141 lines
17 KiB
XML
141 lines
17 KiB
XML
<document id="8E5141A135EA36C240F727ABF003F672" ID-DOI="10.11646/zootaxa.4020.1.7" ID-GBIF-Dataset="82b2f539-5d25-42d6-95c5-e8b16a353e12" ID-ISSN="1175-5326" ID-Zenodo-Dep="237970" ID-ZooBank="BD8D1880-79A9-4672-99AF-5B3D1BEFAFEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461269068152" checkinUser="plazi" docAuthor="Bickel, Daniel J." docDate="2015" docId="172D87FDFFBAFF89FF3DA6BD0153F9BA" docLanguage="en" docName="zt04020p182.pdf" docOrigin="Zootaxa 4020 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systenus naranjensis Bickel, sp. nov." docType="treatment" docVersion="8" lastPageNumber="176" masterDocId="EB14FF85FFBDFF8EFFAAA70A0459FFCC" masterDocTitle="The Costa Rican Systenus Loew (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus" masterLastPageNumber="182" masterPageNumber="169" pageNumber="176" updateTime="1698611248703" updateUser="plazi">
|
||
<mods:mods id="46B379D9A18166DA6D5DF7CD206486AA" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="A291CD73E8C50342BF4C0B5927F06503">
|
||
<mods:title id="0123D7C8996538A07C2F137DEAC42F01">The Costa Rican Systenus Loew (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="81E8DE9B5D7A5923BAF35592A6544362" type="personal">
|
||
<mods:role id="9725B477017E5F9DC78867F68D870880">
|
||
<mods:roleTerm id="EC8127BD0FC755912E65258DA8A3E02F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="7B6467CFEC42A95AFA5E355E953E376B">Bickel, Daniel J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="720ED3F8387335DFEE5CF21B33ECD85E">text</mods:typeOfResource>
|
||
<mods:relatedItem id="857DB9EB8CDFB9856BB87387AA7525A0" type="host">
|
||
<mods:titleInfo id="619EA40D4275BE5A4548FA0E62AE2DB2">
|
||
<mods:title id="2E6AF9D21B92819FF6640301D7539612">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="8BDADEEA19B33D259476DA42117E357A">
|
||
<mods:date id="F9DF700EB206513B58D2945DB28BD50E">2015</mods:date>
|
||
<mods:detail id="5CC1B692E04D320FCFECFA4E478BF89E" type="volume">
|
||
<mods:number id="319BEE2B4CD7A7BC922F16856CA3BE41">4020</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="11D00273D784E477E9FEC0D0F7E6DDE0" type="issue">
|
||
<mods:number id="57AD7C215F153090F1AFBD46FC909182">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="11A270158931702C838AC068F8AEB76B" unit="page">
|
||
<mods:start id="F4B06490A4EF92FDCD09F0F8E8A3CF61">169</mods:start>
|
||
<mods:end id="AA078B3C7ED1125875F84E534D058CA5">182</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="A6F420F42266FE67132CB84668CD967B">journal article</mods:classification>
|
||
<mods:identifier id="B9863E5C1323BF680BE333A41EDDB943" type="DOI">10.11646/zootaxa.4020.1.7</mods:identifier>
|
||
<mods:identifier id="97A2E6B27B139C1F30216D20DF383A8C" type="GBIF-Dataset">82b2f539-5d25-42d6-95c5-e8b16a353e12</mods:identifier>
|
||
<mods:identifier id="095BAC7F2B614AFFFD796A29059A3A9E" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="BAD0D60710094FB1F72BCDD0E42B0E51" type="Zenodo-Dep">237970</mods:identifier>
|
||
<mods:identifier id="5CE6858BE502AAE8858F15F376864455" type="ZooBank">BD8D1880-79A9-4672-99AF-5B3D1BEFAFEE</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="172D87FDFFBAFF89FF3DA6BD0153F9BA" ID-DOI="http://doi.org/10.5281/zenodo.6121807" ID-GBIF-Taxon="119638518" ID-Zenodo-Dep="6121807" LSID="urn:lsid:plazi:treatment:172D87FDFFBAFF89FF3DA6BD0153F9BA" httpUri="http://treatment.plazi.org/id/172D87FDFFBAFF89FF3DA6BD0153F9BA" lastPageNumber="176" pageId="7" pageNumber="176">
|
||
<subSubSection id="D79E6560FFBAFF89FF3DA6BD056BFE3F" pageId="7" pageNumber="176" type="nomenclature">
|
||
<paragraph id="9F3B36EBFFBAFF89FF3DA6BD0609FE1E" blockId="7.[151,592,439,499]" box="[151,592,439,466]" pageId="7" pageNumber="176">
|
||
<heading id="C4738187FFBAFF89FF3DA6BD0609FE1E" bold="true" box="[151,592,439,466]" fontSize="11" level="1" pageId="7" pageNumber="176" reason="1">
|
||
<emphasis id="ADF0EAF9FFBAFF89FF3DA6BD0609FE1E" bold="true" box="[151,592,439,466]" pageId="7" pageNumber="176">
|
||
<taxonomicName id="58844D68FFBAFF89FF3DA6BD05B2FE1E" ID-CoL="9HSKT" authority="Bickel" authorityName="Bickel" box="[151,491,439,466]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="176" phylum="Arthropoda" rank="species" species="naranjensis" status="sp. nov.">
|
||
<emphasis id="ADF0EAF9FFBAFF89FF3DA6BD05CFFE1D" bold="true" box="[151,406,439,465]" italics="true" pageId="7" pageNumber="176">Systenus naranjensis</emphasis>
|
||
Bickel
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B6C35782FFBAFF89FE58A6B20609FE1E" box="[498,592,440,466]" pageId="7" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="9F3B36EBFFBAFF89FF3DA6D0056BFE3F" blockId="7.[151,592,439,499]" box="[151,306,474,499]" pageId="7" pageNumber="176">
|
||
(
|
||
<figureCitation id="07BF2A6EFFBAFF89FF35A6D004BFFE3F" box="[159,230,474,499]" captionStart="FIGURE 2" captionStartId="6.[151,250,1973,1995]" captionTargetBox="[167,1436,373,1951]" captionTargetId="figure@6.[151,1436,373,1952]" captionTargetPageId="6" captionText="FIGURE 2. Systenus emusorum sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. naranjensis sp. nov. (holotype): c. male habitus, left lateral; d. hypopygium, left lateral; e. male head, anterior." httpUri="https://zenodo.org/record/237972/files/figure.png" pageId="7" pageNumber="176">Figs 2</figureCitation>
|
||
c, d, e)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D79E6560FFBAFF89FF3DA528065FFD93" pageId="7" pageNumber="176" type="materials_examined">
|
||
<paragraph id="9F3B36EBFFBAFF89FF3DA528065FFD93" blockId="7.[151,1437,546,1654]" pageId="7" pageNumber="176">
|
||
<emphasis id="ADF0EAF9FFBAFF89FF3DA5280564FDF7" bold="true" box="[151,317,546,571]" pageId="7" pageNumber="176">
|
||
<typeStatus id="403F8849FFBAFF89FF3DA528048BFDF7" box="[151,210,546,571]" pageId="7" pageNumber="176">Type</typeStatus>
|
||
material
|
||
</emphasis>
|
||
.
|
||
<typeStatus id="403F8849FFBAFF89FEE1A52805B3FDF7" box="[331,490,546,571]" pageId="7" pageNumber="176" type="holotype">
|
||
<emphasis id="ADF0EAF9FFBAFF89FEE1A52805B3FDF7" bold="true" box="[331,490,546,571]" pageId="7" pageNumber="176">HOLOTYPE</emphasis>
|
||
</typeStatus>
|
||
♂,
|
||
<collectingCountry id="E793767BFFBAFF89FDB9A52906E2FDF7" box="[531,699,547,571]" name="Costa Rica" pageId="7" pageNumber="176">COSTA RICA</collectingCountry>
|
||
:
|
||
<emphasis id="ADF0EAF9FFBAFF89FD6EA5280709FDF7" bold="true" box="[708,848,546,571]" pageId="7" pageNumber="176">Guanacaste</emphasis>
|
||
:
|
||
<quantity id="587C9B0EFFBAFF89FCF4A52907CEFDF7" box="[862,919,546,571]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="176" unit="km" value="3.0">3 km</quantity>
|
||
SE Rio Naranjo,
|
||
<date id="EB3A102BFFBAFF89FBF6A529009CFDF7" box="[1116,1221,547,571]" pageId="7" pageNumber="176" value="1993-02-01">1.II.1993</date>
|
||
, F. Parker (
|
||
<collectionCode id="F995AE2EFFBAFF89FAE8A52901CEFDF7" LSID="urn:lsid:biocol.org:col:34802" box="[1346,1431,547,571]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="7" pageNumber="176">LACM</collectionCode>
|
||
, ex.
|
||
<collectionCode id="F995AE2EFFBAFF89FF15A54D0549FD93" box="[191,272,583,607]" pageId="7" pageNumber="176">EMUS</collectionCode>
|
||
;
|
||
<collectionCode id="F995AE2EFFBAFF89FEB6A54D0534FD93" LSID="urn:lsid:biocol.org:col:34802" box="[284,365,583,607]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="7" pageNumber="176">LACM</collectionCode>
|
||
Ent 329999).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D79E6560FFBAFF89FF6DA5600016FA2A" pageId="7" pageNumber="176" type="description">
|
||
<paragraph id="9F3B36EBFFBAFF89FF6DA5600016FA2A" blockId="7.[151,1437,546,1654]" pageId="7" pageNumber="176">
|
||
<emphasis id="ADF0EAF9FFBAFF89FF6DA5600508FD4F" bold="true" box="[199,337,618,643]" pageId="7" pageNumber="176">Description</emphasis>
|
||
.
|
||
<emphasis id="ADF0EAF9FFBAFF89FECAA56005C5FD4F" bold="true" box="[352,412,618,643]" pageId="7" pageNumber="176">Male</emphasis>
|
||
: body length:
|
||
<quantity id="587C9B0EFFBAFF89FDE8A56006C2FD4E" box="[578,667,618,643]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="176" unit="mm" value="3.5">3.5 mm</quantity>
|
||
; wing 3.4 x
|
||
<quantity id="587C9B0EFFBAFF89FC82A56107D8FD4E" box="[808,897,619,643]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="176" unit="mm" value="1.2">1.2 mm</quantity>
|
||
. (
|
||
<figureCitation id="07BF2A6EFFBAFF89FC3FA5600783FD4F" box="[917,986,618,643]" captionStart="FIGURE 2" captionStartId="6.[151,250,1973,1995]" captionTargetBox="[167,1436,373,1951]" captionTargetId="figure@6.[151,1436,373,1952]" captionTargetPageId="6" captionText="FIGURE 2. Systenus emusorum sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. naranjensis sp. nov. (holotype): c. male habitus, left lateral; d. hypopygium, left lateral; e. male head, anterior." httpUri="https://zenodo.org/record/237972/files/figure.png" pageId="7" pageNumber="176">Fig. 2</figureCitation>
|
||
c); similar to
|
||
<taxonomicName id="58844D68FFBAFF89FBDBA5610087FD4F" box="[1137,1246,619,643]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="176" phylum="Arthropoda" rank="species" species="tenorio">
|
||
<emphasis id="ADF0EAF9FFBAFF89FBDBA5610087FD4F" box="[1137,1246,619,643]" italics="true" pageId="7" pageNumber="176">S. tenorio</emphasis>
|
||
</taxonomicName>
|
||
except as noted:
|
||
<emphasis id="ADF0EAF9FFBAFF89FF3DA584048DFD6B" box="[151,212,654,679]" italics="true" pageId="7" pageNumber="176">Head</emphasis>
|
||
: frons, face, and clypeus dark metallic green with thick grey pruinosity; palpus brown with short black setae and strong apical seta; proboscis brown; antenna (
|
||
<figureCitation id="07BF2A6EFFBAFF89FD70A5B8077BFD07" box="[730,802,690,715]" captionStart="FIGURE 2" captionStartId="6.[151,250,1973,1995]" captionTargetBox="[167,1436,373,1951]" captionTargetId="figure@6.[151,1436,373,1952]" captionTargetPageId="6" captionText="FIGURE 2. Systenus emusorum sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. naranjensis sp. nov. (holotype): c. male habitus, left lateral; d. hypopygium, left lateral; e. male head, anterior." httpUri="https://zenodo.org/record/237972/files/figure.png" pageId="7" pageNumber="176">Fig. 2</figureCitation>
|
||
e); scape, pedicel and basal third of postpedicel mostly yellow with dark brown dorsobasal area on postpedicel, with distal 2/3 postpedicel dark brown; scape rather long and distally expanded and pedicel short; postpedicel elongate, subrectangular on basal third and tapering triangular distally, covered with short pubescence, about 3 times as long as basal width, and with short apical arista.
|
||
<emphasis id="ADF0EAF9FFBAFF89FAE9A41401CDFCFB" box="[1347,1428,798,823]" italics="true" pageId="7" pageNumber="176">Thorax</emphasis>
|
||
: dorsum dark metallic green with bronze reflections and grey pruinosity; pleura green with dense grey pruinosity; metepimeron yellowish along basal half, becoming infuscated dorsally.
|
||
<emphasis id="ADF0EAF9FFBAFF89FC65A46D0055FCB2" box="[975,1036,871,894]" italics="true" pageId="7" pageNumber="176">Legs:</emphasis>
|
||
all coxae, trochanters, femora and tibiae I & II, and very base of tarsus II yellow, with distal tarsus II brownish; FIII yellow with distal fifth brown; TIII yellow but basal fifth and distal third brown; basitarsus III yellow in basal half, with distal half and remainder of tarsus III brown; CI and CII with white anterior setae and shorter vestiture, CIII with strong white lateral seta at ½; major leg setae black; I: 5.1; 4.9; 2.7/ 1.8/ 1.2/ 0.8/ 0.6; leg I bare of major setation, and entire leg with short ivory colored vestiture, not usual black vestiture (MSSC); II: 5.5; 5.9; 3.4/ 2.5/ 2.0/ 1.3/ 0.8; TII with strong ad seta and weaker pd seta at ¼, and strong ad seta and small pd seta at 2/3, ventral seta at ½ and 2/3, and with apical ring of ad, pd, av and ventral seta; III: 6.0; 7.5; 1.5/ 3.9/ 2.4/ 1.5/ 0.8; TIII with short black, with some 6 black dorsal setae along length, and with apical ring of dorsal, ad and av seta.
|
||
<emphasis id="ADF0EAF9FFBAFF89FC2CA38D07E7FB52" box="[902,958,1159,1182]" italics="true" pageId="7" pageNumber="176">Wing</emphasis>
|
||
: hyaline; vein M diverging from R4+5 and with M slightly bowed one third way between dm-cu crossvein and apex, and with M joining costa just behind wing apex; CuAx ratio: 0.7.
|
||
<emphasis id="ADF0EAF9FFBAFF89FE4CA3DB0609FB26" box="[486,592,1233,1258]" italics="true" pageId="7" pageNumber="176">Abdomen</emphasis>
|
||
: tergite 1 mostly yellow; tergites 2 and 3 yellow, but black dorsally and anteriorly, and with dusting of grey pruinosity; tergites 4–5 mostly dull metallic green, but black dorsally and anteriorly and with dense grey pruinosity; tergite 6 black with little pruinosity; posterior margin of tergite 1 with posterior row of long black setae, tergites otherwise covered with short black vestiture; segment 7 forming elongate peduncle, with sternite 7 glabrous and tergite 7 covered with same black vestiture as preabdomen; hypopygium (
|
||
<figureCitation id="07BF2A6EFFBAFF89FF35A28F04B1FA52" box="[159,232,1413,1438]" captionStart="FIGURE 2" captionStartId="6.[151,250,1973,1995]" captionTargetBox="[167,1436,373,1951]" captionTargetId="figure@6.[151,1436,373,1952]" captionTargetPageId="6" captionText="FIGURE 2. Systenus emusorum sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. naranjensis sp. nov. (holotype): c. male habitus, left lateral; d. hypopygium, left lateral; e. male head, anterior." httpUri="https://zenodo.org/record/237972/files/figure.png" pageId="7" pageNumber="176">Fig. 2</figureCitation>
|
||
d) dark brown with yellow surstylus and cercus; hypandrium as elongate cover; epandrium ventrally extending to narrow curved point; surstylus as elongate curved arm with short curved ventral fork; cercus elongate, swollen basally and distally, and covered with short yellow setae.
|
||
<emphasis id="ADF0EAF9FFBAFF89FCD3A2C70789FA2A" bold="true" box="[889,976,1485,1510]" pageId="7" pageNumber="176">Female</emphasis>
|
||
: unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D79E6560FFBAFF89FF6DA2FB0093F99E" pageId="7" pageNumber="176" type="discussion">
|
||
<paragraph id="9F3B36EBFFBAFF89FF6DA2FB0093F99E" blockId="7.[151,1437,546,1654]" pageId="7" pageNumber="176">
|
||
<emphasis id="ADF0EAF9FFBAFF89FF6DA2FB056DF9C6" bold="true" box="[199,308,1521,1546]" pageId="7" pageNumber="176">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="58844D68FFBAFF89FEE9A2F80676F9C5" box="[323,559,1522,1545]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="176" phylum="Arthropoda" rank="species" species="naranjensis">
|
||
<emphasis id="ADF0EAF9FFBAFF89FEE9A2F80676F9C5" box="[323,559,1522,1545]" italics="true" pageId="7" pageNumber="176">Systenus naranjensis</emphasis>
|
||
</taxonomicName>
|
||
is known from a single male taken at the Rio Naranjo site,
|
||
<collectingCountry id="E793767BFFBAFF89FB72A2F8010DF9C6" box="[1240,1364,1521,1546]" name="Costa Rica" pageId="7" pageNumber="176">Costa Rica</collectingCountry>
|
||
in the month of February. Male leg I (including femur, tibia and tarsus), is almost entirely covered with short ivory colored vestiture but the cuticle is yellow. The coloration of the antennae (
|
||
<figureCitation id="07BF2A6EFFBAFF89FC7DA1330043F99E" box="[983,1050,1593,1618]" captionStart="FIGURE 2" captionStartId="6.[151,250,1973,1995]" captionTargetBox="[167,1436,373,1951]" captionTargetId="figure@6.[151,1436,373,1952]" captionTargetPageId="6" captionText="FIGURE 2. Systenus emusorum sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. naranjensis sp. nov. (holotype): c. male habitus, left lateral; d. hypopygium, left lateral; e. male head, anterior." httpUri="https://zenodo.org/record/237972/files/figure.png" pageId="7" pageNumber="176">Fig. 2</figureCitation>
|
||
e) is diagnostic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D79E6560FFBAFF89FF6DA1570153F9BA" box="[199,1290,1629,1654]" pageId="7" pageNumber="176" type="etymology">
|
||
<paragraph id="9F3B36EBFFBAFF89FF6DA1570153F9BA" blockId="7.[151,1437,546,1654]" box="[199,1290,1629,1654]" pageId="7" pageNumber="176">
|
||
<emphasis id="ADF0EAF9FFBAFF89FF6DA157051FF9BA" bold="true" box="[199,326,1629,1654]" pageId="7" pageNumber="176">Etymology</emphasis>
|
||
. This species is named after the place where the
|
||
<typeStatus id="403F8849FFBAFF89FCC0A1570795F9BA" box="[874,972,1629,1654]" pageId="7" pageNumber="176" type="holotype">holotype</typeStatus>
|
||
was collected, Rio Naranjo.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |