159 lines
23 KiB
XML
159 lines
23 KiB
XML
<document id="A54F88911165210DE6CF324071EBFA8A" ID-DOI="10.11646/palaeoentomology.6.3.11" ID-ISSN="2624-2834" ID-Zenodo-Dep="8209319" ID-ZooBank="339DF6C2-7D16-4BAC-9428-B741F557717C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1691005761504" checkinUser="plazi" docAuthor="BARTEL, CHRISTIAN & DUNLOP, JASON A." docDate="2023" docId="E7240C06FF806863FF72937BFA106B8F" docLanguage="en" docName="palaeoentomology.6.3.11.pdf" docOrigin="Palaeoentomology 6 (3)" docSource="http://dx.doi.org/10.11646/palaeoentomology.6.3.11" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Phalangiidae Latreille 1802" docType="treatment" docVersion="2" lastPageNumber="283" masterDocId="1B1D747EFF856865FFFA9442FF816D4C" masterDocTitle="First eupnoid harvestmen (Arachnida: Opiliones: Eupnoi) from mid-Cretaceous Kachin amber, with notes on sexual dimorphism in Halitherses grimaldii (Arachnida: Opiliones: Dyspnoi)" masterLastPageNumber="291" masterPageNumber="278" pageNumber="283" updateTime="1691006252701" updateUser="ExternalLinkService">
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<mods:title id="3AB42DAB62D4C353FC9D1AB9469B3E81">First eupnoid harvestmen (Arachnida: Opiliones: Eupnoi) from mid-Cretaceous Kachin amber, with notes on sexual dimorphism in Halitherses grimaldii (Arachnida: Opiliones: Dyspnoi)</mods:title>
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<mods:namePart id="B3FC23132A1370409ED76EA2D9E23056">BARTEL, CHRISTIAN</mods:namePart>
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<mods:namePart id="2C81B40A7B19F9773DFD688BB54147B7">DUNLOP, JASON A.</mods:namePart>
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<mods:title id="64854696717EC1157497776945B70F01">Palaeoentomology</mods:title>
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<mods:part id="F5938264972B14EA2EAAEAFA7A9A4D9F">
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<mods:date id="3147A880EF75EDBFB8708B022FE04181">2023</mods:date>
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<mods:number id="316D4B1DD0F4B3DD4A41DF3BE0566186">2023-06-28</mods:number>
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<mods:url id="1AC180D7E4D1D4F0267A630DC908E98E">http://dx.doi.org/10.11646/palaeoentomology.6.3.11</mods:url>
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<mods:classification id="4FA98B9A3049101798CDA4E2AFE563CC">journal article</mods:classification>
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<mods:identifier id="CC6E6599D08DF5398E773AD307EA4786" type="DOI">10.11646/palaeoentomology.6.3.11</mods:identifier>
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<treatment id="E7240C06FF806863FF72937BFA106B8F" LSID="urn:lsid:plazi:treatment:E7240C06FF806863FF72937BFA106B8F" httpUri="http://treatment.plazi.org/id/E7240C06FF806863FF72937BFA106B8F" lastPageId="6" lastPageNumber="283" pageId="5" pageNumber="283">
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<subSubSection id="2797EE9BFF806860FF72937BFE016A1F" box="[136,384,1849,1876]" pageId="5" pageNumber="283" type="nomenclature">
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<paragraph id="6F32BD10FF806860FF72937BFE016A1F" blockId="5.[136,384,1849,1912]" box="[136,384,1849,1876]" pageId="5" pageNumber="283">
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<heading id="347A0A7CFF806860FF72937BFE016A1F" bold="true" box="[136,384,1849,1876]" fontSize="10" level="2" pageId="5" pageNumber="283" reason="3">
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<emphasis id="5DF96102FF806860FF72937BFE016A1F" bold="true" box="[136,384,1849,1876]" pageId="5" pageNumber="283">
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<taxonomicName id="A88DC693FF806860FF72937BFEA56A1F" ID-CoL="6268T" ID-ENA="101160" authorityName="Latreille" authorityYear="1802" box="[136,292,1849,1875]" class="Arachnida" family="Phalangiidae" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="283" phylum="Arthropoda" rank="family">Phalangiidae</taxonomicName>
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<emphasis id="5DF96102FF806860FED1937BFEE46A18" bold="true" box="[299,357,1849,1876]" italics="true" pageId="5" pageNumber="283">indet</emphasis>
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. 1
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="2797EE9BFF806860FF72931CFF5D6A34" box="[136,220,1886,1912]" pageId="5" pageNumber="283" type="description">
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<paragraph id="6F32BD10FF806860FF72931CFF5D6A34" blockId="5.[136,384,1849,1912]" box="[136,220,1886,1912]" pageId="5" pageNumber="283">
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(
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<figureCitation id="F7B6A195FF806860FF6A931CFF556A34" box="[144,212,1886,1912]" captionStart="FIGURE 4" captionStartId="6.[136,235,1068,1092]" captionTargetBox="[154,1435,189,1045]" captionTargetId="figure-390@6.[151,1435,189,1045]" captionTargetPageId="6" captionText="FIGURE 4. BUB4516 Phalangiidae indet. 1. A, Dorsal overview (camera lucida drawing). B, Close-up of the body in dorsal view. C, Close-up of the body in ventral view. Abbreviations: ch, chelicerae; cx, coxa; fe, femur; mt, metatarsus; oc, ocularium; pa, patella; pp, pedipalp; ta, tarsus; ti, tibia; legs numbered from I–IV. Scale bars = 1.0 mm (A) and 0.5 mm (B, C)." figureDoi="http://doi.org/10.5281/zenodo.8209327" httpUri="https://zenodo.org/record/8209327/files/figure.png" pageId="5" pageNumber="283">Fig. 4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="2797EE9BFF806860FF7293E4FD326544" pageId="5" pageNumber="283" type="materials_examined">
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<paragraph id="6F32BD10FF806860FF7293E4FDE16A8D" blockId="5.[136,778,1958,2092]" box="[136,608,1958,1985]" pageId="5" pageNumber="283">
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<emphasis id="5DF96102FF806860FF7293E4FF766A8C" bold="true" box="[136,247,1958,1984]" pageId="5" pageNumber="283">Material.</emphasis>
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coll. Patrick Müller, BUB4516.
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</paragraph>
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<paragraph id="6F32BD10FF806860FF479388FD326544" blockId="5.[136,778,1958,2092]" pageId="5" pageNumber="283">
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<emphasis id="5DF96102FF806860FF479388FE496AA8" bold="true" box="[189,456,1994,2020]" pageId="5" pageNumber="283">Locality and horizon.</emphasis>
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Burmese amber, Hukawng Valley,
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<collectingRegion id="AD4973F2FF806860FF1B93ACFEF56544" box="[225,372,2030,2056]" country="Myanmar" name="Kachin" pageId="5" pageNumber="283">Kachin State</collectingRegion>
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,
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<collectingCountry id="179AFD80FF806860FE8593ADFE716545" box="[383,496,2031,2057]" name="Myanmar" pageId="5" pageNumber="283">Myanmar</collectingCountry>
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; mid-Cretaceous.
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</paragraph>
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</subSubSection>
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<subSubSection id="2797EE9BFF806860FF479C50FAE46AD0" pageId="5" pageNumber="283" type="description">
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<paragraph id="6F32BD10FF806860FF479C50FAE46AD0" blockId="5.[136,778,1958,2092]" lastBlockId="5.[810,1452,1305,2093]" pageId="5" pageNumber="283">
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<emphasis id="5DF96102FF806860FF479C50FECE6560" bold="true" box="[189,335,2066,2092]" pageId="5" pageNumber="283">Description.</emphasis>
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Body relatively small, oval and somewhat granulated, L 1.12, anterior
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<geoCoordinate id="0AB9DBD7FF806860FB209158FAAE6878" box="[1242,1327,1305,1332]" degrees="0.58" direction="west" orientation="longitude" pageId="5" pageNumber="283" precision="555" value="-0.58">W 0.58</geoCoordinate>
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, maximum posterior
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<geoCoordinate id="0AB9DBD7FF806860FC6E917CFC6B681B" box="[916,1002,1341,1368]" degrees="0.66" direction="west" orientation="longitude" pageId="5" pageNumber="283" precision="555" value="-0.66">W 0.66</geoCoordinate>
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. Ocularium very small, slightly elevated and oval with two lateral eye lenses, L 0.15,
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<geoCoordinate id="0AB9DBD7FF806860FAE59120FAF76830" box="[1311,1398,1378,1404]" degrees="0.10" direction="west" orientation="longitude" pageId="5" pageNumber="283" precision="555" value="-0.1">W 0.10</geoCoordinate>
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. Eye lens diameter 0.04. Chelicerae rather large. Cheliceral distal segment with a few small spines. Cheliceral basal segment L 0.19; distal segment L?, with fixed and movable finger. Pedipalps sparsely covered with setae. Pedipalp tarsus elongated bearing a single claw at its tip. Pedipalp length: fe 0.20, pa 0.14, ti 0.15, ta 0.31, total (fe–ta) 0.80. Legs extremely long (leg II and IV longest) and covered with a few setae. Tarsus I–IV bear a single smooth claw. Tarsal formula: 9–10:32:12–14:14–15. Leg length: Leg I cx 0.22, tr 0.10, fe 0.54, pa 0.17, ti 0.61, mt 0.51, ta 1.10, total (cx–ta) 3.25; Leg II cx 0.54, tr 0.12, fe 1.44, pa 0.24, ti 1.56, mt 0.88, ta 3.24, total (cx–ta) 8.02; Leg III cx 0.40, tr 0.10, fe 0.63, pa 0.12, ti 0.63, mt 0.56, ta 1.46, total (cx–ta) 3.90; Leg IV cx 0.45, tr 0.10, fe 0.83, pa 0.24, ti 0.76, mt 0.80, ta 2.05, total (cx–ta) 5.23.
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</paragraph>
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</subSubSection>
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<subSubSection id="2797EE9BFF806863FCA593E4FA106B8F" lastPageId="6" lastPageNumber="284" pageId="5" pageNumber="283" type="discussion">
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<paragraph id="6F32BD10FF806863FCA593E4FA106B8F" blockId="5.[810,1452,1305,2093]" lastBlockId="6.[810,1452,1236,1732]" lastPageId="6" lastPageNumber="284" pageId="5" pageNumber="283">
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<emphasis id="5DF96102FF806860FCA593E4FC526A8C" bold="true" box="[863,979,1958,1984]" pageId="5" pageNumber="283">Remarks.</emphasis>
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BUB4516 (
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<figureCitation id="F7B6A195FF806860FB8D93E4FB7F6A8C" box="[1143,1278,1958,1984]" captionStart="FIGURE 4" captionStartId="6.[136,235,1068,1092]" captionTargetBox="[154,1435,189,1045]" captionTargetId="figure-390@6.[151,1435,189,1045]" captionTargetPageId="6" captionText="FIGURE 4. BUB4516 Phalangiidae indet. 1. A, Dorsal overview (camera lucida drawing). B, Close-up of the body in dorsal view. C, Close-up of the body in ventral view. Abbreviations: ch, chelicerae; cx, coxa; fe, femur; mt, metatarsus; oc, ocularium; pa, patella; pp, pedipalp; ta, tarsus; ti, tibia; legs numbered from I–IV. Scale bars = 1.0 mm (A) and 0.5 mm (B, C)." figureDoi="http://doi.org/10.5281/zenodo.8209327" httpUri="https://zenodo.org/record/8209327/files/figure.png" pageId="5" pageNumber="283">Fig. 4A–C</figureCitation>
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) could be a juvenile/subadult, due to its relatively small body size and the straight pedipalp tarsus. The number of tarsomeres on leg II, which potentially exceeds 30, suggests a later developmental stage. The presence of a pedipalp claw and single claws on all legs again support referral to
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<taxonomicName id="A88DC693FF836863FD4890BAFC86685E" authorityName="Hansen & Sorensen" authorityYear="1904" box="[690,775,1272,1298]" class="Arachnida" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="287" phylum="Arthropoda" rank="subOrder" subOrder="Eupnoi">Eupnoi</taxonomicName>
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. Despite the uncertainty of the developmental stage most of the modern
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<taxonomicName id="A88DC693FF836863FED39102FEFB6816" authorityName="Hansen & Sorensen" authorityYear="1904" box="[297,378,1344,1370]" class="Arachnida" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="287" phylum="Arthropoda" rank="subOrder" subOrder="Eupnoi">Eupnoi</taxonomicName>
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families can be excluded. Members of the family
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<taxonomicName id="A88DC693FF836863FED89126FE0D6832" authorityName="Banks" authorityYear="1893" box="[290,396,1380,1406]" class="Arachnida" family="Caddidae" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="284" phylum="Arthropoda" rank="family">Caddidae</taxonomicName>
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are typically characterized by the presence of a huge, well developed ocularium with large lateral eyes, whereas the fossil has only a small, slightly raised ocularium (
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<figureCitation id="F7B6A195FF836863FEA39193FE5968A7" box="[345,472,1489,1515]" captionStart="FIGURE 4" captionStartId="6.[136,235,1068,1092]" captionTargetBox="[154,1435,189,1045]" captionTargetId="figure-390@6.[151,1435,189,1045]" captionTargetPageId="6" captionText="FIGURE 4. BUB4516 Phalangiidae indet. 1. A, Dorsal overview (camera lucida drawing). B, Close-up of the body in dorsal view. C, Close-up of the body in ventral view. Abbreviations: ch, chelicerae; cx, coxa; fe, femur; mt, metatarsus; oc, ocularium; pa, patella; pp, pedipalp; ta, tarsus; ti, tibia; legs numbered from I–IV. Scale bars = 1.0 mm (A) and 0.5 mm (B, C)." figureDoi="http://doi.org/10.5281/zenodo.8209327" httpUri="https://zenodo.org/record/8209327/files/figure.png" pageId="6" pageNumber="284">Fig. 4A, B</figureCitation>
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). A smooth pedipalp claw further tends to rule out the family
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<taxonomicName id="A88DC693FF836863FDB091B7FC876B43" authorityName="Simon" authorityYear="1879" box="[586,774,1525,1551]" class="Arachnida" family="Sclerosomatidae" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="284" phylum="Arthropoda" rank="family">Sclerosomatidae</taxonomicName>
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, which usually have a pectinate pedipalp claw (
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<bibRefCitation id="0B1CC0E1FF836863FD5F925BFF456B1B" author="Martens, J." pageId="6" pageNumber="284" refId="ref10151" refString="Martens, J. (1978) Spinnentiere, Arachnida: Weberknechte, Opiliones. Die Tierwelt Deutschlands. Vol. 64. G. Fischer Verlag, Jena, 464 pp." type="book" year="1978">Martens, 1978</bibRefCitation>
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). The absence of a
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<emphasis id="5DF96102FF836863FE4B927CFDE06B1B" box="[433,609,1598,1623]" italics="true" pageId="6" pageNumber="284">scutum parvum</emphasis>
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and a heavily sclerotized dorsal body, which are also typical characters for sclerosomatids, additionally supports their exclusion. The Australasian family
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<taxonomicName id="A88DC693FF836863FE3992EBFDE36B8F" authorityName="Forster" authorityYear="1948" box="[451,610,1705,1731]" class="Arachnida" family="Monoscutidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="284" phylum="Arthropoda" rank="family">Monoscutidae</taxonomicName>
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and its two subfamilies Megalopsalidinae and
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<taxonomicName id="A88DC693FF836863FDF1928FFD2B6BAB" box="[523,682,1741,1767]" class="Arachnida" family="Neopilionidae" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="284" phylum="Arthropoda" rank="subFamily" subFamily="Monoscutinae">Monoscutinae</taxonomicName>
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can also be excluded as the fossil lacks the enlarged chelicerae or the shortened legs (femur I is often shorter than the body width) respectively (
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<figureCitation id="F7B6A195FF836863FE2A9378FDA06A18" box="[464,545,1850,1876]" captionStart="FIGURE 4" captionStartId="6.[136,235,1068,1092]" captionTargetBox="[154,1435,189,1045]" captionTargetId="figure-390@6.[151,1435,189,1045]" captionTargetPageId="6" captionText="FIGURE 4. BUB4516 Phalangiidae indet. 1. A, Dorsal overview (camera lucida drawing). B, Close-up of the body in dorsal view. C, Close-up of the body in ventral view. Abbreviations: ch, chelicerae; cx, coxa; fe, femur; mt, metatarsus; oc, ocularium; pa, patella; pp, pedipalp; ta, tarsus; ti, tibia; legs numbered from I–IV. Scale bars = 1.0 mm (A) and 0.5 mm (B, C)." figureDoi="http://doi.org/10.5281/zenodo.8209327" httpUri="https://zenodo.org/record/8209327/files/figure.png" pageId="6" pageNumber="284">Fig. 4</figureCitation>
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). Additionally, the fossil shows neither an elongate pedipalp patella, nor the rather short and thick legs and reduced pedipalp claw often found in the family
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<taxonomicName id="A88DC693FF836863FE4A93E4FDD36A8C" authorityName="Lawrence" authorityYear="1931" box="[432,594,1958,1984]" class="Arachnida" family="Neopilionidae" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="284" phylum="Arthropoda" rank="family">Neopilionidae</taxonomicName>
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. This leaves the families
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<taxonomicName id="A88DC693FF836863FF149388FE0A6AA8" authorityName="Banks" authorityYear="1893" box="[238,395,1994,2020]" class="Arachnida" family="Protolophidae" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="284" phylum="Arthropoda" rank="family">Protolophidae</taxonomicName>
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and
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<taxonomicName id="A88DC693FF836863FE309388FDDF6AA8" authorityName="Latreille" authorityYear="1802" box="[458,606,1994,2020]" class="Arachnida" family="Phalangiidae" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="284" phylum="Arthropoda" rank="family">Phalangiidae</taxonomicName>
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.
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<taxonomicName id="A88DC693FF836863FD979388FC8B6AA8" authorityName="Banks" authorityYear="1893" box="[621,778,1994,2020]" class="Arachnida" family="Protolophidae" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="284" phylum="Arthropoda" rank="family">Protolophidae</taxonomicName>
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are characterized by a pedipalp tarsus which is shorter than the pedipalp tibia (
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<figureCitation id="F7B6A195FF836863FE759C50FE656560" box="[399,484,2066,2092]" captionStart="FIGURE 4" captionStartId="6.[136,235,1068,1092]" captionTargetBox="[154,1435,189,1045]" captionTargetId="figure-390@6.[151,1435,189,1045]" captionTargetPageId="6" captionText="FIGURE 4. BUB4516 Phalangiidae indet. 1. A, Dorsal overview (camera lucida drawing). B, Close-up of the body in dorsal view. C, Close-up of the body in ventral view. Abbreviations: ch, chelicerae; cx, coxa; fe, femur; mt, metatarsus; oc, ocularium; pa, patella; pp, pedipalp; ta, tarsus; ti, tibia; legs numbered from I–IV. Scale bars = 1.0 mm (A) and 0.5 mm (B, C)." figureDoi="http://doi.org/10.5281/zenodo.8209327" httpUri="https://zenodo.org/record/8209327/files/figure.png" pageId="6" pageNumber="284">Fig. 4C</figureCitation>
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) combined with relatively short legs. Females also have slender pedipalps with an apophysis on the pedipalp patella or tibia. Males on the other hand have greatly thickened pedipalps. All of these characters are absent in the fossil. As most of the important characters are consistent with members of the family
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<taxonomicName id="A88DC693FF836863FC5191CBFBBE68EF" authorityName="Latreille" authorityYear="1802" box="[939,1087,1417,1443]" class="Arachnida" family="Phalangiidae" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="284" phylum="Arthropoda" rank="family">Phalangiidae</taxonomicName>
|
||
, the fossil is tentatively placed therein. The lack of significant morphological characters makes a placement at genus level very difficult. Its closest living relatives can probably be found in the genera
|
||
<taxonomicName id="A88DC693FF836863FCD0925BFBF26B78" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[810,1139,1561,1588]" class="Arachnida" family="Phalangiidae" genus="Phalangium" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="284" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5DF96102FF836863FCD0925BFC326B7F" box="[810,947,1561,1587]" italics="true" pageId="6" pageNumber="284">Phalangium</emphasis>
|
||
<bibRefCitation id="0B1CC0E1FF836863FC44925BFBF26B78" author="Linnaeus, C." box="[958,1139,1561,1588]" pageId="6" pageNumber="284" refId="ref10053" refString="Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tom. I. Editio decima, reformata. L. Salvii, Holmiae [= Stockholm], 824 pp. https: // doi. org / 10.5962 / bhl. title. 542" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A88DC693FF836863FB78925BFCE56B1B" authority="Roewer, 1911" authorityName="Roewer" authorityYear="1911" class="Arachnida" family="Phalangiidae" genus="Metaphalangium" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="284" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5DF96102FF836863FB78925BFAC06B7F" box="[1154,1345,1561,1587]" italics="true" pageId="6" pageNumber="284">Metaphalangium</emphasis>
|
||
<bibRefCitation id="0B1CC0E1FF836863FAB79258FCE56B1B" author="Roewer, C. F." pageId="6" pageNumber="284" pagination="1 - 106" refId="ref10765" refString="Roewer, C. F. (1911) Ubersicht der Genera der Subfamilie der Phalangiini der Opiliones Palpatores nebst Beschreibung einigerneuerGattungenundArten. ArchivfurNaturgeschichte, Berlin, Abt. A, Original-Arbeiten, 77 (Supplementheft 2), 1 - 106." type="journal article" year="1911">Roewer, 1911</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A88DC693FF836863FC8B927FFBD56B1B" authority="Herbst, 1798" authorityName="Herbst" authorityYear="1798" box="[881,1108,1597,1623]" class="Arachnida" family="Phalangiidae" genus="Opilio" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="284" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5DF96102FF836863FC8B927FFC386B1B" box="[881,953,1597,1623]" italics="true" pageId="6" pageNumber="284">Opilio</emphasis>
|
||
<bibRefCitation id="0B1CC0E1FF836863FC38927FFBD56B1B" author="Herbst, J. F. W." box="[962,1108,1597,1623]" pageId="6" pageNumber="284" refId="ref9663" refString="Herbst, J. F. W. (1798) Naturgeschichte der Insecten-Gattung Opilio. In: Herbst, J. F. W., Natursystem der ungeflugelten Insekten, vol. 2 [of 4]. Gottlieb August Lange, Berlin, iv + 26 pp., 5 pls." type="book" year="1798">Herbst, 1798</bibRefCitation>
|
||
</taxonomicName>
|
||
or in one of the Asian genera described by
|
||
<emphasis id="5DF96102FF836863FC309220FC576B37" box="[970,982,1634,1659]" italics="true" pageId="6" pageNumber="284">e</emphasis>
|
||
.
|
||
<emphasis id="5DF96102FF836863FC249220FC6D6B37" box="[990,1004,1634,1659]" italics="true" pageId="6" pageNumber="284">g</emphasis>
|
||
.,
|
||
<bibRefCitation id="0B1CC0E1FF836863FBFD9220FB3C6B37" author="Martens, J." box="[1031,1213,1633,1660]" pageId="6" pageNumber="284" pagination="181 - 217" refId="ref10121" refString="Martens, J. (1973) Opiliones aus dem Nepal-Himalaya. II. Phalangiidae und Sclerosomatidae (Arachnida). Senckenbergiana biologica, 54, 181 - 217." type="journal article" year="1973">Martens (1973)</bibRefCitation>
|
||
. In lieu of an adult specimen, the fossil is placed for now as
|
||
<taxonomicName id="A88DC693FF836863FAE192C7FA2D6BD3" authorityName="Latreille" authorityYear="1802" box="[1307,1452,1669,1695]" class="Arachnida" family="Phalangiidae" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="284" phylum="Arthropoda" rank="family">Phalangiidae</taxonomicName>
|
||
<emphasis id="5DF96102FF836863FCD092EBFCE36B8F" box="[810,866,1705,1731]" italics="true" pageId="6" pageNumber="284">indet</emphasis>
|
||
. but described and figured here for completeness.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |