140 lines
16 KiB
XML
140 lines
16 KiB
XML
<document id="01FBE15032D88CA806786A19F472DC1C" ID-DOI="10.11646/zootaxa.4020.1.7" ID-GBIF-Dataset="82b2f539-5d25-42d6-95c5-e8b16a353e12" ID-ISSN="1175-5326" ID-Zenodo-Dep="237970" ID-ZooBank="BD8D1880-79A9-4672-99AF-5B3D1BEFAFEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461269068152" checkinUser="plazi" docAuthor="Bickel, Daniel J." docDate="2015" docId="172D87FDFFB7FF85FF3DA268001EFDF1" docLanguage="en" docName="zt04020p182.pdf" docOrigin="Zootaxa 4020 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systenus parkeri Bickel, sp. nov." docType="treatment" docVersion="8" lastPageNumber="180" masterDocId="EB14FF85FFBDFF8EFFAAA70A0459FFCC" masterDocTitle="The Costa Rican Systenus Loew (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus" masterLastPageNumber="182" masterPageNumber="169" pageNumber="179" updateTime="1698611248703" updateUser="plazi">
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<mods:mods id="4CBBBF1B1E6EB01D9530156F6966217C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="60F2D72EDA6259E65975495079A6DBBE">
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<mods:title id="CA81538CDF9F535C871EBED7EA37D366">The Costa Rican Systenus Loew (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus</mods:title>
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</mods:titleInfo>
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<mods:name id="7286F4F2D63D814AA169732537C37883" type="personal">
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<mods:roleTerm id="2A0317E1C564876DE76F432152D24F97">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="60AF85662BEAA72D525032235EA47120">Bickel, Daniel J.</mods:namePart>
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<mods:typeOfResource id="BA02483084E0D55F25BF1422CA5C8087">text</mods:typeOfResource>
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<mods:title id="6C8F6CA1317AFC831CD792C6C8C44469">Zootaxa</mods:title>
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<mods:part id="7E60AF2581B6D090848FA89E36057A4B">
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<mods:date id="75012E73B83BFBE30404214FEC0713E5">2015</mods:date>
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<mods:detail id="9CB171917C68B0FF91077FDE69A4F41B" type="volume">
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<mods:number id="A676EC343CCC5EB3C9E74E2DC7C0E39D">4020</mods:number>
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<mods:detail id="9E7FA1493CA5C51BFB98380E701EB413" type="issue">
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<mods:number id="63E7E30AF2DEC618988B234C9B5DDAD6">1</mods:number>
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<mods:classification id="1D4E2FC07111706BB824528672D268D1">journal article</mods:classification>
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<mods:identifier id="1BB18AA620567BFFD00C60260553F645" type="DOI">10.11646/zootaxa.4020.1.7</mods:identifier>
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<mods:identifier id="092A0B33641785E59D0E89A376D24CD3" type="GBIF-Dataset">82b2f539-5d25-42d6-95c5-e8b16a353e12</mods:identifier>
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<mods:identifier id="0FA1739224EA68D389C08F78F8B938F1" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="03B9D48ADAE0576D15948CEE979C744D" type="Zenodo-Dep">237970</mods:identifier>
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<mods:identifier id="446402D5CDB7C319BA7DD9CA06EDD626" type="ZooBank">BD8D1880-79A9-4672-99AF-5B3D1BEFAFEE</mods:identifier>
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</mods:mods>
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<treatment id="172D87FDFFB7FF85FF3DA268001EFDF1" ID-DOI="http://doi.org/10.5281/zenodo.6121817" ID-GBIF-Taxon="119638522" ID-Zenodo-Dep="6121817" LSID="urn:lsid:plazi:treatment:172D87FDFFB7FF85FF3DA268001EFDF1" httpUri="http://treatment.plazi.org/id/172D87FDFFB7FF85FF3DA268001EFDF1" lastPageId="11" lastPageNumber="180" pageId="10" pageNumber="179">
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<subSubSection id="D79E6560FFB7FF84FF3DA268056BFA52" pageId="10" pageNumber="179" type="nomenclature">
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<paragraph id="9F3B36EBFFB7FF84FF3DA2680643FAB1" blockId="10.[151,538,1378,1438]" box="[151,538,1378,1405]" pageId="10" pageNumber="179">
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<heading id="C4738187FFB7FF84FF3DA2680643FAB1" bold="true" box="[151,538,1378,1405]" fontSize="11" level="1" pageId="10" pageNumber="179" reason="1">
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<emphasis id="ADF0EAF9FFB7FF84FF3DA2680643FAB1" bold="true" box="[151,538,1378,1405]" pageId="10" pageNumber="179">
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<taxonomicName id="58844D68FFB7FF84FF3DA26805ECFAB1" ID-CoL="9HYXF" authority="Bickel" authorityName="Bickel" box="[151,437,1378,1405]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="179" phylum="Arthropoda" rank="species" species="parkeri" status="sp. nov.">
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<emphasis id="ADF0EAF9FFB7FF84FF3DA2680539FAB0" bold="true" box="[151,352,1378,1404]" italics="true" pageId="10" pageNumber="179">Systenus parkeri</emphasis>
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Bickel
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</taxonomicName>
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<taxonomicNameLabel id="B6C35782FFB7FF84FE17A2690643FAB1" box="[445,538,1379,1405]" pageId="10" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="9F3B36EBFFB7FF84FF3DA28F056BFA52" blockId="10.[151,538,1378,1438]" box="[151,306,1413,1438]" pageId="10" pageNumber="179">
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(
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<figureCitation id="07BF2A6EFFB7FF84FF35A28F04BEFA52" box="[159,231,1413,1438]" captionStart="FIGURE 4" captionStartId="11.[151,250,1770,1792]" captionTargetBox="[160,1436,628,1748]" captionTargetId="figure@11.[151,1436,628,1749]" captionTargetPageId="11" captionText="FIGURE 4. Systenus eboritibia sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. parkeri sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral; e. male postabdomen, right lateral." httpUri="https://zenodo.org/record/237974/files/figure.png" pageId="10" pageNumber="179">Figs 4</figureCitation>
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c, d, e)
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</paragraph>
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</subSubSection>
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<subSubSection id="D79E6560FFB7FF84FF3DA2C7011CF9C6" pageId="10" pageNumber="179" type="materials_examined">
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<paragraph id="9F3B36EBFFB7FF84FF3DA2C7011CF9C6" blockId="10.[151,1437,1485,2014]" pageId="10" pageNumber="179">
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<emphasis id="ADF0EAF9FFB7FF84FF3DA2C70518FA2A" bold="true" box="[151,321,1485,1510]" pageId="10" pageNumber="179">
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<typeStatus id="403F8849FFB7FF84FF3DA2C7048BFA2A" box="[151,210,1485,1510]" pageId="10" pageNumber="179">Type</typeStatus>
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material
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</emphasis>
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.
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<typeStatus id="403F8849FFB7FF84FEF8A2C705A8FA2A" box="[338,497,1485,1510]" pageId="10" pageNumber="179" type="holotype">
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<emphasis id="ADF0EAF9FFB7FF84FEF8A2C705A8FA2A" bold="true" box="[338,497,1485,1510]" pageId="10" pageNumber="179">HOLOTYPE</emphasis>
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</typeStatus>
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♂,
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<collectingCountry id="E793767BFFB7FF84FD8BA2C4069EFA2A" box="[545,711,1486,1510]" name="Costa Rica" pageId="10" pageNumber="179">COSTA RICA</collectingCountry>
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:
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<emphasis id="ADF0EAF9FFB7FF84FD71A2C70729FA2A" bold="true" box="[731,880,1485,1510]" pageId="10" pageNumber="179">Guanacaste:</emphasis>
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<quantity id="587C9B0EFFB7FF84FCD0A2C407EFFA2A" box="[890,950,1485,1510]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="179" unit="km" value="3.0">3 km</quantity>
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SE Rio Naranjo,
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<date id="EB3A102BFFB7FF84FB23A2C40153FA2A" box="[1161,1290,1486,1510]" pageId="10" pageNumber="179" value="1991-11-20">20.XI.1991</date>
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, F.D. Parker (
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<collectionCode id="F995AE2EFFB7FF84FF34A2F804A8F9C6" LSID="urn:lsid:biocol.org:col:34802" box="[158,241,1522,1546]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="10" pageNumber="179">LACM</collectionCode>
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Ent 320003).
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<typeStatus id="403F8849FFB7FF84FE3BA2FB0670F9C6" box="[401,553,1521,1546]" pageId="10" pageNumber="179" type="paratype">
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<emphasis id="ADF0EAF9FFB7FF84FE3BA2FB0670F9C6" bold="true" box="[401,553,1521,1546]" pageId="10" pageNumber="179">PARATYPE</emphasis>
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</typeStatus>
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♂, same but
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<date id="EB3A102BFFB7FF84FD15A2FB070DF9C6" box="[703,852,1521,1546]" pageId="10" pageNumber="179" value="1993-01-22" valueMax="1993-01-25" valueMin="1993-01-22">22–25.I.1993</date>
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(
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<collectionCode id="F995AE2EFFB7FF84FCC9A2F807E1F9C6" LSID="urn:lsid:biocol.org:col:34802" box="[867,952,1522,1546]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="10" pageNumber="179">LACM</collectionCode>
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, ex.
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<collectionCode id="F995AE2EFFB7FF84FC46A2F80066F9C6" box="[1004,1087,1522,1546]" pageId="10" pageNumber="179">EMUS</collectionCode>
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). (
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<collectionCode id="F995AE2EFFB7FF84FBF3A2F800F5F9C6" LSID="urn:lsid:biocol.org:col:34802" box="[1113,1196,1522,1546]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="10" pageNumber="179">LACM</collectionCode>
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Ent 320004).
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</paragraph>
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</subSubSection>
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<subSubSection id="D79E6560FFB7FF85FF6DA11F007BFEA9" lastPageId="11" lastPageNumber="180" pageId="10" pageNumber="179" type="description">
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<paragraph id="9F3B36EBFFB7FF85FF6DA11F007BFEA9" blockId="10.[151,1437,1485,2014]" lastBlockId="11.[151,1436,151,573]" lastPageId="11" lastPageNumber="180" pageId="10" pageNumber="179">
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<emphasis id="ADF0EAF9FFB7FF84FF6DA11F0508F9E2" bold="true" box="[199,337,1557,1582]" pageId="10" pageNumber="179">Description</emphasis>
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.
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<emphasis id="ADF0EAF9FFB7FF84FEF5A11F05C5F9E2" bold="true" box="[351,412,1557,1582]" pageId="10" pageNumber="179">Male</emphasis>
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: body length:
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<quantity id="587C9B0EFFB7FF84FD94A11C06C1F9E1" box="[574,664,1558,1582]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="10" pageNumber="179" unit="mm" value="3.2">3.2 mm</quantity>
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; wing 3.0 x 0.1.0 mm (
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<figureCitation id="07BF2A6EFFB7FF84FC31A11F07B9F9E2" box="[923,992,1557,1582]" captionStart="FIGURE 4" captionStartId="11.[151,250,1770,1792]" captionTargetBox="[160,1436,628,1748]" captionTargetId="figure@11.[151,1436,628,1749]" captionTargetPageId="11" captionText="FIGURE 4. Systenus eboritibia sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. parkeri sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral; e. male postabdomen, right lateral." httpUri="https://zenodo.org/record/237974/files/figure.png" pageId="10" pageNumber="179">Fig. 4</figureCitation>
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c); similar to
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<taxonomicName id="58844D68FFB7FF84FBDEA11C00B8F9E1" box="[1140,1249,1558,1581]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="179" phylum="Arthropoda" rank="species" species="tenorio">
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<emphasis id="ADF0EAF9FFB7FF84FBDEA11C00B8F9E1" box="[1140,1249,1558,1581]" italics="true" pageId="10" pageNumber="179">S. tenorio</emphasis>
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</taxonomicName>
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except as noted:
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<emphasis id="ADF0EAF9FFB7FF84FF3DA133048DF99E" box="[151,212,1593,1618]" italics="true" pageId="10" pageNumber="179">Head</emphasis>
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: proboscis yellow, projecting anteriorly, keel-like; antennal scape, pedicel and basal half of postpedicel yellow and distal tapering half of postpedicel dark brown; scape and pedicel short; postpedicel subrectangular basally and abruptly narrowed to elongate tapering point in distal half, and covered with short pubescence, with antenna overall about 3 times as long as basal width; short apical arista present.
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<emphasis id="ADF0EAF9FFB7FF84FBEBA1AC002FF971" box="[1089,1142,1702,1725]" italics="true" pageId="10" pageNumber="179">Legs</emphasis>
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: CI yellow; CII and CIII mostly dark brown, all trochanters, and remainder of legs yellow; CI and CII with white anterior setae and shorter vestiture, CIII with strong white lateral seta at ½; major leg setae black; I: 4.5; 4.0; 2.0/ 1.4/ 1.0/ 0.4/ 0.4; leg I bare of major setation; tibia I yellow with brownish short vestiture; II: 4.3; 4.7; 2.7/ 1.7/ 1.2/ 0.8/ 0.6; TII with strong ad seta and weaker pd seta at ¼, and strong ad seta and small pd seta at 2/3, and with apical ring of ad, pd, av and ventral seta; III: 4.0; 5.2; 1.0/ 1.6/ 1.0/ 0.6/ 0.5; TIII with ad-pd setal pair near 1/8, without white vestiture, and with 4 black dorsal setae, and with apical ring of dorsal, ad and av seta.
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<emphasis id="ADF0EAF9FFB7FF84FC3EA0740795F859" box="[916,972,1918,1941]" italics="true" pageId="10" pageNumber="179">Wing</emphasis>
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: CuAx ratio: 0.9.
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<emphasis id="ADF0EAF9FFB7FF84FB34A0770151F85A" box="[1182,1288,1917,1942]" italics="true" pageId="10" pageNumber="179">Abdomen</emphasis>
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: tergites 1–4 mostly metallic bronze-green with dusting of grey pruinosity, and tergites 3–4 with anterior margin metallic black; tergites 5–7 mostly dark metallic green; posterior margin of tergite 1 with posterior row of long black setae, tergites otherwise covered with short black vestiture; segment 7 forming elongate peduncle, with sternite 7 glabrous and tergite 7 covered with short black vestiture; sternite 8 dark brown forming cap-like cover over hypopygial foramen on left side of epandrium; hypopygium (
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<figureCitation id="07BF2A6EFFB6FF85FDF4A7D506FFFF34" box="[606,678,223,248]" captionStart="FIGURE 4" captionStartId="11.[151,250,1770,1792]" captionTargetBox="[160,1436,628,1748]" captionTargetId="figure@11.[151,1436,628,1749]" captionTargetPageId="11" captionText="FIGURE 4. Systenus eboritibia sp. nov. (holotype): a. male habitus, left lateral; b. hypopygium, left lateral. S. parkeri sp. nov.: c. male habitus, left lateral; d. hypopygium, left lateral; e. male postabdomen, right lateral." httpUri="https://zenodo.org/record/237974/files/figure.png" pageId="11" pageNumber="180">Figs 4</figureCitation>
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d, e) dark brown with yellow surstylus and cercus; epandrium short and subrectangular; epandrial lobe with long and short apical seta; surstylus basally expanded with numerous pale setae and tapering to curved subrectangular apex; cercus relatively short, subrectangular, broad basally and narrowed to wide recurved apex, and with short yellow hairs.
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<emphasis id="ADF0EAF9FFB6FF85FCE6A64607FAFEA9" bold="true" box="[844,931,332,357]" pageId="11" pageNumber="180">Female</emphasis>
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: unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="D79E6560FFB6FF85FF6DA665071BFE39" pageId="11" pageNumber="180" type="discussion">
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<paragraph id="9F3B36EBFFB6FF85FF6DA665071BFE39" blockId="11.[151,1436,151,573]" pageId="11" pageNumber="180">
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<emphasis id="ADF0EAF9FFB6FF85FF6DA665056DFE44" bold="true" box="[199,308,367,392]" pageId="11" pageNumber="180">Remarks</emphasis>
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.
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<taxonomicName id="58844D68FFB6FF85FEEEA67A05A6FE44" box="[324,511,367,392]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="180" phylum="Arthropoda" rank="species" species="parkeri">
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<emphasis id="ADF0EAF9FFB6FF85FEEEA67A05A6FE44" box="[324,511,367,392]" italics="true" pageId="11" pageNumber="180">Systenus parkeri</emphasis>
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</taxonomicName>
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is known from two males taken at the Rio Naranjo site,
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<collectingCountry id="E793767BFFB6FF85FB27A67A0152FE44" box="[1165,1291,367,392]" name="Costa Rica" pageId="11" pageNumber="180">Costa Rica</collectingCountry>
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, collected in November and January. It has the diagnostic male characters of the postpedicel basally subrectangular and yellow basally and abruptly narrowed to elongate dark brown tapering point in distal half, legs mostly yellow, tibia I yellow, not white, and cercus short, wide and subrectangular.
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</paragraph>
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</subSubSection>
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<subSubSection id="D79E6560FFB6FF85FF6DA6F5001EFDF1" pageId="11" pageNumber="180" type="etymology">
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<paragraph id="9F3B36EBFFB6FF85FF6DA6F5001EFDF1" blockId="11.[151,1436,151,573]" pageId="11" pageNumber="180">
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<emphasis id="ADF0EAF9FFB6FF85FF6DA6F5051FFDD4" bold="true" box="[199,326,511,536]" pageId="11" pageNumber="180">Etymology</emphasis>
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. This species is named in honor of Frank Parker, emeritus Utah State University, who collected extensively in Latin
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<collectingCountry id="E793767BFFB6FF85FED6A52E0585FDF1" box="[380,476,548,573]" name="United States of America" pageId="11" pageNumber="180">America</collectingCountry>
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and provided most of the specimens used in this study.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |