253 lines
28 KiB
XML
253 lines
28 KiB
XML
<document id="79B430526EEC437A3EF6F7BB859AAB0F" ID-DOI="10.11646/zootaxa.5405.2.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="10603393" ID-ZooBank="DD57E713-2FD1-4A05-8E5B-13A0D27A0B5D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1706773369097" checkinUser="plazi" docAuthor="Hong, Yong, Inkhavilay, Khamla & James, Samuel W." docDate="2024" docId="E73187B3813A6F6F799C2F4EE4BF2204" docLanguage="en" docName="zootaxa.5405.2.7.pdf" docOrigin="Zootaxa 5405 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5405.2.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amynthas xayi Hong and James 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="277" masterDocId="1B08FFCB81396F6B790B2B30E07A2565" masterDocTitle="Three new earthworms of the Amynthas corticis-group (Clitellata: Megascolecidae) from the Nam Et-Phouleoi, Laos" masterLastPageNumber="280" masterPageNumber="273" pageNumber="276" updateTime="1707146910219" updateUser="guilherme" zenodo-license-document="CLOSED">
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<mods:mods id="625A586979F5163C8A61B1E0FAA5D3D9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="580D9BDFB4D93A08CBA67BF8B3A03B43">
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<mods:title id="CD930E8106683E4259CAD23DFFE028C4">Three new earthworms of the Amynthas corticis-group (Clitellata: Megascolecidae) from the Nam Et-Phouleoi, Laos</mods:title>
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</mods:titleInfo>
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<mods:name id="26376C0F453251599814FFA698B7E592" type="personal">
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<mods:role id="C8D0CF3D35CCFD8C4DF755CA835164E6">
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<mods:roleTerm id="881A504849E1391CC7A792AED01E1768">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7C200FA2BD273D86863E95904508DABF">Hong, Yong</mods:namePart>
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<mods:affiliation id="50D4944F6FA3CE6572BCDAF2A60B3FA8">Department of Agricultural Biology, College of Agriculture & Life Sciences, Jeonbuk National University, Jeonju 54896, Korea;</mods:affiliation>
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</mods:name>
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<mods:name id="BE24F0DEA6CF3605FB3837727DCB922D" type="personal">
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<mods:role id="561ED533B19642DD3E09F03AFE5528E2">
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<mods:roleTerm id="15F78E81453BFE636ACC9F10D02C8128">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2EFEFA214B95AB03CD8247516C3697E9">Inkhavilay, Khamla</mods:namePart>
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<mods:affiliation id="4CDF9B35C53DD4258C30A061EB45BF5E">Center of Excellence in Biodiversity, Research Academic and Service Office, National University of Laos;</mods:affiliation>
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</mods:name>
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<mods:name id="36B25637548CE081FBBEE1570C3CEBB2" type="personal">
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<mods:roleTerm id="6CE6183DC1A3E598FEEF3447F156BB88">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2843F9567227A3EC5826D517EA896A2F">James, Samuel W.</mods:namePart>
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<mods:affiliation id="0C316D64D34F369BE44781B13608B09E">Department of Biology, University of Iowa, Iowa City, Iowa, USA 52242.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="D9F494366BF80608883C25476F0E3AAF">text</mods:typeOfResource>
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<mods:title id="218F9AAF1A7F239387A9FFF11A69E94E">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="EA16481FD4F061E1603AF8F3CD4B58EF">
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<mods:date id="29476ADFBDA341139971A539BDDD4136">2024</mods:date>
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<mods:detail id="6E1284FDFC3D6A6E7B8F046F9A7D67BF" type="pubDate">
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<mods:number id="F629B85C2FFC231F995EAEC3F7D4895F">2024-01-30</mods:number>
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<mods:detail id="28A0E63FE9A175E873EC245064812195" type="volume">
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<mods:number id="7BAC4647CEDA49870D777C14A1BACE0A">5405</mods:number>
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<mods:detail id="4EBD98EAEDB142D2F805AB010FB5E2B3" type="issue">
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<mods:number id="5A96EC6938FF76C3C84862E7B2BEFC84">2</mods:number>
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<mods:start id="2F4FAA92BCF770023DEBD555A9FE07F8">273</mods:start>
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<mods:location id="5E7B96F33942134377A09E3224E2319C">
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<mods:url id="896CD24DFBEE961EAD4878A018113291">http://dx.doi.org/10.11646/zootaxa.5405.2.7</mods:url>
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</mods:location>
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<mods:classification id="78D84C65AF7170894EDF2896F28EF12F">journal article</mods:classification>
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<mods:identifier id="FB6435D4CD8A8CDEC0128A0746872571" type="DOI">10.11646/zootaxa.5405.2.7</mods:identifier>
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<mods:identifier id="8F5CBDA70B1ED322D8EF07A335813ACD" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D867F028204302E0746C0B5F24C48315" type="Zenodo-Dep">10603393</mods:identifier>
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<mods:identifier id="109A928DDDCEE7FD2C09337864571F14" type="ZooBank">DD57E713-2FD1-4A05-8E5B-13A0D27A0B5D</mods:identifier>
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</mods:mods>
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<treatment id="E73187B3813A6F6F799C2F4EE4BF2204" ID-DOI="http://doi.org/10.5281/zenodo.10603401" ID-Zenodo-Dep="10603401" LSID="urn:lsid:plazi:treatment:E73187B3813A6F6F799C2F4EE4BF2204" httpUri="http://treatment.plazi.org/id/E73187B3813A6F6F799C2F4EE4BF2204" lastPageId="4" lastPageNumber="277" pageId="3" pageNumber="276">
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<subSubSection id="2782652E813A6F68799C2F4EE2F221FC" box="[151,648,1150,1177]" pageId="3" pageNumber="276" type="nomenclature">
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<paragraph id="6F2736A5813A6F68799C2F4EE2F221FC" blockId="3.[151,648,1150,1213]" box="[151,648,1150,1177]" pageId="3" pageNumber="276">
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<heading id="346F81C9813A6F68799C2F4EE2F221FC" bold="true" box="[151,648,1150,1177]" fontSize="11" level="1" pageId="3" pageNumber="276" reason="1">
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<emphasis id="5DECEAB7813A6F68799C2F4EE2F221FC" bold="true" box="[151,648,1150,1177]" pageId="3" pageNumber="276">
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<taxonomicName id="A8984D26813A6F68799C2F4EE25921FC" authority="Hong and James" authorityName="Hong and James" authorityYear="2024" box="[151,547,1150,1177]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="3" pageNumber="276" phylum="Annelida" rank="species" species="xayi" status="sp. nov.">
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||
<emphasis id="5DECEAB7813A6F68799C2F4EE13621FC" bold="true" box="[151,332,1150,1177]" italics="true" pageId="3" pageNumber="276">Amynthas xayi</emphasis>
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||
Hong and James
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||
</taxonomicName>
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||
<taxonomicNameLabel id="46DF57CC813A6F687B202F4FE2F221FC" box="[555,648,1151,1177]" pageId="3" pageNumber="276" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="2782652E813A6F68799C2F93E09121D8" box="[151,235,1187,1213]" pageId="3" pageNumber="276" type="description">
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<paragraph id="6F2736A5813A6F68799C2F93E09121D8" blockId="3.[151,648,1150,1213]" box="[151,235,1187,1213]" pageId="3" pageNumber="276">
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||
(
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||
<figureCitation id="F7A32A20813A6F6879942F93E09821D8" box="[159,226,1187,1213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1324,1348]" captionTargetBox="[494,1047,250,1279]" captionTargetId="figure-51@4.[151,1435,234,1303]" captionTargetPageId="4" captionText="FIGURE 3. Amynthas xayi sp. nov. (A, B). A: ventral view; B: intestinal caeca; C: spermatheca. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10603403" httpUri="https://zenodo.org/record/10603403/files/figure.png" pageId="3" pageNumber="276">Fig. 3</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="2782652E813A6F68799C2FDAE5312014" pageId="3" pageNumber="276" type="materials_examined">
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||
<paragraph id="6F2736A5813A6F68799C2FDAE5312014" blockId="3.[151,1437,1258,2041]" pageId="3" pageNumber="276">
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||
<emphasis id="5DECEAB7813A6F68799C2FDAE1FB2061" bold="true" box="[151,385,1258,1284]" pageId="3" pageNumber="276">Material examined.</emphasis>
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||
<materialsCitation id="DFF03CF8813A6F6878832FDBE57A2028" collectingDate="2005-07-21" collectionCode="BDNUL" collectorName="Phimphet & Xay & Ammaline & Phoang & K. Inkhavilay" country="Laos" county="Viengthong District" elevation="792" latitude="20.257433" location="Thad Ang waterfall area" longLatPrecision="1" longitude="103.14868" municipality="Nam Et-Phouleoi" pageId="3" pageNumber="276" specimenCode="BDNUL 0057" specimenCount="1" stateProvince="Houaphanh Province" typeStatus="holotype">
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||
<typeStatus id="B0238807813A6F6878832FDBE18F2060" box="[392,501,1259,1285]" pageId="3" pageNumber="276" type="holotype">Holotype</typeStatus>
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||
:
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<specimenCount id="799EFD2C813A6F6878F42FDBE20E2060" box="[511,628,1259,1285]" pageId="3" pageNumber="276">1 clitellate</specimenCount>
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||
(
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<specimenCode id="3F3E9EDE813A6F687B882FDBE3532060" box="[643,809,1258,1285]" collectionCode="BDNUL" pageId="3" pageNumber="276">BDNUL 0057</specimenCode>
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):
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<collectingCountry id="178F7635813A6F687A372FDBE30F2060" box="[828,885,1259,1285]" name="Laos" pageId="3" pageNumber="276">Laos</collectingCountry>
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,
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||
<collectingRegion id="AD5CF847813A6F687A8B2FDBE4152060" box="[896,1135,1259,1285]" country="Laos" name="Houaphan" pageId="3" pageNumber="276">Houaphanh Province</collectingRegion>
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||
,
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<collectingCounty id="86464E29813A6F687D712FDBE5212060" box="[1146,1371,1259,1285]" pageId="3" pageNumber="276">Viengthong District</collectingCounty>
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||
,
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||
<collectingMunicipality id="8F43ACDF813A6F687C6D2FDBE167204C" pageId="3" pageNumber="276">Nam Et-Phouleoi</collectingMunicipality>
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||
NPA,
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||
<location id="6A47607E813A6F6878612E3FE2FA204C" LSID="urn:lsid:plazi:treatment:E73187B3813A6F6F799C2F4EE4BF2204:6A47607E813A6F6878612E3FE2FA204C" box="[362,640,1295,1321]" country="Laos" county="Viengthong District" latitude="20.257433" longLatPrecision="1" longitude="103.14868" municipality="Nam Et-Phouleoi" name="Thad Ang waterfall area" pageId="3" pageNumber="276" stateProvince="Houaphanh Province">Thad Ang waterfall area</location>
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||
,
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||
<quantity id="A8609B40813A6F687B862E3FE2AC204C" box="[653,726,1295,1321]" metricMagnitude="2" metricUnit="m" metricValue="7.92" pageId="3" pageNumber="276" unit="m" value="792.0">
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||
<elevation id="E4B5D196813A6F687B862E3FE2AC204C" box="[653,726,1295,1321]" metricMagnitude="2" metricUnit="m" metricValue="7.92" pageId="3" pageNumber="276" unit="m" value="792.0">792 m</elevation>
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||
</quantity>
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||
(
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||
<geoCoordinate id="0AAC5062813A6F687BED2E20E303204E" box="[742,889,1296,1323]" degrees="20" direction="north" minutes="15.446" orientation="latitude" pageId="3" pageNumber="276" precision="1" value="20.257433">20°15.446′N</geoCoordinate>
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||
,
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||
<geoCoordinate id="0AAC5062813A6F687A8E2E20E458204E" box="[901,1058,1296,1323]" degrees="103" direction="east" minutes="08.921" orientation="longitude" pageId="3" pageNumber="276" precision="1" value="103.14868">103°08.921′E</geoCoordinate>
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||
), good forest condition, dark soil and thick litter layer,
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||
<date id="1B261065813A6F68789E2E03E24D2028" box="[405,567,1330,1357]" pageId="3" pageNumber="276" value="2005-07-21">
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||
<collectingDate id="0B62E98D813A6F68789E2E03E24D2028" box="[405,567,1330,1357]" pageId="3" pageNumber="276" value="2005-07-21">July 21, 2005</collectingDate>
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||
</date>
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||
,
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<collectorName id="C26D5373813A6F687B4D2E03E2CE2028" box="[582,692,1331,1357]" pageId="3" pageNumber="276">Phimphet</collectorName>
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,
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<collectorName id="C26D5373813A6F687BC82E03E2892028" box="[707,755,1331,1357]" pageId="3" pageNumber="276">Xay</collectorName>
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,
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<collectorName id="C26D5373813A6F687A0B2E03E3032028" box="[768,889,1331,1357]" pageId="3" pageNumber="276">Ammaline</collectorName>
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,
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<collectorName id="C26D5373813A6F687A8C2E03E3A42028" box="[903,990,1331,1357]" pageId="3" pageNumber="276">Phoang</collectorName>
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, and
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<collectorName id="C26D5373813A6F687D2B2E03E4C42028" box="[1056,1214,1331,1357]" pageId="3" pageNumber="276">K. Inkhavilay</collectorName>
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colls
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</materialsCitation>
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||
.
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<materialsCitation id="DFF03CF8813A6F687C042E03E3732014" collectingDate="2005-07-21" collectionCode="BDNUL" collectorName="Phimphet & Xay & Ammaline & Phoang & K. Inkhavilay" country="Laos" county="Viengthong District" elevation="792" latitude="20.257433" location="Thad Ang" longLatPrecision="1" longitude="103.14868" municipality="Nam Et-Phouleoi" pageId="3" pageNumber="276" specimenCode="BDNUL 0058, 0059" specimenCount="2" stateProvince="Houaphan" typeStatus="paratype">
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<typeStatus id="B0238807813A6F687C042E03E5FA2028" box="[1295,1408,1331,1357]" pageId="3" pageNumber="276" type="paratype">Paratypes</typeStatus>
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||
:
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||
<specimenCount id="799EFD2C813A6F687C852E03E1782014" pageId="3" pageNumber="276">2 clitellates</specimenCount>
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||
(
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||
<specimenCode id="3F3E9EDE813A6F68781A2E67E1CE2014" box="[273,436,1366,1393]" collectionCode="BDNUL" pageId="3" pageNumber="276">BDNUL 0058</specimenCode>
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||
,
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<specimenCode id="3F3E9EDE813A6F6878B42E66E1812014" box="[447,507,1366,1393]" collectionCode="BDNUL" pageId="3" pageNumber="276">0059</specimenCode>
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||
), same data as holotype
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||
</materialsCitation>
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||
.
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<materialsCitation id="DFF03CF8813A6F687A182E67E53D2014" box="[787,1351,1366,1393]" collectingDate="2005-07-21" collectionCode="BDNUL" collectorName="Phimphet & Xay & Ammaline & Phoang & K. Inkhavilay" country="Laos" county="Viengthong District" elevation="792" latitude="20.257433" location="Thad Ang" longLatPrecision="1" longitude="103.14868" municipality="Nam Et-Phouleoi" pageId="3" pageNumber="276" specimenCount="5" stateProvince="Houaphan" typeStatus="paratype">
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||
Non-types:
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||
<specimenCount id="799EFD2C813A6F687A9D2E66E4632014" box="[918,1049,1366,1393]" pageId="3" pageNumber="276">5 clitellates</specimenCount>
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||
, same data as for holotype
|
||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="2782652E813A6F6879CC2E4BE3A320F0" box="[199,985,1403,1429]" pageId="3" pageNumber="276" type="etymology">
|
||
<paragraph id="6F2736A5813A6F6879CC2E4BE3A320F0" blockId="3.[151,1437,1258,2041]" box="[199,985,1403,1429]" pageId="3" pageNumber="276">
|
||
<emphasis id="5DECEAB7813A6F6879CC2E4BE13420F0" bold="true" box="[199,334,1403,1429]" pageId="3" pageNumber="276">Etymology.</emphasis>
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||
The species is named after Mr. Xay, one of the collectors.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2782652E813A6F6879CC2EAEE1192364" pageId="3" pageNumber="276" type="diagnosis">
|
||
<paragraph id="6F2736A5813A6F6879CC2EAEE1192364" blockId="3.[151,1437,1258,2041]" pageId="3" pageNumber="276">
|
||
<emphasis id="5DECEAB7813A6F6879CC2EAEE13820DD" bold="true" box="[199,322,1438,1464]" pageId="3" pageNumber="276">Diagnosis.</emphasis>
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Four pairs of spermathecal pores deep in 5/6–8/9 at 9
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<superScript id="98ED9BED813A6F687A972EAEE3D320C9" attach="left" box="[924,937,1438,1452]" fontSize="6" pageId="3" pageNumber="276">th</superScript>
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||
setal line; male pores superficial in XVIII at centers of round porophores, 0.17–0.29 circumference apart; no genital markings or papillae; caeca pinnate; first dorsal pore in 5/6.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2782652E813A6F6F79CC2D3AE2E0209C" lastPageId="4" lastPageNumber="277" pageId="3" pageNumber="276" type="description">
|
||
<paragraph id="6F2736A5813A6F6879CC2D3AE3A623BC" blockId="3.[151,1437,1258,2041]" pageId="3" pageNumber="276">
|
||
<emphasis id="5DECEAB7813A6F6879CC2D3AE1232341" bold="true" box="[199,345,1546,1572]" pageId="3" pageNumber="276">Description.</emphasis>
|
||
Dimensions
|
||
<quantity id="A8609B40813A6F6878E72D3BE21F2340" box="[492,613,1547,1573]" metricMagnitude="-2" metricUnit="m" metricValue="8.3" metricValueMax="9.0" metricValueMin="7.6" pageId="3" pageNumber="276" unit="mm" value="83.0" valueMax="90.0" valueMin="76.0">76–90 mm</quantity>
|
||
by 3.0–
|
||
<quantity id="A8609B40813A6F687BCA2D3AE36C2340" box="[705,790,1546,1573]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="276" unit="mm" value="3.5">3.5 mm</quantity>
|
||
at segment X,
|
||
<quantity id="A8609B40813A6F687AB52D3AE43F2340" box="[958,1093,1546,1573]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.5" metricValueMin="3.4" pageId="3" pageNumber="276" unit="mm" value="3.45" valueMax="3.5" valueMin="3.4">3.4–3.5 mm</quantity>
|
||
at segment XXX,
|
||
<quantity id="A8609B40813A6F687C1D2D3AE5E72340" box="[1302,1437,1546,1573]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.5" metricValueMin="3.4" pageId="3" pageNumber="276" unit="mm" value="3.45" valueMax="3.5" valueMin="3.4">3.4–3.5 mm</quantity>
|
||
at clitellum; body cylindrical, segments 67–83. Setae distributed around segmental equators, numbering 30–34 at VII, 42–45 at XX, 12–16 between the male pores; setae of II–VIII enlarged, setal formula AA:AB:ZZ:YZ = 2:2:6:3 at XIII. Female pore single at XIV at
|
||
<quantity id="A8609B40813A6F687B4C2D47E2B523F4" box="[583,719,1655,1681]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="3" pageNumber="276" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.3–0.4 mm</quantity>
|
||
on oval. Prostomium epilobic with tongue open, purple-brown anterior dorsal pigmentation, lighter ventrally and posteriorly, clitellum coffee color, formalin preservation. First dorsal pores at 5/6. Clitellum annular XIV–XVI; setae invisible externally.
|
||
</paragraph>
|
||
<paragraph id="6F2736A5813A6F6879CC2DD3E19F220C" blockId="3.[151,1437,1258,2041]" pageId="3" pageNumber="276">
|
||
Male pores superficial in XVIII at 8
|
||
<superScript id="98ED9BED813A6F687B6E2DD2E2082395" attach="left" box="[613,626,1762,1776]" fontSize="6" pageId="3" pageNumber="276">th</superScript>
|
||
setal line, 0.17–0.29 circumference apart ventrally, at centers of round porophores surrounded by 3–4 thick white oval outer rings, at lateral margins of ventrum. Four pairs of spermathecal pores deep in 5/6–8/9 at 9
|
||
<superScript id="98ED9BED813A6F6878B32C1AE1BF225D" attach="left" box="[440,453,1834,1848]" fontSize="6" pageId="3" pageNumber="276">th</superScript>
|
||
setal line, 0.24–0.36 circumference apart ventrally at lateral margin of ventrum. Genital markings and papillae absent.
|
||
</paragraph>
|
||
<paragraph id="6F2736A5813A6F6F79CC2C43E36325B0" blockId="3.[151,1437,1258,2041]" lastBlockId="4.[151,1436,151,213]" lastPageId="4" lastPageNumber="277" pageId="3" pageNumber="276">Septa 5/6, 6/7 thick, 7/8 thin, 8/9/10 absent, 10/11–13/14 thin. Gizzard globular in VIII–X. Intestine beginning in XV, and lymph glands missing. Typhlosole vestigial from XXVII. Intestinal caeca originating in XXVII and extending anteriorly to about XXV, slightly asymmetric pinnate-shaped with 6–7 dorsal, 6–7 ventral lobes. Hearts esophageal in X–XIII; X small. Male sexual system holandric testes and funnels in ventral paired sacs in X and XI. Seminal vesicle two pairs in XI and XII, no dorsal lobes. Prostates in XVIII, 6 main lobes, extending between XVII and XXIII; stout short muscular duct very wide in center.</paragraph>
|
||
<caption id="3BE7662D813D6F6F799C2E1CE5592021" ID-DOI="http://doi.org/10.5281/zenodo.10603403" ID-Zenodo-Dep="10603403" box="[151,1315,1324,1349]" httpUri="https://zenodo.org/record/10603403/files/figure.png" pageId="4" pageNumber="277" startId="4.[151,250,1324,1348]" targetBox="[494,1047,250,1279]" targetPageId="4" targetType="figure">
|
||
<paragraph id="6F2736A5813D6F6F799C2E1CE5592021" blockId="4.[151,1315,1324,1349]" box="[151,1315,1324,1349]" pageId="4" pageNumber="277">
|
||
<emphasis id="5DECEAB7813D6F6F799C2E1CE1692021" bold="true" box="[151,275,1324,1348]" pageId="4" pageNumber="277">FIGURE 3.</emphasis>
|
||
<taxonomicName id="A8984D26813D6F6F78122E1CE1D42021" authority="Hong & Inkhavilay & James, 2024" authorityName="Hong & Inkhavilay & James" authorityYear="2024" box="[281,430,1324,1348]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="4" pageNumber="277" phylum="Annelida" rank="species" species="xayi" status="sp. nov.">
|
||
<emphasis id="5DECEAB7813D6F6F78122E1CE1D42021" box="[281,430,1324,1348]" italics="true" pageId="4" pageNumber="277">Amynthas xayi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5DECEAB7813D6F6F78BF2E1DE27F2020" bold="true" box="[436,517,1325,1349]" pageId="4" pageNumber="277">
|
||
<taxonomicNameLabel id="46DF57CC813D6F6F78BF2E1DE27F2020" box="[436,517,1325,1349]" pageId="4" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A, B). A: ventral view; B: intestinal caeca; C: spermatheca. Scale bars 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6F2736A5813D6F6F79CC2EA7E2E0209C" blockId="4.[151,1437,1431,1889]" pageId="4" pageNumber="277">Ovaries in XIII. Four pairs of spermathecae in VI–IX; ampulla very large lanceolate pouches, irregularly folded or flattened, slender duct shorter than ampulla; diverticulum chamber egg-shaped, iridescent on short stalk attached to duct near body wall; no nephridia on ducts.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2782652E813D6F6F79CC2D33E4BF2204" pageId="4" pageNumber="277" type="discussion">
|
||
<paragraph id="6F2736A5813D6F6F79CC2D33E4BF2204" blockId="4.[151,1437,1431,1889]" pageId="4" pageNumber="277">
|
||
<emphasis id="5DECEAB7813D6F6F79CC2D33E1412378" bold="true" box="[199,315,1539,1565]" pageId="4" pageNumber="277">Remarks.</emphasis>
|
||
The distinctions from the main body of the
|
||
<taxonomicName id="A8984D26813D6F6F7A242D33E3D92378" baseAuthorityName="Kinberg" baseAuthorityYear="1866" box="[815,931,1539,1565]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="4" pageNumber="277" phylum="Annelida" rank="species" species="corticis">
|
||
<emphasis id="5DECEAB7813D6F6F7A242D33E33A2378" box="[815,832,1539,1565]" italics="true" pageId="4" pageNumber="277">A</emphasis>
|
||
.
|
||
<emphasis id="5DECEAB7813D6F6F7A442D34E3D92378" box="[847,931,1540,1565]" italics="true" pageId="4" pageNumber="277">corticis</emphasis>
|
||
</taxonomicName>
|
||
-group are the pinnate caeca and the forward position of the dorsal pores. The Burmese
|
||
<emphasis id="5DECEAB7813D6F6F7B832D17E2822324" box="[648,760,1575,1601]" italics="true" pageId="4" pageNumber="277">andersoni</emphasis>
|
||
-group of 9 octothecate species in
|
||
<bibRefCitation id="0B094B54813D6F6F7D802D17E55A2324" author="Gates, G. E." box="[1163,1312,1575,1601]" pageId="4" pageNumber="277" pagination="1 - 326" refId="ref4125" refString="Gates, G. E. (1972) Burmese earthworms, an introduction to the systematics and biology of megadrile oligochaetes with special reference to Southeast Asia. Transaction of the American Philosophical Society, New Series, 62, 1 - 326. https: // doi. org / 10.2307 / 1006214" type="journal article" year="1972">Gates (1972)</bibRefCitation>
|
||
is entirely composed of species with simple caeca and first dorsal pores in 12/13 or nearby and has a range including northeastern
|
||
<collectingCountry id="178F7635813D6F6F799C2D5FE09C23EC" box="[151,230,1647,1673]" name="Myanmar" pageId="4" pageNumber="277">Burma</collectingCountry>
|
||
, northern
|
||
<collectingCountry id="178F7635813D6F6F78522D5FE1C523EC" box="[345,447,1647,1673]" name="Thailand" pageId="4" pageNumber="277">Thailand</collectingCountry>
|
||
, and
|
||
<collectingCountry id="178F7635813D6F6F78F02D5FE23823EC" box="[507,578,1647,1673]" name="China" pageId="4" pageNumber="277">China</collectingCountry>
|
||
. Thus, it is likely that similar species are present in northern
|
||
<collectingCountry id="178F7635813D6F6F7DFE2D5FE55423EC" box="[1269,1326,1647,1673]" name="Laos" pageId="4" pageNumber="277">Laos</collectingCountry>
|
||
, but none have been found as yet.
|
||
<taxonomicName id="A8984D26813D6F6F78BF2DA3E22123C8" authority="Hong & Inkhavilay & James, 2024" authorityName="Hong & Inkhavilay & James" authorityYear="2024" box="[436,603,1683,1709]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="4" pageNumber="277" phylum="Annelida" rank="species" species="xayi" status="sp. nov.">
|
||
<emphasis id="5DECEAB7813D6F6F78BF2DA3E22123C8" box="[436,603,1683,1709]" italics="true" pageId="4" pageNumber="277">Amynthas xayi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5DECEAB7813D6F6F7B6E2DA3E2BB23C8" bold="true" box="[613,705,1683,1709]" pageId="4" pageNumber="277">
|
||
<taxonomicNameLabel id="46DF57CC813D6F6F7B6E2DA3E2BB23C8" box="[613,705,1683,1709]" pageId="4" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="A8984D26813D6F6F7A542DA3E42F23C8" authority="Hong & Inkhavilay & James, 2024" authorityName="Hong & Inkhavilay & James" authorityYear="2024" box="[863,1109,1683,1709]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="4" pageNumber="277" phylum="Annelida" rank="species" species="piyakarnae" status="sp. nov.">
|
||
<emphasis id="5DECEAB7813D6F6F7A542DA3E42F23C8" box="[863,1109,1683,1709]" italics="true" pageId="4" pageNumber="277">Amynthas piyakarnae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5DECEAB7813D6F6F7D542DA3E4C623C8" bold="true" box="[1119,1212,1683,1709]" pageId="4" pageNumber="277">
|
||
<taxonomicNameLabel id="46DF57CC813D6F6F7D542DA3E4C623C8" box="[1119,1212,1683,1709]" pageId="4" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
with respect to the pinnate intestinal caeca, but it can be separated easily by the following characters: the new species has a lack of genital markings and papillae, but
|
||
<taxonomicName id="A8984D26813D6F6F7B172DEBE36A2390" authority="Hong & Inkhavilay & James, 2024" authorityName="Hong & Inkhavilay & James" authorityYear="2024" box="[540,784,1755,1781]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="4" pageNumber="277" phylum="Annelida" rank="species" species="piyakarnae" status="sp. nov.">
|
||
<emphasis id="5DECEAB7813D6F6F7B172DEBE36A2390" box="[540,784,1755,1781]" italics="true" pageId="4" pageNumber="277">Amynthas piyakarnae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5DECEAB7813D6F6F7A1C2DEBE30B2390" bold="true" box="[791,881,1755,1781]" pageId="4" pageNumber="277">
|
||
<taxonomicNameLabel id="46DF57CC813D6F6F7A1C2DEBE30B2390" box="[791,881,1755,1781]" pageId="4" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has genital markings on circular, paired pre-setal VIII and IX and genital papillae on XVIII and XIX with very short stalks or sessile.
|
||
<taxonomicName id="A8984D26813D6F6F7D6F2DCFE570227C" authority="Hong & Inkhavilay & James, 2024" authorityName="Hong & Inkhavilay & James" authorityYear="2024" box="[1124,1290,1791,1817]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="4" pageNumber="277" phylum="Annelida" rank="species" species="xayi" status="sp. nov.">
|
||
<emphasis id="5DECEAB7813D6F6F7D6F2DCFE570227C" box="[1124,1290,1791,1817]" italics="true" pageId="4" pageNumber="277">Amynthas xayi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5DECEAB7813D6F6F7C182DCFE514227C" bold="true" box="[1299,1390,1791,1817]" pageId="4" pageNumber="277">
|
||
<taxonomicNameLabel id="46DF57CC813D6F6F7C182DCFE514227C" box="[1299,1390,1791,1817]" pageId="4" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has ampulla very large lanceolate pouches, but
|
||
<taxonomicName id="A8984D26813D6F6F7B8E2C13E3032258" authority="Hong & Inkhavilay & James, 2024" authorityName="Hong & Inkhavilay & James" authorityYear="2024" box="[645,889,1827,1853]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="4" pageNumber="277" phylum="Annelida" rank="species" species="piyakarnae" status="sp. nov.">
|
||
<emphasis id="5DECEAB7813D6F6F7B8E2C13E3032258" box="[645,889,1827,1853]" italics="true" pageId="4" pageNumber="277">Amynthas piyakarnae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5DECEAB7813D6F6F7A892C13E3A62258" bold="true" box="[898,988,1827,1853]" pageId="4" pageNumber="277">
|
||
<taxonomicNameLabel id="46DF57CC813D6F6F7A892C13E3A62258" box="[898,988,1827,1853]" pageId="4" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has ampulla oval, small black spots on surface. This new species also has fewer setae in post-clitellar segments, and smaller body size.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |