589 lines
80 KiB
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589 lines
80 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/afrinvertebr.64.113133" ID-Pensoft-Pub="2305-2562-3-303" ID-Pensoft-UUID="1B4311FDBAF2573E8131BD435D38D7EF" ID-ZooBank="000F15D70DD746F7982625F13C43AF4C" ModsDocID="2305-2562-64-3-303" checkinTime="1702376364544" checkinUser="pensoft" docAuthor="Muller, Burgert S., Swart, Vaughn R. & Snyman, Louwrens P." docDate="2023" docId="5F22128A5B6F58F28D7195C12E30AA6B" docLanguage="en" docName="AfrInvert 64(3): 303-322" docOrigin="African Invertebrates 64 (3)" docPubDate="2023-12-11" docSource="http://dx.doi.org/10.3897/afrinvertebr.64.113133" docTitle="Atrichops adamastor Muller & Swart & Snyman 2023, comb. nov." docType="treatment" docVersion="1" id="1B4311FDBAF2573E8131BD435D38D7EF" lastPageNumber="303" masterDocId="1B4311FDBAF2573E8131BD435D38D7EF" masterDocTitle="Afrotropical Atrichops Verrall (Diptera, Athericidae) with description of a new species" masterLastPageNumber="322" masterPageNumber="303" pageNumber="303" updateTime="1702376364544" updateUser="pensoft">
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<mods:title id="3AA71B1B92937D1B37515A20225DE0ED">Afrotropical Atrichops Verrall (Diptera, Athericidae) with description of a new species</mods:title>
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<mods:namePart id="35E638B33979CDF5AB8F5A97D9DB2385">Muller, Burgert S.</mods:namePart>
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<mods:affiliation id="D464B59DFC443E03E14EB68403FEE9CC">Department Terrestrial Invertebrates, National Museum, 36 Aliwal Street, Bloemfontein, 9301, South Africa & Department Zoology & Entomology, University of the Free State, P. O. Box 339, Bloemfontein, 9300, South Africa</mods:affiliation>
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<mods:nameIdentifier id="635800F20D2CD221A450BE14C9724E85" type="email">burgert.muller@nasmus.co.za</mods:nameIdentifier>
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<mods:namePart id="BF94839CC12E44583932D1FEF435290B">Swart, Vaughn R.</mods:namePart>
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<mods:nameIdentifier id="21E9AE3CB88EF98460C25AB8EC23B074" type="ORCID">https://orcid.org/0000-0001-7905-5298</mods:nameIdentifier>
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<mods:affiliation id="F053A10DDF24ADDC5A97F0B58F14D30B">Department Zoology & Entomology, University of the Free State, P. O. Box 339, Bloemfontein, 9300, South Africa</mods:affiliation>
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<mods:namePart id="C823955610FA0ADDB59B08A418656D49">Snyman, Louwrens P.</mods:namePart>
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<mods:affiliation id="C7D317494EFA268C9EAF12D96D7AE7E8">Invertebrate Zoology, Royal Alberta Museum, 9810 103 A Avenue NW, Edmonton, Alberta, T 5 J 0 G 2, Canada</mods:affiliation>
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<mods:title id="5BB17B034AD1E8C95254EB950A6B404B">African Invertebrates</mods:title>
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<mods:date id="0BF0D7A273CC99FAA6C2CE54ECF997D5">2023</mods:date>
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<mods:number id="EBD5075DD35374EBBE5054E9BBC857FE">2023-12-11</mods:number>
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<treatment id="5F22128A5B6F58F28D7195C12E30AA6B" LSID="urn:lsid:plazi:treatment:5F22128A5B6F58F28D7195C12E30AA6B" httpUri="http://treatment.plazi.org/id/5F22128A5B6F58F28D7195C12E30AA6B" lastPageNumber="303" pageId="0" pageNumber="303">
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<taxonomicName id="5044960C9A47D5C3569788A69D934709" LSID="5F22128A-5B6F-58F2-8D71-95C12E30AA6B" authority="(Stuckenberg, 1960)" authorityName="Muller & Swart & Snyman" authorityYear="2023" baseAuthorityName="Stuckenberg" baseAuthorityYear="1960" class="Insecta" family="Athericidae" genus="Atrichops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichops adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor" status="comb. nov.">Atrichops adamastor (Stuckenberg, 1960)</taxonomicName>
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<taxonomicNameLabel id="95AC9C9F98D423B7521A9EA7EC53C7E3" pageId="0" pageNumber="303">comb. nov.</taxonomicNameLabel>
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<subSubSection id="B9B58F8A4DD0600CFD8F14111FFF2F32" pageId="0" pageNumber="303" type="description">
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<figureCitation id="986A5A2C328FF4B7EBF329DC517678BC" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Atrichops spp. ♂ 1 - 3 lateral habitus 4 - 6 head anterior view 1 A. adamastor (BMSA (D) 38908) 2 A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 3 A. stuckenbergi (BMSA (D) 95848) 4 A. adamastor (BMSA (D) 38904) 5 A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 6 A. stuckenbergi (BMSA (D) 95848). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures1-6" httpUri="https://binary.pensoft.net/fig/950648" pageId="0" pageNumber="303">Figs 1</figureCitation>
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<figureCitation id="FCE07A36FB1D1173493933815351B0A9" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Atrichops spp. ♂ 1 - 3 lateral habitus 4 - 6 head anterior view 1 A. adamastor (BMSA (D) 38908) 2 A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 3 A. stuckenbergi (BMSA (D) 95848) 4 A. adamastor (BMSA (D) 38904) 5 A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 6 A. stuckenbergi (BMSA (D) 95848). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures1-6" httpUri="https://binary.pensoft.net/fig/950648" pageId="0" pageNumber="303">, 4</figureCitation>
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<figureCitation id="E6C4CFF573543A953492D2A2A654166F" captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Atrichops spp. ♀ 7 - 9 lateral habitus 10 - 12 head anterior view 7 A. adamastor holotype and specimen labels (MZLU 00205685) 8 A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 9 A. stuckenbergi (CAS) 10 A. adamastor holotype (MZLU 00205685) 11 A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 12 A. stuckenbergi (BMSA (D) 95856). (7, 10 modified from originals provided by Rune Bygebjerg, MZLU, CC BY-NC 2.0 9, 12 modified from originals provided by Rikio Matsumoto, Osaka Museum of Natural History). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures7-12" httpUri="https://binary.pensoft.net/fig/950649" pageId="0" pageNumber="303">, 7</figureCitation>
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<figureCitation id="436A2843483208575215542D222748E4" captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Atrichops spp. ♀ 7 - 9 lateral habitus 10 - 12 head anterior view 7 A. adamastor holotype and specimen labels (MZLU 00205685) 8 A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 9 A. stuckenbergi (CAS) 10 A. adamastor holotype (MZLU 00205685) 11 A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 12 A. stuckenbergi (BMSA (D) 95856). (7, 10 modified from originals provided by Rune Bygebjerg, MZLU, CC BY-NC 2.0 9, 12 modified from originals provided by Rikio Matsumoto, Osaka Museum of Natural History). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures7-12" httpUri="https://binary.pensoft.net/fig/950649" pageId="0" pageNumber="303">, 10</figureCitation>
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<figureCitation id="0E4C2C70C08C73E1C5BBFFF7465AAAFE" captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Atrichops spp. ♂ ♀ dorsal view 13 ♂ A. adamastor (BMSA (D) 38908) 14 ♂ A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 15 ♂ A. stuckenbergi (BMSA (D) 94158) 16 ♀ A. adamastor holotype (MZLU 00205685) 17 ♀ A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 18 ♀ A. stuckenbergi holotype (CAS). (16 modified from original provided by Rune Bygebjerg, MZLU, CC BY-NC 2.0; 18 modified from original provided by Rikio Matsumoto, OMNH). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures13-18" httpUri="https://binary.pensoft.net/fig/950650" pageId="0" pageNumber="303">, 13</figureCitation>
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<figureCitation id="F615783C58E12D812C73CA03781B90C9" captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Atrichops spp. ♂ ♀ dorsal view 13 ♂ A. adamastor (BMSA (D) 38908) 14 ♂ A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 15 ♂ A. stuckenbergi (BMSA (D) 94158) 16 ♀ A. adamastor holotype (MZLU 00205685) 17 ♀ A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 18 ♀ A. stuckenbergi holotype (CAS). (16 modified from original provided by Rune Bygebjerg, MZLU, CC BY-NC 2.0; 18 modified from original provided by Rikio Matsumoto, OMNH). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures13-18" httpUri="https://binary.pensoft.net/fig/950650" pageId="0" pageNumber="303">, 16</figureCitation>
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<figureCitation id="6C94C62C412BC1B1453CC8E5F00C7250" captionStart="Figures 19–24" captionStartId="F4" captionText="Figures 19 - 24. Atrichops spp. ♂ ♀ proepimeral process, lateral view 19 ♂ A. adamastor (BMSA (D) 38904) 20 ♂ A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 21 ♂ A. stuckenbergi (BMSA (D) 94158) 22 ♀ A. adamastor (BMSA (D) 38899) 23 ♀ A. intermedius sp. nov. paratype (NMSA-Dip. 212861) 24 ♀ A. stuckenbergi holotype (CAS). (24 modified from original provided by Rikio Matsumoto, OMNH). Abbreviations: prepm - proepimeral process." figureDoi="10.3897/afrinvertebr.64.113133.figures19-24" httpUri="https://binary.pensoft.net/fig/950652" pageId="0" pageNumber="303">, 19</figureCitation>
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<figureCitation id="B9341AC7EED5A4BE818910FB4318987B" captionStart="Figures 19–24" captionStartId="F4" captionText="Figures 19 - 24. Atrichops spp. ♂ ♀ proepimeral process, lateral view 19 ♂ A. adamastor (BMSA (D) 38904) 20 ♂ A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 21 ♂ A. stuckenbergi (BMSA (D) 94158) 22 ♀ A. adamastor (BMSA (D) 38899) 23 ♀ A. intermedius sp. nov. paratype (NMSA-Dip. 212861) 24 ♀ A. stuckenbergi holotype (CAS). (24 modified from original provided by Rikio Matsumoto, OMNH). Abbreviations: prepm - proepimeral process." figureDoi="10.3897/afrinvertebr.64.113133.figures19-24" httpUri="https://binary.pensoft.net/fig/950652" pageId="0" pageNumber="303">, 22</figureCitation>
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<figureCitation id="3EC797A0504F9754A168186140467629" captionStart="Figures 25–30" captionStartId="F5" captionText="Figures 25 - 30. Atrichops spp. ♂ ♀ wings dorsal view, A. adamastor: 25 ♂ (BMSA (D) 38904) 26 ♀ (BMSA (D) 38904); A. intermedius sp. nov.: 27 ♂ paratype (NMSA-Dip. 81811) 28 ♀ paratype (NMSA-Dip. 212861) A. stuckenbergi: 29 ♂ (BMSA (D) 94161) 30 ♀ (BMSA (D) 95861). Abbreviations: br - basal radial cell; bm - basal medial cell; cua - anterior cubital cell; CuA + CuP - anterior branch of cubital vein + posterior branch of cubital vein; d - discal cell; M 1 - first branch of media; m 1 - first medial cell; M 2 - second branch of media; m 2 - second medial cell; M 3 - third branch of media; m 3 - third medial cell; R 1 - anterior branch of radius; r 1 - first radial cell; R 2 + 3 - second branch of radius; r 2 + 3 - second radial cell; R 4 - upper branch of third branch of radius; R 5 - lower branch of third branch of radius; Sc - subcostal vein; sc - subcostal cell. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures25-30" httpUri="https://binary.pensoft.net/fig/950653" pageId="0" pageNumber="303">, 25</figureCitation>
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<figureCitation id="A6D2272AB2D13823D51893DF54D82601" captionStart="Figures 25–30" captionStartId="F5" captionText="Figures 25 - 30. Atrichops spp. ♂ ♀ wings dorsal view, A. adamastor: 25 ♂ (BMSA (D) 38904) 26 ♀ (BMSA (D) 38904); A. intermedius sp. nov.: 27 ♂ paratype (NMSA-Dip. 81811) 28 ♀ paratype (NMSA-Dip. 212861) A. stuckenbergi: 29 ♂ (BMSA (D) 94161) 30 ♀ (BMSA (D) 95861). Abbreviations: br - basal radial cell; bm - basal medial cell; cua - anterior cubital cell; CuA + CuP - anterior branch of cubital vein + posterior branch of cubital vein; d - discal cell; M 1 - first branch of media; m 1 - first medial cell; M 2 - second branch of media; m 2 - second medial cell; M 3 - third branch of media; m 3 - third medial cell; R 1 - anterior branch of radius; r 1 - first radial cell; R 2 + 3 - second branch of radius; r 2 + 3 - second radial cell; R 4 - upper branch of third branch of radius; R 5 - lower branch of third branch of radius; Sc - subcostal vein; sc - subcostal cell. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures25-30" httpUri="https://binary.pensoft.net/fig/950653" pageId="0" pageNumber="303">, 26</figureCitation>
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<figureCitation id="0D9B82B71656016C9B22D137C73A53C4" captionStart="Figures 31–36" captionStartId="F6" captionText="Figures 31 - 36. Atrichops spp., 31 - 33 ♂ terminalia 31 A. adamastor (BMSA (D) 38904) 32 A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 33 A. stuckenbergi (BMSA (D) 94160), 34 - 36 ♂ epandrium and cerci 34 A. adamastor (BMSA (D) 38904) 35 A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 36 A. stuckenbergi (BMSA (D) 94160). Abbreviations: aed tn - aedeagal tine; cerc - cercus; ej apod - ejaculatory apodeme; epand - epandrium; goncx - gonocoxite; goncx apod - gonocoxal apodeme; gonst - gonostylus; pm - paramere; pm apod - parameral apodeme; pm sh - parameral sheath. Scale bars: 0.15 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures31-36" httpUri="https://binary.pensoft.net/fig/950654" pageId="0" pageNumber="303">, 31</figureCitation>
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<figureCitation id="9BAB2C277EDA6339B719EF93E0992E16" captionStart="Figures 31–36" captionStartId="F6" captionText="Figures 31 - 36. Atrichops spp., 31 - 33 ♂ terminalia 31 A. adamastor (BMSA (D) 38904) 32 A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 33 A. stuckenbergi (BMSA (D) 94160), 34 - 36 ♂ epandrium and cerci 34 A. adamastor (BMSA (D) 38904) 35 A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 36 A. stuckenbergi (BMSA (D) 94160). Abbreviations: aed tn - aedeagal tine; cerc - cercus; ej apod - ejaculatory apodeme; epand - epandrium; goncx - gonocoxite; goncx apod - gonocoxal apodeme; gonst - gonostylus; pm - paramere; pm apod - parameral apodeme; pm sh - parameral sheath. Scale bars: 0.15 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures31-36" httpUri="https://binary.pensoft.net/fig/950654" pageId="0" pageNumber="303">, 34</figureCitation>
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<figureCitation id="3290D526CD61256D2A7EFAF58C18DC12" captionStart="Figures 37–39" captionStartId="F7" captionText="Figures 37 - 39. Atrichops spp., ♀ genital fork and spermatheca 37 A. adamastor (BMSA (D) 38899 38 A. intermedius sp. nov. paratype (NMSA-Dip. 212861) 39 A. stuckenbergi (BMSA (D) 95855). Abbreviations: gen fk - genital fork; spmth - spermatheca. Scale bars: 0.15 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures37-39" httpUri="https://binary.pensoft.net/fig/950655" pageId="0" pageNumber="303">, 37</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="F3F75605CE5ABF110E90DBE50A025815" pageId="0" pageNumber="303" type="reference_group">
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<paragraph id="7389C67D7655DA798AF140D514707F98" pageId="0" pageNumber="303">
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<taxonomicName id="550B82F2B934D02EEA6522DA3A3D9E0C" authorityName="Muller & Swart & Snyman" authorityYear="2023" baseAuthorityName="Muller & Swart & Snyman" baseAuthorityYear="2023" class="Insecta" family="Athericidae" genus="Atherix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atherix adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor" status="comb. nov.">Atherix adamastor</taxonomicName>
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Stuckenberg, 1960: 273, fig. 86;
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<bibRefCitation id="E8D5C6663C515BB9842AC020ABD8D927" author="Stuckenberg, BR" editor="Crosskey, RW" journalOrPublisher="British Museum (Natural History), London" pageId="0" pageNumber="303" pagination="312 - 313" refId="B15" refString="Stuckenberg, BR, 1980. 23. Family Athericidae. In: Crosskey, RW, Ed., Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London: 312 - 313" title="23. Family Athericidae." volumeTitle="Catalogue of the Diptera of the Afrotropical Region." year="1980">Stuckenberg 1980</bibRefCitation>
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: 313;
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<bibRefCitation id="7002DE06E614869369FBFD6B016B512A" author="Nagatomi, A" journalOrPublisher="Memoirs of the Kagoshima University Research Center for the South Pacific" pageId="0" pageNumber="303" pagination="87 - 106" refId="B9" refString="Nagatomi, A, 1984a. Notes on Athericidae (Diptera). Memoirs of the Kagoshima University Research Center for the South Pacific 5: 87 - 106" title="Notes on Athericidae (Diptera)." volume="5" year="1984 a">Nagatomi 1984a</bibRefCitation>
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: 94.
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</paragraph>
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</subSubSection>
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<subSubSection id="FBC120F4E410660940B7E91B1821101A" pageId="0" pageNumber="303" type="identity">
|
||
<paragraph id="B064B4C9775A3A3CE3E8F14B7F068EA8" pageId="0" pageNumber="303">Identity.</paragraph>
|
||
<paragraph id="0B5D92EEE047F11E135F59547B005D5C" pageId="0" pageNumber="303">
|
||
<bibRefCitation id="43979C61D55199112AD16493B6651DF4" author="Stuckenberg, BR" editor="Hanstroem, B" journalOrPublisher="Results of the Lund University Expedition in 1950 - 1951. V. Almqvist & Wiksell, Uppsala" pageId="0" pageNumber="303" pagination="216 - 308" refId="B14" refString="Stuckenberg, BR, 1960. Chapter XII. Diptera (Brachycera): Rhagionidae. In: Hanstroem, B, Brinck, P, Rudebeck, G, Eds., South African animal life. Results of the Lund University Expedition in 1950-1951. V. Almqvist & Wiksell, Uppsala: 216 - 308" title="Chapter XII. Diptera (Brachycera): Rhagionidae." volumeTitle="South African animal life." year="1960">Stuckenberg (1960</bibRefCitation>
|
||
: 273) originally described
|
||
<taxonomicName id="3BEB86E4F9DAFA54F4D22C3E7FEAF6C4" authorityName="Muller & Swart & Snyman" authorityYear="2023" baseAuthorityName="Stuckenberg" baseAuthorityYear="1960" class="Insecta" family="Athericidae" genus="Atrichops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichops adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
|
||
<emphasis id="D4CB845A6C32C06BB6CE0B118025DF11" italics="true" pageId="0" pageNumber="303">Atrichops adamastor</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<taxonomicName id="7E0AEEF2C2D832B2499B8865CF65A776" authorityName="Meigen" authorityYear="1803" class="Insecta" family="Athericidae" genus="Atherix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atherix" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2796B575A30284F69AACE35ADF40B278" italics="true" pageId="0" pageNumber="303">Atherix</emphasis>
|
||
</taxonomicName>
|
||
Meigen, noting that its wings are "unlike those of any other South African species". However, it also differs from all other described southern African athericid species by possessing the key diagnostic characters of
|
||
<taxonomicName id="FCACC3653878A476F51D3EAB111CC15B" authorityName="Verrall" authorityYear="1909" class="Insecta" family="Athericidae" genus="Atrichops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichops" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A85FC577CDFBF946C3FF23E3BE6419CF" italics="true" pageId="0" pageNumber="303">Atrichops</emphasis>
|
||
</taxonomicName>
|
||
, namely a combination of a ventrally projecting knob-like proepimeral process (albeit much reduced), having the face narrower than the frons in female, and antennal bases closely set, nearly touching. It also has wing vein
|
||
<emphasis id="827E21CDB8D26A4AB1703302A8E55DAF" italics="true" pageId="0" pageNumber="303">R4+5</emphasis>
|
||
setulose dorsally, a characteristic only present in
|
||
<taxonomicName id="86756D3C3FD43DC4CF29A6DEE569E5C0" authorityName="Verrall" authorityYear="1909" class="Insecta" family="Athericidae" genus="Atrichops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichops" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="55E1C336F71292FE57937C8497A17BD8" italics="true" pageId="0" pageNumber="303">Atrichops</emphasis>
|
||
</taxonomicName>
|
||
, compared to other Afrotropical genera (see
|
||
<bibRefCitation id="0B76CE4091EBAE17C19F796EB4F8B639" DOI="https://doi.org/10.3853/j.0067-1975.52.2000.1312" author="Stuckenberg, BR" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="303" pagination="151 - 159" refId="B16" refString="Stuckenberg, BR, 2000. A new genus and species of Athericidae (Diptera: Tabanoidea) from Cape York Peninsula. Records of the Australian Museum 52 (2): 151 - 159, DOI: https://doi.org/10.3853/j.0067-1975.52.2000.1312" title="A new genus and species of Athericidae (Diptera: Tabanoidea) from Cape York Peninsula." url="https://doi.org/10.3853/j.0067-1975.52.2000.1312" volume="52" year="2000">Stuckenberg 2000</bibRefCitation>
|
||
: 157). The male terminalia are also typical of
|
||
<taxonomicName id="025156118745AD20CACE862908280023" authorityName="Verrall" authorityYear="1909" class="Insecta" family="Athericidae" genus="Atrichops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichops" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7F5210E90DAE0216A4D96F96C02F8F06" italics="true" pageId="0" pageNumber="303">Atrichops</emphasis>
|
||
</taxonomicName>
|
||
species, with the gonostylus inserted medially on the dorsal surface of the gonocoxite (e.g., Figs
|
||
<figureCitation id="E2E7066AB420D4C0469C90F3AA0130BD" captionStart="Figures 31–36" captionStartId="F6" captionText="Figures 31 - 36. Atrichops spp., 31 - 33 ♂ terminalia 31 A. adamastor (BMSA (D) 38904) 32 A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 33 A. stuckenbergi (BMSA (D) 94160), 34 - 36 ♂ epandrium and cerci 34 A. adamastor (BMSA (D) 38904) 35 A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 36 A. stuckenbergi (BMSA (D) 94160). Abbreviations: aed tn - aedeagal tine; cerc - cercus; ej apod - ejaculatory apodeme; epand - epandrium; goncx - gonocoxite; goncx apod - gonocoxal apodeme; gonst - gonostylus; pm - paramere; pm apod - parameral apodeme; pm sh - parameral sheath. Scale bars: 0.15 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures31-36" httpUri="https://binary.pensoft.net/fig/950654" pageId="0" pageNumber="303">31-33</figureCitation>
|
||
), compared to other genera where it is inserted apically.
|
||
<taxonomicName id="2EB5AA140C96F499A0BDCB4DD6F443FD" authorityName="Muller & Swart & Snyman" authorityYear="2023" baseAuthorityName="Stuckenberg" baseAuthorityYear="1960" class="Insecta" family="Athericidae" genus="Atrichops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichops adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
|
||
<emphasis id="8A193680251C23B377F23EF33D3E4458" italics="true" pageId="0" pageNumber="303">Atrichops adamastor</emphasis>
|
||
</taxonomicName>
|
||
has also been observed blood-feeding on frogs (see biology discussion below), a behaviour typical of the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="480AAFFB9BCEF0764948E5D1E2188F13" pageId="0" pageNumber="303" type="materials_examined">
|
||
<paragraph id="D642B49D0ED9FD6D030E8EC9A34EED46" pageId="0" pageNumber="303">Material examined.</paragraph>
|
||
<paragraph id="782DFDA6E83E231383B6ADD4ED79AF65" pageId="0" pageNumber="303">
|
||
<materialsCitation id="0C73D50E29D1302E9271E20CBAE9DE7B" collectingDate="1950-12-26" collectorName="Swedish South African Expedition, No., Fig." country="South Africa" county="Peninsula" elevation="91" latitude="-34.034473" location="Skoorsteenkop" longLatPrecision="1" longitude="18.3707" municipality="Hout Bay" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape Province" typeStatus="Holotype">
|
||
Type material examined (based on digital photos, additional data from
|
||
<bibRefCitation id="4227DA412676D78EEE9B1FAB12B0942E" author="Brinck, P" editor="Hanstroem, B" journalOrPublisher="Results of the Lund University Expedition in 1950 - 1951. I. Almqvist & Wiksell, Uppsala" pageId="0" pageNumber="303" pagination="62 - 100" refId="B1" refString="Brinck, P, Rudebeck, G, 1955. Chapter II. List of Localities Investigated by the Swedish Expedition to Southern Africa. In: Hanstroem, B, Brinck, P, Rudebeck, G, Eds., South African animal life. Results of the Lund University Expedition in 1950-1951. I. Almqvist & Wiksell, Uppsala: 62 - 100" title="Chapter II. List of Localities Investigated by the Swedish Expedition to Southern Africa." volumeTitle="South African animal life." year="1955">Brinck and Rudebeck (1955</bibRefCitation>
|
||
: 77)):
|
||
<emphasis id="D644AC3806C2F4A5EFFFB2AE482BFD02" bold="true" italics="true" pageId="0" pageNumber="303">
|
||
<typeStatus id="4F82231453639C7FCCF83E62A30A41F0">Holotype</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<collectingCountry id="DCD60A1C6EEBF4DA6AD0A4BA5770984B" name="South Africa">South Africa</collectingCountry>
|
||
•
|
||
<specimenCount id="3B8AF057456154F29CE11162E950D50B" type="female">1♀</specimenCount>
|
||
;
|
||
<collectingRegion id="C986D2C29DC1C546E42172BA1BA3A193" country="South Africa" name="Western Cape">Western Cape Province</collectingRegion>
|
||
,
|
||
<collectingRegion id="3D1C186C01BBC74AE9C87FF9451BD5D2" country="South Africa" name="Western Cape">Cape</collectingRegion>
|
||
<collectingCounty id="0F6DFF4D1FEC8590F6A3D7887D7432EC">Peninsula</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="28EBE38FB418823135AC2658499B8BED">Hout Bay</collectingMunicipality>
|
||
,
|
||
<location id="9CA29D8F4A4A53F2345C26BB5F0A4133" LSID="urn:lsid:plazi:treatment:5F22128A5B6F58F28D7195C12E30AA6B:9CA29D8F4A4A53F2345C26BB5F0A4133" country="South Africa" county="Peninsula" latitude="-34.034473" longLatPrecision="1" longitude="18.3707" municipality="Hout Bay" name="Skoorsteenkop" stateProvince="Western Cape Province">Skoorsteenkop</location>
|
||
; [
|
||
<geoCoordinate id="F5163DD6AF771753041DA5CC99DEB846" degrees="34" direction="south" minutes="02.0684" orientation="latitude" precision="1" value="-34.034473">34°02.0684'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="644C96A3A31AA60E7CB00BAADEA9C4BB" degrees="18" direction="east" minutes="22.2420" orientation="longitude" precision="1" value="18.3707">18°22.2420'E</geoCoordinate>
|
||
]; [
|
||
<quantity id="F27280D5FBD3371BEF40C234CFD7DDAC" metricMagnitude="1" metricUnit="m" metricValue="9.144" unit="ft" value="300.0">
|
||
<elevation id="1E34E0A1319A0028E10CF31375FA0AAA" metricMagnitude="1" metricUnit="m" metricValue="9.144" unit="ft" value="300.0">300 ft</elevation>
|
||
</quantity>
|
||
];
|
||
<collectingDate id="55EB4ADDD7FE288A4EEF06B03FE1C418" value="1950-12-26">26 Dec. 1950</collectingDate>
|
||
; [
|
||
<collectorName id="0AD880E657B50E6080070E0DFDE41FFE">Swedish South African Expedition</collectorName>
|
||
leg.];
|
||
<collectorName id="8B8A689206F25D3FB34C41CDEDCBDAD6">No.</collectorName>
|
||
95, type no. 00790:1, MZLU00205685 (see
|
||
<collectorName id="574580364E97315319C4D8A3A7F69E30">Fig.</collectorName>
|
||
<figureCitation id="4B0D651AFD82A42AE947DF872CC562E7" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Atrichops spp. ♂ 1 - 3 lateral habitus 4 - 6 head anterior view 1 A. adamastor (BMSA (D) 38908) 2 A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 3 A. stuckenbergi (BMSA (D) 95848) 4 A. adamastor (BMSA (D) 38904) 5 A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 6 A. stuckenbergi (BMSA (D) 95848). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures1-6" httpUri="https://binary.pensoft.net/fig/950648" pageId="0" pageNumber="303">1</figureCitation>
|
||
) (MZLU).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<caption id="D3DAD1E081C9F35527ADA9A7C6DDAA7B" doi="10.3897/afrinvertebr.64.113133.figures1-6" httpUri="https://binary.pensoft.net/fig/950648" pageId="0" pageNumber="303" start="Figures 1–6" startId="F1">
|
||
<paragraph id="F1089CC419ED81441973C249941297BB" pageId="0" pageNumber="303">
|
||
<emphasis id="C30D777B5C5F700067DDD611BB80F932" bold="true" pageId="0" pageNumber="303">Figures 1-6.</emphasis>
|
||
<taxonomicName id="3C9316B10DA4163F05B5DB8E87A18EDE" authorityName="Verrall" authorityYear="1909" class="Insecta" family="Athericidae" genus="Atrichops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichops" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AECE6A2862D13B530189703455434E36" italics="true" pageId="0" pageNumber="303">Atrichops</emphasis>
|
||
</taxonomicName>
|
||
spp.
|
||
<specimenCount id="33447938FD93602514DAFF55280E8395">♂</specimenCount>
|
||
<emphasis id="44A8E4622DB98CB55046A3D6B9F7702D" bold="true" pageId="0" pageNumber="303">1-3</emphasis>
|
||
lateral habitus
|
||
<emphasis id="38446A1436B0BA40AD1632A379FFFC18" bold="true" pageId="0" pageNumber="303">4-6</emphasis>
|
||
head anterior view
|
||
<emphasis id="02F984670107858808FE2C8C60EE2B55" bold="true" pageId="0" pageNumber="303">1</emphasis>
|
||
<taxonomicName id="7B2AEA7B164A2E83DF3A8F18910FF9EB" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
|
||
<emphasis id="14820D89343F0404C861A1F7B6A8898C" italics="true" pageId="0" pageNumber="303">A. adamastor</emphasis>
|
||
</taxonomicName>
|
||
(BMSA(D)38908)
|
||
<emphasis id="0C94FC7D6021C87D2681AB395BBC2A1D" bold="true" pageId="0" pageNumber="303">2</emphasis>
|
||
<taxonomicName id="2863C4F91AAA5D0B10F6C6B910AC65CC" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. intermedius" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="intermedius">
|
||
<emphasis id="2C518DC7D85963F617E8E20C22E63818" italics="true" pageId="0" pageNumber="303">A. intermedius</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<typeStatus id="9BF3795C16A6ED43A2EC1DED081E8CCF">holotype</typeStatus>
|
||
(NMSA-Dip. 81808)
|
||
<emphasis id="E6353E894DE2B6D33B3E46C684504D32" bold="true" pageId="0" pageNumber="303">3</emphasis>
|
||
<taxonomicName id="32849E72550F10CE40E66129BBD125E6" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. stuckenbergi" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="stuckenbergi">
|
||
<emphasis id="C637D4CF1B335DC71069E18389CAF9DF" italics="true" pageId="0" pageNumber="303">A. stuckenbergi</emphasis>
|
||
</taxonomicName>
|
||
(BMSA(D) 95848)
|
||
<emphasis id="D448C35EC47D80C71CDE38BA28124A0E" bold="true" pageId="0" pageNumber="303">4</emphasis>
|
||
<taxonomicName id="2E8BEC8A6FBBA7D445D97FD8FF67E661" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
|
||
<emphasis id="4AACB5A552B9CAE549B47940A47364F9" italics="true" pageId="0" pageNumber="303">A. adamastor</emphasis>
|
||
</taxonomicName>
|
||
(BMSA(D)38904)
|
||
<emphasis id="FA72CA54140EAB0BCEEE155C7D7A6987" bold="true" pageId="0" pageNumber="303">5</emphasis>
|
||
<taxonomicName id="9FA31AB61BD4787D8E15D2BF41AE5424" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. intermedius" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="intermedius">
|
||
<emphasis id="AC33F9F2F0B0599E5045DCB9E38A790A" italics="true" pageId="0" pageNumber="303">A. intermedius</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<typeStatus id="C593CF421C14294DED878B9290E2A972">holotype</typeStatus>
|
||
(NMSA-Dip. 81808)
|
||
<emphasis id="5E5773BB1831D78760C266A4C0E7DA6A" bold="true" pageId="0" pageNumber="303">6</emphasis>
|
||
<taxonomicName id="39BD99C0756802BBD8C45F30FD6E34DB" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. stuckenbergi" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="stuckenbergi">
|
||
<emphasis id="0A76603DD313D49BA94AA54B6040EBC1" italics="true" pageId="0" pageNumber="303">A. stuckenbergi</emphasis>
|
||
</taxonomicName>
|
||
(BMSA(D) 95848). Scale bars:
|
||
<quantity id="3BCE4BE1C22E0476D54AAEA2D5FF8563" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="41E2B1984F3566ABC90C4C9B1D69CE90" pageId="0" pageNumber="303" type="materials_examined">
|
||
<paragraph id="0F13F8605D2CD2ABEF19C7017A21D88E" pageId="0" pageNumber="303">Additional material examined.</paragraph>
|
||
<paragraph id="9B382FA504A69D9EF1CFD25EEC240342" pageId="0" pageNumber="303">
|
||
<materialsCitation id="0B44ADD8DF9B95DD8F6AA0BE0F09BCA6" collectingDate="2012-10-16" collectingDateMax="2012-10-18" collectingDateMin="2012-10-16" collectingMethod="Malaise trap" collectorName="Kirk-Spriggs, A. H." country="South Africa" elevation="336" latitude="-33.36347" location="Gamkaskloof" longLatPrecision="1" longitude="21.6275" specimenCount="24" specimenCount-female="20" specimenCount-male="4" stateProvince="Western Cape Province">
|
||
<collectingCountry id="94ADB102858ADC762FBEB65D1B4A44C7" name="South Africa">South Africa</collectingCountry>
|
||
•
|
||
<specimenCount id="57D6F998210ACC4723072A141BC38EE8" type="male">2♂</specimenCount>
|
||
<specimenCount id="317FCD829D086FE79C36F87ED98A78B1" type="female">10♀</specimenCount>
|
||
;
|
||
<collectingRegion id="BF5DF03D8E42CC994AD893BF14627CCE" country="South Africa" name="Western Cape">Western Cape Province</collectingRegion>
|
||
,
|
||
<location id="E0CDC836373B971B920A001AA2D7FAB8" LSID="urn:lsid:plazi:treatment:5F22128A5B6F58F28D7195C12E30AA6B:E0CDC836373B971B920A001AA2D7FAB8" country="South Africa" latitude="-33.36347" longLatPrecision="1" longitude="21.6275" name="Gamkaskloof" stateProvince="Western Cape Province">Gamkaskloof</location>
|
||
(Die Hel) at:;
|
||
<geoCoordinate id="8A13B5FDAC916C4FF963E4A74ED056FA" degrees="33" direction="south" minutes="21.808" orientation="latitude" precision="1" value="-33.36347">33°21.808'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2ECF349A77B694FEDFE7C11E403AEEB7" degrees="21" direction="east" minutes="37.650" orientation="longitude" precision="1" value="21.6275">21°37.650'E</geoCoordinate>
|
||
;
|
||
<elevation id="D157AEE9144DF1669AFC5F756ADA082D" metricMagnitude="2" metricUnit="m" metricValue="3.36" unit="m" value="336.0">
|
||
<quantity id="4BDC17DF2CE8D00D5F9331A8BD16F9FB" metricMagnitude="2" metricUnit="m" metricValue="3.36" unit="m" value="336.0">336 m</quantity>
|
||
asl
|
||
</elevation>
|
||
;
|
||
<collectingDate id="3AA7DCBB92A377CE51AB5D40C94A27E9" value="2012-10-16" valueMax="2012-10-18" valueMin="2012-10-16">16-18 Oct. 2012</collectingDate>
|
||
;
|
||
<collectorName id="9C3820BE41F515A88E61D1B92723FEB6">Kirk-Spriggs, A.H.</collectorName>
|
||
leg.;
|
||
<collectingMethod id="94FB7469405F7B990DB5768F6C858347">Malaise trap</collectingMethod>
|
||
in Karoo and valley
|
||
<taxonomicName id="E0658C9CAB6DBA69ED8455C5E25C80EA" class="Dicotyledoneae" family="Fabaceae" genus="Acacia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Acacia" order="Rosales" pageId="0" pageNumber="303" phylum="Angiospermae" rank="genus">
|
||
<emphasis id="5F6DA34DF59E56246E38FE541CB2F8E3" italics="true" pageId="0" pageNumber="303">Acacia</emphasis>
|
||
</taxonomicName>
|
||
woodland; (
|
||
<specimenCount id="70D7AC44859A56F73F656344419EF3B6" type="male">2♂</specimenCount>
|
||
: BMSA(D)38904, 38908;
|
||
<specimenCount id="F340464AC717D85AE6C557804E133858" type="female">10♀</specimenCount>
|
||
: BMSA(D)38899, 38900, 38901, 38902, 38903, 38905, 38906, 38907, 38909, 40027)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="D16A08667BC9ED26753E4814CD25D82E" doi="10.3897/afrinvertebr.64.113133.figures7-12" httpUri="https://binary.pensoft.net/fig/950649" pageId="0" pageNumber="303" start="Figures 7–12" startId="F2">
|
||
<paragraph id="01735425A1889C931F3948A6745E08EC" pageId="0" pageNumber="303">
|
||
<emphasis id="25B6C84FC930C4B78D874CA9E32535C4" bold="true" pageId="0" pageNumber="303">Figures 7-12.</emphasis>
|
||
<taxonomicName id="31AF938EEE9C2CFD052894F46F9F1532" authorityName="Verrall" authorityYear="1909" class="Insecta" family="Athericidae" genus="Atrichops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichops" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="23132B77C29EEDF3E26F7640C5BEB182" italics="true" pageId="0" pageNumber="303">Atrichops</emphasis>
|
||
</taxonomicName>
|
||
spp.
|
||
<specimenCount id="05FF413010D9BE2C204045EA6EC9EFBB">♀</specimenCount>
|
||
<emphasis id="040FAEE3D3C2C0F4B7782A4F1F7A2EAF" bold="true" pageId="0" pageNumber="303">7-9</emphasis>
|
||
lateral habitus
|
||
<emphasis id="601CCF5D2E9E052E502F6C212E91F996" bold="true" pageId="0" pageNumber="303">10-12</emphasis>
|
||
head anterior view
|
||
<emphasis id="F2B8E2F5929A07CC52DD20CD7A933A6D" bold="true" pageId="0" pageNumber="303">7</emphasis>
|
||
<taxonomicName id="CE18F2E25FDD17FB32B8CF7DBC15D3F8" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
|
||
<emphasis id="2724FC4D7C58DA638D521FC4D178749F" italics="true" pageId="0" pageNumber="303">A. adamastor</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="3FE70E411979224F0A38CF0CDE907718">holotype</typeStatus>
|
||
and specimen labels (MZLU 00205685)
|
||
<emphasis id="65B6558D6D6F05E1EFE808E2B1CD1F1C" bold="true" pageId="0" pageNumber="303">8</emphasis>
|
||
<taxonomicName id="D20A87CCA8B2AEFAC4525070CB69011C" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. intermedius" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="intermedius">
|
||
<emphasis id="D9B4448719D9BA09FAF0E4D0983C68AC" italics="true" pageId="0" pageNumber="303">A. intermedius</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<typeStatus id="49AA6DB05C786A093D131CFF045408B1">paratype</typeStatus>
|
||
(NMSA-Dip. 81809)
|
||
<emphasis id="2A9A5154C2D0A59DBA4D016DB386DD96" bold="true" pageId="0" pageNumber="303">9</emphasis>
|
||
<taxonomicName id="ADD3E10669875AEC52CA254D8BD353A0" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. stuckenbergi" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="stuckenbergi">
|
||
<emphasis id="56A0BCE3C08FF129C8323BC42536AB0A" italics="true" pageId="0" pageNumber="303">A. stuckenbergi</emphasis>
|
||
</taxonomicName>
|
||
(CAS)
|
||
<emphasis id="38DEAB70678D7439F725C0071D81D783" bold="true" pageId="0" pageNumber="303">10</emphasis>
|
||
<taxonomicName id="349C3D9A440BB97CED1AAA84421027F7" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
|
||
<emphasis id="1C2ECE5DA31AF8A4377B181A04F9EB14" italics="true" pageId="0" pageNumber="303">A. adamastor</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="1324A10C539BFB8375ED35134443B325">holotype</typeStatus>
|
||
(MZLU 00205685)
|
||
<emphasis id="02ACD07FAF5685C1A5AC089CFB3A5F1C" bold="true" pageId="0" pageNumber="303">11</emphasis>
|
||
<taxonomicName id="5C3ACBB30A8BB4D8B53CAD19A6ACDF0B" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. intermedius" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="intermedius">
|
||
<emphasis id="64128891DA8D23559C94503A11659E2B" italics="true" pageId="0" pageNumber="303">A. intermedius</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<typeStatus id="1C9831EB14BD2A7BE43A82B70A282BFC">paratype</typeStatus>
|
||
(NMSA-Dip. 81809)
|
||
<emphasis id="1BAC219C28052B86593CE9DD5B0F5258" bold="true" pageId="0" pageNumber="303">12</emphasis>
|
||
<taxonomicName id="7201FED7C4FABD6F131B75547624C1CC" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. stuckenbergi" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="stuckenbergi">
|
||
<emphasis id="670BFE7B8E85BD8666DDF35C03A1A11C" italics="true" pageId="0" pageNumber="303">A. stuckenbergi</emphasis>
|
||
</taxonomicName>
|
||
(BMSA(D)95856). (
|
||
<emphasis id="8A337DE8093A24F96FFB3EFFBB3634EF" bold="true" pageId="0" pageNumber="303">7, 10</emphasis>
|
||
modified from originals provided by Rune Bygebjerg, MZLU, CC BY-NC 2.0
|
||
<emphasis id="674E24E39CF4E5B8C1482A03A6FD547D" bold="true" pageId="0" pageNumber="303">9, 12</emphasis>
|
||
modified from originals provided by Rikio Matsumoto, Osaka Museum of Natural History). Scale bars:
|
||
<quantity id="2F66D3BE840B8DB1D2B3A96D449CC7A2" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="96067F2F367128400D88478EB4470C00" pageId="0" pageNumber="303" type="diagnosis">
|
||
<paragraph id="9E16446CB426CD2DD89B5A7FE49D06FE" pageId="0" pageNumber="303">Diagnosis.</paragraph>
|
||
<paragraph id="73039612D4D1C7C6B08922F6C08CE4FB" pageId="0" pageNumber="303">
|
||
<taxonomicName id="0091DDDCF9D2914B21EB590A050A1404" authorityName="Muller & Swart & Snyman" authorityYear="2023" baseAuthorityName="Stuckenberg" baseAuthorityYear="1960" class="Insecta" family="Athericidae" genus="Atrichops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichops adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
|
||
<emphasis id="C17088C2760D1026BBEF9F9128866819" italics="true" pageId="0" pageNumber="303">Atrichops adamastor</emphasis>
|
||
</taxonomicName>
|
||
can easily be distinguished from
|
||
<taxonomicName id="944F551681FF5151D95A4277B0607512" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. stuckenbergi" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="stuckenbergi">
|
||
<emphasis id="1BC964526AAA4FE6AAF88323DB45DC68" italics="true" pageId="0" pageNumber="303">A. stuckenbergi</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="4D1CA91D8325DDB513C225260C745119" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
|
||
<emphasis id="A9519BD2FA1AE4131BF582CEDFDDB72B" italics="true" pageId="0" pageNumber="303">A. adamastor</emphasis>
|
||
</taxonomicName>
|
||
the ommatrichia densely covers the eye in both sexes, the frons is dark brown, the face is brownish, and the wing is uniformly brown suffused, whereas
|
||
<taxonomicName id="E0BE6B3DA1A6E263B616326F5322013E" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. stuckenbergi" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="stuckenbergi">
|
||
<emphasis id="756B36B425FAF9E1B6984184E117A662" italics="true" pageId="0" pageNumber="303">A. stuckenbergi</emphasis>
|
||
</taxonomicName>
|
||
has the eye sparsely covered in ommatrichia, the frons is shiny black, the face has silver-white pruinosity, and its wing brown suffused mainly on apical half, with discal cell and cell
|
||
<emphasis id="19C0A9708CA527B936302C092AF0CB84" italics="true" pageId="0" pageNumber="303">m3</emphasis>
|
||
less so (e.g., Fig.
|
||
<figureCitation id="9600EBE7AFBE3F2D990D6CDED042475B" captionStart="Figures 25–30" captionStartId="F5" captionText="Figures 25 - 30. Atrichops spp. ♂ ♀ wings dorsal view, A. adamastor: 25 ♂ (BMSA (D) 38904) 26 ♀ (BMSA (D) 38904); A. intermedius sp. nov.: 27 ♂ paratype (NMSA-Dip. 81811) 28 ♀ paratype (NMSA-Dip. 212861) A. stuckenbergi: 29 ♂ (BMSA (D) 94161) 30 ♀ (BMSA (D) 95861). Abbreviations: br - basal radial cell; bm - basal medial cell; cua - anterior cubital cell; CuA + CuP - anterior branch of cubital vein + posterior branch of cubital vein; d - discal cell; M 1 - first branch of media; m 1 - first medial cell; M 2 - second branch of media; m 2 - second medial cell; M 3 - third branch of media; m 3 - third medial cell; R 1 - anterior branch of radius; r 1 - first radial cell; R 2 + 3 - second branch of radius; r 2 + 3 - second radial cell; R 4 - upper branch of third branch of radius; R 5 - lower branch of third branch of radius; Sc - subcostal vein; sc - subcostal cell. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures25-30" httpUri="https://binary.pensoft.net/fig/950653" pageId="0" pageNumber="303">26</figureCitation>
|
||
vs Fig.
|
||
<figureCitation id="6662E3946A7FAE874063EB5F22A6DF25" captionStart="Figures 25–30" captionStartId="F5" captionText="Figures 25 - 30. Atrichops spp. ♂ ♀ wings dorsal view, A. adamastor: 25 ♂ (BMSA (D) 38904) 26 ♀ (BMSA (D) 38904); A. intermedius sp. nov.: 27 ♂ paratype (NMSA-Dip. 81811) 28 ♀ paratype (NMSA-Dip. 212861) A. stuckenbergi: 29 ♂ (BMSA (D) 94161) 30 ♀ (BMSA (D) 95861). Abbreviations: br - basal radial cell; bm - basal medial cell; cua - anterior cubital cell; CuA + CuP - anterior branch of cubital vein + posterior branch of cubital vein; d - discal cell; M 1 - first branch of media; m 1 - first medial cell; M 2 - second branch of media; m 2 - second medial cell; M 3 - third branch of media; m 3 - third medial cell; R 1 - anterior branch of radius; r 1 - first radial cell; R 2 + 3 - second branch of radius; r 2 + 3 - second radial cell; R 4 - upper branch of third branch of radius; R 5 - lower branch of third branch of radius; Sc - subcostal vein; sc - subcostal cell. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures25-30" httpUri="https://binary.pensoft.net/fig/950653" pageId="0" pageNumber="303">30</figureCitation>
|
||
). It is most similar to
|
||
<taxonomicName id="691A70530097215CA822AEE50EA14FCB" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. intermedius" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="intermedius">
|
||
<emphasis id="7E800315744A109E4C7EE01ACA66DC0A" italics="true" pageId="0" pageNumber="303">A. intermedius</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., a South African contemporary (see
|
||
<taxonomicName id="8F3F7B541CDC25066F614D5513A94170" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. intermedius" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="intermedius">
|
||
<emphasis id="513D95F25EF10220A1C0EF34409BE910" italics="true" pageId="0" pageNumber="303">A. intermedius</emphasis>
|
||
</taxonomicName>
|
||
diagnosis).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E7A6535D8A4EB2C7E3625F7D446458A1" pageId="0" pageNumber="303" type="remarks">
|
||
<paragraph id="25C8EAE6567BA1908CEC193C75D539F0" pageId="0" pageNumber="303">Remarks.</paragraph>
|
||
<paragraph id="B3E673DF98F2B3F7268B7BCA4AA2DE43" pageId="0" pageNumber="303">
|
||
The species was described by
|
||
<bibRefCitation id="5030F658FD835F6569CAE60C520A3920" author="Stuckenberg, BR" editor="Hanstroem, B" journalOrPublisher="Results of the Lund University Expedition in 1950 - 1951. V. Almqvist & Wiksell, Uppsala" pageId="0" pageNumber="303" pagination="216 - 308" refId="B14" refString="Stuckenberg, BR, 1960. Chapter XII. Diptera (Brachycera): Rhagionidae. In: Hanstroem, B, Brinck, P, Rudebeck, G, Eds., South African animal life. Results of the Lund University Expedition in 1950-1951. V. Almqvist & Wiksell, Uppsala: 216 - 308" title="Chapter XII. Diptera (Brachycera): Rhagionidae." volumeTitle="South African animal life." year="1960">Stuckenberg (1960</bibRefCitation>
|
||
: 273), based only on a single female. The description is sufficient and no re-description is necessary. A description of its previously undescribed (but known) male follows. Mention is made of female characters should they differ from the male.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B035C577206FEB61586CE296C9C3D7E2" pageId="0" pageNumber="303" type="description">
|
||
<paragraph id="BDFD9DC70ADB9591AD440C18704E4D4D" pageId="0" pageNumber="303">Description.</paragraph>
|
||
<paragraph id="088ED217F76159D749CD7E562BD8F9AB" pageId="0" pageNumber="303">
|
||
<emphasis id="0BF9C1F7B17D4CF12C5D6DE218FC61F8" bold="true" italics="true" pageId="0" pageNumber="303">Measurements</emphasis>
|
||
(♂ n = 2, ♀ n = 10): Wing span: ♂ 4.8-5.2 mm (avg. 5.0 mm); ♀ 5.0-6.0 mm (avg. 5.4 mm); body length: ♂ 4.6 mm (avg. 4.6 mm); ♀ 3.7-4.9 mm (avg. 4.5 mm); wing span to body length ratio (avg.): ♂ 1.1; ♀ 1.2.
|
||
</paragraph>
|
||
<paragraph id="5FAE0FD60B2F27827AE9C0F2D5A0C24E" pageId="0" pageNumber="303">
|
||
<emphasis id="86AAB2706CD4BBDC7314ACC1EEA411B0" bold="true" pageId="0" pageNumber="303">
|
||
Male.
|
||
<emphasis id="BFBCC2E7F6B2093DFAE187F13EE5D6EE" italics="true" pageId="0" pageNumber="303">Head</emphasis>
|
||
</emphasis>
|
||
(Fig.
|
||
<figureCitation id="2486A9F18D8A920CB29575464AEB6B5C" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Atrichops spp. ♂ 1 - 3 lateral habitus 4 - 6 head anterior view 1 A. adamastor (BMSA (D) 38908) 2 A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 3 A. stuckenbergi (BMSA (D) 95848) 4 A. adamastor (BMSA (D) 38904) 5 A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 6 A. stuckenbergi (BMSA (D) 95848). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures1-6" httpUri="https://binary.pensoft.net/fig/950648" pageId="0" pageNumber="303">4</figureCitation>
|
||
): Colour brown, with some light yellowish pruinosity on face; eye densely covered in ommatrichia; narrowly dichoptic, ♀ widely dichoptic; ommatidia on lower
|
||
<normalizedToken id="FB11204966810AADC88522C3CF55133E" originalValue="½">1/2</normalizedToken>
|
||
of eye smaller than upper
|
||
<normalizedToken id="45CF380E6A7D60152DD5536B5B67CA5B" originalValue="½">1/2</normalizedToken>
|
||
(♀ ommatidia uniform, comparatively larger than in ♂); lateral edge of eye with slight indentation, nearly absent in ♀; ocellar tubercle base level with frons, ♀ base with sunken appearance; ocellar tubercle with short dark setulae, vertex dark brown, almost appearing black, with somewhat longer setulae than ocellar tubercle; ocelli similar in size; ocellar tubercle in front of dorsal margin of eye, margin less indented than in ♀; vertex narrower than in ♀; dorsal inner edge of eye surrounding ocellar tubercle without discernible paired dark markings (present in ♀); occiput same dark brown as frons, when viewed at angle appearing shiny blackish; upper occiput with short dark setulae on dorsal margin and on rest of upper surface, lower occiput with erect, long and pale setulae, these continue ventrally on head to before mouthparts, bearing dark ventral setulae; frons dark brown, almost black, somewhat shiny, narrow area above antenna lighter brown; frons at narrowest as wide as anterior ocellus, widening towards antennal base, ♀ frons at least 2
|
||
<normalizedToken id="738DF61220464BD6DDD230A1CE64612F" originalValue="×">x</normalizedToken>
|
||
width of ocellar tubercle, narrowing only slightly towards antennal base (Fig.
|
||
<figureCitation id="7D0E09E324E4F6DDB5D30F4F74C513C5" captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Atrichops spp. ♀ 7 - 9 lateral habitus 10 - 12 head anterior view 7 A. adamastor holotype and specimen labels (MZLU 00205685) 8 A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 9 A. stuckenbergi (CAS) 10 A. adamastor holotype (MZLU 00205685) 11 A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 12 A. stuckenbergi (BMSA (D) 95856). (7, 10 modified from originals provided by Rune Bygebjerg, MZLU, CC BY-NC 2.0 9, 12 modified from originals provided by Rikio Matsumoto, Osaka Museum of Natural History). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures7-12" httpUri="https://binary.pensoft.net/fig/950649" pageId="0" pageNumber="303">10</figureCitation>
|
||
); frons with well-developed dark setulae on surface, but only on lateral margins, very similar to ocellar and vertical setulae; ♀ frons more setulose than ♂; face lateral edges bare; gena bare; face light brown, clypeus darker brown, ♀ clypeus orange-brown; clypeus bare; face separated from clypeus by prominent transverse suture, in ♀ separated by transverse emargination, never forming suture; clypeus less prominent than in ♀; face much wider than in ♀, with prominent longitudinal emargination on side of clypeus, giving appearance that face bulges laterally; face and clypeus not visible in profile (visible in ♀); antennal bases close together, almost touching, gap somewhat larger than in ♀; scape brown, dorsally infuscate around setulae; pedicel comparatively darker than scape, dorsal surface infuscate; scape and pedicel setulae dark; 1st flagellomere reniform, appearing almost 2
|
||
<normalizedToken id="F464742F67EFD9ABA10688CC6EEBFE39" originalValue="×">x</normalizedToken>
|
||
height of pedicel, concolourous with pedicel, basal margins lighter brown; 2nd flagellomere arista-like, dark brown; scape and pedicel setulae similar in size; palpus dark brown, well-developed, ca 0.5
|
||
<normalizedToken id="D67C13A3FF5617BF282C2916059C37CB" originalValue="×">x</normalizedToken>
|
||
length of proboscis, with long, dark setulae; proboscis dark brown with orange-brown base.
|
||
</paragraph>
|
||
<paragraph id="1844739C45AA324EDEE601F5EEDB6416" pageId="0" pageNumber="303">
|
||
<emphasis id="89DE26ADF9E06B82AD66156535E86720" bold="true" italics="true" pageId="0" pageNumber="303">Thorax</emphasis>
|
||
(Figs
|
||
<figureCitation id="DB8EA27E2706A6418FEB0723CB30BACE" captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Atrichops spp. ♂ ♀ dorsal view 13 ♂ A. adamastor (BMSA (D) 38908) 14 ♂ A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 15 ♂ A. stuckenbergi (BMSA (D) 94158) 16 ♀ A. adamastor holotype (MZLU 00205685) 17 ♀ A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 18 ♀ A. stuckenbergi holotype (CAS). (16 modified from original provided by Rune Bygebjerg, MZLU, CC BY-NC 2.0; 18 modified from original provided by Rikio Matsumoto, OMNH). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures13-18" httpUri="https://binary.pensoft.net/fig/950650" pageId="0" pageNumber="303">13</figureCitation>
|
||
,
|
||
<figureCitation id="66242DEDD7C5F30950A3AE6FCA556DE4" captionStart="Figures 19–24" captionStartId="F4" captionText="Figures 19 - 24. Atrichops spp. ♂ ♀ proepimeral process, lateral view 19 ♂ A. adamastor (BMSA (D) 38904) 20 ♂ A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 21 ♂ A. stuckenbergi (BMSA (D) 94158) 22 ♀ A. adamastor (BMSA (D) 38899) 23 ♀ A. intermedius sp. nov. paratype (NMSA-Dip. 212861) 24 ♀ A. stuckenbergi holotype (CAS). (24 modified from original provided by Rikio Matsumoto, OMNH). Abbreviations: prepm - proepimeral process." figureDoi="10.3897/afrinvertebr.64.113133.figures19-24" httpUri="https://binary.pensoft.net/fig/950652" pageId="0" pageNumber="303">19</figureCitation>
|
||
): Scutum with short dark setulae, posterior setulae longer than anterior setulae; scutellum with well-developed dark setulae; postpronotal lobe lighter brown than scutum, with fine dark setulae; scutum uniformly dark brown; scutellum uniformly dark brown; pleura generally brown in colour, with anepisternum, katepisternum, anatergite and katatergite somewhat darker; proepimeron with reduced flap-like process near anterior edge; notopleuron with well-developed dark setulae; area surrounding posterior spiracle dark brown, postspiracular scale dark brown almost black; proepisternum and pronotum yellow; anterior spiracle bare posteriorly; proepimeron, proepisternum bare, anepisternum with short dark setulae; katatergite with long pale setulae; rest of pleura bare; postmetacoxal bridge narrow.
|
||
</paragraph>
|
||
<caption id="F4DD2C281B3A957DE19CAF21169B5B20" doi="10.3897/afrinvertebr.64.113133.figures13-18" httpUri="https://binary.pensoft.net/fig/950650" pageId="0" pageNumber="303" start="Figures 13–18" startId="F3">
|
||
<paragraph id="34512F5C22FCB08FFE0DFA383C885879" pageId="0" pageNumber="303">
|
||
<emphasis id="F1B30FCFE4AA1E96CE87CDC783D54212" bold="true" pageId="0" pageNumber="303">Figures 13-18.</emphasis>
|
||
<taxonomicName id="56E72D67A836D6DF293CE171F73664E6" authorityName="Verrall" authorityYear="1909" class="Insecta" family="Athericidae" genus="Atrichops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichops" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="genus">Atrichops</taxonomicName>
|
||
spp. ♂♀ dorsal view
|
||
<emphasis id="172A8A6FB9CAA05DBCACA7E0EE605897" bold="true" pageId="0" pageNumber="303">13</emphasis>
|
||
♂
|
||
<taxonomicName id="2317DCD0ADE477FD62ED133547399A30" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
|
||
<emphasis id="39F5CA65B45C0B2830E1DBE343342209" italics="true" pageId="0" pageNumber="303">A. adamastor</emphasis>
|
||
</taxonomicName>
|
||
(BMSA(D)38908)
|
||
<emphasis id="FA948107EC870ED29B941C664063E63B" bold="true" pageId="0" pageNumber="303">14</emphasis>
|
||
♂
|
||
<taxonomicName id="88589F85F7C37B83CF6B77A677EC666F" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. intermedius" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="intermedius">
|
||
<emphasis id="7E48A1C431362949F1FC501FD7F9459F" italics="true" pageId="0" pageNumber="303">A. intermedius</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. holotype (NMSA-Dip. 81808)
|
||
<emphasis id="553129C8C8C146C90A6C04F0F03B3A5D" bold="true" pageId="0" pageNumber="303">15</emphasis>
|
||
♂
|
||
<taxonomicName id="B419E1D78312004701BA4A0DA3490DB8" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. stuckenbergi" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="stuckenbergi">
|
||
<emphasis id="7B76D9B6D94EEB95CB2F34D8E8344BFD" italics="true" pageId="0" pageNumber="303">A. stuckenbergi</emphasis>
|
||
</taxonomicName>
|
||
(BMSA(D)94158)
|
||
<emphasis id="B0F0939727AE5062D0C2108E12F3D0BE" bold="true" pageId="0" pageNumber="303">16</emphasis>
|
||
♀
|
||
<taxonomicName id="EEF56A8C46BC19C349701195624727F5" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
|
||
<emphasis id="BC7964027C70BDAA1699B07F8D934161" italics="true" pageId="0" pageNumber="303">A. adamastor</emphasis>
|
||
</taxonomicName>
|
||
holotype (MZLU 00205685)
|
||
<emphasis id="E9DD82879DBD1B447940DFD650AC5033" bold="true" pageId="0" pageNumber="303">17</emphasis>
|
||
♀
|
||
<taxonomicName id="3C99005E64F09D6714B4B3BDA6587416" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. intermedius" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="intermedius">
|
||
<emphasis id="263D2F4FD8D0E8C2B8C0A988B5A77DC5" italics="true" pageId="0" pageNumber="303">A. intermedius</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. paratype (NMSA-Dip. 81809)
|
||
<emphasis id="76D5030FB1F957DDA7BA7DB1EB6D9D7F" bold="true" pageId="0" pageNumber="303">18</emphasis>
|
||
♀
|
||
<taxonomicName id="FD5164654ED549E23F8E10E1EF3FF185" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. stuckenbergi" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="stuckenbergi">
|
||
<emphasis id="4DD93B502DBC2F8CF64831A5F0CB954D" italics="true" pageId="0" pageNumber="303">A. stuckenbergi</emphasis>
|
||
</taxonomicName>
|
||
holotype (CAS). (
|
||
<emphasis id="8F0242EC0969D61648051D689D379CCE" bold="true" pageId="0" pageNumber="303">16</emphasis>
|
||
modified from original provided by Rune Bygebjerg, MZLU, CC BY-NC 2.0;
|
||
<emphasis id="376BC14E494303ED42FA0D04D4F9A0F1" bold="true" pageId="0" pageNumber="303">18</emphasis>
|
||
modified from original provided by Rikio Matsumoto, OMNH). Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="9A4945DAD59FCD6B9DD32B2B5DB35FF2" doi="10.3897/afrinvertebr.64.113133.figures19-24" httpUri="https://binary.pensoft.net/fig/950652" pageId="0" pageNumber="303" start="Figures 19–24" startId="F4">
|
||
<paragraph id="576DA1B20F6BF8F84466E9C50AAFDBA7" pageId="0" pageNumber="303">
|
||
<emphasis id="73BF7E9B757D33C18CB31E06E873C830" bold="true" pageId="0" pageNumber="303">Figures 19-24.</emphasis>
|
||
<taxonomicName id="B9EACBE30271C4A1799035151E44F2F5" authorityName="Verrall" authorityYear="1909" class="Insecta" family="Athericidae" genus="Atrichops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichops" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B8197F66ADC93A2F90CFFE2E48F3842F" italics="true" pageId="0" pageNumber="303">Atrichops</emphasis>
|
||
</taxonomicName>
|
||
spp. ♂♀ proepimeral process, lateral view
|
||
<emphasis id="2796152E5254E3177AF1EC3A31845BE4" bold="true" pageId="0" pageNumber="303">19</emphasis>
|
||
♂
|
||
<taxonomicName id="8ACC20D3DA47FACAEAE3E0C2B0E9A7AC" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
|
||
<emphasis id="DB6947E577FD8AA74C451A560B41F7F2" italics="true" pageId="0" pageNumber="303">A. adamastor</emphasis>
|
||
</taxonomicName>
|
||
(BMSA(D)38904)
|
||
<emphasis id="B729366CE3A4E6843459FD9104CEFECF" bold="true" pageId="0" pageNumber="303">20</emphasis>
|
||
♂
|
||
<taxonomicName id="43DA4B90C419E557EA8B6FE1E4DC55CF" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. intermedius" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="intermedius">
|
||
<emphasis id="EE5B943A8051A632990A214D4FEAEDC8" italics="true" pageId="0" pageNumber="303">A. intermedius</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. paratype (NMSA-Dip. 81811)
|
||
<emphasis id="0F5FF9C8CACAA53BB039B9492C596441" bold="true" pageId="0" pageNumber="303">21</emphasis>
|
||
♂
|
||
<taxonomicName id="09BD46D2A81CD857AE0D589BE0E9B1B6" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. stuckenbergi" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="stuckenbergi">
|
||
<emphasis id="E41E84E3E50B54DD49197109A84A30C2" italics="true" pageId="0" pageNumber="303">A. stuckenbergi</emphasis>
|
||
</taxonomicName>
|
||
(BMSA(D)94158)
|
||
<emphasis id="628284905900E34004ED2DFD93798057" bold="true" pageId="0" pageNumber="303">22</emphasis>
|
||
♀
|
||
<taxonomicName id="A7D619F501D3FD3B216393DEE5D47613" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
|
||
<emphasis id="38AC277F6814D5EC3AE88EC46D7DBED8" italics="true" pageId="0" pageNumber="303">A. adamastor</emphasis>
|
||
</taxonomicName>
|
||
(BMSA(D)38899)
|
||
<emphasis id="2A94F08AFF102F235B69DDDEB3FE5200" bold="true" pageId="0" pageNumber="303">23</emphasis>
|
||
♀
|
||
<taxonomicName id="E19DA941336D8DD1DCB28EAC6A6032A3" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. intermedius" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="intermedius">
|
||
<emphasis id="33D7F6D2D180462FD59426681CF9097B" italics="true" pageId="0" pageNumber="303">A. intermedius</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. paratype (NMSA-Dip. 212861)
|
||
<emphasis id="1EF877E10B22A241502FD5F14942A3C5" bold="true" pageId="0" pageNumber="303">24</emphasis>
|
||
♀
|
||
<taxonomicName id="69366B4111B91B79902572D824CFE40B" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. stuckenbergi" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="stuckenbergi">
|
||
<emphasis id="D1BB710E0CCE7190578CB7616EA8926D" italics="true" pageId="0" pageNumber="303">A. stuckenbergi</emphasis>
|
||
</taxonomicName>
|
||
holotype (CAS). (
|
||
<emphasis id="085DF08DCE55231C4A4F651F302D169C" bold="true" pageId="0" pageNumber="303">24</emphasis>
|
||
modified from original provided by Rikio Matsumoto, OMNH). Abbreviations: prepm - proepimeral process.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="48D9B9B7547FA4BC274BFD81EC15FDBA" pageId="0" pageNumber="303">
|
||
<emphasis id="F661881DBA1660A03523CE4E92B8BC22" bold="true" italics="true" pageId="0" pageNumber="303">Legs</emphasis>
|
||
(Fig.
|
||
<figureCitation id="14C5051376CD12221C1DDE9B574CEA4B" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Atrichops spp. ♂ 1 - 3 lateral habitus 4 - 6 head anterior view 1 A. adamastor (BMSA (D) 38908) 2 A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 3 A. stuckenbergi (BMSA (D) 95848) 4 A. adamastor (BMSA (D) 38904) 5 A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 6 A. stuckenbergi (BMSA (D) 95848). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures1-6" httpUri="https://binary.pensoft.net/fig/950648" pageId="0" pageNumber="303">1</figureCitation>
|
||
): Coxae yellow, anterior surface of mid- and hind coxae brown; fore coxa with short dark setulae on surface, more pronounced apically; mid coxa with long dark setulae on surface, hind coxa with dark setulae on anterior and lateral apical edges, and with well-developed anterior apical point; trochanters with some scattered short dark setulae, trochanters yellow, apical margins with darker markings, more pronounced than in ♀; fore and mid femora, and basal
|
||
<normalizedToken id="51CF0F57E8EBB211E40B705F82C7C5B0" originalValue="¼">1/4</normalizedToken>
|
||
of hind femur yellow, apical
|
||
<normalizedToken id="5A602728F8DE8696D5ACB7B4F36C8687" originalValue="¾">3/4</normalizedToken>
|
||
of hind femur dark brown (♀ with all femora yellow); fore, mid and hind femora with small anterior apical dark mark; fore, mid and hind tibiae dark brown (♀ with fore and mid tibiae yellow); all tarsi dark brown (♀ with yellow fore and mid tarsi); fore tarsal claws symmetrical, empodium at least 2
|
||
<normalizedToken id="35FE84305C700D88CBA75F6CE9E5C850" originalValue="×">x</normalizedToken>
|
||
size of pulvilli on fore legs; fore femur covered with dark setulae on all surfaces, posteroventral setulae forming row, mid femur with row of longer anteroventral setae and hind femur with similar dark setulae on ventral and dorsal surfaces; hind leg stouter than fore and mid legs; fore tibia and tarsi densely covered with long setulae along dorsal and ventral surfaces, at least as long or longer than width of segments (♀ without long hairs); hind tarsomeres 1.01-1.02 (♂) and 0.87-1.02 (♀) times as long as hind tibia.
|
||
</paragraph>
|
||
<paragraph id="606FEA6A4F1BEA12BD248ED4C51CC0A4" pageId="0" pageNumber="303">
|
||
<emphasis id="7BA415948B194C543F1720E6B7D083F4" bold="true" italics="true" pageId="0" pageNumber="303">Wing</emphasis>
|
||
(Fig.
|
||
<figureCitation id="0FE8CDE9DF039F1C2035BC8409B4E59E" captionStart="Figures 25–30" captionStartId="F5" captionText="Figures 25 - 30. Atrichops spp. ♂ ♀ wings dorsal view, A. adamastor: 25 ♂ (BMSA (D) 38904) 26 ♀ (BMSA (D) 38904); A. intermedius sp. nov.: 27 ♂ paratype (NMSA-Dip. 81811) 28 ♀ paratype (NMSA-Dip. 212861) A. stuckenbergi: 29 ♂ (BMSA (D) 94161) 30 ♀ (BMSA (D) 95861). Abbreviations: br - basal radial cell; bm - basal medial cell; cua - anterior cubital cell; CuA + CuP - anterior branch of cubital vein + posterior branch of cubital vein; d - discal cell; M 1 - first branch of media; m 1 - first medial cell; M 2 - second branch of media; m 2 - second medial cell; M 3 - third branch of media; m 3 - third medial cell; R 1 - anterior branch of radius; r 1 - first radial cell; R 2 + 3 - second branch of radius; r 2 + 3 - second radial cell; R 4 - upper branch of third branch of radius; R 5 - lower branch of third branch of radius; Sc - subcostal vein; sc - subcostal cell. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures25-30" httpUri="https://binary.pensoft.net/fig/950653" pageId="0" pageNumber="303">25</figureCitation>
|
||
): Suffused brown on entire surface, with much darker pterostigma over area of veins
|
||
<emphasis id="4D178D67946FC7CAA5833A47D3318697" italics="true" pageId="0" pageNumber="303">R1</emphasis>
|
||
and
|
||
<emphasis id="5C2CBB5BE06C5AE2CEB0913A3802DFE9" italics="true" pageId="0" pageNumber="303">R2+3</emphasis>
|
||
and cell
|
||
<emphasis id="3C9CCF4F3C048966B9F13C4D03A56DFF" italics="true" pageId="0" pageNumber="303">r1</emphasis>
|
||
; vein
|
||
<emphasis id="BA03A2D5E1EEF133B32E772D2B2B9FC9" italics="true" pageId="0" pageNumber="303">R4+5</emphasis>
|
||
with setulae dorsally, extending to vein
|
||
<emphasis id="22E69F8FB9165144C9ADCF874DF9AE59" italics="true" pageId="0" pageNumber="303">R5</emphasis>
|
||
; veins dark brown; cell
|
||
<emphasis id="88856443393811D26AB6ABE3F37E354C" italics="true" pageId="0" pageNumber="303">br</emphasis>
|
||
and discal cell uniform in colour; costa with distinct downward flexure over pterostigma; cell
|
||
<emphasis id="38DFA52BF12EBFCA69E57B4FBE32FF89" italics="true" pageId="0" pageNumber="303">cua</emphasis>
|
||
closed at short distance from wing margin, cell
|
||
<emphasis id="6D39BBB7E1D765247D30B541CB5DAA10" italics="true" pageId="0" pageNumber="303">m3</emphasis>
|
||
open, veins
|
||
<emphasis id="58C5D25F93D57A1137CE8D711E7374E8" italics="true" pageId="0" pageNumber="303">M1</emphasis>
|
||
,
|
||
<emphasis id="DC4DD95AA973050DFC7F438E461CD85A" italics="true" pageId="0" pageNumber="303">M2</emphasis>
|
||
,
|
||
<emphasis id="24E87945B16F624E34EF430C60183B0F" italics="true" pageId="0" pageNumber="303">M3</emphasis>
|
||
present; halter with yellow stalk and dark knob in both sexes and ♀ holotype (Fig.
|
||
<figureCitation id="652909132A871520592EDB7940A173CB" captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Atrichops spp. ♀ 7 - 9 lateral habitus 10 - 12 head anterior view 7 A. adamastor holotype and specimen labels (MZLU 00205685) 8 A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 9 A. stuckenbergi (CAS) 10 A. adamastor holotype (MZLU 00205685) 11 A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 12 A. stuckenbergi (BMSA (D) 95856). (7, 10 modified from originals provided by Rune Bygebjerg, MZLU, CC BY-NC 2.0 9, 12 modified from originals provided by Rikio Matsumoto, Osaka Museum of Natural History). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures7-12" httpUri="https://binary.pensoft.net/fig/950649" pageId="0" pageNumber="303">7</figureCitation>
|
||
). This differs from original description of
|
||
<taxonomicName id="804DE2FE1A841B43F9AB512868A7F4C3" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
|
||
<emphasis id="A94A24CBFB02F819F7723600C90A97BF" italics="true" pageId="0" pageNumber="303">A. adamastor</emphasis>
|
||
</taxonomicName>
|
||
♀ holotype with
|
||
<bibRefCitation id="68914D0B7EEADEB8A303034224984013" author="Stuckenberg, BR" editor="Hanstroem, B" journalOrPublisher="Results of the Lund University Expedition in 1950 - 1951. V. Almqvist & Wiksell, Uppsala" pageId="0" pageNumber="303" pagination="216 - 308" refId="B14" refString="Stuckenberg, BR, 1960. Chapter XII. Diptera (Brachycera): Rhagionidae. In: Hanstroem, B, Brinck, P, Rudebeck, G, Eds., South African animal life. Results of the Lund University Expedition in 1950-1951. V. Almqvist & Wiksell, Uppsala: 216 - 308" title="Chapter XII. Diptera (Brachycera): Rhagionidae." volumeTitle="South African animal life." year="1960">Stuckenberg (1960</bibRefCitation>
|
||
: 274) noting "Halteres with pale yellowish knob."
|
||
</paragraph>
|
||
<paragraph id="61925AEF28F0AB49F2DAD5173F26DF2F" pageId="0" pageNumber="303">
|
||
<emphasis id="35D70BE95E1CA6D62FB36CF88086464F" bold="true" italics="true" pageId="0" pageNumber="303">Abdomen</emphasis>
|
||
: Dark brown, with anterodorsal margins of tergites 1-3 as well as entirety of sternites 1-5 yellow; tergites and sternites with short dark setulae, longer dark setulae on lateral margins of tergites, as well as long pale setulae on sternites 1-3; tergite 1 without median suture.
|
||
</paragraph>
|
||
<paragraph id="CF18736DA14472E6463FFD465E75B719" pageId="0" pageNumber="303">
|
||
<emphasis id="3D8AAF611F03F281076A396714274FB9" bold="true" italics="true" pageId="0" pageNumber="303">Terminalia</emphasis>
|
||
(Figs
|
||
<figureCitation id="9F9C82E096F18E8CE43EB64F410ADA4B" captionStart="Figures 31–36" captionStartId="F6" captionText="Figures 31 - 36. Atrichops spp., 31 - 33 ♂ terminalia 31 A. adamastor (BMSA (D) 38904) 32 A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 33 A. stuckenbergi (BMSA (D) 94160), 34 - 36 ♂ epandrium and cerci 34 A. adamastor (BMSA (D) 38904) 35 A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 36 A. stuckenbergi (BMSA (D) 94160). Abbreviations: aed tn - aedeagal tine; cerc - cercus; ej apod - ejaculatory apodeme; epand - epandrium; goncx - gonocoxite; goncx apod - gonocoxal apodeme; gonst - gonostylus; pm - paramere; pm apod - parameral apodeme; pm sh - parameral sheath. Scale bars: 0.15 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures31-36" httpUri="https://binary.pensoft.net/fig/950654" pageId="0" pageNumber="303">31</figureCitation>
|
||
,
|
||
<figureCitation id="B994BE280991B85101B53110CB7C6348" captionStart="Figures 31–36" captionStartId="F6" captionText="Figures 31 - 36. Atrichops spp., 31 - 33 ♂ terminalia 31 A. adamastor (BMSA (D) 38904) 32 A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 33 A. stuckenbergi (BMSA (D) 94160), 34 - 36 ♂ epandrium and cerci 34 A. adamastor (BMSA (D) 38904) 35 A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 36 A. stuckenbergi (BMSA (D) 94160). Abbreviations: aed tn - aedeagal tine; cerc - cercus; ej apod - ejaculatory apodeme; epand - epandrium; goncx - gonocoxite; goncx apod - gonocoxal apodeme; gonst - gonostylus; pm - paramere; pm apod - parameral apodeme; pm sh - parameral sheath. Scale bars: 0.15 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures31-36" httpUri="https://binary.pensoft.net/fig/950654" pageId="0" pageNumber="303">34</figureCitation>
|
||
): Epandrium and cercus dark brown; hypoproct dark brown; epandrium, hypandrium and cercus with dark setulae; gonostylus finger-like with rounded apex; gonocoxite narrowing from middle towards base, apically rounded with long setulae; parameral apodeme long, extending to base of gonocoxite in ventral view; gonocoxal apodeme markedly shorter than gonocoxite.
|
||
</paragraph>
|
||
<paragraph id="C31A570B8C796433BE2EB4FFC1060578" pageId="0" pageNumber="303">
|
||
<emphasis id="2ADCC52464EB3E142AA077653AC4692F" bold="true" pageId="0" pageNumber="303">
|
||
Female.
|
||
<emphasis id="FE9DC842FA2F0D8D427F2264713578E8" italics="true" pageId="0" pageNumber="303">Terminalia</emphasis>
|
||
</emphasis>
|
||
:
|
||
<bibRefCitation id="5DD32E2F368D3789F33F488F061C8E99" author="Stuckenberg, BR" editor="Hanstroem, B" journalOrPublisher="Results of the Lund University Expedition in 1950 - 1951. V. Almqvist & Wiksell, Uppsala" pageId="0" pageNumber="303" pagination="216 - 308" refId="B14" refString="Stuckenberg, BR, 1960. Chapter XII. Diptera (Brachycera): Rhagionidae. In: Hanstroem, B, Brinck, P, Rudebeck, G, Eds., South African animal life. Results of the Lund University Expedition in 1950-1951. V. Almqvist & Wiksell, Uppsala: 216 - 308" title="Chapter XII. Diptera (Brachycera): Rhagionidae." volumeTitle="South African animal life." year="1960">Stuckenberg (1960)</bibRefCitation>
|
||
did not describe proepimeral process, which is reduced flap-like process (Fig.
|
||
<figureCitation id="48BDB63EFB2C51F8C32B2FD0C33E52FF" captionStart="Figures 19–24" captionStartId="F4" captionText="Figures 19 - 24. Atrichops spp. ♂ ♀ proepimeral process, lateral view 19 ♂ A. adamastor (BMSA (D) 38904) 20 ♂ A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 21 ♂ A. stuckenbergi (BMSA (D) 94158) 22 ♀ A. adamastor (BMSA (D) 38899) 23 ♀ A. intermedius sp. nov. paratype (NMSA-Dip. 212861) 24 ♀ A. stuckenbergi holotype (CAS). (24 modified from original provided by Rikio Matsumoto, OMNH). Abbreviations: prepm - proepimeral process." figureDoi="10.3897/afrinvertebr.64.113133.figures19-24" httpUri="https://binary.pensoft.net/fig/950652" pageId="0" pageNumber="303">22</figureCitation>
|
||
), much as in ♂. Cercus dark brown with dark setulae; genital fork (Fig.
|
||
<figureCitation id="7B1DFCB77CAC7F005E772CC9145FB874" captionStart="Figures 37–39" captionStartId="F7" captionText="Figures 37 - 39. Atrichops spp., ♀ genital fork and spermatheca 37 A. adamastor (BMSA (D) 38899 38 A. intermedius sp. nov. paratype (NMSA-Dip. 212861) 39 A. stuckenbergi (BMSA (D) 95855). Abbreviations: gen fk - genital fork; spmth - spermatheca. Scale bars: 0.15 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures37-39" httpUri="https://binary.pensoft.net/fig/950655" pageId="0" pageNumber="303">37</figureCitation>
|
||
) with slender distal apodeme, median lobe with deep apical emargination, paired apical lobes with somewhat square appearance, arms each gradually rounded; 3 sclerotised, somewhat oblong, spermathecae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0AEAC869A943CF3FAE55FC2541F36903" pageId="0" pageNumber="303" type="distribution">
|
||
<paragraph id="748F4D022C967D18E3770387D95A75CA" pageId="0" pageNumber="303">Distribution.</paragraph>
|
||
<paragraph id="9797F4F440B86D180A50280BBEAA13DE" pageId="0" pageNumber="303">South Africa (Western Cape Province).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E7E40E033EF601D954E9F48D1B57B93C" pageId="0" pageNumber="303" type="biology_ecology">
|
||
<paragraph id="C0B943BBCE5CFC1443391E25BC3FFA8A" pageId="0" pageNumber="303">Biology.</paragraph>
|
||
<paragraph id="584A2B7FA118D1AB3839118A78E2209F" pageId="0" pageNumber="303">
|
||
The species was observed and photographed feeding on the Cape River Frog,
|
||
<taxonomicName id="1BBAC1733012A73AB9EDD13223BD9BB9" class="Amphibia" family="Ranidae" genus="Amieta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Amieta fuscigula" order="Anura" pageId="0" pageNumber="303" phylum="Chordata" rank="species" species="fuscigula">
|
||
<emphasis id="4B9745E2ED2AA0E8D72D1850ECD68B22" italics="true" pageId="0" pageNumber="303">Amieta fuscigula</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken id="F4C4C5D73A698742781AC248BC101282" originalValue="Duméril">Dumeril</normalizedToken>
|
||
& Bibron in the Garden Route Botanical Garden, George, South Africa, by Colin Ralston (iNaturalist observation 9344668). While this behaviour has been informally observed before, it is the first published photographic observation of the species exhibiting typical
|
||
<taxonomicName id="B49634EF88D5A2EF3BCE69D0C6B47064" authorityName="Verrall" authorityYear="1909" class="Insecta" family="Athericidae" genus="Atrichops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichops" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="488A1E8B56A9C63629B6DEDFF946AADC" italics="true" pageId="0" pageNumber="303">Atrichops</emphasis>
|
||
</taxonomicName>
|
||
behaviour. It also emphasizes the importance of citizen science in highlighting species interactions that might otherwise go unrecorded or unnoticed. Although an attempt was made to sample material from the Botanical Garden for study, it was unsuccessful, possibly due to heavy rains earlier in the season.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |