1931">Hoffmann’s (1931)</bibRefCitation>
‘‘second section’’ of 
<taxonomicName id="58E52ED0FFC42E67097D9D3A9430FB80" authorityName="C. L. Koch" authorityYear="1836" box="[264,360,1123,1144]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC42E67097D9D3A9430FB80" box="[264,360,1123,1144]" italics="true" pageId="55" pageNumber="55">Vaejovis</emphasis>
</taxonomicName>
, later termed the 
<taxonomicName id="58E52ED0FFC42E670A4E9D3A95FEFB6E" baseAuthorityName="Thorell" baseAuthorityYear="1876" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="species" species="intrepidus">
<emphasis id="AD918941FFC42E670A4E9D3A95FEFB6E" italics="true" pageId="55" pageNumber="55">intrepidus</emphasis>
</taxonomicName>
group by 
<bibRefCitation id="FB7428A2FFC42E6709339DD994AFFB6E" author="Sissom, W. D." box="[326,503,1152,1174]" pageId="55" pageNumber="55" pagination="179 - 187" refId="ref39026" refString="Sissom, W. D. 1989. Redescription of Vaejovis occidentalis Hoffmann with a revised diagnosis for Vaejovis subcristatus Pocock (Scorpiones: Vaejovidae). Revue Arachnologique 8: 179 - 187." type="journal article" year="1989">Sissom (1989)</bibRefCitation>
, for which 
<bibRefCitation id="FB7428A2FFC42E6708F59DC494F2FB4B" author="Soleglad, M. E. &amp; V. Fet" box="[128,426,1181,1203]" pageId="55" pageNumber="55" pagination="1 - 115" refId="ref39584" refString="Soleglad, M. E., and V. Fet. 2008. Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae). Euscorpius 71: 1 - 115." type="journal article" year="2008">Soleglad and Fet (2008)</bibRefCitation>
devised the name 
<taxonomicName id="58E52ED0FFC42E6708F59DE395AAFB29" authorityName="Soleglad and Fet" authorityYear="2008" box="[128,242,1210,1233]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC42E6708F59DE395AAFB29" box="[128,242,1210,1233]" italics="true" pageId="55" pageNumber="55">Thorellius</emphasis>
</taxonomicName>
, without quantitatively testing its monophyly or composition. 
<taxonomicName id="58E52ED0FFC42E67099F9D8E9704FB16" authorityName="Soleglad and Fet" authorityYear="2008" box="[490,604,1239,1262]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC42E67099F9D8E9704FB16" box="[490,604,1239,1262]" italics="true" pageId="55" pageNumber="55">Thorellius</emphasis>
</taxonomicName>
, as defined by 
<bibRefCitation id="FB7428A2FFC42E6709609DAC9714FAF3" author="Soleglad, M. E. &amp; V. Fet" box="[277,588,1269,1291]" pageId="55" pageNumber="55" pagination="1 - 115" refId="ref39584" refString="Soleglad, M. E., and V. Fet. 2008. Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae). Euscorpius 71: 1 - 115." type="journal article" year="2008">Soleglad and Fet (2008)</bibRefCitation>
, was consistently polyphyletic, and the group of species hereby assigned to it consistently monophyletic, in the phylogenetic analyses of González-Santillán and Prendini (in press) based on morphology and those based on morphology and DNA. The following species are therefore transferred to other genera in the present contribution: 
<taxonomicName id="58E52ED0FFC42E6709DD9C8797DEFA0D" authorityName="González-Santillán &amp; Prendini" authorityYear="2013" baseAuthorityName="Ponce-Saavedra and Sissom" baseAuthorityYear="2004" box="[424,646,1502,1525]" class="Arachnida" family="Vaejovidae" genus="Balsateres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="species" species="cisnerosi" status="comb. nov.">
<emphasis id="AD918941FFC42E6709DD9C8797DEFA0D" box="[424,646,1502,1525]" italics="true" pageId="55" pageNumber="55">Balsateres cisnerosi</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="B6A2343AFFC42E6708F59CA5945DF9EA" box="[128,261,1532,1554]" pageId="55" pageNumber="55" rank="species">comb. nov.</taxonomicNameLabel>
, previously assigned to 
<taxonomicName id="58E52ED0FFC42E670A699CA594F3F9C8" authority="(Soleglad and Fet, 2008)" baseAuthorityName="Soleglad and Fet" baseAuthorityYear="2008" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC42E670A699CA597D6F9EB" box="[540,654,1532,1555]" italics="true" pageId="55" pageNumber="55">Thorellius</emphasis>
(
<bibRefCitation id="FB7428A2FFC42E6708F29F4394FAF9C8" author="Soleglad, M. E. &amp; V. Fet" box="[135,418,1562,1584]" pageId="55" pageNumber="55" pagination="1 - 115" refId="ref39584" refString="Soleglad, M. E., and V. Fet. 2008. Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae). Euscorpius 71: 1 - 115." type="journal article" year="2008">Soleglad and Fet, 2008</bibRefCitation>
)
</taxonomicName>
and the 
<taxonomicName id="58E52ED0FFC42E670A6B9F4097D5F9C8" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[542,653,1561,1584]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="species" species="intrepidus">
<emphasis id="AD918941FFC42E670A6B9F4097D5F9C8" box="[542,653,1561,1584]" italics="true" pageId="55" pageNumber="55">intrepidus</emphasis>
</taxonomicName>
group (
<bibRefCitation id="FB7428A2FFC42E6708A39F6E97DAF9B5" author="Santibanez-Lopez, C. &amp; W. D. Sissom" box="[214,642,1591,1613]" pageId="55" pageNumber="55" pagination="49 - 58" refId="ref38931" refString="Santibanez-Lopez, C., and W. D. Sissom. 2010. A new species of the Vaejovis eusthenura group in Oaxaca, Mexico (Scorpiones: Vaejovidae). Zootaxa 2493: 49 - 58." type="journal article" year="2010">Santibáñez-López and Sissom, 2010</bibRefCitation>
); 
<taxonomicName id="58E52ED0FFC42E6708F59F0C94F5F992" authorityName="González-Santillán &amp; Prendini" authorityYear="2013" baseAuthorityName="Hoffmann" baseAuthorityYear="1931" box="[128,429,1619,1642]" class="Arachnida" family="Vaejovidae" genus="Mesomexovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="species" species="occidentalis" status="comb. nov.">
<emphasis id="AD918941FFC42E6708F59F0C94F5F992" box="[128,429,1619,1642]" italics="true" pageId="55" pageNumber="55">Mesomexovis occidentalis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="B6A2343AFFC42E6709B79F0D9717F992" box="[450,591,1620,1642]" pageId="55" pageNumber="55" rank="species">comb. nov.</taxonomicNameLabel>
, and 
<taxonomicName id="58E52ED0FFC42E6708F59F2B94F0F970" authorityName="González-Santillán &amp; Prendini" authorityYear="2013" baseAuthorityName="Pocock" baseAuthorityYear="1898" box="[128,424,1649,1672]" class="Arachnida" family="Vaejovidae" genus="Mesomexovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="species" species="subcristatus" status="comb. nov.">
<emphasis id="AD918941FFC42E6708F59F2B94F0F970" box="[128,424,1649,1672]" italics="true" pageId="55" pageNumber="55">Mesomexovis subcristatus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="B6A2343AFFC42E6709C29F289765F97F" box="[439,573,1649,1671]" pageId="55" pageNumber="55" rank="species">comb. nov.</taxonomicNameLabel>
, previously assigned to the 
<taxonomicName id="58E52ED0FFC42E67090B9FD794B5F95D" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[382,493,1678,1701]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="species" species="intrepidus">
<emphasis id="AD918941FFC42E67090B9FD794B5F95D" box="[382,493,1678,1701]" italics="true" pageId="55" pageNumber="55">intrepidus</emphasis>
</taxonomicName>
group (
<bibRefCitation id="FB7428A2FFC42E670A3E9FD6944EF939" author="Hoffmann, C. C." pageId="55" pageNumber="55" pagination="291 - 408" refId="ref37778" refString="Hoffmann, C. C. 1931. Monografias para la entomologia medica de Mexico. Monografia Num. 2, Los escorpiones de Mexico. Primera parte: Diplocentridae, Chactidae, Vaejovidae. Anales del Instituto de Biologia Universidad Nacional Autonoma de Mexico 2: 291 - 408." type="journal article" year="1931">Hoffmann, 1931</bibRefCitation>
; 
<bibRefCitation id="FB7428A2FFC42E67095A9FF5948BF93A" author="Sissom, W. D." box="[303,467,1708,1730]" pageId="55" pageNumber="55" pagination="503 - 553" refId="ref39182" refString="Sissom, W. D. 2000. Family Vaejovidae. In V. Fet, W. D. Sissom, G. Lowe, and M. E. Braunwalder. Catalog of the scorpions of the world (1758 - 1998): 503 - 553. New York Entomological Society: New York." type="book chapter" year="2000">Sissom, 2000</bibRefCitation>
) and then to 
<taxonomicName id="58E52ED0FFC42E6708F59F919778F927" authority="(Soleglad and Fet, 2008)" baseAuthorityName="Soleglad and Fet" baseAuthorityYear="2008" box="[128,544,1736,1759]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="AD918941FFC42E6708F59F9195AAF927" box="[128,242,1736,1759]" italics="true" pageId="55" pageNumber="55">Thorellius</emphasis>
(
<bibRefCitation id="FB7428A2FFC42E6709779F909740F927" author="Soleglad, M. E. &amp; V. Fet" box="[258,536,1737,1759]" pageId="55" pageNumber="55" pagination="1 - 115" refId="ref39584" refString="Soleglad, M. E., and V. Fet. 2008. Contributions to scorpion systematics. III. Subfamilies Smeringurinae and Syntropinae (Scorpiones: Vaejovidae). Euscorpius 71: 1 - 115." type="journal article" year="2008">Soleglad and Fet, 2008</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="D7FF06D8FFC42E670AAF998A96D2FD7E" pageId="55" pageNumber="55" type="materials_examined">
<paragraph id="9F5A5553FFC42E670AAF998A96D2FD7E" blockId="55.[701,1228,211,647]" pageId="55" pageNumber="55">
MATERIAL EXAMINED: 
<materialsCitation id="2F8D5F0EFFC42E670C7C998D962CFE98" ID-GBIF-Occurrence="3757437374" pageId="55" pageNumber="55" typeStatus="holotype">
<taxonomicName id="58E52ED0FFC42E670C7C998D96C0FEF0" authority="(Hoffmann, 1931)" baseAuthorityName="Hoffmann" baseAuthorityYear="1931" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="species" species="atrox">
<emphasis id="AD918941FFC42E670C7C998D9193FF12" box="[1033,1227,212,235]" italics="true" pageId="55" pageNumber="55">Thorellius atrox</emphasis>
(
<bibRefCitation id="FB7428A2FFC42E670AB099AB96C8FEF0" author="Hoffmann, C. C." box="[709,912,242,264]" pageId="55" pageNumber="55" pagination="291 - 408" refId="ref37778" refString="Hoffmann, C. C. 1931. Monografias para la entomologia medica de Mexico. Monografia Num. 2, Los escorpiones de Mexico. Primera parte: Diplocentridae, Chactidae, Vaejovidae. Anales del Instituto de Biologia Universidad Nacional Autonoma de Mexico 2: 291 - 408." type="journal article" year="1931">Hoffmann, 1931</bibRefCitation>
)
</taxonomicName>
: 
<emphasis id="AD918941FFC42E670BDC99AB91D5FEF0" bold="true" box="[937,1165,242,264]" pageId="55" pageNumber="55">
<collectingCountry id="E7F215C3FFC42E670BDC99AB9179FEF0" box="[937,1057,242,264]" name="Mexico" pageId="55" pageNumber="55">MEXICO</collectingCountry>
: 
<collectingRegion id="5D219BB1FFC42E670C4699AB91D0FEF0" box="[1075,1160,242,264]" country="Mexico" name="Colima" pageId="55" pageNumber="55">Colima</collectingRegion>
:
</emphasis>
<emphasis id="AD918941FFC42E670CEF99AA96DBFEDE" italics="true" pageId="55" pageNumber="55">
Municipio de 
<collectingRegion id="5D219BB1FFC42E670B45985696DBFEDE" box="[816,899,271,294]" country="Mexico" name="Colima" pageId="55" pageNumber="55">Colima</collectingRegion>
</emphasis>
: 
<collectingRegion id="5D219BB1FFC42E670BE4985696B2FEDD" box="[913,1002,271,293]" country="Mexico" name="Colima" pageId="55" pageNumber="55">Colima</collectingRegion>
, in house, 
<typeStatus id="405EEBF1FFC42E670C1098569193FEDD" box="[1125,1227,271,293]" pageId="55" pageNumber="55" type="holotype">holotype</typeStatus>
♂ of 
<taxonomicName id="58E52ED0FFC42E670B78987497ACFEA7" authority="Hoffmann, 1931" authorityName="Hoffmann" authorityYear="1931" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="subSpecies" species="intrepidus" subSpecies="atrox">
<emphasis id="AD918941FFC42E670B7898749162FEBB" box="[781,1082,300,323]" italics="true" pageId="55" pageNumber="55">Vaejovis intrepidus atrox</emphasis>
<bibRefCitation id="FB7428A2FFC42E670C3F987497ACFEA7" author="Hoffmann, C. C." pageId="55" pageNumber="55" pagination="291 - 408" refId="ref37778" refString="Hoffmann, C. C. 1931. Monografias para la entomologia medica de Mexico. Monografia Num. 2, Los escorpiones de Mexico. Primera parte: Diplocentridae, Chactidae, Vaejovidae. Anales del Instituto de Biologia Universidad Nacional Autonoma de Mexico 2: 291 - 408." type="journal article" year="1931">Hoffmann, 1931</bibRefCitation>
</taxonomicName>
(
<collectionCode id="F9F4CD96FFC42E670B709813963FFE98" box="[773,871,330,352]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="55" pageNumber="55">AMNH</collectionCode>
),
</materialsCitation>
<materialsCitation id="2F8D5F0EFFC42E670B0998139194FE98" ID-GBIF-Occurrence="3757437380" box="[892,1228,327,353]" pageId="55" pageNumber="55" typeStatus="paratype">
18, 
<specimenCount id="89E39EDAFFC42E670BC49813968BFE99" box="[945,979,327,353]" pageId="55" pageNumber="55" type="female">1♀</specimenCount>
<typeStatus id="405EEBF1FFC42E670BAE98129114FE98" box="[987,1100,331,352]" pageId="55" pageNumber="55" type="paratype">paratypes</typeStatus>
(
<collectionCode id="F9F4CD96FFC42E670C28981391E7FE98" box="[1117,1215,330,352]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="55" pageNumber="55">AMNH</collectionCode>
).
</materialsCitation>
<materialsCitation id="2F8D5F0EFFC42E670AC8983E9146FE2E" ID-GBIF-Occurrence="3757437305" pageId="55" pageNumber="55">
<taxonomicName id="58E52ED0FFC42E670AC8983E912DFE86" authority="(Pocock, 1898)" baseAuthorityName="Pocock" baseAuthorityYear="1898" box="[701,1141,359,382]" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="species" species="cristimanus">
<emphasis id="AD918941FFC42E670AC8983E96E1FE85" box="[701,953,359,382]" italics="true" pageId="55" pageNumber="55">Thorellius cristimanus</emphasis>
(Pocock, 1898)
</taxonomicName>
: 
<emphasis id="AD918941FFC42E670CF49831963AFE63" bold="true" pageId="55" pageNumber="55">
MEXICO: 
<collectingRegion id="5D219BB1FFC42E670B7298DC9604FE63" box="[775,860,389,411]" country="Mexico" name="Colima" pageId="55" pageNumber="55">Colima</collectingRegion>
:
</emphasis>
<emphasis id="AD918941FFC42E670B1A98DF913AFE63" box="[879,1122,388,411]" italics="true" pageId="55" pageNumber="55">Municipio de Tonila</emphasis>
: Tonila, 3.2 km S, 
<date id="EB5B7393FFC42E670B4998FB969BFE40" box="[828,963,418,440]" pageId="55" pageNumber="55" value="1965-08-28">28.viii.1965</date>
, W.J. Gertsch and 
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. Hastings, 18, 
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(
<collectionCode id="F9F4CD96FFC42E670BDA98999149FE2E" box="[943,1041,448,470]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="55" pageNumber="55">AMNH</collectionCode>
).
</materialsCitation>
<materialsCitation id="2F8D5F0EFFC42E670C5998E6917DFDB4" ID-GBIF-Occurrence="3757437429" collectionCode="AMNH" country="Mexico" elevation="275" latitude="19.028" location="Mine La Salada" longLatPrecision="1" longitude="-103.783936" municipality="Municipio de Armeria" pageId="55" pageNumber="55" specimenCount="1" specimenCount-female="1" stateProvince="Colima">
<taxonomicName id="58E52ED0FFC42E670C5998E69681FE0B" authority="(Thorell, 1876)" baseAuthorityName="Thorell" baseAuthorityYear="1876" class="Arachnida" family="Vaejovidae" genus="Thorellius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="55" phylum="Arthropoda" rank="species" species="intrepidus">
<emphasis id="AD918941FFC42E670C5998E6964FFE0C" italics="true" pageId="55" pageNumber="55">Thorellius intrepidus</emphasis>
(
<bibRefCitation id="FB7428A2FFC42E670B5998849688FE0B" author="Thorell, T." box="[812,976,477,499]" pageId="55" pageNumber="55" pagination="1 - 15" refId="ref39938" refString="Thorell, T. 1876. On the classification of scorpions. Annals and Magazine of Natural History 4: 1 - 15." type="journal article" year="1876">Thorell, 1876</bibRefCitation>
)
</taxonomicName>
: 
<emphasis id="AD918941FFC42E670B9D98849194FE0B" bold="true" box="[1000,1228,477,499]" pageId="55" pageNumber="55">
<collectingCountry id="E7F215C3FFC42E670B9D98849138FE0B" box="[1000,1120,477,499]" name="Mexico" pageId="55" pageNumber="55">MEXICO</collectingCountry>
: 
<collectingRegion id="5D219BB1FFC42E670C079884919FFE0B" box="[1138,1223,477,499]" country="Mexico" name="Colima" pageId="55" pageNumber="55">Colima</collectingRegion>
:
</emphasis>
<emphasis id="AD918941FFC42E670AC898A296E3FDE8" box="[701,955,506,529]" italics="true" pageId="55" pageNumber="55">
<collectingMunicipality id="7F3ECF29FFC42E670AC898A296E3FDE8" box="[701,955,506,529]" pageId="55" pageNumber="55">Municipio de Armería</collectingMunicipality>
</emphasis>
: 
<location id="9A3A0388FFC42E670BB998A291D4FDE9" LSID="urn:lsid:plazi:treatment:174CE445FFC62E6708B79D4896D2FD7E:9A3A0388FFC42E670BB998A291D4FDE9" box="[972,1164,507,529]" country="Mexico" latitude="19.028" longLatPrecision="1" longitude="-103.783936" municipality="Municipio de Armeria" name="Mine La Salada" pageId="55" pageNumber="55" stateProvince="Colima">Mine La Salada</location>
, NW 
<location id="9A3A0388FFC42E670AC89B41961CFDD6" LSID="urn:lsid:plazi:treatment:174CE445FFC62E6708B79D4896D2FD7E:9A3A0388FFC42E670AC89B41961CFDD6" box="[701,836,536,558]" country="Mexico" latitude="19.028" longLatPrecision="1" longitude="-103.783936" municipality="Municipio de Armeria" name="Ixtlahuacan" pageId="55" pageNumber="55" stateProvince="Colima">Ixtlahuacan</location>
, 
<geoCoordinate id="FAD13394FFC42E670B389B419680FDD6" box="[845,984,535,558]" degrees="19" direction="north" minutes="01.680" orientation="latitude" pageId="55" pageNumber="55" precision="1" value="19.028">
19 
<emphasis id="AD918941FFC42E670B129B41962AFDD5" bold="true" box="[871,882,536,557]" pageId="55" pageNumber="55">°</emphasis>
01.680 
<emphasis id="AD918941FFC42E670BCC9B4E9699FDD6" box="[953,961,535,558]" italics="true" pageId="55" pageNumber="55">′</emphasis>
N
</geoCoordinate>
<geoCoordinate id="FAD13394FFC42E670BAE9B419122FDD6" box="[987,1146,535,558]" degrees="103" direction="west" minutes="47.036" orientation="longitude" pageId="55" pageNumber="55" precision="1" value="-103.783936">
103 
<emphasis id="AD918941FFC42E670C769B419156FDD5" bold="true" box="[1027,1038,536,557]" pageId="55" pageNumber="55">°</emphasis>
47.036 
<emphasis id="AD918941FFC42E670C209B4E9105FDD6" box="[1109,1117,535,558]" italics="true" pageId="55" pageNumber="55">′</emphasis>
W
</geoCoordinate>
, 
<quantity id="581DF8B6FFC42E670CF69B41919FFDD6" box="[1155,1223,536,558]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="55" pageNumber="55" unit="m" value="275.0">
<elevation id="14C8B260FFC42E670CF69B41919FFDD6" box="[1155,1223,536,558]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="55" pageNumber="55" unit="m" value="275.0">275 m</elevation>
</quantity>
, 
<location id="9A3A0388FFC42E670AC89B6F97A1FDB4" LSID="urn:lsid:plazi:treatment:174CE445FFC62E6708B79D4896D2FD7E:9A3A0388FFC42E670AC89B6F97A1FDB4" box="[701,761,566,588]" country="Mexico" latitude="19.028" longLatPrecision="1" longitude="-103.783936" municipality="Municipio de Armeria" name="Mine" pageId="55" pageNumber="55" stateProvince="Colima">Mine</location>
staff, 18, 
<specimenCount id="89E39EDAFFC42E670BF69B6F96FDFDB4" box="[899,933,562,588]" pageId="55" pageNumber="55" type="female">1♀</specimenCount>
(
<collectionCode id="F9F4CD96FFC42E670BCC9B6F9141FDB4" box="[953,1049,566,588]" collectionName="USA, New York, New York, American Museum of Natural History" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="55" pageNumber="55">AMNH</collectionCode>
).
</materialsCitation>
<materialsCitation id="2F8D5F0EFFC42E670C449B6F96D2FD7E" ID-GBIF-Occurrence="3757437327" collectionCode="AMNH" latitude="18.908333" location="Tecoman" longLatPrecision="21" longitude="-103.87444" municipality="Municipio de Tecoman" pageId="55" pageNumber="55" specimenCount="1" specimenCount-female="1">
<emphasis id="AD918941FFC42E670C449B6F967DFD90" italics="true" pageId="55" pageNumber="55">
<collectingMunicipality id="7F3ECF29FFC42E670C449B6F967DFD90" pageId="55" pageNumber="55">Municipio de Tecomán</collectingMunicipality>
</emphasis>
: 
<location id="9A3A0388FFC42E670B4D9B0A96F9FD91" LSID="urn:lsid:plazi:treatment:174CE445FFC62E6708B79D4896D2FD7E:9A3A0388FFC42E670B4D9B0A96F9FD91" box="[824,929,595,617]" latitude="18.908333" longLatPrecision="21" longitude="-103.87444" municipality="Municipio de Tecoman" name="Tecoman" pageId="55" pageNumber="55">Tecomán</location>
, 
<geoCoordinate id="FAD13394FFC42E670BDA9B0A9174FD90" box="[943,1068,594,617]" degrees="18" direction="north" minutes="54" orientation="latitude" pageId="55" pageNumber="55" precision="15" seconds="30" value="18.908333">
18 
<emphasis id="AD918941FFC42E670BBD9B0A968BFD90" bold="true" box="[968,979,595,616]" pageId="55" pageNumber="55">°</emphasis>
54 
<emphasis id="AD918941FFC42E670B9E9B0B96ABFD91" box="[1003,1011,594,617]" italics="true" pageId="55" pageNumber="55">′</emphasis>
30 
<emphasis id="AD918941FFC42E670C789B0B914DFD91" box="[1037,1045,594,617]" italics="true" pageId="55" pageNumber="55">&quot;</emphasis>
N
</geoCoordinate>
<geoCoordinate id="FAD13394FFC42E670C439B0A919FFD90" box="[1078,1223,594,617]" degrees="103" direction="west" minutes="52" orientation="longitude" pageId="55" pageNumber="55" precision="15" seconds="28" value="-103.87444">
103 
<emphasis id="AD918941FFC42E670C299B0A913FFD90" bold="true" box="[1116,1127,595,616]" pageId="55" pageNumber="55">°</emphasis>
52 
<emphasis id="AD918941FFC42E670CF59B0B91D0FD91" box="[1152,1160,594,617]" italics="true" pageId="55" pageNumber="55">′</emphasis>
28 
<emphasis id="AD918941FFC42E670CD79B0B91F2FD91" box="[1186,1194,594,617]" italics="true" pageId="55" pageNumber="55">&quot;</emphasis>
W
</geoCoordinate>
, 18, 
<specimenCount id="89E39EDAFFC42E670A859B299649FD7F" box="[752,785,621,647]" pageId="55" pageNumber="55" type="female">1♀</specimenCount>
(
<collectionCode id="F9F4CD96FFC42E670B559B299626FD7E" box="[800,894,624,646]" collectionName="USA, New York, New York, American Museum of Natural History" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="55" pageNumber="55">AMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>