treatments-xml/data/C4/99/B0/C499B059A1A25442872FAA446F6CBEEA.xml
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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.187.73338" ID-GBIF-Dataset="4cbda95a-5b4a-45e9-b53a-9b90db01ee17" ID-PMC="PMC8688410" ID-Pensoft-Pub="1314-2003-187-23" ID-Pensoft-UUID="6AF5EAA0500558E5838FD916171188B8" ID-PubMed="35002365" ModsDocID="1314-2003-187-23" checkinTime="1639455686940" checkinUser="pensoft" docAuthor="Morales, Eduardo A., Wetzel, Carlos E. &amp; Ector, Luc" docDate="2021" docId="C499B059A1A25442872FAA446F6CBEEA" docLanguage="en" docName="PhytoKeys 187: 23-70" docOrigin="PhytoKeys 187" docPubDate="2021-12-13" docSource="http://dx.doi.org/10.3897/phytokeys.187.73338" docTitle="Pseudostaurosira occulta E. Morales, C. E. Wetzel &amp; Ector 2021, sp. nov." docType="treatment" docVersion="4" id="6AF5EAA0500558E5838FD916171188B8" lastPageNumber="23" masterDocId="6AF5EAA0500558E5838FD916171188B8" masterDocTitle="New and poorly known &quot; araphid &quot; diatom species (Bacillariophyta) from regions near Lake Titicaca, South America and a discussion on the continued use of morphological characters in &quot; araphid &quot; diatom taxonomy" masterLastPageNumber="70" masterPageNumber="23" pageNumber="23" updateTime="1668140567392" updateUser="ExternalLinkService">
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<mods:title>New and poorly known &quot; araphid &quot; diatom species (Bacillariophyta) from regions near Lake Titicaca, South America and a discussion on the continued use of morphological characters in &quot; araphid &quot; diatom taxonomy</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Morales, Eduardo A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5998-4831</mods:nameIdentifier>
<mods:affiliation>Water Laboratory, University of Evora, P. I. T. E. Rua da Barba Rala No. 1, 7005 - 345 Evora, Portugal &amp; Institute of Earth Sciences - ICT, University of Evora, Rua Romao Ramalho n °. 59, 7000 - 671 Evora, Portugal</mods:affiliation>
<mods:nameIdentifier type="email">edu_mora123@outlook.com</mods:nameIdentifier>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Wetzel, Carlos E.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5330-0494</mods:nameIdentifier>
<mods:affiliation>Observatory for Climate, Environment and Biodiversity (OCEB), Environmental Research and Innovation (ERIN) Department, Luxembourg Institute of Science and Technology (LIST), 41 rue du Brill, 4422 Belvaux, Luxembourg</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ector, Luc</mods:namePart>
<mods:affiliation>Observatory for Climate, Environment and Biodiversity (OCEB), Environmental Research and Innovation (ERIN) Department, Luxembourg Institute of Science and Technology (LIST), 41 rue du Brill, 4422 Belvaux, Luxembourg</mods:affiliation>
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<mods:title>PhytoKeys</mods:title>
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<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-12-13</mods:number>
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<mods:detail type="volume">
<mods:number>187</mods:number>
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<mods:start>23</mods:start>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.187.73338</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.187.73338</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-187-23</mods:identifier>
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<treatment ID-GBIF-Taxon="191535334" LSID="urn:lsid:plazi:treatment:C499B059A1A25442872FAA446F6CBEEA" httpUri="http://treatment.plazi.org/id/C499B059A1A25442872FAA446F6CBEEA" lastPageNumber="23" pageId="0" pageNumber="23">
<subSubSection pageId="0" pageNumber="23" type="nomenclature">
<paragraph pageId="0" pageNumber="23">
<taxonomicName LSID="C499B059-A1A2-5442-872F-AA446F6CBEEA" authority="E. Morales, C. E. Wetzel &amp; Ector" authorityName="E. Morales, C. E. Wetzel &amp; Ector" authorityYear="2021" class="Fragilariophyceae" family="Staurosiraceae" genus="Pseudostaurosira" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Pseudostaurosira occulta" order="Fragilariales" pageId="0" pageNumber="23" phylum="Bacillariophyta" rank="species" species="occulta" status="sp. nov.">Pseudostaurosira occulta E. Morales, C.E. Wetzel &amp; Ector</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="23">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="description">
<paragraph pageId="0" pageNumber="23">
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-V LM images of little known and new &quot; araphid &quot; diatoms from the Bolivian Altiplano A-D Pseudostaurosira vulpina stat. nov. E-I P. frankenae sp. nov. (Fig. 6 E is the holotype) J-O P. occulta sp. nov. (Fig. 6 K corresponds to the holotype) P-V P. oblonga sp. nov. (Fig. 6 R corresponds to the holotype). Scale bar: 10 µm." figureDoi="10.3897/phytokeys.187.73338.figure6" httpUri="https://binary.pensoft.net/fig/622443" pageId="0" pageNumber="23">Figs 6J-O (LM), 9A-F (SEM)</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="holotype">
<paragraph pageId="0" pageNumber="23">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="23">
Slide BR-4679, Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-V LM images of little known and new &quot; araphid &quot; diatoms from the Bolivian Altiplano A-D Pseudostaurosira vulpina stat. nov. E-I P. frankenae sp. nov. (Fig. 6 E is the holotype) J-O P. occulta sp. nov. (Fig. 6 K corresponds to the holotype) P-V P. oblonga sp. nov. (Fig. 6 R corresponds to the holotype). Scale bar: 10 µm." figureDoi="10.3897/phytokeys.187.73338.figure6" httpUri="https://binary.pensoft.net/fig/622443" pageId="0" pageNumber="23">6K</figureCitation>
, Meise Botanic Garden, Belgium.
<emphasis bold="true" pageId="0" pageNumber="23">Isotype.</emphasis>
Slide DBOL-0249a, Diatomotheca Boliviensis (before HCUCB), Cochabamba, Bolivia.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="type locality">
<paragraph pageId="0" pageNumber="23">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="23">
Bolivia. Sajama Province, Department of Oruro, Sajama River, epipsammon,
<geoCoordinate degrees="17" direction="south" minutes="30" orientation="latitude" precision="15" seconds="33" value="-17.509167">17°30'33&quot;S</geoCoordinate>
;
<geoCoordinate degrees="68" direction="west" minutes="20" orientation="longitude" precision="15" seconds="35" value="-68.343056">68°20'35&quot;W</geoCoordinate>
, 4000 m elev.,
<emphasis italics="true" pageId="0" pageNumber="23">leg.</emphasis>
G.
<normalizedToken originalValue="Chávez">Chavez</normalizedToken>
, 05.07.2009.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="description">
<paragraph pageId="0" pageNumber="23">Description.</paragraph>
<paragraph pageId="0" pageNumber="23">
Frustules rectangular in girdle view, joined together by interlocking spines. Valves lanceolate, isopolar with semi-gradual transition from valve face to mantle. Valve apices subrostrate with broadly rounded, somewhat squarish ends (Figs
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-V LM images of little known and new &quot; araphid &quot; diatoms from the Bolivian Altiplano A-D Pseudostaurosira vulpina stat. nov. E-I P. frankenae sp. nov. (Fig. 6 E is the holotype) J-O P. occulta sp. nov. (Fig. 6 K corresponds to the holotype) P-V P. oblonga sp. nov. (Fig. 6 R corresponds to the holotype). Scale bar: 10 µm." figureDoi="10.3897/phytokeys.187.73338.figure6" httpUri="https://binary.pensoft.net/fig/622443" pageId="0" pageNumber="23">6J-O</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-F SEM images of Pseudostaurosira occulta sp. nov. A, B external views of valves showing apical pore fields (black arrow in A) and other features C internal view of valve showing depressions containing the striae (dashed arrow) D internal close-up showing apical pore field depression (black arrow) and striae depression (dashed line) E close-up on valve apex showing flaps on apical pore field (black arrow) and open girdle element (white arrow) F close-up on cell-cell connection showing apical pore field covered with flaps (black arrow) and open girdle element (white arrow). Scale bars: 2 µm (D, E); 3 µm (F); 5 µm (A-C)." figureDoi="10.3897/phytokeys.187.73338.figure9" httpUri="https://binary.pensoft.net/fig/622446" pageId="0" pageNumber="23">9A-D</figureCitation>
). Axial area lanceolate (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-V LM images of little known and new &quot; araphid &quot; diatoms from the Bolivian Altiplano A-D Pseudostaurosira vulpina stat. nov. E-I P. frankenae sp. nov. (Fig. 6 E is the holotype) J-O P. occulta sp. nov. (Fig. 6 K corresponds to the holotype) P-V P. oblonga sp. nov. (Fig. 6 R corresponds to the holotype). Scale bar: 10 µm." figureDoi="10.3897/phytokeys.187.73338.figure6" httpUri="https://binary.pensoft.net/fig/622443" pageId="0" pageNumber="23">6J-O</figureCitation>
), externally and internally faintly depressed with respect to virgae (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-F SEM images of Pseudostaurosira occulta sp. nov. A, B external views of valves showing apical pore fields (black arrow in A) and other features C internal view of valve showing depressions containing the striae (dashed arrow) D internal close-up showing apical pore field depression (black arrow) and striae depression (dashed line) E close-up on valve apex showing flaps on apical pore field (black arrow) and open girdle element (white arrow) F close-up on cell-cell connection showing apical pore field covered with flaps (black arrow) and open girdle element (white arrow). Scale bars: 2 µm (D, E); 3 µm (F); 5 µm (A-C)." figureDoi="10.3897/phytokeys.187.73338.figure9" httpUri="https://binary.pensoft.net/fig/622446" pageId="0" pageNumber="23">9A-D</figureCitation>
). Vimines short and wide (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-F SEM images of Pseudostaurosira occulta sp. nov. A, B external views of valves showing apical pore fields (black arrow in A) and other features C internal view of valve showing depressions containing the striae (dashed arrow) D internal close-up showing apical pore field depression (black arrow) and striae depression (dashed line) E close-up on valve apex showing flaps on apical pore field (black arrow) and open girdle element (white arrow) F close-up on cell-cell connection showing apical pore field covered with flaps (black arrow) and open girdle element (white arrow). Scale bars: 2 µm (D, E); 3 µm (F); 5 µm (A-C)." figureDoi="10.3897/phytokeys.187.73338.figure9" httpUri="https://binary.pensoft.net/fig/622446" pageId="0" pageNumber="23">9A, B</figureCitation>
). Striae composed of round to elliptic areolae, decreasing in size towards the axial area (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-F SEM images of Pseudostaurosira occulta sp. nov. A, B external views of valves showing apical pore fields (black arrow in A) and other features C internal view of valve showing depressions containing the striae (dashed arrow) D internal close-up showing apical pore field depression (black arrow) and striae depression (dashed line) E close-up on valve apex showing flaps on apical pore field (black arrow) and open girdle element (white arrow) F close-up on cell-cell connection showing apical pore field covered with flaps (black arrow) and open girdle element (white arrow). Scale bars: 2 µm (D, E); 3 µm (F); 5 µm (A-C)." figureDoi="10.3897/phytokeys.187.73338.figure9" httpUri="https://binary.pensoft.net/fig/622446" pageId="0" pageNumber="23">9A, B</figureCitation>
); wide elliptical areolae present toward valve face/mantle transition before and after the spine, sometimes accompanied by a narrower additional areola on valve mantle (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-F SEM images of Pseudostaurosira occulta sp. nov. A, B external views of valves showing apical pore fields (black arrow in A) and other features C internal view of valve showing depressions containing the striae (dashed arrow) D internal close-up showing apical pore field depression (black arrow) and striae depression (dashed line) E close-up on valve apex showing flaps on apical pore field (black arrow) and open girdle element (white arrow) F close-up on cell-cell connection showing apical pore field covered with flaps (black arrow) and open girdle element (white arrow). Scale bars: 2 µm (D, E); 3 µm (F); 5 µm (A-C)." figureDoi="10.3897/phytokeys.187.73338.figure9" httpUri="https://binary.pensoft.net/fig/622446" pageId="0" pageNumber="23">9E, F</figureCitation>
). Striae contained in a single depression in internal view (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-F SEM images of Pseudostaurosira occulta sp. nov. A, B external views of valves showing apical pore fields (black arrow in A) and other features C internal view of valve showing depressions containing the striae (dashed arrow) D internal close-up showing apical pore field depression (black arrow) and striae depression (dashed line) E close-up on valve apex showing flaps on apical pore field (black arrow) and open girdle element (white arrow) F close-up on cell-cell connection showing apical pore field covered with flaps (black arrow) and open girdle element (white arrow). Scale bars: 2 µm (D, E); 3 µm (F); 5 µm (A-C)." figureDoi="10.3897/phytokeys.187.73338.figure9" httpUri="https://binary.pensoft.net/fig/622446" pageId="0" pageNumber="23">9C, D</figureCitation>
). Well-developed volae, arising from the areolar inner periphery and projecting inwards (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-F SEM images of Pseudostaurosira occulta sp. nov. A, B external views of valves showing apical pore fields (black arrow in A) and other features C internal view of valve showing depressions containing the striae (dashed arrow) D internal close-up showing apical pore field depression (black arrow) and striae depression (dashed line) E close-up on valve apex showing flaps on apical pore field (black arrow) and open girdle element (white arrow) F close-up on cell-cell connection showing apical pore field covered with flaps (black arrow) and open girdle element (white arrow). Scale bars: 2 µm (D, E); 3 µm (F); 5 µm (A-C)." figureDoi="10.3897/phytokeys.187.73338.figure9" httpUri="https://binary.pensoft.net/fig/622446" pageId="0" pageNumber="23">9B-D</figureCitation>
). Flaps small and present on some valve face areolae (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-F SEM images of Pseudostaurosira occulta sp. nov. A, B external views of valves showing apical pore fields (black arrow in A) and other features C internal view of valve showing depressions containing the striae (dashed arrow) D internal close-up showing apical pore field depression (black arrow) and striae depression (dashed line) E close-up on valve apex showing flaps on apical pore field (black arrow) and open girdle element (white arrow) F close-up on cell-cell connection showing apical pore field covered with flaps (black arrow) and open girdle element (white arrow). Scale bars: 2 µm (D, E); 3 µm (F); 5 µm (A-C)." figureDoi="10.3897/phytokeys.187.73338.figure9" httpUri="https://binary.pensoft.net/fig/622446" pageId="0" pageNumber="23">9A, B</figureCitation>
), more commonly on larger mantle areolae (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-F SEM images of Pseudostaurosira occulta sp. nov. A, B external views of valves showing apical pore fields (black arrow in A) and other features C internal view of valve showing depressions containing the striae (dashed arrow) D internal close-up showing apical pore field depression (black arrow) and striae depression (dashed line) E close-up on valve apex showing flaps on apical pore field (black arrow) and open girdle element (white arrow) F close-up on cell-cell connection showing apical pore field covered with flaps (black arrow) and open girdle element (white arrow). Scale bars: 2 µm (D, E); 3 µm (F); 5 µm (A-C)." figureDoi="10.3897/phytokeys.187.73338.figure9" httpUri="https://binary.pensoft.net/fig/622446" pageId="0" pageNumber="23">9D-F</figureCitation>
). Spines originating from vimines at the valve face/mantle junction; solid, with round to elliptical base (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-F SEM images of Pseudostaurosira occulta sp. nov. A, B external views of valves showing apical pore fields (black arrow in A) and other features C internal view of valve showing depressions containing the striae (dashed arrow) D internal close-up showing apical pore field depression (black arrow) and striae depression (dashed line) E close-up on valve apex showing flaps on apical pore field (black arrow) and open girdle element (white arrow) F close-up on cell-cell connection showing apical pore field covered with flaps (black arrow) and open girdle element (white arrow). Scale bars: 2 µm (D, E); 3 µm (F); 5 µm (A-C)." figureDoi="10.3897/phytokeys.187.73338.figure9" httpUri="https://binary.pensoft.net/fig/622446" pageId="0" pageNumber="23">9A, B</figureCitation>
), as wide as the vimines they sit on (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-F SEM images of Pseudostaurosira occulta sp. nov. A, B external views of valves showing apical pore fields (black arrow in A) and other features C internal view of valve showing depressions containing the striae (dashed arrow) D internal close-up showing apical pore field depression (black arrow) and striae depression (dashed line) E close-up on valve apex showing flaps on apical pore field (black arrow) and open girdle element (white arrow) F close-up on cell-cell connection showing apical pore field covered with flaps (black arrow) and open girdle element (white arrow). Scale bars: 2 µm (D, E); 3 µm (F); 5 µm (A-C)." figureDoi="10.3897/phytokeys.187.73338.figure9" httpUri="https://binary.pensoft.net/fig/622446" pageId="0" pageNumber="23">9A, B, E</figureCitation>
); with cylindrical body and shallow concave sides (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-F SEM images of Pseudostaurosira occulta sp. nov. A, B external views of valves showing apical pore fields (black arrow in A) and other features C internal view of valve showing depressions containing the striae (dashed arrow) D internal close-up showing apical pore field depression (black arrow) and striae depression (dashed line) E close-up on valve apex showing flaps on apical pore field (black arrow) and open girdle element (white arrow) F close-up on cell-cell connection showing apical pore field covered with flaps (black arrow) and open girdle element (white arrow). Scale bars: 2 µm (D, E); 3 µm (F); 5 µm (A-C)." figureDoi="10.3897/phytokeys.187.73338.figure9" httpUri="https://binary.pensoft.net/fig/622446" pageId="0" pageNumber="23">9E, F</figureCitation>
), and spatulate tip with lateral projections and a serrate border pattern (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-F SEM images of Pseudostaurosira occulta sp. nov. A, B external views of valves showing apical pore fields (black arrow in A) and other features C internal view of valve showing depressions containing the striae (dashed arrow) D internal close-up showing apical pore field depression (black arrow) and striae depression (dashed line) E close-up on valve apex showing flaps on apical pore field (black arrow) and open girdle element (white arrow) F close-up on cell-cell connection showing apical pore field covered with flaps (black arrow) and open girdle element (white arrow). Scale bars: 2 µm (D, E); 3 µm (F); 5 µm (A-C)." figureDoi="10.3897/phytokeys.187.73338.figure9" httpUri="https://binary.pensoft.net/fig/622446" pageId="0" pageNumber="23">9D-F</figureCitation>
). Stipules incipient and subtending a small circular depression on the spine upper body (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-F SEM images of Pseudostaurosira occulta sp. nov. A, B external views of valves showing apical pore fields (black arrow in A) and other features C internal view of valve showing depressions containing the striae (dashed arrow) D internal close-up showing apical pore field depression (black arrow) and striae depression (dashed line) E close-up on valve apex showing flaps on apical pore field (black arrow) and open girdle element (white arrow) F close-up on cell-cell connection showing apical pore field covered with flaps (black arrow) and open girdle element (white arrow). Scale bars: 2 µm (D, E); 3 µm (F); 5 µm (A-C)." figureDoi="10.3897/phytokeys.187.73338.figure9" httpUri="https://binary.pensoft.net/fig/622446" pageId="0" pageNumber="23">9D, F</figureCitation>
). Apical pore fields covered by external flaps (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-F SEM images of Pseudostaurosira occulta sp. nov. A, B external views of valves showing apical pore fields (black arrow in A) and other features C internal view of valve showing depressions containing the striae (dashed arrow) D internal close-up showing apical pore field depression (black arrow) and striae depression (dashed line) E close-up on valve apex showing flaps on apical pore field (black arrow) and open girdle element (white arrow) F close-up on cell-cell connection showing apical pore field covered with flaps (black arrow) and open girdle element (white arrow). Scale bars: 2 µm (D, E); 3 µm (F); 5 µm (A-C)." figureDoi="10.3897/phytokeys.187.73338.figure9" httpUri="https://binary.pensoft.net/fig/622446" pageId="0" pageNumber="23">9A, E, F</figureCitation>
). Internally, apical pore field opening into roundish depression, revealing several rows of round poroids (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-F SEM images of Pseudostaurosira occulta sp. nov. A, B external views of valves showing apical pore fields (black arrow in A) and other features C internal view of valve showing depressions containing the striae (dashed arrow) D internal close-up showing apical pore field depression (black arrow) and striae depression (dashed line) E close-up on valve apex showing flaps on apical pore field (black arrow) and open girdle element (white arrow) F close-up on cell-cell connection showing apical pore field covered with flaps (black arrow) and open girdle element (white arrow). Scale bars: 2 µm (D, E); 3 µm (F); 5 µm (A-C)." figureDoi="10.3897/phytokeys.187.73338.figure9" httpUri="https://binary.pensoft.net/fig/622446" pageId="0" pageNumber="23">9D</figureCitation>
). Well-developed blister-like depositions present on abvalvar edge of mantle also covering both apices (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-F SEM images of Pseudostaurosira occulta sp. nov. A, B external views of valves showing apical pore fields (black arrow in A) and other features C internal view of valve showing depressions containing the striae (dashed arrow) D internal close-up showing apical pore field depression (black arrow) and striae depression (dashed line) E close-up on valve apex showing flaps on apical pore field (black arrow) and open girdle element (white arrow) F close-up on cell-cell connection showing apical pore field covered with flaps (black arrow) and open girdle element (white arrow). Scale bars: 2 µm (D, E); 3 µm (F); 5 µm (A-C)." figureDoi="10.3897/phytokeys.187.73338.figure9" httpUri="https://binary.pensoft.net/fig/622446" pageId="0" pageNumber="23">9D-F</figureCitation>
). Girdle elements variable in number, open, lacking pores, ligulated, with larger valvocopula (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-F SEM images of Pseudostaurosira occulta sp. nov. A, B external views of valves showing apical pore fields (black arrow in A) and other features C internal view of valve showing depressions containing the striae (dashed arrow) D internal close-up showing apical pore field depression (black arrow) and striae depression (dashed line) E close-up on valve apex showing flaps on apical pore field (black arrow) and open girdle element (white arrow) F close-up on cell-cell connection showing apical pore field covered with flaps (black arrow) and open girdle element (white arrow). Scale bars: 2 µm (D, E); 3 µm (F); 5 µm (A-C)." figureDoi="10.3897/phytokeys.187.73338.figure9" httpUri="https://binary.pensoft.net/fig/622446" pageId="0" pageNumber="23">9E, F</figureCitation>
).
</paragraph>
<caption doi="10.3897/phytokeys.187.73338.figure9" httpUri="https://binary.pensoft.net/fig/622446" pageId="0" pageNumber="23" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="23">
<emphasis bold="true" pageId="0" pageNumber="23">Figure 9.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="23">A-F</emphasis>
SEM images of
<taxonomicName authorityName="E. Morales, C. E. Wetzel &amp; Ector" authorityYear="2021" class="Fragilariophyceae" family="Fragilariaceae" genus="Pseudostaurosira" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Pseudostaurosira occulta" order="Fragilariales" pageId="0" pageNumber="23" phylum="Bacillariophyta" rank="species" species="occulta">
<emphasis italics="true" pageId="0" pageNumber="23">Pseudostaurosira occulta</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="23">A, B</emphasis>
external views of valves showing apical pore fields (black arrow in A) and other features
<emphasis bold="true" pageId="0" pageNumber="23">C</emphasis>
internal view of valve showing depressions containing the striae (dashed arrow)
<emphasis bold="true" pageId="0" pageNumber="23">D</emphasis>
internal close-up showing apical pore field depression (black arrow) and striae depression (dashed line)
<emphasis bold="true" pageId="0" pageNumber="23">E</emphasis>
close-up on valve apex showing flaps on apical pore field (black arrow) and open girdle element (white arrow)
<emphasis bold="true" pageId="0" pageNumber="23">F</emphasis>
close-up on cell-cell connection showing apical pore field covered with flaps (black arrow) and open girdle element (white arrow). Scale bars: 2
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="23">D, E</emphasis>
); 3
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="23">F</emphasis>
); 5
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="23">A-C</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="23">
Dimensions (n&gt; 30): Length 6.7-35.6
<normalizedToken originalValue="μm">μm</normalizedToken>
; width 3.3-3.8
<normalizedToken originalValue="μm">μm</normalizedToken>
; striae 14-16 in 10
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="etymology">
<paragraph pageId="0" pageNumber="23">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="23">The species epithet alludes to the fact that this diatom has remained unidentified thus far and has been confused with morphologically similar taxa (see Discussion).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="23" type="distribution">
<paragraph pageId="0" pageNumber="23">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="23">Found in the Sajama River.</paragraph>
</subSubSection>
</treatment>
</document>