217 lines
19 KiB
XML
217 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zitteliana.96.e84187" ID-Pensoft-Pub="2747-8106-96-179" ID-Pensoft-UUID="FE0861D71BB454999EE6637C0D9B2B0C" ID-ZooBank="35B619086E6548B09A177281C2253391" ModsDocID="2747-8106-96-179" checkinTime="1670952513976" checkinUser="pensoft" docAuthor="Gruendel, Joachim, Keupp, Helmut, Lang, Fritz & Nuetzel, Alexander" docDate="2022" docId="36055E12A8B95738BB927475F5DAE368" docLanguage="en" docName="Zitteliana 96: 179-221" docOrigin="Zitteliana 96" docPubDate="2022-12-12" docSource="http://dx.doi.org/10.3897/zitteliana.96.e84187" docTitle="Ptygmatis ursicina" docType="treatment" docVersion="1" id="FE0861D71BB454999EE6637C0D9B2B0C" lastPageNumber="179" masterDocId="FE0861D71BB454999EE6637C0D9B2B0C" masterDocTitle="Late Jurassic (Upper Kimmeridgian) Heterobranchia (Gastropoda) of the coral-facies of Saal near Kelheim and the viciniy of Nattheim (Germany)" masterLastPageNumber="221" masterPageNumber="179" pageNumber="179" updateTime="1670952513976" updateUser="pensoft">
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<mods:titleInfo>
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<mods:title>Late Jurassic (Upper Kimmeridgian) Heterobranchia (Gastropoda) of the coral-facies of Saal near Kelheim and the viciniy of Nattheim (Germany)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gruendel, Joachim</mods:namePart>
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<mods:affiliation>Institut fuer Geowissenschaften, Fachrichtung Palaeontologie, Freie Universitaet Berlin, Malteserstrasse 74 - 100, 12249 Berlin, Germany</mods:affiliation>
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<mods:nameIdentifier type="email">joachim.gruendel@lingua-pura.de</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Keupp, Helmut</mods:namePart>
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<mods:affiliation>Institut fuer Geowissenschaften, Fachrichtung Palaeontologie, Freie Universitaet Berlin, Malteserstrasse 74 - 100, 12249 Berlin, Germany</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Lang, Fritz</mods:namePart>
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<mods:affiliation>Drosselweg 16, 96114 Hirschaid, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Nuetzel, Alexander</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8852-7688</mods:nameIdentifier>
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<mods:affiliation>SNSB-Bayerische Staatssammlung fuer Palaeontologie und Geologie, Richard-Wagner-Str. 10, 80333 Muenchen, Germany & Department of Earth and Environmental Sciences, Paleontology and Geobiology, Ludwig-Maximilians-Universitaet Muenchen, Richard-Wagner-Str. 10, 80333 Muenchen, Germany</mods:affiliation>
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<mods:title>Zitteliana</mods:title>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-12-12</mods:number>
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<mods:detail type="volume">
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<mods:number>96</mods:number>
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<mods:extent unit="page">
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<mods:start>179</mods:start>
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<mods:end>221</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zitteliana.96.e84187</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zitteliana.96.e84187</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2747-8106-96-179</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:36055E12A8B95738BB927475F5DAE368" httpUri="http://treatment.plazi.org/id/36055E12A8B95738BB927475F5DAE368" lastPageNumber="179" pageId="0" pageNumber="179">
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<subSubSection pageId="0" pageNumber="179" type="nomenclature">
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<paragraph pageId="0" pageNumber="179">
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<taxonomicName LSID="36055E12-A8B9-5738-BB92-7475F5DAE368" authority="(Thurmann, 1861 in Thurmann and Etallon 1861 - 1864)" baseAuthorityName="Thurmann, 1861 in Thurmann and Etallon 1861 -" baseAuthorityYear="1864" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis ursicina" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="ursicina">
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Ptygmatis? ursicina (Thurmann, 1861 in Thurmann and
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<normalizedToken originalValue="Étallon">Etallon</normalizedToken>
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1861-1864)
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="179" type="description">
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<paragraph pageId="0" pageNumber="179">
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<figureCitation captionStart="Plate 12" captionStartId="F12" captionText="Plate 12. (1 - 3) Ptygmatis clio (d'Orbigny, 1852). (1 - 2) SNSB-BSPG 2021 XV 59, Saal (collection Lang); (1) section of fragment, height 21 mm; (2) detail, height 3.7 mm. (3) SNSB-BSPG 2021 XV 60, Saal (collection Lang), lateral view, height 108 mm. (4) Ptygmatis? polyspira (Quenstedt, 1881 - 1884), holotype, collection University Tuebingen (collection Quenstedt); Nattheim, lateral view, height 32 mm (original to Quenstedt 1881 - 1884, pl. 207, fig. 3. (11 - 12) Bactroptyxis teres (Muenster in Goldfuss, 1844), collection Sauerborn, Nattheim area. (5 - 7) Ptygmatis? ursicina (Thurmann in Thurmann & Etallon, 1861). (5 - 6) SNSB-BSPG 2021 XV 30, Saal (collection Lang); (5) last whorl with aperture, height 4 mm; (6) lateral view, height 9.5 mm. (7) SNSB-BSPG 2021 XV 31, Saal (collection Lang), aperture and ornament of base, width 2.5 mm." figureDoi="10.3897/zitteliana.96.e84187.plate12" httpUri="https://binary.pensoft.net/fig/780390" pageId="0" pageNumber="179">Plate 12: figs 5-7</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="179" type="reference_group">
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<paragraph pageId="0" pageNumber="179">
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<taxonomicName class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea visurgis" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="visurgis">?1844 - Nerinea visurgis</taxonomicName>
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<normalizedToken originalValue="Röm">Roem</normalizedToken>
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. - Goldfuss: 44, pl. 176, fig. 6.
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</paragraph>
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<paragraph pageId="0" pageNumber="179">
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<taxonomicName class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea visurgis" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="visurgis">1852 - Nerinea visurgis</taxonomicName>
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Roemer, 1836 -
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<normalizedToken originalValue="d’Orbigny">d'Orbigny</normalizedToken>
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: 122, pl. 268, figs 5-7.
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</paragraph>
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<paragraph pageId="0" pageNumber="179">
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<taxonomicName class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea ursicina" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="ursicina">*1861-1864 - Nerinea ursicina</taxonomicName>
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Th. - Thurmann and
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<normalizedToken originalValue="Étallon">Etallon</normalizedToken>
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: 103, pl. 8, fig. 50.
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</paragraph>
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<paragraph pageId="0" pageNumber="179">
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<taxonomicName class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea pseudospeciosa" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="pseudospeciosa">?1872 - Nerinea pseudospeciosa</taxonomicName>
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P. de Loriol, 1871 - Loriol, Royer and Tombeck: 89, pl. 6, fig. 7.
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</paragraph>
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<paragraph pageId="0" pageNumber="179">
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<taxonomicName class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea ursicina" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="ursicina">1889 - Nerinea ursicina</taxonomicName>
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Thurmann - Loriol and Koby: 37, pl. 6, figs 1-8.
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</paragraph>
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<paragraph pageId="0" pageNumber="179">
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<taxonomicName class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea ursicinensis" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="ursicinensis">1898 - Nerinea ursicinensis</taxonomicName>
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Thurmann - Cossmann: 37, pl. 3, figs 11, 12.
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</paragraph>
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<paragraph pageId="0" pageNumber="179">
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<taxonomicName class="Gastropoda" family="Ptygmatididae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea ursicinensis" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="ursicinensis">1927 - Nerinea ursicinensis</taxonomicName>
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Thurmann - Maire: 142, pl. 7, figs 15, 16.
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</paragraph>
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<paragraph pageId="0" pageNumber="179">
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<taxonomicName class="Gastropoda" family="Ptygmatididae" genus="Cossmannea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cossmannea (Eunerinea) ursicina" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="ursicina" subGenus="Eunerinea">1997 - Cossmannea (Eunerinea) ursicina</taxonomicName>
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(Thurmann, 1861) - Fischer and Weber: 40, pl. 9, fig. 2.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="179" type="materials_examined">
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<paragraph pageId="0" pageNumber="179">Material.</paragraph>
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<paragraph pageId="0" pageNumber="179">Nineteen juvenile specimens from Saal (collection Lang), of which two are illustrated (SNSB-BSPG 2021 XV 30, 31).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="179" type="description">
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<paragraph pageId="0" pageNumber="179">Description.</paragraph>
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<paragraph pageId="0" pageNumber="179">The largest specimen from Saal is 9 mm high. The shell is slender to very slender. The whorls are regularly increasing in width. The whorl face is concave, with a strong, sometimes ramp-like bulge. The bulge is formed at the suture by both whorls. The suture is situated somewhat above the middle of the bulge. The whorl face between the bulges is initially ornamented with one spiral cord, later whorls with 4-5 spiral cords: two of these spiral cords may be stronger than the others. The bulges and stronger spiral cords are possibly nodular (unclear due to preservation). The base is flat, with an almost rectangular transition to the whorl face. The transition has a strong, protruding bulge that is largely covered by the following whorls of the spire. The base is densely covered with weak spiral cords. The aperture has an approximately rectangular outline and distinct oblique siphonal canal. The aperture has a single parietal plait, two columellar plaits, and a palatal plait.</paragraph>
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<caption doi="10.3897/zitteliana.96.e84187.plate12" httpUri="https://binary.pensoft.net/fig/780390" pageId="0" pageNumber="179" start="Plate 12" startId="F12">
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<paragraph pageId="0" pageNumber="179">
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<emphasis bold="true" pageId="0" pageNumber="179">Plate 12.</emphasis>
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<emphasis bold="true" pageId="0" pageNumber="179">(1-3)</emphasis>
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<taxonomicName baseAuthorityName="d'Orbigny" baseAuthorityYear="1852" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis clio" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="clio">
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<emphasis italics="true" pageId="0" pageNumber="179">Ptygmatis clio</emphasis>
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</taxonomicName>
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(
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<normalizedToken originalValue="d’Orbigny">d'Orbigny</normalizedToken>
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, 1852).
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<emphasis bold="true" pageId="0" pageNumber="179">(1-2)</emphasis>
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SNSB-BSPG 2021 XV 59, Saal (collection Lang);
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<emphasis bold="true" pageId="0" pageNumber="179">(1)</emphasis>
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section of fragment, height 21 mm;
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<emphasis bold="true" pageId="0" pageNumber="179">(2)</emphasis>
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detail, height 3.7 mm.
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<emphasis bold="true" pageId="0" pageNumber="179">(3)</emphasis>
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SNSB-BSPG 2021 XV 60, Saal (collection Lang), lateral view, height 108 mm.
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<emphasis bold="true" pageId="0" pageNumber="179">(4)</emphasis>
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<taxonomicName baseAuthorityName="Quenstedt" baseAuthorityYear="1884" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis polyspira" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="polyspira">Ptygmatis? polyspira</taxonomicName>
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(Quenstedt, 1881-1884), holotype, collection University
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<normalizedToken originalValue="Tübingen">Tuebingen</normalizedToken>
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(collection Quenstedt); Nattheim, lateral view, height 32 mm (original to
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<bibRefCitation author="Quenstedt, FA" journalOrPublisher="Fues, Leipzig" pageId="0" pageNumber="179" refId="B82" refString="Quenstedt, FA, 1881-1884. Petrefaktenkunde Deutschlands. Erste Abteilung, Band 7: Gasteropoden. Fues, Leipzig" title="Petrefaktenkunde Deutschlands. Erste Abteilung, Band 7: Gasteropoden." year="1881 - 1884">Quenstedt 1881-1884</bibRefCitation>
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, pl. 207, fig. 3. (11-12)
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<taxonomicName baseAuthorityName="Muenster in Goldfuss" baseAuthorityYear="1844" class="Gastropoda" family="Ptygmatididae" genus="Bactroptyxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bactroptyxis teres" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="teres">
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<emphasis italics="true" pageId="0" pageNumber="179">Bactroptyxis teres</emphasis>
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</taxonomicName>
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(
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<normalizedToken originalValue="Münster">Muenster</normalizedToken>
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in Goldfuss, 1844), collection Sauerborn, Nattheim area.
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<emphasis bold="true" pageId="0" pageNumber="179">(5-7)</emphasis>
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<taxonomicName baseAuthorityName="Thurmann, 1861 in Thurmann and Etallon 1861 -" baseAuthorityYear="1864" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis ursicina" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="ursicina">Ptygmatis? ursicina</taxonomicName>
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(Thurmann in Thurmann &
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<normalizedToken originalValue="Étallon">Etallon</normalizedToken>
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, 1861).
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<emphasis bold="true" pageId="0" pageNumber="179">(5-6)</emphasis>
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SNSB-BSPG 2021 XV 30, Saal (collection Lang);
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<emphasis bold="true" pageId="0" pageNumber="179">(5)</emphasis>
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last whorl with aperture, height 4 mm;
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<emphasis bold="true" pageId="0" pageNumber="179">(6)</emphasis>
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lateral view, height 9.5 mm.
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<emphasis bold="true" pageId="0" pageNumber="179">(7)</emphasis>
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SNSB-BSPG 2021 XV 31, Saal (collection Lang), aperture and ornament of base, width 2.5 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="179" type="remarks">
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<paragraph pageId="0" pageNumber="179">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="179">
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The identity of the present juvenile shells with much larger growing taxa is uncertain (see synonymy list). Thurmann in
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<bibRefCitation author="Thurmann, J" journalOrPublisher="Memoires de la Societe du museum d'histoire naturelle de Strasbourg" pageId="0" pageNumber="179" refId="B100" refString="Thurmann, J, Etallon, A, 1861-1864. Lethea bruntrutana ou etudes paleontologiques et stratigraphiques sur le Jura bernois et en particulier les environs de Porrentruy. Neue Denkschrift der allgemeinen Schweizerischen Gesellschaft fuer die Naturwissenschaften 18-20: 1-500." title="Lethea bruntrutana ou etudes paleontologiques et stratigraphiques sur le Jura bernois et en particulier les environs de Porrentruy. Neue Denkschrift der allgemeinen Schweizerischen Gesellschaft fuer die Naturwissenschaften 18 - 20: 1 - 500." year="1861 - 1864">
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Thurmann and
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<normalizedToken originalValue="Étallon">Etallon</normalizedToken>
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(1861-1864)
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</bibRefCitation>
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reported that
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<taxonomicName class="Gastropoda" family="Itieriidae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea ursicina" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="ursicina">
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<emphasis italics="true" pageId="0" pageNumber="179">Nerinea ursicina</emphasis>
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</taxonomicName>
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has distinct knobs on the bulges and partly also on the spiral cords between the bulges. Such a knobby ornament could not be substantiated for the present material, due perhaps to preservation.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="179" type="relationships">
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<paragraph pageId="0" pageNumber="179">Relationships.</paragraph>
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<paragraph pageId="0" pageNumber="179">
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Differences to
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<taxonomicName baseAuthorityName="Quenstedt" baseAuthorityYear="1884" class="Gastropoda" family="Ptygmatididae" genus="Ptygmatis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptygmatis polyspira" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="polyspira">Ptygmatis? polyspira</taxonomicName>
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(Quenstedt, 1881-1884) are herein discussed in the treatment of this taxon.
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<taxonomicName class="Gastropoda" family="Itieriidae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea subscalaris" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="subscalaris">
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<emphasis italics="true" pageId="0" pageNumber="179">Nerinea subscalaris</emphasis>
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</taxonomicName>
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<normalizedToken originalValue="Münster">Muenster</normalizedToken>
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in Goldfuss (1844) has higher whorls in relation to whorl width and it has more oblique sutures.
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<taxonomicName class="Gastropoda" family="Itieriidae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea lafayettensis" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="lafayettensis">Nerinea? lafayettensis</taxonomicName>
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Imlay, 1945 has, among other differences, only two spiral cords between the bulges.
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<taxonomicName class="Gastropoda" family="Itieriidae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea speciosa" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="speciosa">
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<emphasis italics="true" pageId="0" pageNumber="179">Nerinea speciosa</emphasis>
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</taxonomicName>
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Voltz sensu
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<bibRefCitation author="Maire, V" journalOrPublisher="Bulletin de la Societe grayloise d'emulation Lettress, Sciences, Beaux-arts" pageId="0" pageNumber="179" refId="B61" refString="Maire, V, 1927. Etudes geologiques et paleontologiques sur l'arrondissement de Gray. Les Gastropodes du Jurassique superieur graylois (fin): 85-173." title="Etudes geologiques et paleontologiques sur l'arrondissement de Gray. Les Gastropodes du Jurassique superieur graylois (fin): 85 - 173." year="1927">Maire (1927)</bibRefCitation>
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has a broader shell with lower whorls and its bulges are not as much protruding.
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<taxonomicName class="Gastropoda" family="Itieriidae" genus="Nerinea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerinea mariae" pageId="0" pageNumber="179" phylum="Mollusca" rank="species" species="mariae">
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<emphasis italics="true" pageId="0" pageNumber="179">Nerinea mariae</emphasis>
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</taxonomicName>
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<normalizedToken originalValue="d’Orbigny">d'Orbigny</normalizedToken>
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sensu
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<bibRefCitation author="Cossmann, M" journalOrPublisher="The author and Comptoir Geologique, Paris" pageId="0" pageNumber="179" refId="B12" refString="Cossmann, M, 1898. Contribution a la Paleontologie francaise des terrains jurassiques: Nerinees. Memoires de la Societe gelogique de France, ser. Paleontologie (2) 8: 1-179." title="Contribution a la Paleontologie francaise des terrains jurassiques: Nerinees. Memoires de la Societe gelogique de France, ser. Paleontologie (2) 8: 1 - 179." year="1898">Cossmann (1898)</bibRefCitation>
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is slenderer and has higher whorls.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |