161 lines
21 KiB
XML
161 lines
21 KiB
XML
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<mods:title id="8A919F6F5B3C85C157DA8BB6551D41D7">Neogene and Quaternary fossil remains of beaked whales (Cetacea, Odontoceti, Ziphiidae) from deep-sea deposits off Crozet and Kerguelen islands, Southern Ocean</mods:title>
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<mods:namePart id="CCA8F5902A9AF2AEFCA3A23DF4BA420E">Lambert, Olivier</mods:namePart>
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<mods:namePart id="9823A142AC96FBD53DE2D507496CE0A0">Muizon, Christian de</mods:namePart>
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<mods:namePart id="FAC03EF7D90B970916B18A1B9EC44A82">Duhamel, Guy</mods:namePart>
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<mods:namePart id="B051CA0B543D4471DD24B53AF4E78CE7">Plicht, Johannes Van Der</mods:namePart>
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<mods:title id="FD353EB3CEE8EECE0A6966F1173AF7A2">Geodiversitas</mods:title>
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<mods:date id="E100BDF8233D7DB158BC0954E2C8B064">2018</mods:date>
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<mods:number id="4DD51CF3334114B717DE4CA9500E588B">2018-03-29</mods:number>
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<mods:number id="EC1D5A43515D282B1ADBEF4E2A85917A">40</mods:number>
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<paragraph id="A45036AFBD4B7A7CFF5EA9B6FD7DF95A" blockId="10.[146,761,1745,1772]" box="[146,761,1745,1772]" pageId="10" pageNumber="143">
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<taxonomicName id="63EF4D2CBD4B7A7CFF5EA9B6FD7DF95A" authority="Bianucci, Lambert & Post, 2007" authorityName="Bianucci, Lambert & Post" authorityYear="2007" box="[146,761,1745,1772]" class="Mammalia" family="Hyperoodontidae" genus="Africanacetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="10" pageNumber="143" phylum="Chordata" rank="species" species="ceratopsis">
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<emphasis id="969BEABDBD4B7A7CFF5EA9B6FE15F95A" bold="true" box="[146,401,1745,1772]" italics="true" pageId="10" pageNumber="143">Africanacetus ceratopsis</emphasis>
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<bibRefCitation id="C07E4B5EBD4B7A7CFE54A9B5FD7DF95A" author="BIANUCCI G. & LAMBERT O. & POST K." box="[408,761,1746,1772]" pageId="10" pageNumber="143" pagination="5 - 62" refId="ref15041" refString="BIANUCCI G., LAMBERT O. & POST K. 2007. - A high diversity in fossil beaked whales (Odontoceti, Ziphiidae) recovered by trawling from the sea floor off South Africa. Geodiversitas 29: 5 - 62." type="journal article" year="2007">Bianucci, Lambert & Post, 2007</bibRefCitation>
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</taxonomicName>
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<paragraph id="A45036AFBD4B7A7CFF48A87DFDC5F85D" blockId="10.[131,775,1818,2027]" pageId="10" pageNumber="143">
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REFERRED SPECIMENS AND LOCALITIES. — Partial cranium MNHN.F.COI12 including rostrum, premaxillary sac fossae, part of both supraorbital regions, and vertex (
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<figureCitation id="3CD42A2ABD4B7A7CFDD9A829FDCAF8D0" box="[533,590,1870,1894]" captionStart="FIG" captionStartId="11.[132,143,1985,2003]" captionTargetBox="[193,1403,215,1948]" captionTargetId="figure-23@11.[745,1320,757,1308]" captionTargetPageId="11" captionText="FIG. 5. — Partial cranium of Africanacetus ceratopsis (MNHN.F.COI12): A, dorsal view; B, right lateral view; C, detail of the vertex in dorsal view; D, anterodorsal view. Scale bars: 100 mm." figureDoi="http://doi.org/10.5281/zenodo.5745704" httpUri="https://zenodo.org/record/5745704/files/figure.png" pageId="10" pageNumber="143">Fig. 5</figureCitation>
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);
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<materialsCitation id="14873CF2BD4B7A7CFDAFA829FC87F837" latitude="-49.8" location="Skiff Bank" longLatPrecision="1101" longitude="66.46667" pageId="10" pageNumber="143" specimenCount="1" stateProvince="Iles Kerguelen">
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geographic coordinates
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<geoCoordinate id="C1DB5068BD4B7A7CFF02A80EFE99F837" box="[206,285,1897,1921]" degrees="49" direction="south" minutes="48" orientation="latitude" pageId="10" pageNumber="143" precision="925" value="-49.8">49°48’S</geoCoordinate>
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,
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<geoCoordinate id="C1DB5068BD4B7A7CFEEAA80EFEF1F837" box="[294,373,1897,1921]" degrees="66" direction="east" minutes="28" orientation="longitude" pageId="10" pageNumber="143" precision="925" value="66.46667">66°28’E</geoCoordinate>
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(
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<location id="A1306074BD4B7A7CFE4CA80EFE6DF837" LSID="urn:lsid:plazi:treatment:2C4687B9BD4B7A7AFF5EA9B6FE5AF89A:A1306074BD4B7A7CFE4CA80EFE6DF837" box="[384,489,1897,1921]" latitude="-49.8" longLatPrecision="1101" longitude="66.46667" name="Skiff Bank" pageId="10" pageNumber="143" stateProvince="Iles Kerguelen">Skiff Bank</location>
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, SWW to
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<collectingRegion id="662BF84DBD4B7A7CFD81A80EFD78F837" box="[589,764,1897,1921]" country="French Southern Territories" name="Iles Kerguelen" pageId="10" pageNumber="143">Kerguelen Islands</collectingRegion>
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)
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</materialsCitation>
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; depth
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<quantity id="63179B4ABD4B7A7CFF73A8E4FE89F82D" box="[191,269,1923,1947]" metricMagnitude="3" metricUnit="m" metricValue="1.68" pageId="10" pageNumber="143" unit="m" value="1680.0">1680 m</quantity>
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. Partial cranium MNHN.F.COI2 including rostrum, premaxillary sac fossae, and part of both supraorbital regions (
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<figureCitation id="3CD42A2ABD4B7A7CFD08A8F9FD7CF800" box="[708,760,1950,1974]" captionStart="FIG" captionStartId="12.[132,143,1402,1420]" captionTargetBox="[141,1442,217,1358]" captionTargetId="figure-295@12.[698,1267,745,1286]" captionTargetPageId="12" captionText="FIG. 6. — Partial cranium of Africanacetus ceratopsis Bianucci, Lambert & Post, 2007 (MNHN.F.COI2): A, dorsal view; B, anterior view; C, right lateral view. Scale bar: 100 mm." figureDoi="http://doi.org/10.5281/zenodo.5745712" httpUri="https://zenodo.org/record/5745712/files/figure.png" pageId="10" pageNumber="143">Fig. 6</figureCitation>
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);
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<materialsCitation id="14873CF2BD4B7A7CFF48A8DFFED1F85D" latitude="-47.133" location="Kerguelen Islands" longLatPrecision="1" longitude="65.74983" pageId="10" pageNumber="143" specimenCount="1" stateProvince="Iles Kerguelen">
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geographic coordinates
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<geoCoordinate id="C1DB5068BD4B7A7CFEA4A8DFFE6AF866" box="[360,494,1976,2000]" degrees="47" direction="south" minutes="07" orientation="latitude" pageId="10" pageNumber="143" precision="1" seconds="58.8" value="-47.133">47°07’58.8”S</geoCoordinate>
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,
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<geoCoordinate id="C1DB5068BD4B7A7CFE3BA8DFFDF9F866" box="[503,637,1976,2000]" degrees="65" direction="east" minutes="44" orientation="longitude" pageId="10" pageNumber="143" precision="1" seconds="59.4" value="65.74983">65°44’59.4”E</geoCoordinate>
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(
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<quantity id="63179B4ABD4B7A7CFD45A8DFFD51F866" box="[649,725,1976,2000]" metricMagnitude="5" metricUnit="m" metricValue="2.9" pageId="10" pageNumber="143" unit="km" value="290.0">290 km</quantity>
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NW to
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<collectingRegion id="662BF84DBD4B7A7CFF52A8B4FECAF85D" box="[158,334,2003,2027]" country="French Southern Territories" name="Iles Kerguelen" pageId="10" pageNumber="143">Kerguelen Islands</collectingRegion>
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)
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</materialsCitation>
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; depth
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<quantity id="63179B4ABD4B7A7CFE52A8B4FE6FF85C" box="[414,491,2003,2027]" metricMagnitude="3" metricUnit="m" metricValue="1.6219999999999999" pageId="10" pageNumber="143" unit="m" value="1622.0">1622 m</quantity>
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(
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<figureCitation id="3CD42A2ABD4B7A7CFE37A8B4FDB7F85D" box="[507,563,2003,2027]" captionStart="FIG" captionStartId="4.[132,143,1125,1143]" captionTargetBox="[323,1264,214,1081]" captionTargetId="graphics-412@4.[324,1264,214,1081]" captionTargetPageId="4" captionText="FIG. 1. — Locality maps: A, schematic map providing the position of Crozet and Kerguelen islands in the Southern Ocean; B, schematic map of the Crozet and Kerguelen islands area, including the main localities where fossil ziphiid specimens («) described in this work were found via longline fishing,as well as two ODP sites (l). Light grey shading for areas with a water depth lower than 1000 m; dark grey shading for islands and continents." figureDoi="http://doi.org/10.5281/zenodo.5745696" httpUri="https://zenodo.org/record/5745696/files/figure.png" pageId="10" pageNumber="143">Fig. 1</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="ECF56524BD4B7A7AFCE1AFBFFE5AF89A" lastPageId="12" lastPageNumber="145" pageId="10" pageNumber="143" type="description">
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<paragraph id="A45036AFBD4B7A7CFCE1AFBFFB46FF46" blockId="10.[810,1457,216,2027]" box="[813,1218,216,242]" pageId="10" pageNumber="143">BRIEF DESCRIPTION AND COMPARISON</paragraph>
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<paragraph id="A45036AFBD4B7A7CFCE1AF90FB37FAFB" blockId="10.[810,1457,216,2027]" pageId="10" pageNumber="143">
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The rostrum is not complete anteriorly in both specimens, although it is probably close to the original length in MNHN.F.COI2. Probably half of the rostrum in missing in COI12. The rostrum is robust and nearly as wide as high at half of its preserved length on COI2. The mesorostral groove is completely filled with the vomer, the latter reaching a dorsal level much higher than the bordering premaxilla. A median suture is visible on the dorsal surface along the proximal part of the vomer on COI2. The lateral margin of the rostrum of COI2 is a sharp crest in its proximal part; this crest forms a thin blade markedly diverging towards the prominential notch; the blade is somewhat thicker on the left side. No crest is observed on COI12 (except at the rostrum base, close to the prominential notch) and in dorsal view the rostrum is significantly narrower and more slender than on the other specimen. In lateral view, no indication for individual alveoli could be found on the maxilla. Deeper on COI12, the prominential notch is followed posteriorly by a dome-shaped, moderately elevated rostral maxillary crest (
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<emphasis id="969BEABDBD4B7A7CFCFAAC31FCEEFCD9" box="[822,874,854,879]" italics="true" pageId="10" pageNumber="143">sensu</emphasis>
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<bibRefCitation id="C07E4B5EBD4B7A7CFCBDAC32FBE3FCD9" author="MEAD J. G. & FORDYCE R. E." box="[881,1127,853,879]" pageId="10" pageNumber="143" pagination="1 - 248" refId="ref16482" refString="MEAD J. G. & FORDYCE R. E. 2009. - The therian skull: a lexicon with emphasis on the odontocetes. Smithsonian Contributions to Zoology 627: 1 - 248. https: // doi. org / 10.5479 / si. 00810282.627" type="journal article" year="2009">Mead & Fordyce 2009</bibRefCitation>
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). The crest is higher and more posteriorly located on the right side. Compared to COI2, right and left crests are more elevated, massive, and rounded on COI12. A small dorsal infraorbital foramen is present at the top of each crest (on COI2) and a large foramen pierces the medial base of the crest, being followed anteriorly by a shallow depression. Anterolateral to the rostral maxillary crest, the antorbital notch is widely open anteriorly, barely limited laterally by a short anterior extension of the antorbital part of the maxilla and the lacrimojugal complex. Medial to the antorbital notch a small maxillary tubercle protrudes anteriorly. The maxillary tubercle is more salient in COI12, in relation to the larger maxillary crest in this specimen. Right and left premaxillary foramina are difficult to distinguish, hidden by a concretion. Each premaxillary sac fossa is thick, with the dorsal surface raising dorsomedially.
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</paragraph>
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<paragraph id="A45036AFBD4B7A7CFC8FAA34FB3FF95A" blockId="10.[810,1457,216,2027]" pageId="10" pageNumber="143">The ascending process of the premaxilla of MNHN.F.COI12 strongly raises dorsally, with its anterior surface being almost subvertical when reaching the level of the premaxillary crest. Each ascending process is distinctly constricted transversely at the level of the maximum width of the bony nares. The vertex is well preserved on COI12, with intact premaxillary crests. The latter are robust and the right crest diverges posterolaterally, whereas the left crest is directed more laterally. In dorsal view, the nasals are anteroposteriorly long and transversely wide; their anterolateral angle reaches the medial margin of the corresponding premaxillary crest, without being integrated in it. In dorsal view, the median suture of the nasal is salient anteriorly, dorsal to the presphenoid.</paragraph>
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<paragraph id="A45036AFBD4B7A7AFC8FA995FE5AF89A" blockId="10.[810,1457,216,2027]" lastBlockId="12.[131,777,1523,1836]" lastPageId="12" lastPageNumber="145" pageId="10" pageNumber="143">
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The thick vomer in the mesorostral groove, the dome-like rostral maxillary crest, and the dorsomedially raising premaxillary sac fossae are similarly observed in specimens of
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<taxonomicName id="63EF4D2CBD4B7A7CFA94A855FC48F8DD" authorityName="Bianucci, Lambert & Post" authorityYear="2007" class="Mammalia" family="Hyperoodontidae" genus="Africanacetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="10" pageNumber="143" phylum="Chordata" rank="species" species="ceratopsis">
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<emphasis id="969BEABDBD4B7A7CFA94A855FC48F8DD" italics="true" pageId="10" pageNumber="143">Africanacetus ceratopsis</emphasis>
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</taxonomicName>
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from
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<collectingCountry id="DCF8763FBD4B7A7CFBC0A836FB15F8DD" box="[1036,1169,1873,1899]" name="South Africa" pageId="10" pageNumber="143">South Africa</collectingCountry>
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(
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<bibRefCitation id="C07E4B5EBD4B7A7CFB6CA836FAF5F8DA" author="BIANUCCI G. & LAMBERT O. & POST K." box="[1184,1393,1873,1900]" pageId="10" pageNumber="143" pagination="5 - 62" refId="ref15041" refString="BIANUCCI G., LAMBERT O. & POST K. 2007. - A high diversity in fossil beaked whales (Odontoceti, Ziphiidae) recovered by trawling from the sea floor off South Africa. Geodiversitas 29: 5 - 62." type="journal article" year="2007">
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Bianucci
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<emphasis id="969BEABDBD4B7A7CFACFA835FAB1F8DD" box="[1283,1333,1873,1899]" italics="true" pageId="10" pageNumber="143">et al.</emphasis>
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2007
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</bibRefCitation>
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). The dimensions of the rostrum and facial region (
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<tableCitation id="E96D0314BD4B7A7CFAC2A816FAD9F83A" box="[1294,1373,1905,1932]" captionStart="TABLE" captionStartId="14.[132,143,219,237]" captionText="TABLE 5. — Measurements (in mm) on the partial crania of Africanacetus ceratopsis Bianucci,Lambert & Post,2007 (MNHN.F.COI2, COI12, and NMR 991-00001993 [holotype] [from Bianucci et al. 2007]). Abbreviations: +, not complete; –, no data." pageId="10" pageNumber="143">Table 5</tableCitation>
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) match well the South African sample of
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<taxonomicName id="63EF4D2CBD4B7A7CFBB4A8F5FB69F81D" authorityName="Bianucci, Lambert & Post" authorityYear="2007" box="[1144,1261,1938,1964]" class="Mammalia" family="Hyperoodontidae" genus="Africanacetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="10" pageNumber="143" phylum="Chordata" rank="species" species="ceratopsis">
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<emphasis id="969BEABDBD4B7A7CFBB4A8F5FB69F81D" box="[1144,1261,1938,1964]" italics="true" pageId="10" pageNumber="143">A. ceratopsis</emphasis>
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</taxonomicName>
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and the specimens of
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<taxonomicName id="63EF4D2CBD4B7A7CFC8BA8D6FC74F87A" box="[839,1008,1969,1996]" class="Mammalia" family="Hyperoodontidae" genus="Africanacetus" kingdom="Animalia" order="Cetacea" pageId="10" pageNumber="143" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="969BEABDBD4B7A7CFC8BA8D6FC4AF87D" box="[839,974,1969,1995]" italics="true" pageId="10" pageNumber="143">Africanacetus</emphasis>
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sp.
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</taxonomicName>
|
||
from the Banzare Bank, southernmost part of the Kerguelen Plateau, described by Gol’din & Vishnyakova (2012). The rostral maxillary crests are significantly lower on MNHN.F.COI2 and more rounded on MNHN.F.COI12. For part of the dimensions, the
|
||
<typeStatus id="7B54880DBD4D7A7AFE23A955FDD4F9FA" box="[495,592,1586,1612]" pageId="12" pageNumber="145" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="63EF4D2CBD4D7A7AFDB7A954FF4FF9DA" authorityName="Ichishima, Augustin, Toyofuku & Kitazato" authorityYear="2017" class="Mammalia" family="Hyperoodontidae" genus="Africanacetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="12" pageNumber="145" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis id="969BEABDBD4D7A7AFDB7A954FF4FF9DA" italics="true" pageId="12" pageNumber="145">Africanacetus gracilis</emphasis>
|
||
</taxonomicName>
|
||
is moderately to markedly smaller, especially for the measurements at rostrum base (see
|
||
<bibRefCitation id="C07E4B5EBD4D7A7AFE3BA915FD53F93A" author="ICHISHIMA H. & AUGUSTIN A. H. & TOYOFUKU T. & KITAZATO H." box="[503,727,1650,1676]" pageId="12" pageNumber="145" refId="ref15939" refString="ICHISHIMA H., AUGUSTIN A. H., TOYOFUKU T. & KITAZATO H. 2017. - A new species of Africanacetus (Odontoceti: Ziphiidae) found on the deep ocean floor off the coast of Brazil. Deep Sea Research Part II. https: // doi. org / 10.1016 / j. dsr 2.2016.12.002" type="book" year="2017">
|
||
Ichishima
|
||
<emphasis id="969BEABDBD4D7A7AFDA4A914FD1EF93A" box="[616,666,1650,1676]" italics="true" pageId="12" pageNumber="145">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
). As in the latter, the antorbital notch of both specimens is more anteriorly located than in the
|
||
<typeStatus id="7B54880DBD4D7A7AFE03A9D5FDAAF97A" box="[463,558,1714,1740]" pageId="12" pageNumber="145" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="63EF4D2CBD4D7A7AFD94A9D5FD53F97A" authorityName="Bianucci, Lambert & Post" authorityYear="2007" box="[600,727,1714,1740]" class="Mammalia" family="Hyperoodontidae" genus="Africanacetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="12" pageNumber="145" phylum="Chordata" rank="species" species="ceratopsis">
|
||
<emphasis id="969BEABDBD4D7A7AFD94A9D5FD53F97A" box="[600,727,1714,1740]" italics="true" pageId="12" pageNumber="145">A. ceratopsis</emphasis>
|
||
</taxonomicName>
|
||
and several referred specimens; such differences may be explained by sexual dimorphism or any other intraspecific, populationrelated morphological variation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |