treatments-xml/data/93/BB/21/93BB2111CE6F022ED76A3F804C344D57.xml
2024-06-21 12:44:20 +02:00

391 lines
34 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.701.13261" ID-GBIF-Dataset="a9b15909-ea61-40d7-ba04-325b8af91669" ID-PMC="PMC5674090" ID-Pensoft-Pub="1313-2970-701-1" ID-PubMed="29118596" ID-ZBK="23BDD7CE1C7E4D2092A8ED47267579FD" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-701-1" ModsDocOrigin="ZooKeys 701" ModsDocTitle="Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species" checkinTime="1506095869158" checkinUser="pensoft" docAuthor="Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A. &amp; Dowling, Ashley P. G." docDate="2017" docId="93BB2111CE6F022ED76A3F804C344D57" docLanguage="en" docName="ZooKeys 701: 1-496" docOrigin="ZooKeys 701" docSource="http://dx.doi.org/10.3897/zookeys.701.13261" docTitle="Torrenticola sierrensis Marshall 1943" docType="treatment" docVersion="4" lastPageNumber="1" masterDocId="3D440132FFABFFFC1505FFA0FFDDEA09" masterDocTitle="Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species" masterLastPageNumber="496" masterPageNumber="1" pageNumber="1" updateTime="1668164828938" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, J. Ray</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, Danielle M.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Skvarla, Michael J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nelson, Whitney A.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dowling, Ashley P. G.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>701</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>496</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.701.13261</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.701.13261</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-701-1</mods:identifier>
<mods:identifier type="ZBK">23BDD7CE1C7E4D2092A8ED47267579FD</mods:identifier>
<mods:identifier type="ZooBank">23BDD7CE1C7E4D2092A8ED47267579FD</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="134839644" LSID="urn:lsid:plazi:treatment:93BB2111CE6F022ED76A3F804C344D57" httpUri="http://treatment.plazi.org/id/93BB2111CE6F022ED76A3F804C344D57" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authority="Marshall, 1943" authorityName="Marshall" authorityYear="1943" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola sierrensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sierrensis">Torrenticola sierrensis (Marshall, 1943)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Atractides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractides sierrensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sierrensis">Atractides sierrensis</taxonomicName>
Marshall, 1943: 307.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Atractides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atractides mercedensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mercedensis">Atractides mercedensis</taxonomicName>
Marshall, 1943: 310.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola sierrensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sierrensis">Torrenticola sierrensis</taxonomicName>
Mitchell, 1954: 40.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">LECTOTYPE (1 ♀): USA, California, Santa Cruz County, Waddell Creek, 25-26 Aug 1932, by PR Needham, RM320007</paragraph>
<paragraph pageId="0" pageNumber="1">PARALECTOTYPE (1 ♂): California, Santa Cruz County, Waddell Creek, 1 Dec 1932, by PR Needham, RM320008.</paragraph>
<paragraph pageId="0" pageNumber="1">TOPOTYPES (1 ♀; 1 ♂): from USA, California, Santa Cruz County, Waddell Creek, 30-31 Aug 1933, by PR Needham, RM330016.</paragraph>
<paragraph pageId="0" pageNumber="1">
OTHER MATERIAL (54 ♀; 34 ♂): British Columbia, Canada: 3 ♀ and 1 ♂ from Ryan Rest Area off Highway 3, east of Yahk Moyie River, 15 Aug 2012, by IM Smith, IMS120071
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ from Vancouver Island, Lake Cowichan, Cowichan River, above Skutz Falls, 9 Jul 1979, by IM Smith, IMS790035
<normalizedToken originalValue="•">*</normalizedToken>
California, USA: 1 ♀ from Del Norte County, Six Rivers National Forest, Middle Fork Smith River (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="41.855553">41°51'20&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-123.88611">123°53'10&quot;W</geoCoordinate>
), 15 Aug 2013, by JR Fisher, JRF 13-0815-002
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ from El Dorado County, Upper Truckee River (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="38.84889">38°50'56&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-120.027504">120°1'39&quot;W</geoCoordinate>
), 29 Aug 2013, by JR Fisher, JRF 13-0829-003
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ and 1 ♂ from El Dorado County, Upper Truckee River (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="38.84889">38°50'56&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-120.027504">120°1'39&quot;W</geoCoordinate>
), 29 Aug 2013, by JR Fisher, JRF 13-0829-004
<normalizedToken originalValue="•">*</normalizedToken>
2 ♀ and 2 ♂ from Mariposa County, El Portal, Indian Flat campground, Merced River, 9-10 Jun 1976, by IM Smith, IMS760087
<normalizedToken originalValue="•">*</normalizedToken>
2 ♀ from Mariposa County, Yosemite Valley, East Fork of Merced River, 22 Aug 1933, by PR Needham, RM330012
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ from Mendocino County, Jackson Demonstration State Park, North Fork of Big River (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="39.34611">39°20'46&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-123.50972">123°30'35&quot;W</geoCoordinate>
), 22 Aug 2013, by JR Fisher, JRF 13-0822-002
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ from Mendocino County, Paul M. Dimmick Recreation Area, North Fork of Navarro River, beside Route 128, 4 Aug 1987, by IM Smith, IMS870127A
<normalizedToken originalValue="•">*</normalizedToken>
1 ♂ from Mendocino County, Rancheria Creek, beside Route 128, 7.3 kilometers south of Boonville, 4 Aug 1987, by IM Smith, IMS870126A
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ and 1 ♂ from Monterey County, Big Sur River, beside Route 1 near Pfeiffer-Big Sur State Park, 28-29 Jul 1987, by IM Smith, IMS870116A
<normalizedToken originalValue="•">*</normalizedToken>
3 ♀ from Monterey County, Big Sur, Pfeiffer-Big Sure State Park, Big Sur Creek, 7 Jun 1976, by IM Smith, IMS760074
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ from Monterey County, Los Padres National Forest, Salmon Creek (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="35.815834">35°48'57&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-121.358055">121°21'29&quot;W</geoCoordinate>
), 6 Sep 2013, by JR Fisher, JRF 13-0906-003
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ from Monterey County, Pfeiffer State Park, Big Sur River (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="36.245">36°14'42&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-121.77862">121°46'43&quot;W</geoCoordinate>
), 4 Sep 2013, by JR Fisher, JRF 13-0904-004
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ and 1 ♂ from Monterey County, Salmon Creek, beside Route 1, south of Gorda, 28 Jul 1987, by IM Smith, IMS870114A
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ from Nevada County, beside Route 89, north of Hobart Mills, 13 Jun 1976, by IM Smith, IMS760109
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ from Nevada County, Tahoe National Forest, Sagehen Creek (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="39.43389">39°26'2&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-120.20472">120°12'17&quot;W</geoCoordinate>
), 26 Aug 2013, by JR Fisher, JRF 13-0826-006
<normalizedToken originalValue="•">*</normalizedToken>
2 ♀ from Trinity County, Shasta-Trinity National Forest, Wilson Creek (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="40.42139">40°25'17&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-123.05139">123°3'5&quot;W</geoCoordinate>
), 20 Aug 2013, by JR Fisher, JRF 13-0820-003
<normalizedToken originalValue="•">*</normalizedToken>
1 ♂ from Shasta County, Battle Creek, beside Route 44, 5.6 kilometers west of Viola, 10 Aug 1987, by IM Smith, IMS870139A
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ from Trinity County, South Fork of Trinity River, beside Route 36 at Forest Glen campground, 6 Aug 1987, by IM Smith, IMS870131
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ and 1 ♂ from Tulare County, Brush Creek, Brush Creek Flat, beside SM99 between Roads End Station &amp; Johnsondale, 31 Jul 1987, by IM Smith, IMS870122
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ and 3 ♂ from Tulare County, Kern River, Brush Creek Flat, beside SM99 between Roads End Station &amp; Johnsondale, 31 Jul 1987, by IM Smith, IMS870121
<normalizedToken originalValue="•">*</normalizedToken>
1 ♂ from Tulare County, Stony Creek, Stony Creek Picnic Area, east of Sequoia National Park, 1 Aug 1987, by IM Smith, IMS870124A
<normalizedToken originalValue="•">*</normalizedToken>
Idaho, USA: 1 ♀ and 1 ♂ from Custer County, Challis National Forest, Stanley Creek (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="44.253334">44°15'12&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-115.00528">115°0'19&quot;W</geoCoordinate>
), 30 Jul 2012, by JR Fisher, WA Nelson, &amp; JC
<normalizedToken originalValue="ONeill">O'Neill</normalizedToken>
, ROW 12-0730-005
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ and 1 ♂ from Custer County, Salmon River at picnic area, beside Route 93, north of Morgan Creek, 2 Jul 1985, by IM Smith, IMS850064
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ from Fremont County, Targhee National Forest, Rock Creek (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="44.11222">44°6'44&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-111.251114">111°15'4&quot;W</geoCoordinate>
), by JR Fisher, WA Nelson, &amp; JC
<normalizedToken originalValue="ONeill">O'Neill</normalizedToken>
, ROW 12-0725-001
<normalizedToken originalValue="•">*</normalizedToken>
Montana, USA: 1 ♀ from Ravalli County, Bitterroot National Forest, West Fork Bitterroot River (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="45.910557">45°54'38&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-114.16195">114°9'43&quot;W</geoCoordinate>
), 6 Aug 2012, by JR Fisher, WA Nelson, &amp; JC
<normalizedToken originalValue="ONeill">O'Neill</normalizedToken>
, ROW 12-0806-003
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ from Ravalli County, Medicine Springs, Spring Gulch campground, East Fork of Bitterroot River, beside Route 93, 1 Jul 1985, by IM Smith, IMS850060
<normalizedToken originalValue="•">*</normalizedToken>
Nevada, USA: 1 ♀ and 1 ♂ from Humboldt County, Paradise Valley, Cabin Creek beside road, 10.7 kilometers north of Hinkey Summit, 11 Aug 1987, by IM Smith, IMS870141A
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ and 2 ♂ from Humboldt County, Paradise Valley, Dutch John Creek beside road, 8.3 kilometers north of Hinkey Summit, 11 Aug 1987, by IM Smith, IMS870142A
<normalizedToken originalValue="•">*</normalizedToken>
Oregon, USA: 1 ♀ and 2 ♂ from Coos County, Siskiyou National Forest, Road 33 between Powers &amp; Agness, 2 Jul 1983, by IM Smith, IMS830015
<normalizedToken originalValue="•">*</normalizedToken>
2 ♀ from Curry County, Port Orford, Butler Bar campground, Elk River, 25 Jun 1976, by IM Smith, IMS760162
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ and 1 ♂ from Curry County, Port Orford, Humbug Mountain State Park Picnic Area, beside Route 1, Brush Creek, 1 Jul 1983, by IM Smith, IMS830012
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ and 2 ♂ from Curry County, Port Orford, Humbug Mountain State Park Picnic Area, Brush Creek, beside Route 1, 3 Jul 1983, by IM Smith, IMS830020B
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ from Curry County, Quosatana Creek (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="42.489166">42°29'21&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-124.23389">124°14'2&quot;W</geoCoordinate>
), 14 Aug 2013, by JR Fisher, JRF 13-0814-003
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ from Curry County, Rogue River National Forest, Elk River (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="42.71278">42°42'46&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-124.31139">124°18'41&quot;W</geoCoordinate>
), 13 Aug 2013, by JR Fisher, JRF 13-0813-003
<normalizedToken originalValue="•">*</normalizedToken>
3 ♀ from Lane County, Gate Creek (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="44.146667">44°8'48&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-122.57222">122°34'20&quot;W</geoCoordinate>
), 11 Aug 2013, by JC
<normalizedToken originalValue="ONeill">O'Neill</normalizedToken>
, &amp; WA Nelson, JNOW 13-0811-001
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ and 1 ♂ from Curry County, Sixes, Sixes River, beside road at mouth of Edson Creek, 4 Jul 1983, by IM Smith, IMS830021B
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ and 1 ♂ from Lincoln County, Blackberry campground near Tidewater, Alsea River, 28 Jun 1983, by IM Smith, IMS830009
<normalizedToken originalValue="•">*</normalizedToken>
2 ♀ and 2 ♂ from Lincoln County, Blackberry campground near Tidewater, Alsea River, 29 Jun 1983, by IM Smith, IMS830010
<normalizedToken originalValue="•">*</normalizedToken>
2 ♀ and 2 ♂ from Lincoln County, Siuslaw National Forest, Five Rivers Creek (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="44.331387">44°19'53&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-123.849724">123°50'59&quot;W</geoCoordinate>
), 8 Aug 2013, by JC
<normalizedToken originalValue="ONeill">O'Neill</normalizedToken>
, &amp; WA Nelson, JNOW 13-0808-001
<normalizedToken originalValue="•">*</normalizedToken>
Washington, USA: 2 ♀ and 2 ♂ from Cowlitz County (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="46.373333">46°22'24&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-122.57916">122°34'45&quot;W</geoCoordinate>
), 16 Jul 2013, by JC
<normalizedToken originalValue="ONeill">O'Neill</normalizedToken>
, &amp; WA Nelson, JNOW 13-0716-001
<normalizedToken originalValue="•">*</normalizedToken>
1 ♀ and 1 ♂ from Grays Harbor County, Stewarts Creek (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="47.26361">47°15'49&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-123.92">123°55'12&quot;W</geoCoordinate>
), 25 Jul 2013, by JC
<normalizedToken originalValue="ONeill">O'Neill</normalizedToken>
, &amp; WA Nelson, JNOW 13-0725-001
<normalizedToken originalValue="•">*</normalizedToken>
Wyoming, USA: 1 ♀ and 2 ♂ from Sublette County, Boulder Creek, beside Route 191, just north of Boulder, 15 Aug 1987, by IM Smith, IMS870151A &amp; IMS870151B.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type deposition">
<paragraph pageId="0" pageNumber="1">Type deposition.</paragraph>
<paragraph pageId="0" pageNumber="1">Types (1 ♀; 1 ♂) deposited in the CNC.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola sierrensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sierrensis">Torrenticola sierrensis</taxonomicName>
are similar to other members of the Tricolor Complex (
<taxonomicName lsidName="T. bittikoferae" pageId="0" pageNumber="1" rank="species" species="bittikoferae">T. bittikoferae</taxonomicName>
,
<taxonomicName lsidName="T. hoosieri" pageId="0" pageNumber="1" rank="species" species="hoosieri">T. hoosieri</taxonomicName>
,
<taxonomicName lsidName="T. larvata" pageId="0" pageNumber="1" rank="species" species="larvata">T. larvata</taxonomicName>
,
<taxonomicName lsidName="T. pearsoni" pageId="0" pageNumber="1" rank="species" species="pearsoni">T. pearsoni</taxonomicName>
,
<taxonomicName lsidName="T. olliei" pageId="0" pageNumber="1" rank="species" species="olliei">T. olliei</taxonomicName>
,
<taxonomicName lsidName="T. tricolor" pageId="0" pageNumber="1" rank="species" species="tricolor">T. tricolor</taxonomicName>
,
<taxonomicName lsidName="T. trimaculata" pageId="0" pageNumber="1" rank="species" species="trimaculata">T. trimaculata</taxonomicName>
,
<taxonomicName lsidName="T. unimaculata" pageId="0" pageNumber="1" rank="species" species="unimaculata">T. unimaculata</taxonomicName>
,
<taxonomicName lsidName="T. cardia" pageId="0" pageNumber="1" rank="species" species="cardia">T. cardia</taxonomicName>
,
<taxonomicName lsidName="T. kringi" pageId="0" pageNumber="1" rank="species" species="kringi">T. kringi</taxonomicName>
,
<taxonomicName lsidName="T. dimorpha" pageId="0" pageNumber="1" rank="species" species="dimorpha">T. dimorpha</taxonomicName>
, and
<taxonomicName lsidName="T. mohawk" pageId="0" pageNumber="1" rank="species" species="mohawk">T. mohawk</taxonomicName>
) in having a short, conical rostrum.
<taxonomicName lsidName="T. sierrensis" pageId="0" pageNumber="1" rank="species" species="sierrensis">T. sierrensis</taxonomicName>
can be differentiated from most Tricolor Complex by being distributed in the west and having a wider genital field (♀ = 180-213 in
<taxonomicName lsidName="T. sierrensis" pageId="0" pageNumber="1" rank="species" species="sierrensis">T. sierrensis</taxonomicName>
, 145-180 in others; ♂ = 135-175 in
<taxonomicName lsidName="T. sierrensis" pageId="0" pageNumber="1" rank="species" species="sierrensis">T. sierrensis</taxonomicName>
, 92-120 in others), except for
<taxonomicName lsidName="T. olliei" pageId="0" pageNumber="1" rank="species" species="olliei">T. olliei</taxonomicName>
(♀ = 190-203, ♂ = 130-138).
<taxonomicName lsidName="T. sierrensis" pageId="0" pageNumber="1" rank="species" species="sierrensis">T. sierrensis</taxonomicName>
can be differentiated from
<taxonomicName lsidName="T. olliei" pageId="0" pageNumber="1" rank="species" species="olliei">T. olliei</taxonomicName>
by having a more elongate rostrum (length/width = 1.95-2.14 in
<taxonomicName lsidName="T. sierrensis" pageId="0" pageNumber="1" rank="species" species="sierrensis">T. sierrensis</taxonomicName>
, 1.56-1.81 in
<taxonomicName lsidName="T. olliei" pageId="0" pageNumber="1" rank="species" species="olliei">T. olliei</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Re-description.</paragraph>
<paragraph pageId="0" pageNumber="1">Female (Figure 241) (n = 9) (holotype measurements in parentheses when available) with characters of the genus with following specifications.</paragraph>
<paragraph pageId="0" pageNumber="1">Dorsum - (700-880 (865) long; 550-740 (710) wide) circular with orange or reddish purple coloration separated into anterior and posterior portions with faint orange medially (occasionally colorless). Anterio-medial platelets (162.5-197.5 (187.5) long; 65-82.5 (81.25) wide). Anterio-lateral platelets (192.5-250 (225) long; 82.5-100 (92.5) wide) free from dorsal plate. Dgl-4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl-4 435-520 (500)). Dorsal plate proportions: dorsum length/width 1.19-1.27 (1.22); dorsal width/distance between Dgl-4 1.22-1.43 (1.42); anterio-medial platelet length/width 2.10-2.50 (2.31); anterio-lateral platelet length/width 2.33-2.50 (2.43); anterio-lateral/anterio-medial length 1.18-1.27 (1.20).</paragraph>
<paragraph pageId="0" pageNumber="1">Gnathosoma - Subcapitulum 320-365 (365) long (ventral); 227-265 (260) long (dorsal); 140-165 (152.5) tall) colorless. Rostrum (120-140 (140) long; 60-70 (70) wide) short and conical. Chelicerae (320-355 long) with curved fangs (61-77.5 long). Subcapitular proportions: ventral length/height 2.19-2.39 (2.39); rostrum length/width 1.96-2.08 (2.00). Pedipalps with tuberculate ventral extensions with dentate tips on femora and genua. Palpomeres: trochanter (45-55 (55) long); femur (115-135 (135) long); genu (80-92.5 (92.5) long); tibia (107.5-117.5 (117.5) long; 32.5-35 (35) wide); tarsus (21.25-35 32.5) long). Palpomere proportions: femur/genu 1.35-1.55 (1.46); tibia/femur 0.86-0.95 (0.87); tibia length/width 3.07-3.48 (3.36).</paragraph>
<paragraph pageId="0" pageNumber="1">Venter - (775-1040 (1015) long; 619-820 (770) wide) colorless. Gnathosomal bay (155-205 (195) long; 95-112.5 (112.5) wide). Cxgl-4 subapical. Medial suture (17.5-37.5 (22.5) long). Genital plates (210-235 (222.5) long; 180-212.5 (212.5) wide). Additional measurements: Cx-1 (192.5-370 (370) long (total); 136-185 (172.5) long (medial)); Cx-3 (250-475 (475) wide); anterior venter (192.5-237.5 (212.5) long). Ventral proportions: gnathosomal bay length/width 1.50-1.91 (1.73); anterior venter/genital field length 0.90-1.06 (0.96); anterior venter length/genital field width 1.00-1.13 (1.00); anterior venter/medial suture 6.33-12.14 (9.44).</paragraph>
<paragraph pageId="0" pageNumber="1">Male (Figure 242) (n = 7) (allotypic measurements in parentheses when available) with characters of the genus with following specifications.</paragraph>
<paragraph pageId="0" pageNumber="1">Dorsum - (590-735 (715) long; 460-590 (580) wide) circular with orange or reddish purple coloration separated into anterior and posterior portions with faint orange medially (occasionally colorless). Anterio-medial platelets (137.5-172.5 (172.5) long; 60-75 (75) wide). Anterio-lateral platelets (185-222.5 (215) long; 72.5-95 (92.5) wide) free from dorsal plate. Dgl-4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl-4 355-460 (460)). Dorsal plate proportions: dorsum length/width 1.17-1.28 (1.23); dorsal width/distance between Dgl-4 1.26-1.33 (1.26); anterio-medial platelet length/width 2.24-2.48 (2.30); anterio-lateral platelet length/width 2.12-2.55 (2.32); anterio-lateral/anterio-medial length 1.22-1.38 (1.25).</paragraph>
<paragraph pageId="0" pageNumber="1">Gnathosoma - Subcapitulum (270-330 (325) long (ventral); 200-235 (235) long (dorsal); 118.75-140 (135) tall) colorless. Rostrum (100-122.5 (122.5) long; 50-60 (60) wide) short and conical. Chelicerae (262-317.5 (315) long) with curved fangs (50-67.5 (60) long). Subcapitular proportions: ventral length/height 2.23-2.46 (2.41); rostrum length/width 1.95-2.14 (2.04). Pedipalps with tuberculate ventral extensions with dentate tips on femora and genua. Palpomeres: trochanter (40-50 (50) long); femur (92.5-120 (120) long); genu (65-80 (78.75) long); tibia (82.5-107.5 (102.5) long; 27.5-32.5 (32.5) wide); tarsus (25-32.5 (32.5) long). Palpomere proportions: femur/genu 1.42-1.52 (1.52); tibia/femur 0.85-0.95 (0.85); tibia length/width 3.00-3.31 (3.15).</paragraph>
<paragraph pageId="0" pageNumber="1">Venter - (690-895 (875) long; 538-667 (665) wide) colorless. Gnathosomal bay (112.5-180 (175) long; 80-100 (92.5) wide). Cxgl-4 subapical. Medial suture (72.5-92.5 (90) long). Genital plates (172.5-220 (217.5) long; 135-175 (175) wide). Additional measurements: Cx-1 (269-360 (360) long (total); 137-182.5 (182.5) long (medial)); Cx-3 (364-455 (455) wide); anterior venter (255-295 (290) long). Ventral proportions: gnathosomal bay length/width 1.41-1.89 (1.89); anterior venter/genital field length 1.32-1.51 (1.33); anterior venter length/genital field width 1.66-1.89 (1.66); anterior venter/medial suture 3.03-3.59 (3.22).</paragraph>
<paragraph pageId="0" pageNumber="1">Immatures unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Although
<bibRefCitation author="Marshall, R" pageId="0" pageNumber="1" url="http://www.jstor.org/stable/3223036" year="1943">Marshall (1943)</bibRefCitation>
did not specify an etymology for the specific epithet (
<taxonomicName class="Arachnida" family="Arrenuridae" genus="Lebertioidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lebertioidea sierrensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sierrensis">sierrensis</taxonomicName>
), surely it refers to the type locality of this species within the Sierra Nevada in California.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Western (Figure 240), but likely not found in the southwest.
<taxonomicName lsidName="T. sierrensis" pageId="0" pageNumber="1" rank="species" species="sierrensis">T. sierrensis</taxonomicName>
was previously known only from a few localities in California; we extend the range in western North America, but not in the southwest.
</paragraph>
<caption pageId="0" pageNumber="1">
<paragraph pageId="0" pageNumber="1">
Figure 240.
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola sierrensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sierrensis">Torrenticola sierrensis</taxonomicName>
distribution.
</paragraph>
</caption>
<caption pageId="0" pageNumber="1">
<paragraph pageId="0" pageNumber="1">
Figure 241.
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola sierrensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sierrensis">Torrenticola sierrensis</taxonomicName>
female: A dorsal plates B venter (legs removed) C subcapitulum D pedipalp (setae not accurately depicted). Scale = 100
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
<caption pageId="0" pageNumber="1">
<paragraph pageId="0" pageNumber="1">
Figure 242.
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola sierrensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sierrensis">Torrenticola sierrensis</taxonomicName>
male: A dorsal plates B venter (legs removed) C subcapitulum D pedipalp (setae not accurately depicted). Scale = 100
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola sierrensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sierrensis">Torrenticola sierrensis</taxonomicName>
group with other members of the Tricolor Complex with high support in all analyses and group with
<taxonomicName lsidName="T. olliei" pageId="0" pageNumber="1" rank="species" species="olliei">T. olliei</taxonomicName>
to form the western portion of this complex. Specimens within this complex are 0-4.5% different in COI sequence from each other and greater than 6% different from
<taxonomicName lsidName="T. olliei" pageId="0" pageNumber="1" rank="species" species="olliei">T. olliei</taxonomicName>
. This is higher sequence variability in COI than in most species hypotheses presented herein. However, given the topology in the COI tree (Figure 10) and morphological similarity, it seems apparent that the variability represents a continuum across a large distribution, rather than isolated species. This species hypothesis is supported by phylogenetic affinity and by the morphological characters outlined in the diagnosis.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Upon examining the types of
<taxonomicName lsidName="T. sierrensis" pageId="0" pageNumber="1" rank="species" species="sierrensis">T. sierrensis</taxonomicName>
and
<taxonomicName lsidName="T. mercedensis" pageId="0" pageNumber="1" rank="species" species="mercedensis">T. mercedensis</taxonomicName>
(Marshall, 1943), all characters for both species overlap with members of only one clade in our analyses. Therefore, it is apparent that these represent the same species hypothesis and must be synonymized. As First Revisers (ICZN Article 24.2), we select &quot;
<taxonomicName class="Arachnida" family="Arrenuridae" genus="Lebertioidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lebertioidea sierrensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sierrensis">sierrensis</taxonomicName>
&quot; as the senior synonym over &quot;
<taxonomicName class="Arachnida" family="Arrenuridae" genus="Lebertioidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lebertioidea mercedensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mercedensis">mercedensis</taxonomicName>
&quot; due to its broader range implication (&quot;
<taxonomicName class="Arachnida" family="Arrenuridae" genus="Lebertioidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lebertioidea mercedensis" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mercedensis">mercedensis</taxonomicName>
&quot; refers to the Merced River), even though
<taxonomicName lsidName="T. sierrensis" pageId="0" pageNumber="1" rank="species" species="sierrensis">T. sierrensis</taxonomicName>
is distributed more widely than just the Sierra Nevada ranges.
</paragraph>
</subSubSection>
</treatment>
</document>