treatments-xml/data/E5/1B/78/E51B780E9C8C5A10A00E73204F39FAD2.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1198.116265" ID-Pensoft-Pub="1313-2970-1198-295" ID-Pensoft-UUID="0E3939F66FB455AEA5F94377938D22EE" ID-ZooBank="1566523605DF4C4FA6E641A404EE6545" ModsDocID="1313-2970-1198-295" checkinTime="1714220719304" checkinUser="pensoft" docAuthor="Nasir, Nurul Syafiqah, Lee, Jie Ying, Marzuki, Mohammad Effendi, Vermeulen, Jaap J., Mohd-Azlan, Jayasilan &amp; Khalik, Mohd Zacaery" docDate="2024" docId="E51B780E9C8C5A10A00E73204F39FAD2" docLanguage="en" docName="ZooKeys 1198: 295-340" docOrigin="ZooKeys 1198" docPubDate="2024-04-26" docSource="http://dx.doi.org/10.3897/zookeys.1198.116265" docTitle="Diplommatina rumbangensis Nasir &amp; Lee &amp; Marzuki &amp; Vermeulen &amp; Mohd-Azlan &amp; Khalik 2024, sp. nov." docType="treatment" docUuid="C34336B3-6421-49FD-84AC-154E8A1C4691" docUuidSource="ZooBank" docVersion="1" id="0E3939F66FB455AEA5F94377938D22EE" lastPageNumber="295" masterDocId="0E3939F66FB455AEA5F94377938D22EE" masterDocTitle="Checklist of land snail species of Gua Rumbang, Sarawak, Malaysian Borneo (Mollusca, Gastropoda), with a description of a new species, Diplommatina rumbangensis sp. nov." masterLastPageNumber="340" masterPageNumber="295" pageNumber="295" updateTime="1714220719304" updateUser="pensoft">
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<mods:title id="6F6931609EE0E354EE3EE72030726AC5">Checklist of land snail species of Gua Rumbang, Sarawak, Malaysian Borneo (Mollusca, Gastropoda), with a description of a new species, Diplommatina rumbangensis sp. nov.</mods:title>
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<mods:namePart id="4DB608B2651BBE8E01763FBED957A2D6">Nasir, Nurul Syafiqah</mods:namePart>
<mods:affiliation id="E44BA10BB393FC814B4ABD361E636C51">Faculty of Resource Science and Technology, Universiti Malaysia Sarawak, 94300, Kota Samarahan, Sarawak, Malaysia</mods:affiliation>
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<mods:namePart id="5BAE79648B845D41DD55CF5340C2D8BA">Lee, Jie Ying</mods:namePart>
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<mods:affiliation id="5ECAF2B99A1513E7F75E15808137EAF7">Faculty of Resource Science and Technology, Universiti Malaysia Sarawak, 94300, Kota Samarahan, Sarawak, Malaysia</mods:affiliation>
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<mods:namePart id="299E056B3F1D0876279B03A4ADD385DD">Marzuki, Mohammad Effendi</mods:namePart>
<mods:affiliation id="08757E806C4219ED5D5804E64D79E565">Faculty of Resource Science and Technology, Universiti Malaysia Sarawak, 94300, Kota Samarahan, Sarawak, Malaysia</mods:affiliation>
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<mods:namePart id="EE18FA9D65F323FD5D699BC1BE22F0C5">Vermeulen, Jaap J.</mods:namePart>
<mods:affiliation id="02D6EC4A70FF5E41687D27C00D3ABA2E">Institute of Biodiversity and Environmental Conservation, Universiti Malaysia Sarawak, 94300, Kota Samarahan, Sarawak, Malaysia</mods:affiliation>
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<mods:namePart id="1B37BD1F5847CEBFF9322FA4C1D85C9B">Mohd-Azlan, Jayasilan</mods:namePart>
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<mods:affiliation id="C2C85DF8474C1739B2F85A7C5C68FEFB">Faculty of Resource Science and Technology, Universiti Malaysia Sarawak, 94300, Kota Samarahan, Sarawak, Malaysia</mods:affiliation>
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<mods:namePart id="EDF3BCC947A39B8AAC601F14878E2C50">Khalik, Mohd Zacaery</mods:namePart>
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<mods:date id="1166E0E5129BC19E8398F863F32ED87E">2024</mods:date>
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<paragraph id="BA69AB71FB1B295076F2951CF3C1273A" pageId="0" pageNumber="295">
<taxonomicName id="4EE691C0AD84770397172D0111D4C44D" LSID="https://zoobank.org/C34336B3-6421-49FD-84AC-154E8A1C4691" authority="Nasir &amp; Lee &amp; Marzuki &amp; Vermeulen &amp; Mohd-Azlan &amp; Khalik, 2024" authorityName="Nasir &amp; Lee &amp; Marzuki &amp; Vermeulen &amp; Mohd-Azlan &amp; Khalik" authorityYear="2024" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina rumbangensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="rumbangensis" status="sp. nov.">Diplommatina rumbangensis</taxonomicName>
<taxonomicNameLabel id="5BE898D672BFA55EB1777C5E2E4DAB54" pageId="0" pageNumber="295">sp. nov.</taxonomicNameLabel>
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<paragraph id="6C51B1AB6AF116EF3AF1CC5FDD572203" pageId="0" pageNumber="295">
<figureCitation id="208F582123D569DB30C6D57544D5D720" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Scanning Electron Microscope images of Diplommatina rumbangensis sp. nov. A-F Paratypes (ME 14471). A apertural view B side view C apical view D enlargement of apical view showing the apex with radial ribs E umbilical view F top whorls view showing the tubular projection and semi-circular scars." figureDoi="10.3897/zookeys.1198.116265.figure2" httpUri="https://binary.pensoft.net/fig/1034241" pageId="0" pageNumber="295">Figs 2A-F</figureCitation>
<figureCitation id="D2A1FF0EE60172652618755AEAC0B92F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A Diplommatina adversa (H. Adams &amp; A. Adams, 1851) ME 14476 B Diplommatina baritensis E. A. Smith, 1893 ME 14472 C Diplommatina concinna H. Adams, 1872 ME 14473 D Diplommatina maduana maduana Laidlaw, 1949 ME 14474 E Diplommatina rumbangensis sp. nov. holotype MZU. MOL. 22.132 F Diplommatina subglaber subisensis (Vermeulen, 1993) ME 14475." figureDoi="10.3897/zookeys.1198.116265.figure8" httpUri="https://binary.pensoft.net/fig/1034247" pageId="0" pageNumber="295">, 8E</figureCitation>
</paragraph>
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<subSubSection id="977BA37DDD91258E56EDA4099A1682B9" pageId="0" pageNumber="295" type="type material">
<paragraph id="68613EF2C1FEDBE8FCB5B3DE7182E400" pageId="0" pageNumber="295">Type material examined.</paragraph>
<paragraph id="841A1370AF14CE02653438ACB7B05B44" pageId="0" pageNumber="295">
<emphasis id="C71A8B3FEB5E992325668250AD449CD1" bold="true" pageId="0" pageNumber="295">
<emphasis id="E834F83BCD3FFCB65DC33822DF8BD4A8" italics="true" pageId="0" pageNumber="295">Holotype</emphasis>
.
</emphasis>
Malaysia • (SH = 5.25 mm, SW = 2.52 mm); Sarawak, Gua Rumbang, near Kampung Semadang, along Sarawak Kanan River, ~ 11 miles Southwest Padawan, Kuching Division; 
<geoCoordinate id="4FFE11E2DE1468C785DD42C6A89FA905" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="AC9CFC28C276E121B6F242F9DD80A5E0" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022; N.S. Nasir and M. E. Marzuki leg.; MZU.MOL.22.132. 
<emphasis id="5E7E6B06B8AB7EC0AC87FDE8FE26E405" bold="true" pageId="0" pageNumber="295">
<emphasis id="C22660547F5DE5DC79A0E57C99BD1F13" italics="true" pageId="0" pageNumber="295">Paratypes</emphasis>
.
</emphasis>
Malaysia • 4 ex. (SH = 5.1 mm, SW = 2.7 mm; SH = 5.06 mm, SW = 2.5 mm; SH = 5.37 mm, SW = 2.58 mm; SH = 5.01 mm, SW = 2.65 mm); same data as holotype; MZU.MOL.22.491, ME 14983, ME 15021. Both holotype and paratypes were deposited in Zoological Museum UNIMAS (ZMU) and additional paratypes in the private collection of the third author.
</paragraph>
<caption id="53AA87C23BF025D36166F9A9BB6DFCE9" doi="10.3897/zookeys.1198.116265.figure2" httpUri="https://binary.pensoft.net/fig/1034241" pageId="0" pageNumber="295" start="Figure 2" startId="F2">
<paragraph id="8E2936E827C958FA0440F0DD0E3772DE" pageId="0" pageNumber="295">
<emphasis id="DC675ADAE20269ACE4C79B18EAA64417" bold="true" pageId="0" pageNumber="295">Figure 2.</emphasis>
Scanning Electron Microscope images of 
<taxonomicName id="C0DA2585F8FC10C17F8C95D7CBD743A9" authorityName="Nasir &amp; Lee &amp; Marzuki &amp; Vermeulen &amp; Mohd-Azlan &amp; Khalik" authorityYear="2024" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina rumbangensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="rumbangensis">
<emphasis id="94CBBD07FB5DB8100D865085E73D19B2" italics="true" pageId="0" pageNumber="295">Diplommatina rumbangensis</emphasis>
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sp. nov. 
<emphasis id="8C4F32FFC5EE01E08D8C4BDAAD24BDAE" bold="true" pageId="0" pageNumber="295">A-F</emphasis>
Paratypes (ME 14471). 
<emphasis id="5E9A9C5988E8D0A3F7BE4A5F18791CFD" bold="true" pageId="0" pageNumber="295">A</emphasis>
apertural view 
<emphasis id="4D676FBEDA0265E1D9920DA068AC6D55" bold="true" pageId="0" pageNumber="295">B</emphasis>
side view 
<emphasis id="E8E5AD37048E6A60C3147BFD12507474" bold="true" pageId="0" pageNumber="295">C</emphasis>
apical view 
<emphasis id="E36691AB610772BD81A1ADB05B007723" bold="true" pageId="0" pageNumber="295">D</emphasis>
enlargement of apical view showing the apex with radial ribs 
<emphasis id="98FB2B1837602A5F50DF22B227959082" bold="true" pageId="0" pageNumber="295">E</emphasis>
umbilical view 
<emphasis id="45B0CE9D020C38423DBEBDDB54D42323" bold="true" pageId="0" pageNumber="295">F</emphasis>
top whorls view showing the tubular projection and semi-circular scars.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="9AFB64AE2B85EDA28CBA1DC055EE0554" pageId="0" pageNumber="295" type="description">
<paragraph id="1058EA251D44E31913F7BA0825612217" pageId="0" pageNumber="295">Description.</paragraph>
<paragraph id="1E3AF7029C7004CC40FA8242CE2428A6" pageId="0" pageNumber="295">
Shell dextral, fusiform to moderately conical, reddish orange, shining and translucent, with the penultimate whorl widest, convex, well rounded. Suture impressed. Protoconch with 
<normalizedToken id="9573FE8A37706338B9B5037E6F547796" originalValue="1½">11/2</normalizedToken>
whorls, punctate with small pits, without radial and spiral lines. Constriction nearly level with the edge between the parietal and columellar side of the peristome, with two parietales, two upper longitudinal palatales which are not covered by the peristome on the outer surface of the shell, one transversal palatalis and one columellaris. The columellaris positioned at the start of the constriction together with the longitudinal palatales. Tuba approximately 
<normalizedToken id="84747EC1E255D465205E8F10E177DE11" originalValue="¾">3/4</normalizedToken>
whorl. 
<emphasis id="8DAD9606FDA632ACF10C997BF42E8C92" bold="true" italics="true" pageId="0" pageNumber="295">Sculpture</emphasis>
: Radial ribs on the top whorls only, widely spaced, inconspicuous, but halfway the whorl with an almost tubular projection, in adults sometimes abraded to a semi-circular scar. Spiral striation inconspicuous, on top whorls only. 
<emphasis id="70302C18F21E811D04D26BAFE7B6104D" bold="true" italics="true" pageId="0" pageNumber="295">Aperture</emphasis>
: Hardly tilted regarding the coiling axis; columellaris distinct, directed downwards. Peristome double, expanding; palatal side hardly sinuous, without edge; basal side with an edge; basal edge hardly sinuous, rounded; inner peristome somewhat expanding beyond the outer, with a palatal lip, free and erect on the columellar side, expanding on the parietal side. 
<emphasis id="4208E28D5B5CE633374431E3F5D964C4" bold="true" italics="true" pageId="0" pageNumber="295">Umbilicus</emphasis>
: Open, narrow. 
<emphasis id="CEE1DB9CFB263E584B4F663FFB276017" bold="true" italics="true" pageId="0" pageNumber="295">Dimensions</emphasis>
: Height 5.01-5.37 mm; width 2.5-2.7 mm; number of whorls 
<normalizedToken id="D9CFA9F7520ABFB48804B56918613534" originalValue="6¾">63/4</normalizedToken>
-7; height and width aperture 1.91-2.05 mm; 2.05-2.09 mm.
</paragraph>
</subSubSection>
<subSubSection id="0D49B18A12501E8D844BE1A775FC452F" pageId="0" pageNumber="295" type="differential diagnosis">
<paragraph id="8DA21A9D45D811188BFC6F5CE6D740B1" pageId="0" pageNumber="295">Differential diagnosis.</paragraph>
<paragraph id="4C73601F1A3BFE71E32E112582702427" pageId="0" pageNumber="295">
<taxonomicName id="AB81AC7626F1213F506FAC0AF41C65F7" authorityName="Nasir &amp; Lee &amp; Marzuki &amp; Vermeulen &amp; Mohd-Azlan &amp; Khalik" authorityYear="2024" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina rumbangensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="rumbangensis">
<emphasis id="9D0F7642850063C540650A356B192759" italics="true" pageId="0" pageNumber="295">Diplommatina rumbangensis</emphasis>
</taxonomicName>
sp. nov. has two distinct upper longitudinal palatales at the shell constriction that are not covered by the peristome on the outer surface of the shell (Fig. 
<figureCitation id="DAC330F5A2117135EC32742805053B65" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Sketch of Diplommatina spinosa and Diplommatina bicoronata bicoronata extracted from Vermeulen (1993), and Diplommatina rumbangensis sp. nov. showing the position of the longitudinal palatalis (red), the transversal palatalis (blue), and the constriction area (grey)." figureDoi="10.3897/zookeys.1198.116265.figure3" httpUri="https://binary.pensoft.net/fig/1034242" pageId="0" pageNumber="295">3</figureCitation>
). This differs from 
<taxonomicName id="D85B51724C143DFDBFAE5D43600260D6" authorityName="Godwin-Austen" authorityYear="1889" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina spinosa" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="spinosa">
<emphasis id="8AF64D5034BA6DD6D9AF05867E79EF39" italics="true" pageId="0" pageNumber="295">Diplommatina spinosa</emphasis>
</taxonomicName>
Godwin-Austen, 1889 which has one longitudinal palatalis only. Additionally, 
<taxonomicName id="DB47F5294E094CFEF00A702552BC3D10" lsidName="D. spinosa" pageId="0" pageNumber="295" rank="species" species="spinosa">
<emphasis id="09CADBFD5CE37F105209054892645B62" italics="true" pageId="0" pageNumber="295">D. spinosa</emphasis>
</taxonomicName>
exhibits a tubular projection that extends to the penultimate whorl, whereas in 
<taxonomicName id="56B91350EF699A3226C6D102ACFED58F" lsidName="D. rumbangensis" pageId="0" pageNumber="295" rank="species" species="rumbangensis">
<emphasis id="D0A7C4AD1A230D17FABF7FB4319F6A7E" italics="true" pageId="0" pageNumber="295">D. rumbangensis</emphasis>
</taxonomicName>
sp. nov., this projection only reaches halfway through the ultimate whorl. 
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<emphasis id="CA67C739796950ABACA589E8ACB1268C" italics="true" pageId="0" pageNumber="295">Diplommatina spinosa</emphasis>
</taxonomicName>
is distributed in the Kuching and Serian divisions. It also differs from 
<taxonomicName id="C87E491FD907CD123C76DDCB3B5614F6" authorityName="E. von Martens" authorityYear="1884" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina bicoronata subsp. bicoronata" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="subSpecies" species="bicoronata" subSpecies="bicoronata">
<emphasis id="38C07F893792C75C4D69E63FA24378CE" italics="true" pageId="0" pageNumber="295">Diplommatina bicoronata bicoronata</emphasis>
</taxonomicName>
von Martens, 1884 which is endemic to Kalimantan, Indonesia by having two or three longitudinal palatales with two of them covered by the peristome. 
<taxonomicName id="D50C4C067AD0BC0528B550ADCFC60BA7" authorityName="E. von Martens" authorityYear="1884" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina bicoronata subsp. bicoronata" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="subSpecies" species="bicoronata" subSpecies="bicoronata">
<emphasis id="E942E5269CEFFD9D4906AB73B1C088DB" italics="true" pageId="0" pageNumber="295">Diplommatina bicoronata bicoronata</emphasis>
</taxonomicName>
also has radial ribs near the suture of the whorls which is absent in 
<taxonomicName id="318C9FE21D0FA3952FCC7A1AEDD7BA5C" lsidName="D. rumbangensis" pageId="0" pageNumber="295" rank="species" species="rumbangensis">
<emphasis id="546299719AF45BB55DAE7BA613B98D49" italics="true" pageId="0" pageNumber="295">D. rumbangensis</emphasis>
</taxonomicName>
sp. nov. This latter species can be distinguished from 
<taxonomicName id="315C4273D16BCDDD962A3F5B98FCA4A6" authorityName="Vermeulen" authorityYear="1993" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina aurisdiaboli" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="aurisdiaboli">
<emphasis id="E18082BE38C3553DB4B994A9115869E4" italics="true" pageId="0" pageNumber="295">Diplommatina aurisdiaboli</emphasis>
</taxonomicName>
Vermeulen, 1993 by the absence of palatal lip expanding up the suture of the previous whorl.
</paragraph>
<caption id="EC6D39D6365704150FCF430012141124" doi="10.3897/zookeys.1198.116265.figure3" httpUri="https://binary.pensoft.net/fig/1034242" pageId="0" pageNumber="295" start="Figure 3" startId="F3">
<paragraph id="8D3E3F1C1BC1657BBA5D32B422ECEA15" pageId="0" pageNumber="295">
<emphasis id="E6E95E08E731741C7F28FE65D24F422C" bold="true" pageId="0" pageNumber="295">Figure 3.</emphasis>
Sketch of 
<taxonomicName id="8F56BEC25442FF0971766BAAD8B1E2DC" authorityName="Godwin-Austen" authorityYear="1889" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina spinosa" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="spinosa">
<emphasis id="DD16074134A87A64191DB6DD946A565B" italics="true" pageId="0" pageNumber="295">Diplommatina spinosa</emphasis>
</taxonomicName>
and 
<taxonomicName id="878E25D299A484F63CF8DB58C0ACA976" authorityName="E. von Martens" authorityYear="1884" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina bicoronata subsp. bicoronata" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="subSpecies" species="bicoronata" subSpecies="bicoronata">
<emphasis id="8C1729820F5B150C562B0B880D785218" italics="true" pageId="0" pageNumber="295">Diplommatina bicoronata bicoronata</emphasis>
</taxonomicName>
extracted from 
<bibRefCitation id="9BE360394D86C67076BF537523885828" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" pagination="3 - 69" publicationUrl="http://natuurtijdschriften.nl/record/597005" refId="B52" refString="Vermeulen, JJ, 1993. Notes on the non-marine mollusks of the island of Borneo 5. The genus Diplommatina (Gastropoda Prosobranchia: Diplommatinidae). Basteria 57: 3 - 69, http://natuurtijdschriften.nl/record/597005" title="Notes on the non-marine mollusks of the island of Borneo 5. The genus Diplommatina (Gastropoda Prosobranchia: Diplommatinidae)." url="http://natuurtijdschriften.nl/record/597005" volume="57" year="1993">Vermeulen (1993)</bibRefCitation>
, and 
<taxonomicName id="26C21221CFCDE47EA6C64474E21638A7" authorityName="Nasir &amp; Lee &amp; Marzuki &amp; Vermeulen &amp; Mohd-Azlan &amp; Khalik" authorityYear="2024" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina rumbangensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="rumbangensis">
<emphasis id="3B4057A0924BCF4F96D1A0D64C6F1A60" italics="true" pageId="0" pageNumber="295">Diplommatina rumbangensis</emphasis>
</taxonomicName>
sp. nov. showing the position of the longitudinal palatalis (red), the transversal palatalis (blue), and the constriction area (grey).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="0970E63B9F0EC45C700CBDF5A74803AC" pageId="0" pageNumber="295" type="etymology">
<paragraph id="766F0EF2BA6CF40F1ACB036017A2EACC" pageId="0" pageNumber="295">Etymology.</paragraph>
<paragraph id="CF19E9E934B14F58EB08F091DEE9D3CE" pageId="0" pageNumber="295">
The epithet 
<emphasis id="0FE30C0B70FF82A7B2BF707541DA8413" italics="true" pageId="0" pageNumber="295">rumbangensis</emphasis>
refers to the type locality Gua Rumbang.
</paragraph>
</subSubSection>
<subSubSection id="7066FDD5661E8E5644842E32367FBFDC" pageId="0" pageNumber="295" type="distribution">
<paragraph id="3600E69A540FCC03F400E6683BDC94D1" pageId="0" pageNumber="295">Geographic distribution and habitat.</paragraph>
<paragraph id="5DB1F28FD8CB70331940AC52EFA1DF81" pageId="0" pageNumber="295">
<taxonomicName id="058B5C59C05FEC6845ADECBF1D14358B" authorityName="Nasir &amp; Lee &amp; Marzuki &amp; Vermeulen &amp; Mohd-Azlan &amp; Khalik" authorityYear="2024" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina rumbangensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="rumbangensis">
<emphasis id="800FCD401A329FB1FE44346C3D86E870" italics="true" pageId="0" pageNumber="295">Diplommatina rumbangensis</emphasis>
</taxonomicName>
sp. nov. is known from the type locality only. The living animals were not observed.
</paragraph>
</subSubSection>
<subSubSection id="9145BFF4D761B62892C2B3404CD0538E" pageId="0" pageNumber="295" type="remarks">
<paragraph id="A14071E7E27243BD59C6832CF392BB84" pageId="0" pageNumber="295">Remarks.</paragraph>
<paragraph id="057424059FA12F0570E1C7CBD1EED6E3" pageId="0" pageNumber="295">
The spine or tubular projection of the shells of 
<taxonomicName id="AE4A14E0227723C2FAC8D78F2A7F250E" authorityName="Nasir &amp; Lee &amp; Marzuki &amp; Vermeulen &amp; Mohd-Azlan &amp; Khalik" authorityYear="2024" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina rumbangensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="rumbangensis">
<emphasis id="4E2A94393C71AC0D326511A6063A0E29" italics="true" pageId="0" pageNumber="295">Diplommatina rumbangensis</emphasis>
</taxonomicName>
sp. nov. are mostly broken, leaving semi-circular scars.
</paragraph>
</subSubSection>
<subSubSection id="4B561733395BA6C1C47448A26B82EDF9" pageId="0" pageNumber="295" type="checklist">
<paragraph id="498EB7B3B0D94EAEAEC27E8357DB21DB" pageId="0" pageNumber="295">Checklist</paragraph>
<paragraph id="97F97978194A6F8792E112359DCE01D2" pageId="0" pageNumber="295">
<emphasis id="75A7214AB0EBDCF4CD2845E93A13CDAE" bold="true" pageId="0" pageNumber="295">
Class 
<taxonomicName id="6E4CAE0580A1108C5B285B3823191107" authorityName="Benson" authorityYear="1849" class="Gastropoda" kingdom="Animalia" lsidName="" pageId="0" pageNumber="295" rank="class">Gastropoda</taxonomicName>
Cuvier, 1795
</emphasis>
,
</paragraph>
<paragraph id="2127C7F5D06ABDD12CA18808B743684A" pageId="0" pageNumber="295">
<emphasis id="787BD6CDCBDF7B8E2F3EFFE1E19654EA" bold="true" pageId="0" pageNumber="295">
Subclass 
<taxonomicName id="7B0921424E7F5E17213865EFE5AFEFA1" authorityName="Benson" authorityYear="1849" class="Gastropoda" kingdom="Animalia" lsidName="" pageId="0" pageNumber="295" rank="subClass" subClass="Caenogastropoda">Caenogastropoda</taxonomicName>
Cox, 1960
</emphasis>
</paragraph>
<paragraph id="5308B9B48007A7EB11DF73AD83476476" pageId="0" pageNumber="295">
<emphasis id="219823E3801C35BFDEFEDB82D2E7FC02" bold="true" pageId="0" pageNumber="295">
Family 
<taxonomicName id="77B366AD4CF7D42E8C33A52EEED37D74" authorityName="W. T. Blanford" authorityYear="1864" class="Gastropoda" family="Alycaeidae" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="family">Alycaeidae</taxonomicName>
W. T. Blanford, 1864
</emphasis>
</paragraph>
<paragraph id="54E4D4DC59B54F2CBE57B35854A2F11C" pageId="0" pageNumber="295">
<emphasis id="15DE24E64A9E257C4AD266C896B616BE" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="8D6519AF9EF5D5B5B52327ACF3E1C1CA" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1889" class="Gastropoda" family="Alycaeidae" genus="Chamalycaeus" kingdom="Animalia" lsidName="Chamalycaeus specus" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="specus">
<emphasis id="F98C9CD743CA112D4E044905728D9823" italics="true" pageId="0" pageNumber="295">Chamalycaeus specus</emphasis>
</taxonomicName>
(Godwin-Austen, 1889)
</emphasis>
</paragraph>
<paragraph id="2ED47FC44D783775D38151EAE7FC26D3" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="F3E2F3D799DC8566A58CF453457C9617" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A Chamalycaeus specus (Godwin-Austen, 1889) ME 14470 B Pincerna globosa (H. Adams, 1871) ME 14468 C Stomacosmethis hosei (Godwin-Austen, 1889) ME 14469 D Solenomphala scalaris (Heude, 1882) ME 14512." figureDoi="10.3897/zookeys.1198.116265.figure4" httpUri="https://binary.pensoft.net/fig/1034243" pageId="0" pageNumber="295">4A</figureCitation>
</paragraph>
<caption id="80105529E9388D3F23EF3297F62870C1" doi="10.3897/zookeys.1198.116265.figure4" httpUri="https://binary.pensoft.net/fig/1034243" pageId="0" pageNumber="295" start="Figure 4" startId="F4">
<paragraph id="2431FD21572182C0BB019550FF615903" pageId="0" pageNumber="295">
<emphasis id="25BBD68B113238DECFF2F67FA8EB6ABF" bold="true" pageId="0" pageNumber="295">Figure 4.</emphasis>
<emphasis id="678F2304D6C90F904CA06B3D370D56B9" bold="true" pageId="0" pageNumber="295">A</emphasis>
<taxonomicName id="1C6E2159C5473F8A937B0D00E0DC6C09" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1889" class="Gastropoda" family="Alycaeidae" genus="Chamalycaeus" kingdom="Animalia" lsidName="Chamalycaeus specus" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="specus">
<emphasis id="84F850155F02DBE04DE65E8F2F81C522" italics="true" pageId="0" pageNumber="295">Chamalycaeus specus</emphasis>
</taxonomicName>
(Godwin-Austen, 1889) ME 14470 
<emphasis id="84394562772E84A0FE7FC73B9BA80697" bold="true" pageId="0" pageNumber="295">B</emphasis>
<taxonomicName id="452D9430A1F84D6CFB4C1D95F2781419" baseAuthorityName="H. Adams" baseAuthorityYear="1871" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" lsidName="Pincerna globosa" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="globosa">
<emphasis id="9F28AD1730014D53233674D137E2369C" italics="true" pageId="0" pageNumber="295">Pincerna globosa</emphasis>
</taxonomicName>
(H. Adams, 1871) ME 14468 
<emphasis id="74CFEE45EDBAEB1485B673F7B4B7E39F" bold="true" pageId="0" pageNumber="295">C</emphasis>
<taxonomicName id="051E05A2BAC24E2FCD8016ED08524CF1" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1889" class="Gastropoda" family="Alycaeidae" genus="Stomacosmethis" kingdom="Animalia" lsidName="Stomacosmethis hosei" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="hosei">
<emphasis id="BDE93951CCBCC6918EDF2DFCDE7D4192" italics="true" pageId="0" pageNumber="295">Stomacosmethis hosei</emphasis>
</taxonomicName>
(Godwin-Austen, 1889) ME 14469 
<emphasis id="D175791503EE64BBD4CB805F6F04AC80" bold="true" pageId="0" pageNumber="295">D</emphasis>
<taxonomicName id="5822D6BC9130EAE7209996EAD34BA8F6" baseAuthorityName="Heude" baseAuthorityYear="1882" class="Gastropoda" family="Assimineidae" genus="Solenomphala" kingdom="Animalia" lsidName="Solenomphala scalaris" order="Littorinimorpha" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="scalaris">
<emphasis id="DA24C02C94E96BA9FDC0486EC4C41120" italics="true" pageId="0" pageNumber="295">Solenomphala scalaris</emphasis>
</taxonomicName>
(Heude, 1882) ME 14512.
</paragraph>
</caption>
<paragraph id="CD6DC948930589658E8FD52881E93B8A" pageId="0" pageNumber="295">
<emphasis id="85B984A65427A22DAD506662494A08D3" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;In limestone caves at Jambusan, Borneo&quot;.
</paragraph>
<paragraph id="79C3A56D6668F66FA1F2D2759E615621" pageId="0" pageNumber="295">
<emphasis id="0561AA3FD6B1109332D79C1017C23A2B" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="E5084CC5A194616A79064E5FD9512647" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="7149FB73BCED8EEFEE9D32DEA3C872FA" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Apr. 2016-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14470, ME 14982, ME 15020, MZU.MOL.16.116.
</paragraph>
<paragraph id="7795B957A94B5E25DC3B933711BF1EE0" pageId="0" pageNumber="295">
<emphasis id="3F356411B86CC78B90BF1E1BEE012AC9" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widespread in Borneo. Endemic to Borneo (
<bibRefCitation id="8FDDE1D6BAB007C416C6F8E36EE0AA75" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
; 
<bibRefCitation id="578367BF711C9413AB64BA76DA8146F5" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
).
</paragraph>
<paragraph id="43DF3DF786C9BF61E7FF753AEC8DFD20" pageId="0" pageNumber="295">
<emphasis id="9293E1118B680D3D498E2193C131077F" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="AA39B2B2FAEC7CE31EC6913E22857377" pageId="0" pageNumber="295">
<emphasis id="6C2C9C2FDD3A94D0A1B6964D38FCFF51" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="672A112F4B814EA9494D63E047032D01" baseAuthorityName="H. Adams" baseAuthorityYear="1871" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" lsidName="Pincerna globosa" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="globosa">
<emphasis id="824FF8012E33D38F48AADEBA58C03EE5" italics="true" pageId="0" pageNumber="295">Pincerna globosa</emphasis>
</taxonomicName>
(H. Adams, 1871)
</emphasis>
</paragraph>
<paragraph id="06998465C5F70A716AB907C10BAA1AE3" pageId="0" pageNumber="295">
Figs 
<figureCitation id="2563F35711DB44699A27CEFE78DD889C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A Chamalycaeus specus (Godwin-Austen, 1889) ME 14470 B Pincerna globosa (H. Adams, 1871) ME 14468 C Stomacosmethis hosei (Godwin-Austen, 1889) ME 14469 D Solenomphala scalaris (Heude, 1882) ME 14512." figureDoi="10.3897/zookeys.1198.116265.figure4" httpUri="https://binary.pensoft.net/fig/1034243" pageId="0" pageNumber="295">4B</figureCitation>
, 
<figureCitation id="2501A7A88197CDFB76204C19A500C12C" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Living snails from Gua Rumbang. A Pincerna globosa (H. Adams, 1871) MZU. MOL. 22.135 B Stomacosmethis hosei (Godwin-Austen, 1889) MZU. MOL. 22.136 C Craspedotropis borneensis (Godwin-Austen, 1889) MZU. MOL. 23.141 D Japonia borneensis (E. A. Smith, 1893) MZU. MOL. 23.138. Images not to scale." figureDoi="10.3897/zookeys.1198.116265.figure19" httpUri="https://binary.pensoft.net/fig/1034258" pageId="0" pageNumber="295">19A</figureCitation>
</paragraph>
<paragraph id="0D898B5D0044F012B4B5D4162C892B82" pageId="0" pageNumber="295">
<emphasis id="17D50BAE37D78BCAB3294258FCFCE640" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Busan, near Sarawak, Borneo&quot;.
</paragraph>
<paragraph id="A56C06FA18583ABB8ECDEA5DCFD0C77E" pageId="0" pageNumber="295">
<emphasis id="16B93E1FEA69753999B32FFC31801B32" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="030B0688D46AD0061C8F4985B0EDDB2E" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="2952C70181196532F865CE3B14A0AF16" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Apr. 2016-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14468, ME 15018, MZU.MOL.16.109, MZU.MOL.22.135, MZU.MOL.23.139.
</paragraph>
<paragraph id="A4EBCF20D4666E53507BF7C987DF88AA" pageId="0" pageNumber="295">
<emphasis id="C6E89B79924F74BCEAC27E67AEF77B04" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak to Niah, further to the north. Also found in Sabah and West Kalimantan. Endemic to Borneo (
<bibRefCitation id="C3BA8CD8DB36BED2CBDBECB1780A80B8" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
; 
<bibRefCitation id="236A847F75891A48123C760FEBE6A06F" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
).
</paragraph>
<paragraph id="F5F9EEDD3B814FAD58BE6EE121126165" pageId="0" pageNumber="295">
<emphasis id="2592787F6A481090F7A1210402CFCE13" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were observed foraging on leaf surface of trees at the base of limestone cliffs.
</paragraph>
<paragraph id="395872809F3D9E2408C423EF24008106" pageId="0" pageNumber="295">
<emphasis id="A9C42612275195CD3499900447B04E73" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="6AED5F6664B7EC9C076C2B16958E4B62" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1889" class="Gastropoda" family="Alycaeidae" genus="Stomacosmethis" kingdom="Animalia" lsidName="Stomacosmethis hosei" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="hosei">
<emphasis id="5AC2E90409C8CD52B49EDB3221FE1D5E" italics="true" pageId="0" pageNumber="295">Stomacosmethis hosei</emphasis>
</taxonomicName>
(Godwin-Austen, 1889)
</emphasis>
</paragraph>
<paragraph id="561451D7CB185E86576A355528D5FE3E" pageId="0" pageNumber="295">
Figs 
<figureCitation id="BCE9E59B27B00DB4E3D29701988334D3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A Chamalycaeus specus (Godwin-Austen, 1889) ME 14470 B Pincerna globosa (H. Adams, 1871) ME 14468 C Stomacosmethis hosei (Godwin-Austen, 1889) ME 14469 D Solenomphala scalaris (Heude, 1882) ME 14512." figureDoi="10.3897/zookeys.1198.116265.figure4" httpUri="https://binary.pensoft.net/fig/1034243" pageId="0" pageNumber="295">4C</figureCitation>
, 
<figureCitation id="6A3A589D2DD2815E1313CC8456C90600" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Living snails from Gua Rumbang. A Pincerna globosa (H. Adams, 1871) MZU. MOL. 22.135 B Stomacosmethis hosei (Godwin-Austen, 1889) MZU. MOL. 22.136 C Craspedotropis borneensis (Godwin-Austen, 1889) MZU. MOL. 23.141 D Japonia borneensis (E. A. Smith, 1893) MZU. MOL. 23.138. Images not to scale." figureDoi="10.3897/zookeys.1198.116265.figure19" httpUri="https://binary.pensoft.net/fig/1034258" pageId="0" pageNumber="295">19B</figureCitation>
</paragraph>
<paragraph id="D5DBE5C34C5B1F39DEEACC3014BBFC06" pageId="0" pageNumber="295">
<emphasis id="C63744D8B2090F4D441E5C2B8DA9127D" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Busan Hills, Borneo&quot;.
</paragraph>
<paragraph id="C98F27FDA7AB064CAA7B4827E269EBC0" pageId="0" pageNumber="295">
<emphasis id="C163F8A282ABD24650F4942F5649DFDF" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="7598958134B46370DFEE3651BD2C8A20" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="63E371B470AB24DDEC7DAB6E79256344" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Apr. 2016-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14469, ME 14981, ME 15019, MZU.MOL.16.108, MZU.MOL.22.136
</paragraph>
<paragraph id="8A4F0EEF94629C57DF70FF226C86DE70" pageId="0" pageNumber="295">
<emphasis id="EFAED7407933DEAF96513B7262C7EA51" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak. Endemic to western Sarawak (
<bibRefCitation id="03C9E2F75847AACFBEA3778F60BD710C" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="8D1C7A0A47152A69D6E684F7C36B284B" pageId="0" pageNumber="295">
<emphasis id="24CF918AAC976FF70ED5D172C491F65F" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails found on wet limestone surfaces covered with mosses. Empty shells were found among leaf litter at the base of limestone hills.
</paragraph>
<paragraph id="047F1D5087A9FE2B35CB20CE4A46CDCA" pageId="0" pageNumber="295">
<emphasis id="807A354BB88D6C459CAB08CD600ED5E0" bold="true" pageId="0" pageNumber="295">
Family 
<taxonomicName id="004EAB1A8A99B9C3B1C6B3F72C0AE80D" authorityName="H. Adams &amp; A. Adams" authorityYear="1856" class="Gastropoda" family="Assimineidae" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="295" phylum="Mollusca" rank="family">Assimineidae</taxonomicName>
H. Adams &amp; A. Adams, 1856
</emphasis>
</paragraph>
<paragraph id="3654DBB1B0F995468E36DDB0680FF1E0" pageId="0" pageNumber="295">
<emphasis id="EFCD09663DA85EE473484BA10F3EDD9E" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="D0B98E784B9A0D1998BB169299BCDE75" baseAuthorityName="Heude" baseAuthorityYear="1882" class="Gastropoda" family="Assimineidae" genus="Solenomphala" kingdom="Animalia" lsidName="Solenomphala scalaris" order="Littorinimorpha" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="scalaris">
<emphasis id="9958056B7DAA086BF2225948A91FE760" italics="true" pageId="0" pageNumber="295">Solenomphala scalaris</emphasis>
</taxonomicName>
(Heude, 1882)
</emphasis>
</paragraph>
<paragraph id="5B9ED09051F3E909AF25680D3986330C" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="A3364C79D1F541E88F4DCE27C275BF84" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A Chamalycaeus specus (Godwin-Austen, 1889) ME 14470 B Pincerna globosa (H. Adams, 1871) ME 14468 C Stomacosmethis hosei (Godwin-Austen, 1889) ME 14469 D Solenomphala scalaris (Heude, 1882) ME 14512." figureDoi="10.3897/zookeys.1198.116265.figure4" httpUri="https://binary.pensoft.net/fig/1034243" pageId="0" pageNumber="295">4D</figureCitation>
</paragraph>
<paragraph id="82BFC2C471F0DEBA1FFFED863EA8BDD5" pageId="0" pageNumber="295">
<emphasis id="94A94D60292D58CD7BE5D04AE96E1DF1" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Ad parietes humidos in civitate Chang-hai sat copiosa&quot; [= Shanghai, China].
</paragraph>
<paragraph id="8122750A62DC7D5903E8929653DB7CF3" pageId="0" pageNumber="295">
<emphasis id="FE111FA47339B4CD2B29A4248CC0D986" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="12D267A2A4A6792ED32D3EABFE3F91F0" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="CBC380E569C0DBA9EB215DCC7FEB7665" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14512.
</paragraph>
<paragraph id="DC8BA9013C04F32C71ABE631A96BB1CF" pageId="0" pageNumber="295">
<emphasis id="72B3E227A2C6D3DF203DEC61AD51A2EA" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed in Borneo. Also found in China and Peninsular Malaysia (
<bibRefCitation id="AF67EA05A65AC970AE255AA118AE0A88" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
; 
<bibRefCitation id="05891A20170B7A074ED37B77A7FBC9C3" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
).
</paragraph>
<paragraph id="02987E32A17DC8C1C7AAB658F3735D4B" pageId="0" pageNumber="295">
<emphasis id="1D128CEB2D414699867AB51C758F0682" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Only empty shells were found. An introduced species. Records show that it occurs in the damp areas around human settlements (
<bibRefCitation id="B81CA1FEF8D5F3C56616E810DD03A1F4" author="Chan, SY" journalOrPublisher="Club Conchylia Informationen" pageId="0" pageNumber="295" pagination="35 - 46" refId="B8" refString="Chan, SY, 1997. Non marine mollusks from Selangor, West Malaysia, with a comparison note on introduced species. Club Conchylia Informationen 29: 35 - 46" title="Non marine mollusks from Selangor, West Malaysia, with a comparison note on introduced species." volume="29" year="1997">Chan 1997</bibRefCitation>
).
</paragraph>
<paragraph id="EDE15F993B7C49721F068ED540487043" pageId="0" pageNumber="295">
<emphasis id="B550FFBBEAB5D00F59A832A4B21BE6D6" bold="true" pageId="0" pageNumber="295">
Family 
<taxonomicName id="231F87ED7E4B1AACF643CCA9878835E4" authorityName="Gray" authorityYear="1847" class="Gastropoda" family="Cyclophoridae" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="family">Cyclophoridae</taxonomicName>
Gray, 1847
</emphasis>
</paragraph>
<paragraph id="333DE97F5A82EE28E5C078FE6BC774BE" pageId="0" pageNumber="295">
<emphasis id="60F5BD46B7EA9237EB59E018B1B20238" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="1E8C7F9F425263851159DDCCDCFD8FF7" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1889" class="Gastropoda" family="Cyclophoridae" genus="Craspedotropis" kingdom="Animalia" lsidName="Craspedotropis borneensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="borneensis">
<emphasis id="6D03343ACE85E11D8451501C0F54659B" italics="true" pageId="0" pageNumber="295">Craspedotropis borneensis</emphasis>
</taxonomicName>
(Godwin-Austen, 1889)
</emphasis>
</paragraph>
<paragraph id="94570B351470187E5F43DFC0250E3B6A" pageId="0" pageNumber="295">
Figs 
<figureCitation id="E5A917D8663D0ADDE7A3EA0AE388AAD9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A Craspedotropis borneensis (Godwin-Austen, 1889) MZU. MOL. 22.137 B Cyclophorus perdix borneensis (Metcalfe, 1852) MZU. MOL. 22.184 C Ditropopsis everetti (E. A Smith, 1895) ME 14467." figureDoi="10.3897/zookeys.1198.116265.figure5" httpUri="https://binary.pensoft.net/fig/1034244" pageId="0" pageNumber="295">5A</figureCitation>
, 
<figureCitation id="0999AEEE3D128051D0FA38C873651B9B" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Living snails from Gua Rumbang. A Pincerna globosa (H. Adams, 1871) MZU. MOL. 22.135 B Stomacosmethis hosei (Godwin-Austen, 1889) MZU. MOL. 22.136 C Craspedotropis borneensis (Godwin-Austen, 1889) MZU. MOL. 23.141 D Japonia borneensis (E. A. Smith, 1893) MZU. MOL. 23.138. Images not to scale." figureDoi="10.3897/zookeys.1198.116265.figure19" httpUri="https://binary.pensoft.net/fig/1034258" pageId="0" pageNumber="295">19C</figureCitation>
</paragraph>
<caption id="AD52017F2AA88FA4C15F0E1BDCAB6A1C" doi="10.3897/zookeys.1198.116265.figure5" httpUri="https://binary.pensoft.net/fig/1034244" pageId="0" pageNumber="295" start="Figure 5" startId="F5">
<paragraph id="9B73C25E97B7CA2E6282A9956C12D0CD" pageId="0" pageNumber="295">
<emphasis id="E2BD040B254B38A52DE8E69D8578CBE1" bold="true" pageId="0" pageNumber="295">Figure 5.</emphasis>
<emphasis id="629130AE8EA0E25867BB97727EA42CC9" bold="true" pageId="0" pageNumber="295">A</emphasis>
<taxonomicName id="ECF002D29B1909548F6277EA4AD39E59" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1889" class="Gastropoda" family="Cyclophoridae" genus="Craspedotropis" kingdom="Animalia" lsidName="Craspedotropis borneensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="borneensis">
<emphasis id="A8EACB726B2C2536E1DDA9FF9F79A26E" italics="true" pageId="0" pageNumber="295">Craspedotropis borneensis</emphasis>
</taxonomicName>
(Godwin-Austen, 1889) MZU.MOL.22.137 
<emphasis id="4F605FD9554F756910C570D7B13C4CE7" bold="true" pageId="0" pageNumber="295">B</emphasis>
<taxonomicName id="04F87070065D11124718E1D548CFC059" baseAuthorityName="Metcalfe" baseAuthorityYear="1852" class="Gastropoda" family="Cyclophoridae" genus="Cyclophorus" kingdom="Animalia" lsidName="Cyclophorus perdix subsp. borneensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="subSpecies" species="perdix" subSpecies="borneensis">
<emphasis id="AC1F06E88F40A0F9244A0528FEBF3C75" italics="true" pageId="0" pageNumber="295">Cyclophorus perdix borneensis</emphasis>
</taxonomicName>
(Metcalfe, 1852) MZU.MOL.22.184 
<emphasis id="40470117EA9AD3DF4C7B8782CCB383D2" bold="true" pageId="0" pageNumber="295">C</emphasis>
<taxonomicName id="F8E846ECFD8F86AB030F5E2619D95C6D" baseAuthorityName="E. A. Smith" baseAuthorityYear="1895" class="Gastropoda" family="Cyclophoridae" genus="Ditropopsis" kingdom="Animalia" lsidName="Ditropopsis everetti" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="everetti">
<emphasis id="B4CA0D476E8F5041D26CF177FEFC228E" italics="true" pageId="0" pageNumber="295">Ditropopsis everetti</emphasis>
</taxonomicName>
(E. A Smith, 1895) ME 14467.
</paragraph>
</caption>
<paragraph id="9BA32FD5699803F246C0345BFA88C804" pageId="0" pageNumber="295">
<emphasis id="CB55999C0291B1859B4D7E2EC521677B" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Busan Hills, Borneo&quot; [= Jambusan Hills, Sarawak].
</paragraph>
<paragraph id="E9D8F7663ACDBE18AB83CAFB4E80AEA2" pageId="0" pageNumber="295">
<emphasis id="F6B5D22009646C96F2C1B5BCFECA7EAA" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="767929E07CF1388B81A192EAA5E1BC98" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="56F71554F0695813DB859CFDDBDF40E9" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14980, ME 15016. MZU.MOL.22.137, MZU.MOL.23.141.
</paragraph>
<paragraph id="604F0BEBE4FF1D0170E06C484BD4298B" pageId="0" pageNumber="295">
<emphasis id="B60DF6854392C8F5DB6EB1BE8598E4B8" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak. Endemic to western Sarawak.
</paragraph>
<paragraph id="1EC20BAF9BA9E7AEF80CF48DFCCE36D8" pageId="0" pageNumber="295">
<emphasis id="F74EB7A3CF9196472AF2D9D70ACB1D22" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were found foraging on limestone surfaces and among leaf-litter and topsoil at the base of limestone cliffs. Living individuals of this species are always covered by dirt, which makes it difficult to be spotted.
</paragraph>
<paragraph id="292FF111B01E86A3F814968C49CA422B" pageId="0" pageNumber="295">
<emphasis id="B64533D0EEA936DDB182C5A5A3B35729" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="B492957947583DB94B629CBD01053DC5" baseAuthorityName="Metcalfe" baseAuthorityYear="1852" class="Gastropoda" family="Cyclophoridae" genus="Cyclophorus" kingdom="Animalia" lsidName="Cyclophorus perdix subsp. borneensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="subSpecies" species="perdix" subSpecies="borneensis">
<emphasis id="DA99BB42A7F4BB3F8811A8CEC74D663F" italics="true" pageId="0" pageNumber="295">Cyclophorus perdix borneensis</emphasis>
</taxonomicName>
(Metcalfe, 1852)
</emphasis>
</paragraph>
<paragraph id="2F9CA5E6808DBC530128A4E092DF2D5D" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="6680D98802E8E81005A65E1D1C29EE2B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A Craspedotropis borneensis (Godwin-Austen, 1889) MZU. MOL. 22.137 B Cyclophorus perdix borneensis (Metcalfe, 1852) MZU. MOL. 22.184 C Ditropopsis everetti (E. A Smith, 1895) ME 14467." figureDoi="10.3897/zookeys.1198.116265.figure5" httpUri="https://binary.pensoft.net/fig/1034244" pageId="0" pageNumber="295">5B</figureCitation>
</paragraph>
<paragraph id="E4D27BF5372B18D20518E569463BA7F8" pageId="0" pageNumber="295">
<emphasis id="FECB34F6617102A787E794358DAECD27" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
<normalizedToken id="3A1C5FF5491B2D1174B721EA5937B4DD" originalValue="“Borneo”">&quot;Borneo&quot;</normalizedToken>
.
</paragraph>
<paragraph id="E5ED32ABB0394BED2313DC0F93709F47" pageId="0" pageNumber="295">
<emphasis id="55DDC9F3C57267E53957E410B9550831" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="DF6CD2FC88C0D9AC22537E200547763B" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="F36680092059C5CF02F932779875CD52" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14461, ME 15010, MZU.MOL.22.184.
</paragraph>
<paragraph id="700A4D96B6586ED31DD2432426AA2D22" pageId="0" pageNumber="295">
<emphasis id="06BCEC6682BF8F4F8347FF48B8002A63" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed in Borneo and Sarawak but rare in Sabah (
<bibRefCitation id="DBCAD5AED16037D85F6065FB85818236" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
; 
<bibRefCitation id="F687C1F5C4C0025283A079486BF3AEDD" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
). Also found in West Malaysia (
<bibRefCitation id="D0AE607F64A2A70C1DBDEE9AE18BA270" author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="295" pagination="261 - 271" refId="B49" refString="Stoliczka, F, 1872. On the land shells of Penang Island, with descriptions of the animals and anatomical notes; Part first, Cyclostomacea. Journal of the Asiatic Society of Bengal 41 (2): 261 - 271" title="On the land shells of Penang Island, with descriptions of the animals and anatomical notes; Part first, Cyclostomacea." volume="41" year="1872">Stoliczka 1872</bibRefCitation>
; 
<bibRefCitation id="999F1CF69B93105E968655361958386B" DOI="https://doi.org/10.5962/bhl.part.14301" author="Morgan, J" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="0" pageNumber="295" pagination="353 - 429" refId="B33" refString="Morgan, J, 1885. Mollusques terrestres &amp; fluviatiles du Royaume de Perak et des pays voisins (presqu'ile Malaise). Bulletin de la Societe Zoologique de France 10: 353 - 429, DOI: https://doi.org/10.5962/bhl.part.14301" title="Mollusques terrestres &amp; fluviatiles du Royaume de Perak et des pays voisins (presqu'ile Malaise)." url="https://doi.org/10.5962/bhl.part.14301" volume="10" year="1885">Morgan 1885</bibRefCitation>
).
</paragraph>
<paragraph id="485601641728E81C630629F9203737BE" pageId="0" pageNumber="295">
<emphasis id="4EFFBE16B0E8E6489F709A37BEE0F1E4" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="7655B34C0B1BE4A9F2AC6DA905F4CBE4" pageId="0" pageNumber="295">
<emphasis id="4234B2DE79FDBF51887BCDB76A240642" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="EB0F8EF1994C98230808AD293642129A" baseAuthorityName="E. A. Smith" baseAuthorityYear="1895" class="Gastropoda" family="Cyclophoridae" genus="Ditropopsis" kingdom="Animalia" lsidName="Ditropopsis everetti" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="everetti">
<emphasis id="440A12272706562DE78D669D3FBCE92F" italics="true" pageId="0" pageNumber="295">Ditropopsis everetti</emphasis>
</taxonomicName>
(E. A. Smith, 1895)
</emphasis>
</paragraph>
<paragraph id="5BD65C0DB7F4C13B68CB19B96DAFE56D" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="85B325066FB3F54336E9BB891C6F936C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A Craspedotropis borneensis (Godwin-Austen, 1889) MZU. MOL. 22.137 B Cyclophorus perdix borneensis (Metcalfe, 1852) MZU. MOL. 22.184 C Ditropopsis everetti (E. A Smith, 1895) ME 14467." figureDoi="10.3897/zookeys.1198.116265.figure5" httpUri="https://binary.pensoft.net/fig/1034244" pageId="0" pageNumber="295">5C</figureCitation>
</paragraph>
<paragraph id="3BA59D06CA605343A7F7EBB05D24926B" pageId="0" pageNumber="295">
<emphasis id="501A333BEF05698ABE77CFECDF3A6DA9" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Rumbang, Sarawak&quot; [= Rumbang Hills, Padawan, Sarawak].
</paragraph>
<paragraph id="C9F889D433A5FFA468A8681918A0E461" pageId="0" pageNumber="295">
<emphasis id="B5F86F4C9F656BB7F4177A197D581D1E" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="B4A832AE384821B0F318610B06D6056A" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="D565040F939B9BC8E93A7E1D20337E7C" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14467, ME 15017.
</paragraph>
<paragraph id="7FD176565FBFCB9998F860E4396191B2" pageId="0" pageNumber="295">
<emphasis id="FBEBDD84C780556029B0064C7AAD9295" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak. Endemic to western Sarawak (
<bibRefCitation id="88284E1583616CEF17E147F4866B2A9B" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="9BB03A97ACAEC8DC74A2B5EF0C8C3ACA" pageId="0" pageNumber="295">
<emphasis id="2C5AD5DBD6B92E87A2B640FB4E6A0225" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="55496A0F7A986B7776992A62D5194DBE" pageId="0" pageNumber="295">
<emphasis id="F3F39EED819B216DF9ACD68FDB6B05CA" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="F377CD3A54251911B13BE1A5C0A27383" class="Gastropoda" family="Cyclophoridae" genus="Japonia" kingdom="Animalia" lsidName="Japonia barbata" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="barbata">
<emphasis id="92B987DF52AE2DC03031C76799737971" italics="true" pageId="0" pageNumber="295">Japonia barbata</emphasis>
</taxonomicName>
(L. Pfeiffer, 1855)
</emphasis>
</paragraph>
<paragraph id="AB1A5381F38C2F03A3FE052B0812536C" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="85F08DD259608873BBB8B790EB54D466" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A Japonia barbata (L. Pfeiffer, 1855) MZU. MOL. 22.149 B Japonia bauensis Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 0014465 C Japonia borneensis (E. A. Smith, 1893) MZU. MOL. 22.155 D Japonia mundyana (Godwin-Austen, 1889) MZU. MOL. 22.133." figureDoi="10.3897/zookeys.1198.116265.figure6" httpUri="https://binary.pensoft.net/fig/1034245" pageId="0" pageNumber="295">6A</figureCitation>
</paragraph>
<paragraph id="17E9B6ED420329C58FFBCD8B54709A22" pageId="0" pageNumber="295">
<emphasis id="E015F57CC1AED6CF23FC26A2F7FE7BAF" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Borneo, Sarawak&quot;.
</paragraph>
<paragraph id="33D71BF3FE7F60728C7870D1A51D96F5" pageId="0" pageNumber="295">
<emphasis id="B63DF95490AB769AEA4B392C0C20C4CF" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="76ABE63BD6E95CCBD36884A3D0B69DA9" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="CD41C61AAB6244A23F05A306328B14A6" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14464, ME 15012, MZU.MOL.22.149.
</paragraph>
<paragraph id="2645716B01399B59F1D00CDBB867067C" pageId="0" pageNumber="295">
<emphasis id="B4AE9B2DC8BA58DA26F9AA776A6BF154" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak to Mukah, also in central Sarawak. Endemic to Sarawak (
<bibRefCitation id="242F7523D9E761541248462A21551E7A" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="5B442F7B421B41CE7F56BA04F3919B91" pageId="0" pageNumber="295">
<emphasis id="891612A20FD8C8D620D5EC56131C6FCE" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="337B90CB29DEEDF323D4FB06A909B7EF" pageId="0" pageNumber="295">
<emphasis id="059816B83C4A52821F8A77FEFD4B0E2F" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="BCADC618B1A38596A8B00568734DEFC1" authorityName="Marzuki, T. S. Liew &amp; Mohd-Azlan" authorityYear="2021" class="Gastropoda" family="Cyclophoridae" genus="Japonia" kingdom="Animalia" lsidName="Japonia bauensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="bauensis">
<emphasis id="48C7D5153678B934FBEDE443170FFC85" italics="true" pageId="0" pageNumber="295">Japonia bauensis</emphasis>
</taxonomicName>
Marzuki, Liew &amp; Mohd-Azlan, 2021
</emphasis>
</paragraph>
<paragraph id="8D7F72039461903AFC60C8DB304B8377" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="4691CA4DE2053A25C9D9B7CFD0991F59" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A Japonia barbata (L. Pfeiffer, 1855) MZU. MOL. 22.149 B Japonia bauensis Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 0014465 C Japonia borneensis (E. A. Smith, 1893) MZU. MOL. 22.155 D Japonia mundyana (Godwin-Austen, 1889) MZU. MOL. 22.133." figureDoi="10.3897/zookeys.1198.116265.figure6" httpUri="https://binary.pensoft.net/fig/1034245" pageId="0" pageNumber="295">6B</figureCitation>
</paragraph>
<caption id="672C0ACD1B7DE450FC8592D65E2A90FA" doi="10.3897/zookeys.1198.116265.figure6" httpUri="https://binary.pensoft.net/fig/1034245" pageId="0" pageNumber="295" start="Figure 6" startId="F6">
<paragraph id="C48A45A729808E7B0BFB0D059A3DF57E" pageId="0" pageNumber="295">
<emphasis id="9D9AD16161F917C51BAB8DA21B6C1A5C" bold="true" pageId="0" pageNumber="295">Figure 6.</emphasis>
<emphasis id="FED56A2C71EF0C013762C9B1B6EA2C51" bold="true" pageId="0" pageNumber="295">A</emphasis>
<taxonomicName id="E508661B7100ACD9BE16E225401FEEBD" class="Gastropoda" family="Cyclophoridae" genus="Japonia" kingdom="Animalia" lsidName="Japonia barbata" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="barbata">
<emphasis id="9FE4D98749C6E98C4F1F0690C4049F7F" italics="true" pageId="0" pageNumber="295">Japonia barbata</emphasis>
</taxonomicName>
(L. Pfeiffer, 1855) MZU.MOL.22.149 
<emphasis id="0136B054207A9F78A3621ABDBCB2C4FA" bold="true" pageId="0" pageNumber="295">B</emphasis>
<taxonomicName id="7FDFA86D5B36F37AD9A033AAC858761B" authorityName="Marzuki, T. S. Liew &amp; Mohd-Azlan" authorityYear="2021" class="Gastropoda" family="Cyclophoridae" genus="Japonia" kingdom="Animalia" lsidName="Japonia bauensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="bauensis">
<emphasis id="2E13C08EB786630B962F22E43CA0A187" italics="true" pageId="0" pageNumber="295">Japonia bauensis</emphasis>
</taxonomicName>
Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 0014465 
<emphasis id="1F2664C633E6EB18738B21F7AB7BC42A" bold="true" pageId="0" pageNumber="295">C</emphasis>
<taxonomicName id="4951A7AF1228E70559A848F0386D1FE7" baseAuthorityName="E. A. Smith" baseAuthorityYear="1893" class="Gastropoda" family="Cyclophoridae" genus="Japonia" kingdom="Animalia" lsidName="Japonia borneensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="borneensis">
<emphasis id="3E51C3C7D77C634AA7D09678B1A476DE" italics="true" pageId="0" pageNumber="295">Japonia borneensis</emphasis>
</taxonomicName>
(E. A. Smith, 1893) MZU.MOL.22.155 
<emphasis id="A788492C79A9DECC13F47D0FAC0BFD8A" bold="true" pageId="0" pageNumber="295">D</emphasis>
<taxonomicName id="469D40C486303122B47B126D92DFA6CD" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1889" class="Gastropoda" family="Cyclophoridae" genus="Japonia" kingdom="Animalia" lsidName="Japonia mundyana" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="mundyana">
<emphasis id="9F8CA08EB25666267A30439EFAA0D7E6" italics="true" pageId="0" pageNumber="295">Japonia mundyana</emphasis>
</taxonomicName>
(Godwin-Austen, 1889) MZU.MOL.22.133.
</paragraph>
</caption>
<paragraph id="497DC8CA09C35C7A223067E5C29B4480" pageId="0" pageNumber="295">
<emphasis id="DA2468804413980EACFD8B39237B69B6" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Bau and Serian-Padawan limestone hill clusters.&quot;
</paragraph>
<paragraph id="1AFF6252CE7B0644B9E17F35E093F9CA" pageId="0" pageNumber="295">
<emphasis id="B7A238AB937A037A625AC5FF0900FDBB" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="2257D364D1EB5F6F73DDF5526D396541" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="48B42B0DE7E82ABFBBCBAD6246074A41" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14465, ME 15013, MZU.MOL.22.156.
</paragraph>
<paragraph id="BC77BFF9E138E141D3CE1A549A1CBE9B" pageId="0" pageNumber="295">
<emphasis id="7D985C3ADDAC16A15ECC3B46BB23022D" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in Sarawak. Endemic to Sarawak (
<bibRefCitation id="31A9308DB4E6F17C873884B353FF895C" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="7E7013B3A72951802CDBDFBA0554A02B" pageId="0" pageNumber="295">
<emphasis id="B67F2889B7EB14C9031610C1C7B75AFE" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="19B6DE631506FE796013F669CA52F570" pageId="0" pageNumber="295">
<emphasis id="EB94D6508CBDA0C45757686A19314545" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="44D7EAA428270F0DC866AB278D996243" baseAuthorityName="E. A. Smith" baseAuthorityYear="1893" class="Gastropoda" family="Cyclophoridae" genus="Japonia" kingdom="Animalia" lsidName="Japonia borneensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="borneensis">
<emphasis id="F64F2AEE1C9425C8FB492C317AC7220D" italics="true" pageId="0" pageNumber="295">Japonia borneensis</emphasis>
</taxonomicName>
(E. A. Smith, 1893)
</emphasis>
</paragraph>
<paragraph id="4AF57962D99462D7F9445B16401C46CD" pageId="0" pageNumber="295">
Figs 
<figureCitation id="B92E2BF9700E48BAF2BA880B83847AC7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A Japonia barbata (L. Pfeiffer, 1855) MZU. MOL. 22.149 B Japonia bauensis Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 0014465 C Japonia borneensis (E. A. Smith, 1893) MZU. MOL. 22.155 D Japonia mundyana (Godwin-Austen, 1889) MZU. MOL. 22.133." figureDoi="10.3897/zookeys.1198.116265.figure6" httpUri="https://binary.pensoft.net/fig/1034245" pageId="0" pageNumber="295">6C</figureCitation>
, 
<figureCitation id="F9A674BCBE21B789EDC87934F0AB5B3A" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Living snails from Gua Rumbang. A Pincerna globosa (H. Adams, 1871) MZU. MOL. 22.135 B Stomacosmethis hosei (Godwin-Austen, 1889) MZU. MOL. 22.136 C Craspedotropis borneensis (Godwin-Austen, 1889) MZU. MOL. 23.141 D Japonia borneensis (E. A. Smith, 1893) MZU. MOL. 23.138. Images not to scale." figureDoi="10.3897/zookeys.1198.116265.figure19" httpUri="https://binary.pensoft.net/fig/1034258" pageId="0" pageNumber="295">19D</figureCitation>
</paragraph>
<paragraph id="0461E01D26B4C9B0017E443E915A868B" pageId="0" pageNumber="295">
<emphasis id="B9F4E8AF50EE47CF7A57301067FD4980" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Westliches Borneo bei Bengkajang&quot; [= Western Borneo near Bengkayang].
</paragraph>
<paragraph id="58B91E97496A7FB7ED8D671CB2C6D3F1" pageId="0" pageNumber="295">
<emphasis id="0EA96DA0C8BDE672BD80A8AC36DF5D9E" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="4A478AC35CC5576CAF6BD7CD960FC74C" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="E55426643C6510C927AD428913CE8184" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Apr. 2016-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; MZU.MOL.23.138, MZU.MOL.22.155, MZU.MOL.16.110.
</paragraph>
<paragraph id="5A3BE891216FD86CAA58744519BD15F9" pageId="0" pageNumber="295">
<emphasis id="F798690DCC21C11568DAB16CE6539D43" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed in Borneo (
<bibRefCitation id="9FE773F137AA53A341B8DAD8CD7DE841" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
).
</paragraph>
<paragraph id="75E936ED0F0E6348ACB287A46439F9F5" pageId="0" pageNumber="295">
<emphasis id="F93B55C9702663AFD32E295A1531489C" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were found foraging in leaf-litter and plant debris at the base of limestone cliffs.
</paragraph>
<paragraph id="9144478D15AF5404CE2B4AB80FE26E6B" pageId="0" pageNumber="295">
<emphasis id="C5142456F77DBD969A9E308ECD9D0C0E" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="D51479FCE79C4B984F9E65792B8C7F30" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1889" class="Gastropoda" family="Cyclophoridae" genus="Japonia" kingdom="Animalia" lsidName="Japonia mundyana" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="mundyana">
<emphasis id="3CE2D43FC67E3257C16A3139F3BA9A1D" italics="true" pageId="0" pageNumber="295">Japonia mundyana</emphasis>
</taxonomicName>
(Godwin-Austen, 1889)
</emphasis>
</paragraph>
<paragraph id="D423E2B62F2862DBE365B747C5CE4036" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="B3BEA493DC118DD0AB17584E2D367AB3" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A Japonia barbata (L. Pfeiffer, 1855) MZU. MOL. 22.149 B Japonia bauensis Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 0014465 C Japonia borneensis (E. A. Smith, 1893) MZU. MOL. 22.155 D Japonia mundyana (Godwin-Austen, 1889) MZU. MOL. 22.133." figureDoi="10.3897/zookeys.1198.116265.figure6" httpUri="https://binary.pensoft.net/fig/1034245" pageId="0" pageNumber="295">6D</figureCitation>
</paragraph>
<paragraph id="E50B736EDF1F568DF609450FBF3E974F" pageId="0" pageNumber="295">
<emphasis id="0AE3425E59B98EDED11FC4E97F704E06" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Busan Hills, Borneo&quot; [= Jambusan Hills, Bau, Sarawak].
</paragraph>
<paragraph id="AF9C06A2450CC0C08C6C85C3A8304840" pageId="0" pageNumber="295">
<emphasis id="D735DCA214B38D616E75F90DCCA89228" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="6B933287FFFE7624E353E895FF58AB0D" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="B8F739848C19DFBE069CAC26497D3BBE" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 15014, MZU.MOL.22.133.
</paragraph>
<paragraph id="F9C44EDEEC0F4CF254EA05B4F20BB92B" pageId="0" pageNumber="295">
<emphasis id="806756C93B4B4C3CAFD8F68F6E978488" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak. Endemic to western Sarawak (
<bibRefCitation id="4E9E2BC5B9D99D6419D4DD70F646F66A" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="254BA68958A28A01030D1001827DF152" pageId="0" pageNumber="295">
<emphasis id="5B67F8F12B94EAAA8D553C5C9FAEFF15" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snail was observed foraging on leaf surface of plants at the base of limestone cliffs.
</paragraph>
<paragraph id="AFAFEE0D0A254945BA1F1B40264F9238" pageId="0" pageNumber="295">
<emphasis id="24760204E840CCB8CB52B9BE1D384702" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="AD54641B90FCA076BB1EB43D1F4552C1" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1853" class="Gastropoda" family="Cyclophoridae" genus="Leptopoma" kingdom="Animalia" lsidName="Leptopoma sericatum" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="sericatum">
<emphasis id="F4C982B3C3E6CEB06EB97767EB411E1B" italics="true" pageId="0" pageNumber="295">Leptopoma sericatum</emphasis>
</taxonomicName>
(L. Pfeiffer, 1851)
</emphasis>
</paragraph>
<paragraph id="BB2DEFDAB51AF760F67D487E95D2CE84" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="2406D6FDB393C5F73398CC1FC528A4EF" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A Leptopoma sericatum (L. Pfeiffer, 1851) MZU. MOL. 23.135 B Opisthoporus biciliatus (Mousson, 1849) ME 14463 C Pterocyclos tenuilabiatus (Metcalfe, 1852) MZU. MOL. 22.129." figureDoi="10.3897/zookeys.1198.116265.figure7" httpUri="https://binary.pensoft.net/fig/1034246" pageId="0" pageNumber="295">7A</figureCitation>
</paragraph>
<caption id="1E130A641004FCBC730162F251AC0894" doi="10.3897/zookeys.1198.116265.figure7" httpUri="https://binary.pensoft.net/fig/1034246" pageId="0" pageNumber="295" start="Figure 7" startId="F7">
<paragraph id="F351AACD630E3E37EAD5D6AF4772DF42" pageId="0" pageNumber="295">
<emphasis id="36E5955B663A688627DF98DA7605E972" bold="true" pageId="0" pageNumber="295">Figure 7.</emphasis>
<emphasis id="719B0DBD68FB702AA7F3CCBCC475B37A" bold="true" pageId="0" pageNumber="295">A</emphasis>
<taxonomicName id="F4FBFDA63BD27D360EA5EF2B01008B56" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1853" class="Gastropoda" family="Cyclophoridae" genus="Leptopoma" kingdom="Animalia" lsidName="Leptopoma sericatum" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="sericatum">
<emphasis id="1AA1D74CFAF669C5F98FBCE261CFF66E" italics="true" pageId="0" pageNumber="295">Leptopoma sericatum</emphasis>
</taxonomicName>
(L. Pfeiffer, 1851) MZU. MOL.23.135 
<emphasis id="67334B2EC61B740C19CD486A89948EC7" bold="true" pageId="0" pageNumber="295">B</emphasis>
<taxonomicName id="BEB5767CCFB21655F4E95B869A9A1B3E" baseAuthorityName="Mousson" baseAuthorityYear="1849" class="Gastropoda" family="Cyclophoridae" genus="Opisthoporus" kingdom="Animalia" lsidName="Opisthoporus biciliatus" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="biciliatus">
<emphasis id="F7EB710AFF367AF703E590F63DFF7EA2" italics="true" pageId="0" pageNumber="295">Opisthoporus biciliatus</emphasis>
</taxonomicName>
(Mousson, 1849) ME 14463 
<emphasis id="058619F26C408217F3D1306C1D6D76A3" bold="true" pageId="0" pageNumber="295">C</emphasis>
<taxonomicName id="2D06032D9F938B97FF4EE3D86FF0D5EB" baseAuthorityName="Metcalfe" baseAuthorityYear="1852" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos tenuilabiatus" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="tenuilabiatus">
<emphasis id="B3946FDADDF159112E1836A1B5E434E4" italics="true" pageId="0" pageNumber="295">Pterocyclos tenuilabiatus</emphasis>
</taxonomicName>
(Metcalfe, 1852) MZU.MOL.22.129.
</paragraph>
</caption>
<paragraph id="A1270C524FE2BB4A87058217B7D6D3C2" pageId="0" pageNumber="295">
<emphasis id="1CEB8C45458CBED51394F10F62C71680" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
<normalizedToken id="A68E23C1810371F6F46734ECAB202404" originalValue="“Borneo”">&quot;Borneo&quot;</normalizedToken>
.
</paragraph>
<paragraph id="D1E19783CE637A4C4791D638177A519C" pageId="0" pageNumber="295">
<emphasis id="EB09EA5A5587A236408011A936ED6441" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="430BDA94DFA4BCB35AB91715BF836987" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="B50124EFBD6B538299B54F929E4C5376" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 15015, MZU.MOL.23.135.
</paragraph>
<paragraph id="D1D101E445DBDF3188D61169C97C8B51" pageId="0" pageNumber="295">
<emphasis id="32602F592CAA54672DAB1A89B366B91F" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed in Borneo (
<bibRefCitation id="3878D6C2A78B91D7AA533298AF54D570" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
; 
<bibRefCitation id="B97346A57BBD8AC5A71CBF266280EA7A" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
).
</paragraph>
<paragraph id="93811B419E5BA401E88C99E56431BA81" pageId="0" pageNumber="295">
<emphasis id="AB408D8970F42F9B6B6C6C4DF4F9E2C6" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="04F27342EA3BA0827388C885A5894D23" pageId="0" pageNumber="295">
<emphasis id="CD680212540F9FC339FD0D8FAFEAC346" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="2AB7BBDA935522CCBAFC9EBBCB731FCA" baseAuthorityName="Mousson" baseAuthorityYear="1849" class="Gastropoda" family="Cyclophoridae" genus="Opisthoporus" kingdom="Animalia" lsidName="Opisthoporus biciliatus" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="biciliatus">
<emphasis id="C20901BD7554F15AF71622FF87423E8B" italics="true" pageId="0" pageNumber="295">Opisthoporus biciliatus</emphasis>
</taxonomicName>
(Mousson, 1849)
</emphasis>
</paragraph>
<paragraph id="84696B5061337E4262FFA733E4A05D91" pageId="0" pageNumber="295">
Figs 
<figureCitation id="59203C20B4BDB481A48D3A118147D08B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A Leptopoma sericatum (L. Pfeiffer, 1851) MZU. MOL. 23.135 B Opisthoporus biciliatus (Mousson, 1849) ME 14463 C Pterocyclos tenuilabiatus (Metcalfe, 1852) MZU. MOL. 22.129." figureDoi="10.3897/zookeys.1198.116265.figure7" httpUri="https://binary.pensoft.net/fig/1034246" pageId="0" pageNumber="295">7B</figureCitation>
, 
<figureCitation id="D7B466EF6BC4683AB5B50C6B2032230F" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Living snails from Gua Rumbang. A Opisthoporus biciliatus (Mousson, 1849) ME 14979 B Diplommatina adversa (H. Adams &amp; A. Adams, 1851) MZU. MOL. 23.140 C Diplommatina baritensis E. A. Smith, 1893 MZU. MOL. 23.137 D Plectostoma anisopterum ME 14479 (Vermeulen, 1994). All images not to scale." figureDoi="10.3897/zookeys.1198.116265.figure20" httpUri="https://binary.pensoft.net/fig/1034259" pageId="0" pageNumber="295">20A</figureCitation>
</paragraph>
<paragraph id="3D8419E7A5D578E00E2FC841E010B5C5" pageId="0" pageNumber="295">
<emphasis id="14368DE015417BDEE14B0A7E0A2FB1AD" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
<normalizedToken id="89C67048BBEB34BC593AE89307B41532" originalValue="“Java”">&quot;Java&quot;</normalizedToken>
[= Borneo (
<bibRefCitation id="EFE245E7E8F2765419E4B156EBB8051C" DOI="https://doi.org/10.1111/j.1096-3642.1851.tb01132.x" author="Metcalfe, W" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="295" pagination="70 - 74" refId="B32" refString="Metcalfe, W, 1852. An enumeration of species of recent shells, received by W. J. Hamilton, Esq., from Borneo, in November 1850, with descriptions of the new species. Proceedings of the Zoological Society of London 19: 70 - 74, DOI: https://doi.org/10.1111/j.1096-3642.1851.tb01132.x" title="An enumeration of species of recent shells, received by W. J. Hamilton, Esq., from Borneo, in November 1850, with descriptions of the new species." url="https://doi.org/10.1111/j.1096-3642.1851.tb01132.x" volume="19" year="1852">Metcalfe 1852</bibRefCitation>
].
</paragraph>
<paragraph id="A1547B059BA0BBA0B10BFC7EF33F11D5" pageId="0" pageNumber="295">
<emphasis id="4773110C4092DD56132B884AB477D207" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="27EA5B0AC650732823026D6AB1009A57" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="C9B88C59101A2EF92BD0E247C8933247" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14463, ME 14979, ME 15011, MZU.MOL.22.152.
</paragraph>
<paragraph id="CD567D0F8586416C9EE2A11E6F6ECEDC" pageId="0" pageNumber="295">
<emphasis id="E20471C861A1D326C30DE6C9E804BADA" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak to Mukah, also in central Sarawak. Endemic to Sarawak (
<bibRefCitation id="D6856FCAE1B1E4DF3D430C478D6DF806" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="F06DF8A502C7E2A2AE3A99E31A2D6E69" pageId="0" pageNumber="295">
<emphasis id="45B9A16E75D793E87F10974B5A16C304" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="C233FCDF8ABABE5387025BF645D10523" pageId="0" pageNumber="295">
<emphasis id="DA4FC6A15783F87222F6637AE04A4A2E" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="3110B77714028F539026E3CF10F72E38" baseAuthorityName="Metcalfe" baseAuthorityYear="1852" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos tenuilabiatus" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="tenuilabiatus">
<emphasis id="4BFE98DDCBB4FF817BDA9F10C3378805" italics="true" pageId="0" pageNumber="295">Pterocyclos tenuilabiatus</emphasis>
</taxonomicName>
(Metcalfe, 1852)
</emphasis>
</paragraph>
<paragraph id="759B77DA983049E88D6E237FA15B0054" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="361329D706E518170199F26D9490791B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A Leptopoma sericatum (L. Pfeiffer, 1851) MZU. MOL. 23.135 B Opisthoporus biciliatus (Mousson, 1849) ME 14463 C Pterocyclos tenuilabiatus (Metcalfe, 1852) MZU. MOL. 22.129." figureDoi="10.3897/zookeys.1198.116265.figure7" httpUri="https://binary.pensoft.net/fig/1034246" pageId="0" pageNumber="295">7C</figureCitation>
</paragraph>
<paragraph id="67EB65CEBD29D95D73EE6AE7545D593A" pageId="0" pageNumber="295">
<emphasis id="D7D61433D48D8AA36AE5A3DCC919D3A6" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
<normalizedToken id="74B8F391C014A441784C0692401EF20E" originalValue="“Borneo”">&quot;Borneo&quot;</normalizedToken>
.
</paragraph>
<paragraph id="E9D1390399C2449547ABE652B46F4521" pageId="0" pageNumber="295">
<emphasis id="D2186416A51295EEB922AD7CD92E3DE1" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="B0F81E2E41C213BADDD7B5CEEACD10D9" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="AE760EA6A380E099DE42EC7D10997B84" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14462, ME 14978, MZU.MOL.22.129.
</paragraph>
<paragraph id="137BC4EC08EBFF1108F5BAAB390D1F03" pageId="0" pageNumber="295">
<emphasis id="DEC080FD742DEC91B07A5A3AA7556F20" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed in Borneo. Endemic to Borneo (
<bibRefCitation id="9DFE66415CCDD554BC4A69A9748A8EE0" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
; 
<bibRefCitation id="5BD085279759823B145432E78AA07F70" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
).
</paragraph>
<paragraph id="717A28FDDC374E0E4D5EF771D2216BC5" pageId="0" pageNumber="295">
<emphasis id="DAC6D80FAD27676986430BF44152613A" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="5014D182A10489FA26F1DABA205FEE24" pageId="0" pageNumber="295">
<emphasis id="698FC2AEABA3A67DC8BA3EB63E2DE300" bold="true" pageId="0" pageNumber="295">
Family 
<taxonomicName id="C1A23FB0FD22AD2036B25F1AEA9CB2F9" authorityName="L. Pfeiffer" authorityYear="1856" class="Gastropoda" family="Diplommatinidae" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="family">Diplommatinidae</taxonomicName>
L. Pfeiffer, 1856
</emphasis>
</paragraph>
<paragraph id="FB06EDC9DB4BF2372EDC914E920975FD" pageId="0" pageNumber="295">
<emphasis id="E3DDE73FFD2CCADCE05320754DDD780C" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="E5CE04009CC3D27E1398A43530C6CF6A" baseAuthorityName="H. Adams &amp; A. Adams" baseAuthorityYear="1851" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina adversa" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="adversa">
<emphasis id="8E76374213E1A28DAC0A0DA80812F498" italics="true" pageId="0" pageNumber="295">Diplommatina adversa</emphasis>
</taxonomicName>
(H. Adams &amp; A. Adams, 1851)
</emphasis>
</paragraph>
<paragraph id="E790D809EBA24A494411F17FE67A99A1" pageId="0" pageNumber="295">
Figs 
<figureCitation id="74C52F043C200612AA067F1A84F00C4F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A Diplommatina adversa (H. Adams &amp; A. Adams, 1851) ME 14476 B Diplommatina baritensis E. A. Smith, 1893 ME 14472 C Diplommatina concinna H. Adams, 1872 ME 14473 D Diplommatina maduana maduana Laidlaw, 1949 ME 14474 E Diplommatina rumbangensis sp. nov. holotype MZU. MOL. 22.132 F Diplommatina subglaber subisensis (Vermeulen, 1993) ME 14475." figureDoi="10.3897/zookeys.1198.116265.figure8" httpUri="https://binary.pensoft.net/fig/1034247" pageId="0" pageNumber="295">8A</figureCitation>
, 
<figureCitation id="78708BC6786947BADF381347E6F3AC82" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Living snails from Gua Rumbang. A Opisthoporus biciliatus (Mousson, 1849) ME 14979 B Diplommatina adversa (H. Adams &amp; A. Adams, 1851) MZU. MOL. 23.140 C Diplommatina baritensis E. A. Smith, 1893 MZU. MOL. 23.137 D Plectostoma anisopterum ME 14479 (Vermeulen, 1994). All images not to scale." figureDoi="10.3897/zookeys.1198.116265.figure20" httpUri="https://binary.pensoft.net/fig/1034259" pageId="0" pageNumber="295">20B</figureCitation>
</paragraph>
<caption id="A9102891E10BFBEE590E4F9F144D4723" doi="10.3897/zookeys.1198.116265.figure8" httpUri="https://binary.pensoft.net/fig/1034247" pageId="0" pageNumber="295" start="Figure 8" startId="F8">
<paragraph id="37ED41B2E93847D7B4E0F333C821DC84" pageId="0" pageNumber="295">
<emphasis id="0311492C2DFA769214C6AC8FD34DC88F" bold="true" pageId="0" pageNumber="295">Figure 8.</emphasis>
<emphasis id="CA44AFDE4769F3EF4A7A39FD5969A454" bold="true" pageId="0" pageNumber="295">A</emphasis>
<taxonomicName id="C007CAC315B0D1ABE9EF90E8A399A255" baseAuthorityName="H. Adams &amp; A. Adams" baseAuthorityYear="1851" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina adversa" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="adversa">
<emphasis id="8D9902AAE1808E3B3071B335CC0D0901" italics="true" pageId="0" pageNumber="295">Diplommatina adversa</emphasis>
</taxonomicName>
(H. Adams &amp; A. Adams, 1851) ME 14476 
<emphasis id="36FF1F9C440836B19D06EE2EF38229BF" bold="true" pageId="0" pageNumber="295">B</emphasis>
<taxonomicName id="E12A6E797BF800DE91534BEE5E432586" authorityName="E. A. Smith" authorityYear="1893" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina baritensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="baritensis">
<emphasis id="EBDC9203E8929035C851D85F702485A4" italics="true" pageId="0" pageNumber="295">Diplommatina baritensis</emphasis>
</taxonomicName>
E. A. Smith, 1893 ME 14472 
<emphasis id="7CBAD25433916FBC938BE26E7A7C26C7" bold="true" pageId="0" pageNumber="295">C</emphasis>
<taxonomicName id="F5ADD7141DAF7205B75BEB7A9B8A0B93" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina concinna" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="concinna">
<emphasis id="03751A1CFB8B57D08F8534FF48CB1B9A" italics="true" pageId="0" pageNumber="295">Diplommatina concinna</emphasis>
</taxonomicName>
H. Adams, 1872 ME 14473 
<emphasis id="AA65F1A05EEC515A4C69D2F6871AF96B" bold="true" pageId="0" pageNumber="295">D</emphasis>
<taxonomicName id="F5F4A42CFC5985FEBF5189E68F97EEA3" authorityName="Laidlaw" authorityYear="1949" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina maduana subsp. maduana" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="subSpecies" species="maduana" subSpecies="maduana">
<emphasis id="796B5DCB11AAB4F0C93D38A71E123D51" italics="true" pageId="0" pageNumber="295">Diplommatina maduana maduana</emphasis>
</taxonomicName>
Laidlaw, 1949 ME 14474 
<emphasis id="F558874C596B52343244DB7D59A460C9" bold="true" pageId="0" pageNumber="295">E</emphasis>
<taxonomicName id="0A04C36B134D30F7DE1BFC425EA06BDB" authorityName="Nasir &amp; Lee &amp; Marzuki &amp; Vermeulen &amp; Mohd-Azlan &amp; Khalik" authorityYear="2024" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina rumbangensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="rumbangensis">
<emphasis id="7549F4E33279E205CD643596FF236B14" italics="true" pageId="0" pageNumber="295">Diplommatina rumbangensis</emphasis>
</taxonomicName>
sp. nov. holotype MZU.MOL.22.132 
<emphasis id="6BA6B7A4FA462DE2C2492006F2EB2AE0" bold="true" pageId="0" pageNumber="295">F</emphasis>
<taxonomicName id="BA09D8E8AE77C9679F28AB7D029CAA3E" baseAuthorityName="Vermeulen" baseAuthorityYear="1993" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina subglaber subsp. subisensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="subSpecies" species="subglaber" subSpecies="subisensis">
<emphasis id="ECFA64C969AD043CC611C1CA3044C4AA" italics="true" pageId="0" pageNumber="295">Diplommatina subglaber subisensis</emphasis>
</taxonomicName>
(Vermeulen, 1993) ME 14475.
</paragraph>
</caption>
<paragraph id="A509542E111514FF86AF31ABF06C4628" pageId="0" pageNumber="295">
<emphasis id="6A8EC2F8139D517028F1683EC9D1C6DE" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
<normalizedToken id="EFF82DE564D1911182A2ABFE036C9137" originalValue="“Singapore”">&quot;Singapore&quot;</normalizedToken>
.
</paragraph>
<paragraph id="FB19B4E2FFD35DC7F5024ED39A9395E4" pageId="0" pageNumber="295">
<emphasis id="4A872046F2BA62BA50ED6B85C8F49864" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="291F76B55E12B30565E28EA4ACD0A28B" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="73E492C459F1C08D6AD36AEB7DBCE49A" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14476, MZU.MOL.22.140, MZU.MOL.23.140.
</paragraph>
<paragraph id="CD2476DCCC5AA97E31C12E30FD308EA3" pageId="0" pageNumber="295">
<emphasis id="5CB070F7AF4511A60CE17DC519F864B7" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak. In Sabah found in Ulu Segama only (
<bibRefCitation id="9DAF7EF8830053CB659F735D092D0ED5" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
; 
<bibRefCitation id="1E3547C6041861B65590254327BE1235" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
). Also found in West Malaysia and Singapore (
<bibRefCitation id="04F42BFCFAF71BF50418CA7999568DEA" author="Laidlaw, FF" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="295" pagination="199 - 215" publicationUrl="https://lkcnhm.nus.edu.sg/app/uploads/2017/06/19brm199-215.pdf" refId="B25" refString="Laidlaw, FF, 1949. The Malayan species of Diplommatina (Cyclophoridae). Bulletin of the Raffles Museum 19: 199 - 215, https://lkcnhm.nus.edu.sg/app/uploads/2017/06/19brm199-215.pdf" title="The Malayan species of Diplommatina (Cyclophoridae)." url="https://lkcnhm.nus.edu.sg/app/uploads/2017/06/19brm199-215.pdf" volume="19" year="1949">Laidlaw 1949</bibRefCitation>
).
</paragraph>
<paragraph id="63B5EE75B1232078E926A5B7CEB6F7DD" pageId="0" pageNumber="295">
<emphasis id="4CB5C4B63C1BEF8B4B398CBF884FC929" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were found foraging in leaf-litter and plant debris at the base of limestone cliffs.
</paragraph>
<paragraph id="958A75A381707956A409766F7A89DD33" pageId="0" pageNumber="295">
<emphasis id="1D8BD706D5307A48F64D88664A379F08" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="5C3CA7EA0DD79E13175B0A70FBCB215A" authorityName="E. A. Smith" authorityYear="1893" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina baritensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="baritensis">
<emphasis id="80CEACEE78C6A9382675B339112440CB" italics="true" pageId="0" pageNumber="295">Diplommatina baritensis</emphasis>
</taxonomicName>
E. A Smith, 1893
</emphasis>
</paragraph>
<paragraph id="808582E5AE2C6ED0088E93D3706BCFF5" pageId="0" pageNumber="295">
Figs 
<figureCitation id="F7707C1E924E1A4C490CE8E7E971790D" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A Diplommatina adversa (H. Adams &amp; A. Adams, 1851) ME 14476 B Diplommatina baritensis E. A. Smith, 1893 ME 14472 C Diplommatina concinna H. Adams, 1872 ME 14473 D Diplommatina maduana maduana Laidlaw, 1949 ME 14474 E Diplommatina rumbangensis sp. nov. holotype MZU. MOL. 22.132 F Diplommatina subglaber subisensis (Vermeulen, 1993) ME 14475." figureDoi="10.3897/zookeys.1198.116265.figure8" httpUri="https://binary.pensoft.net/fig/1034247" pageId="0" pageNumber="295">8B</figureCitation>
, 
<figureCitation id="11464DCBA1DE70247859A2E731017421" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Living snails from Gua Rumbang. A Opisthoporus biciliatus (Mousson, 1849) ME 14979 B Diplommatina adversa (H. Adams &amp; A. Adams, 1851) MZU. MOL. 23.140 C Diplommatina baritensis E. A. Smith, 1893 MZU. MOL. 23.137 D Plectostoma anisopterum ME 14479 (Vermeulen, 1994). All images not to scale." figureDoi="10.3897/zookeys.1198.116265.figure20" httpUri="https://binary.pensoft.net/fig/1034259" pageId="0" pageNumber="295">20C</figureCitation>
</paragraph>
<paragraph id="2ED4AEFCDB2CE76F5EA472792BA4EBCC" pageId="0" pageNumber="295">
<emphasis id="28B7DFF3E2764C0944593848F4A155E0" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Barit Mountain, N.W. Borneo&quot;.
</paragraph>
<paragraph id="90B1D66317A3B2210A0516CE52CA0AE1" pageId="0" pageNumber="295">
<emphasis id="FDFF14106B94C822E3C3D08A009D0B9C" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="98786375CC762E465B3FDE2D5D0A8FB3" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="4518CA75862451558FE9A0ACDAEC3441" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14472, ME 14984, ME 15022, MZU.MOL.22.169, MZU.MOL.23.137.
</paragraph>
<paragraph id="D2FCC7E60960887D96F2DFD875DE1530" pageId="0" pageNumber="295">
<emphasis id="9F390AB10EBA87B5DCFF26841572CCC7" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Padawan limestone hills at western Sarawak and Mulu limestone hills in northern Sarawak (
<bibRefCitation id="2BEB6D47CA192B88AE5BE151B7C597A7" author="Smith, EA" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="295" pagination="97 - 127" publicationUrl="https://www.biodiversitylibrary.org/page/30982902" refId="B48" refString="Smith, EA, 1895. On a collection of land-shells from Sarawak, British North Borneo, Palawan, and other neighboring islands. Proceedings of the Zoological Society of London 63 (1): 97 - 127, https://www.biodiversitylibrary.org/page/30982902" title="On a collection of land-shells from Sarawak, British North Borneo, Palawan, and other neighboring islands." url="https://www.biodiversitylibrary.org/page/30982902" volume="63" year="1895">Smith 1895</bibRefCitation>
; 
<bibRefCitation id="44F7376BD390FBEEBB740E3D86CD48E9" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" pagination="3 - 69" publicationUrl="http://natuurtijdschriften.nl/record/597005" refId="B52" refString="Vermeulen, JJ, 1993. Notes on the non-marine mollusks of the island of Borneo 5. The genus Diplommatina (Gastropoda Prosobranchia: Diplommatinidae). Basteria 57: 3 - 69, http://natuurtijdschriften.nl/record/597005" title="Notes on the non-marine mollusks of the island of Borneo 5. The genus Diplommatina (Gastropoda Prosobranchia: Diplommatinidae)." url="http://natuurtijdschriften.nl/record/597005" volume="57" year="1993">Vermeulen 1993</bibRefCitation>
).
</paragraph>
<paragraph id="840D06E637BD8E8AA9AE2C81885BC5BE" pageId="0" pageNumber="295">
<emphasis id="79CC02495EF4539338F5E8A18074E34A" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were found foraging in leaf-litter and plant debris at the base of limestone cliffs.
</paragraph>
<paragraph id="A430522086C563A1A42A6203D6DBAE2B" pageId="0" pageNumber="295">
<emphasis id="39410E2EC7C659DD88072FADDF542C1B" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="CE60EB181EE927DE91756F739EA225ED" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina concinna" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="concinna">
<emphasis id="00371E9626EF8242C32B80437D8A68A8" italics="true" pageId="0" pageNumber="295">Diplommatina concinna</emphasis>
</taxonomicName>
H. Adams, 1872
</emphasis>
</paragraph>
<paragraph id="5CBB6C33FD623960C5532361FA2E8D64" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="36C44016321FB4854BE7FB88B5EF3474" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A Diplommatina adversa (H. Adams &amp; A. Adams, 1851) ME 14476 B Diplommatina baritensis E. A. Smith, 1893 ME 14472 C Diplommatina concinna H. Adams, 1872 ME 14473 D Diplommatina maduana maduana Laidlaw, 1949 ME 14474 E Diplommatina rumbangensis sp. nov. holotype MZU. MOL. 22.132 F Diplommatina subglaber subisensis (Vermeulen, 1993) ME 14475." figureDoi="10.3897/zookeys.1198.116265.figure8" httpUri="https://binary.pensoft.net/fig/1034247" pageId="0" pageNumber="295">8C</figureCitation>
</paragraph>
<paragraph id="EB755339F3DD37490B6D7B0F63304C3E" pageId="0" pageNumber="295">
<emphasis id="47E321DAA2A0F30F5751A42EE4E42821" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
<normalizedToken id="92527760E7E8C23A40A34EA4DBD9EE62" originalValue="“Borneo”">&quot;Borneo&quot;</normalizedToken>
.
</paragraph>
<paragraph id="8E4107BCDC061436F86FB3A2A17A87D0" pageId="0" pageNumber="295">
<emphasis id="6A4DAAAB4BC88C7721D851277BCA9D38" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="66CAE17F8295A8A5ECA1D6DB7299A752" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="FCB42BF00FBA2BEB1BE46A430F78A166" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14473, MZU.MOL.22.83.
</paragraph>
<paragraph id="1A903E2DCF95F3B9F6F46D725D22197E" pageId="0" pageNumber="295">
<emphasis id="743ADC0EB0C837A423285C9455271749" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak to Niah, in northern Sarawak. Also found in Bunguran, Indonesia (
<bibRefCitation id="8C7F3D12D67CEBD07042F6D97E77947C" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
) and Singapore (
<bibRefCitation id="133B37AD69CACB8B9BDD4AB03778D205" author="Chan, SY" journalOrPublisher="Club Conchylia Informationen" pageId="0" pageNumber="295" publicationUrl="https://lkcnhm.nus.edu.sg/wp-content/uploads/sites/10/2020/11/sbr2020_200.pdf" refId="B9" refString="Chan, SY, 2020. Annual diplomate snails, Diplommatina concinna, on Bukit Timah Hill. Singapore Biodiversity Records 2020: 200. https://lkcnhm.nus.edu.sg/wp-content/uploads/sites/10/2020/11/sbr2020_200.pdf" title="Annual diplomate snails, Diplommatina concinna, on Bukit Timah Hill. Singapore Biodiversity Records 2020: 200." url="https://lkcnhm.nus.edu.sg/wp-content/uploads/sites/10/2020/11/sbr2020_200.pdf" year="2020">Chan 2020</bibRefCitation>
).
</paragraph>
<paragraph id="BC3C30D27E2E5E0089EDD9A3AD8F0823" pageId="0" pageNumber="295">
<emphasis id="E14F780ACE0FA8056F9658CE8A8EFEB5" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were found foraging in leaf-litter and plant debris at the base of limestone cliffs.
</paragraph>
<paragraph id="18B885B840EE7F10F3D9DCE8436CBCBD" pageId="0" pageNumber="295">
<emphasis id="4CCFBBEA90E5197CB3747669BFB03C62" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="20CB18B5C732C5239D493C3334C50D9D" authorityName="Laidlaw" authorityYear="1949" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina maduana subsp. maduana" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="subSpecies" species="maduana" subSpecies="maduana">
<emphasis id="0CA33462A2820C973476908DE229A918" italics="true" pageId="0" pageNumber="295">Diplommatina maduana maduana</emphasis>
</taxonomicName>
Laidlaw, 1949
</emphasis>
</paragraph>
<paragraph id="79016C5C11F3A72C00548732C3B41A3A" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="E4ED43BB13D4D752D0AF0817A7057A33" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A Diplommatina adversa (H. Adams &amp; A. Adams, 1851) ME 14476 B Diplommatina baritensis E. A. Smith, 1893 ME 14472 C Diplommatina concinna H. Adams, 1872 ME 14473 D Diplommatina maduana maduana Laidlaw, 1949 ME 14474 E Diplommatina rumbangensis sp. nov. holotype MZU. MOL. 22.132 F Diplommatina subglaber subisensis (Vermeulen, 1993) ME 14475." figureDoi="10.3897/zookeys.1198.116265.figure8" httpUri="https://binary.pensoft.net/fig/1034247" pageId="0" pageNumber="295">8D</figureCitation>
</paragraph>
<paragraph id="2ABDE7914AA464F2AB8E54617261386B" pageId="0" pageNumber="295">
<emphasis id="13486355F4EBC472F696C5304B2655CF" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Gua Madu, Kelantan&quot;.
</paragraph>
<paragraph id="2ABD8EB4ECFCDC9FCD550308B8376C89" pageId="0" pageNumber="295">
<emphasis id="B1DCCF64A6D335FD168AE0221BA97466" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="DC7F2D901F7720757229FD4A6B947F4B" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="72EEE36EFBCC806DCCB5A2CD89350AE1" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14474, ME 15023, MZU.MOL.22.190.
</paragraph>
<paragraph id="ED3BCD3434045422CF711CFEA85D04D2" pageId="0" pageNumber="295">
<emphasis id="6DA4BDFA0B560941328067054D2D1A5C" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak to Mulu hills in northern Sarawak (
<bibRefCitation id="250C53A5CB199144DE5B4C682E487134" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
). Also found in West Malaysia (
<bibRefCitation id="A9EF7B7CDDBADB3179CC66A5FF045387" author="Laidlaw, FF" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="295" pagination="199 - 215" publicationUrl="https://lkcnhm.nus.edu.sg/app/uploads/2017/06/19brm199-215.pdf" refId="B25" refString="Laidlaw, FF, 1949. The Malayan species of Diplommatina (Cyclophoridae). Bulletin of the Raffles Museum 19: 199 - 215, https://lkcnhm.nus.edu.sg/app/uploads/2017/06/19brm199-215.pdf" title="The Malayan species of Diplommatina (Cyclophoridae)." url="https://lkcnhm.nus.edu.sg/app/uploads/2017/06/19brm199-215.pdf" volume="19" year="1949">Laidlaw 1949</bibRefCitation>
).
</paragraph>
<paragraph id="1CA5D3BD6C59EB60C2F82AB1C01EE54B" pageId="0" pageNumber="295">
<emphasis id="E9A9491C84E4E60EBDA8C6C9B59ED46C" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were found foraging in leaf-litter and plant debris at the base of limestone cliffs.
</paragraph>
<paragraph id="B70E99A951DA967CB2C57995BE8B4A08" pageId="0" pageNumber="295">
<emphasis id="1AE99A5FB27713CE8F3F74014EE45F26" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="2D6E36D48BDB222B8F4C498B6FCAA46D" baseAuthorityName="Vermeulen" baseAuthorityYear="1993" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina subglaber subsp. subisensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="subSpecies" species="subglaber" subSpecies="subisensis">
<emphasis id="037EA71DDF325912EFEF459E3D5B8F61" italics="true" pageId="0" pageNumber="295">Diplommatina subglaber subisensis</emphasis>
</taxonomicName>
(Vermeulen, 1993)
</emphasis>
</paragraph>
<paragraph id="A1B425CD6740A4D9DA9EAA5A2D4C1C1E" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="0AA145805AB1150E078712E98770CE87" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A Diplommatina adversa (H. Adams &amp; A. Adams, 1851) ME 14476 B Diplommatina baritensis E. A. Smith, 1893 ME 14472 C Diplommatina concinna H. Adams, 1872 ME 14473 D Diplommatina maduana maduana Laidlaw, 1949 ME 14474 E Diplommatina rumbangensis sp. nov. holotype MZU. MOL. 22.132 F Diplommatina subglaber subisensis (Vermeulen, 1993) ME 14475." figureDoi="10.3897/zookeys.1198.116265.figure8" httpUri="https://binary.pensoft.net/fig/1034247" pageId="0" pageNumber="295">8F</figureCitation>
</paragraph>
<paragraph id="3FD1D5389742E1EE6976E99E97629CFB" pageId="0" pageNumber="295">
<emphasis id="1D31EDCD5B5DEFC1A16C4527F1A9EE34" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Sarawak 4th Div.: G. Subis (Batu Niah)&quot;.
</paragraph>
<paragraph id="79C1E5C53260B066328D015AD02E755C" pageId="0" pageNumber="295">
<emphasis id="EF28D3ABA27715A08B931E771944D511" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="070895F6A3022587AEF3E0E62AB559E0" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="A9957D9C869E544B789F6C4CB4EFC0E7" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14475.
</paragraph>
<paragraph id="2E546FD0C4BD2FF49A01471CA4C298EE" pageId="0" pageNumber="295">
<emphasis id="20CD49AFE148661E23D2560DFD062331" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Padawan limestone hills in western Sarawak and Niah limestone hills in northern Sarawak (
<bibRefCitation id="FCBE840FA68BEDF95938F1C29CD49F03" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" pagination="3 - 69" publicationUrl="http://natuurtijdschriften.nl/record/597005" refId="B52" refString="Vermeulen, JJ, 1993. Notes on the non-marine mollusks of the island of Borneo 5. The genus Diplommatina (Gastropoda Prosobranchia: Diplommatinidae). Basteria 57: 3 - 69, http://natuurtijdschriften.nl/record/597005" title="Notes on the non-marine mollusks of the island of Borneo 5. The genus Diplommatina (Gastropoda Prosobranchia: Diplommatinidae)." url="http://natuurtijdschriften.nl/record/597005" volume="57" year="1993">Vermeulen 1993</bibRefCitation>
, 
<bibRefCitation id="4EF0EE8CF9AB35D83DD2113C66EF12AD" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" publicationUrl="http://natuurtijdschriften.nl/record/597075" refId="B54" refString="Vermeulen, JJ, 1996. Notes on the non-marine molluscs of the island of Borneo 8. The genus Arina additions to the genera Diplommatina and Opisthostoma. Basteria 60: 87-138. http://natuurtijdschriften.nl/record/597075" title="Notes on the non-marine molluscs of the island of Borneo 8. The genus Arina additions to the genera Diplommatina and Opisthostoma. Basteria 60: 87 - 138." url="http://natuurtijdschriften.nl/record/597075" year="1996">1996</bibRefCitation>
).
</paragraph>
<paragraph id="34CA64D87C2D356D76BC62044C1CDBE4" pageId="0" pageNumber="295">
<emphasis id="A46AB2D30A422B429BF67A92C20D53EC" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="2AC3E524C6C8D6268A435790D0C542BE" pageId="0" pageNumber="295">
<emphasis id="4E65A0B67CB050BDE4BBC291FFA5387C" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="9BF0D4F3F44F6EF29D5FAFE6D33FE42D" authorityName="Vermeulen" authorityYear="1991" class="Gastropoda" family="Diplommatinidae" genus="Opisthostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthostoma brachyacrum subsp. lambii" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="subSpecies" species="brachyacrum" subSpecies="lambii">
<emphasis id="30E145B69AD089A7D4A3CEF17CE8BCD9" italics="true" pageId="0" pageNumber="295">Opisthostoma brachyacrum lambii</emphasis>
</taxonomicName>
(Vermeulen, 1991)
</emphasis>
</paragraph>
<paragraph id="0CE3336C6A9C5125C0589325DB6D42FB" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="C80883BBA668D1112A8F01018A012015" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A Opisthostoma brachyacrum lambii Vermeulen, 1991 ME 14480 B Opisthostoma tridens Vermeulen, 1991 ME 14481 C Plectostoma anisopterum (Vermeulen, 1994) ME 14479 D Plectostoma austeni (E. A. Smith, 1894 a) ME 14478 Gua Rumbang E Plectostoma pumilio (E. A. Smith, 1894 a) ME 14477." figureDoi="10.3897/zookeys.1198.116265.figure9" httpUri="https://binary.pensoft.net/fig/1034248" pageId="0" pageNumber="295">9A</figureCitation>
</paragraph>
<caption id="5522CA02EEA8D707CBFD49F6087A1793" doi="10.3897/zookeys.1198.116265.figure9" httpUri="https://binary.pensoft.net/fig/1034248" pageId="0" pageNumber="295" start="Figure 9" startId="F9">
<paragraph id="6D5A4272330B55EE2890A933BD47D8DF" pageId="0" pageNumber="295">
<emphasis id="CC63D757D505AA99011ED3D2625EC6D4" bold="true" pageId="0" pageNumber="295">Figure 9.</emphasis>
<emphasis id="1C7285C7EFF6695AF01378ECD6448377" bold="true" pageId="0" pageNumber="295">A</emphasis>
<taxonomicName id="B75BA780F94641F18CBF2213EA10FC15" authorityName="Vermeulen" authorityYear="1991" class="Gastropoda" family="Diplommatinidae" genus="Opisthostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthostoma brachyacrum subsp. lambii" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="subSpecies" species="brachyacrum" subSpecies="lambii">
<emphasis id="C4B05DFE537459F47AB75B53F6855FF4" italics="true" pageId="0" pageNumber="295">Opisthostoma brachyacrum lambii</emphasis>
</taxonomicName>
Vermeulen, 1991 ME 14480 
<emphasis id="26ABE7B75B59A98921E5CB400B6DE0B7" bold="true" pageId="0" pageNumber="295">B</emphasis>
<taxonomicName id="D958A7D8C1893089B4D40F9628480B70" authorityName="Vermeulen" authorityYear="1991" class="Gastropoda" family="Diplommatinidae" genus="Opisthostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthostoma tridens" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="tridens">
<emphasis id="4CF78461AAB898D0AFE84BF0D3FA325F" italics="true" pageId="0" pageNumber="295">Opisthostoma tridens</emphasis>
</taxonomicName>
Vermeulen, 1991 ME 14481 
<emphasis id="08FEC194975010757E6DA4A35BDB7804" bold="true" pageId="0" pageNumber="295">C</emphasis>
<taxonomicName id="7A227045ECD1B310293FCA714BE7E0A3" baseAuthorityName="Vermeulen" baseAuthorityYear="1994" class="Gastropoda" family="Diplommatinidae" genus="Plectostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectostoma anisopterum" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="anisopterum">
<emphasis id="FDDD68E798A45E4306CDF75339B47806" italics="true" pageId="0" pageNumber="295">Plectostoma anisopterum</emphasis>
</taxonomicName>
(Vermeulen, 1994) ME 14479 
<emphasis id="1DE752EF8CA5C800683308F999373BEA" bold="true" pageId="0" pageNumber="295">D</emphasis>
<taxonomicName id="87A64BE434DFFD071C762F2CD8950B8D" baseAuthorityName="E. A. Smith" baseAuthorityYear="1894" class="Gastropoda" family="Diplommatinidae" genus="Plectostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectostoma austeni" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="austeni">
<emphasis id="C57006E84754A124C1859966634C7EE5" italics="true" pageId="0" pageNumber="295">Plectostoma austeni</emphasis>
</taxonomicName>
(E. A. Smith, 1894a) ME 14478 Gua Rumbang 
<emphasis id="6A09ABE77A13AE3D571D08008E042C08" bold="true" pageId="0" pageNumber="295">E</emphasis>
<taxonomicName id="010258F8CADEC79D1C7C999ECC7D6BDE" baseAuthorityName="E. A. Smith" baseAuthorityYear="1894" class="Gastropoda" family="Diplommatinidae" genus="Plectostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectostoma pumilio" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="pumilio">
<emphasis id="316481AF9522892660F264B21730F299" italics="true" pageId="0" pageNumber="295">Plectostoma pumilio</emphasis>
</taxonomicName>
(E. A. Smith, 1894a) ME 14477.
</paragraph>
</caption>
<paragraph id="E364B59D37214DF7F275966F52575252" pageId="0" pageNumber="295">
<emphasis id="D2B609863F944C91894CE2F2CB7A12C3" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Sarawak 1st Div.: W of Kpg. Lobang Batu 12.5 km S of Tebakang&quot;.
</paragraph>
<paragraph id="840E97680533F5A9B043621E7D2638AE" pageId="0" pageNumber="295">
<emphasis id="F4CC1359F83C2BE2593A04C3B5746109" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="C5BCBDCB4510F972CE54782159DB9168" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="3B074BE93B56E6F935356C1CC23AD94A" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14480, ME 15027, MZU.MOL.22.211.
</paragraph>
<paragraph id="9913F0C65A40CE2EDA34249187C8A9E0" pageId="0" pageNumber="295">
<emphasis id="581210C75F78990E40E97B33099A94E7" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak. Also found in Sabah. Endemic to Borneo (
<bibRefCitation id="1C28F92097A4B0AAF6C9FA7DA6FFABEF" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="265430FC4EB42DDB23CFB3185B9AB6E6" pageId="0" pageNumber="295">
<emphasis id="A6370C6B4B270EBDE7A1CA114949F411" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were found foraging in leaf-litter and plant debris at the base of limestone cliffs.
</paragraph>
<paragraph id="3BDD55159F88A12FA27ED4EB03E8DA73" pageId="0" pageNumber="295">
<emphasis id="4C09C23162387ED799D00D7F7BB947ED" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="C4FD3C3CA6FAAFC585549EFC34D9355D" authorityName="Vermeulen" authorityYear="1991" class="Gastropoda" family="Diplommatinidae" genus="Opisthostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthostoma tridens" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="tridens">
<emphasis id="9D02B05DCC79A11BA2D8F668DD08569C" italics="true" pageId="0" pageNumber="295">Opisthostoma tridens</emphasis>
</taxonomicName>
Vermeulen, 1991
</emphasis>
</paragraph>
<paragraph id="94C3E285FBD3D9C8DBDE33BE24556357" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="F9686B31765364F69F6B281799636309" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A Opisthostoma brachyacrum lambii Vermeulen, 1991 ME 14480 B Opisthostoma tridens Vermeulen, 1991 ME 14481 C Plectostoma anisopterum (Vermeulen, 1994) ME 14479 D Plectostoma austeni (E. A. Smith, 1894 a) ME 14478 Gua Rumbang E Plectostoma pumilio (E. A. Smith, 1894 a) ME 14477." figureDoi="10.3897/zookeys.1198.116265.figure9" httpUri="https://binary.pensoft.net/fig/1034248" pageId="0" pageNumber="295">9B</figureCitation>
</paragraph>
<paragraph id="9B34BA992A74357A2358077AACB43BDB" pageId="0" pageNumber="295">
<emphasis id="7FD84A01C403241906052258A6F7E922" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Sarawak 1st Div.: Kpg. Beratok along road Kuching-Serian&quot;.
</paragraph>
<paragraph id="6BDEF54D3963C7B02196C2761AA6427E" pageId="0" pageNumber="295">
<emphasis id="BE09F1968A2D8822A46FC5A1D1FFC4A9" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="0424729F7914A2EF71D45164AE61A349" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="B116C3C1B6D1E62371874C8F2F8B4772" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14481, ME 15028, MZU.MOL.22.443.
</paragraph>
<paragraph id="BD88BE21298F7A0F90039D7EB76F7006" pageId="0" pageNumber="295">
<emphasis id="4041BFE08FD0AC5DDC2F6001A1A583C0" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak. Endemic to western Sarawak region (
<bibRefCitation id="3E6FB02E955F8E2CBC581E983D078633" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="C9C7314C67AE7B0817DEF3B6FA359DA0" pageId="0" pageNumber="295">
<emphasis id="8C0FD69529E5210BAF2F1F10AD318F7F" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails found on wet vertical limestone surfaces covered with mosses.
</paragraph>
<paragraph id="0BAF7CA2E9002CC8FE1765E65E5D1BE1" pageId="0" pageNumber="295">
<emphasis id="0E8C64578B17DCFA4C9B54F6C837BB0F" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="BE6B335FB85AD1013050ABF0C7C33E26" baseAuthorityName="Vermeulen" baseAuthorityYear="1994" class="Gastropoda" family="Diplommatinidae" genus="Plectostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectostoma anisopterum" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="anisopterum">
<emphasis id="D38AD1E9812B08F6836FD55E06DEC576" italics="true" pageId="0" pageNumber="295">Plectostoma anisopterum</emphasis>
</taxonomicName>
(Vermeulen, 1994)
</emphasis>
</paragraph>
<paragraph id="791D9CFAE117E9B2F0E81ADF8C50215E" pageId="0" pageNumber="295">
Figs 
<figureCitation id="A57C9232CFB2A1B2AC162346978F0A84" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A Opisthostoma brachyacrum lambii Vermeulen, 1991 ME 14480 B Opisthostoma tridens Vermeulen, 1991 ME 14481 C Plectostoma anisopterum (Vermeulen, 1994) ME 14479 D Plectostoma austeni (E. A. Smith, 1894 a) ME 14478 Gua Rumbang E Plectostoma pumilio (E. A. Smith, 1894 a) ME 14477." figureDoi="10.3897/zookeys.1198.116265.figure9" httpUri="https://binary.pensoft.net/fig/1034248" pageId="0" pageNumber="295">9C</figureCitation>
, 
<figureCitation id="6DC3B0AF51DAB70B6C9600595221B531" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Living snails from Gua Rumbang. A Opisthoporus biciliatus (Mousson, 1849) ME 14979 B Diplommatina adversa (H. Adams &amp; A. Adams, 1851) MZU. MOL. 23.140 C Diplommatina baritensis E. A. Smith, 1893 MZU. MOL. 23.137 D Plectostoma anisopterum ME 14479 (Vermeulen, 1994). All images not to scale." figureDoi="10.3897/zookeys.1198.116265.figure20" httpUri="https://binary.pensoft.net/fig/1034259" pageId="0" pageNumber="295">20D</figureCitation>
</paragraph>
<paragraph id="AF25798D5E504C36D2AB015C1E727B12" pageId="0" pageNumber="295">
<emphasis id="4E55593E38A48DC2C9A0F852BE51B2BD" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;G. Saak 1 mile W. of Begu, 24 miles S. of Kuching&quot;.
</paragraph>
<paragraph id="6FF0E10CED15031EBBA99938EE7EE8A0" pageId="0" pageNumber="295">
<emphasis id="568073CC52962942E26C78782BCF18BE" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="66BF757D6B5ABD2864CF09922C41333C" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="A8E959A702875A66388414A2DBB03124" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Apr. 2016-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14479, ME 14987, ME 15026, MZU.MOL.16.112, MZU.MOL.22.134, MZU.MOL.23.142.
</paragraph>
<paragraph id="09CEB820AFF0FEEDE45365CD1B6143CE" pageId="0" pageNumber="295">
<emphasis id="C72B49DA900DD6F83A2D3AB825963097" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities in Serian-Padawan limestone hills in western Sarawak. Endemic to western Sarawak (
<bibRefCitation id="1DCAE3861DAEF253B6A52B15C92E448A" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" pagination="75 - 191" publicationUrl="http://natuurtijdschriften.nl/record/597042" refId="B53" refString="Vermeulen, JJ, 1994. Notes on the non-marine mollusks of the island of Borneo 6. The genus Opisthostoma (Gastropoda Prosobranchia: Diplommatinidae), part 2. Basteria 58: 75 - 191, http://natuurtijdschriften.nl/record/597042" title="Notes on the non-marine mollusks of the island of Borneo 6. The genus Opisthostoma (Gastropoda Prosobranchia: Diplommatinidae), part 2." url="http://natuurtijdschriften.nl/record/597042" volume="58" year="1994">Vermeulen 1994</bibRefCitation>
).
</paragraph>
<paragraph id="6401DC13A5137A32BC71AF62ECD1F8AE" pageId="0" pageNumber="295">
<emphasis id="7394B0206DBAC66974697DE376193EF4" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails found on wet vertical limestone surfaces covered with mosses. It can also be found among boulders.
</paragraph>
<paragraph id="2935B71D3F802B1713523A3B378E4E28" pageId="0" pageNumber="295">
<emphasis id="E5F179976AAB94CE0DCCC34801F3AC1C" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="F0C9B02F8422D40D6644452CB37F970D" baseAuthorityName="E. A. Smith" baseAuthorityYear="1894" class="Gastropoda" family="Diplommatinidae" genus="Plectostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectostoma austeni" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="austeni">
<emphasis id="60A1AC99061C5EEA01003F6245D8AB08" italics="true" pageId="0" pageNumber="295">Plectostoma austeni</emphasis>
</taxonomicName>
(E. A. Smith, 1894a)
</emphasis>
</paragraph>
<paragraph id="5A1770CCEC723FFFF201EC060FE68B95" pageId="0" pageNumber="295">
Figs 
<figureCitation id="2C562C12C25FE684A8885AC0BE6440C0" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A Opisthostoma brachyacrum lambii Vermeulen, 1991 ME 14480 B Opisthostoma tridens Vermeulen, 1991 ME 14481 C Plectostoma anisopterum (Vermeulen, 1994) ME 14479 D Plectostoma austeni (E. A. Smith, 1894 a) ME 14478 Gua Rumbang E Plectostoma pumilio (E. A. Smith, 1894 a) ME 14477." figureDoi="10.3897/zookeys.1198.116265.figure9" httpUri="https://binary.pensoft.net/fig/1034248" pageId="0" pageNumber="295">9D</figureCitation>
, 
<figureCitation id="737676A2C50A2DE78921045C72240FB9" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Living snails from Gua Rumbang. A Plectostoma pumilio (E. A. Smith, 1894 a) ME 14477 B Plectostoma austeni (E. A. Smith, 1894 a) ME 14477 C Rahula raricostulata (E. A. Smith, 1893) MZU. MOL. 22.139 D Semi adult Macrochlamys infans (Reeve, 1854) MZU. MOL. 23.144. Images not to scale." figureDoi="10.3897/zookeys.1198.116265.figure21" httpUri="https://binary.pensoft.net/fig/1034260" pageId="0" pageNumber="295">21B</figureCitation>
</paragraph>
<paragraph id="66556224EAB9F6005ED4D76F35C38587" pageId="0" pageNumber="295">
<emphasis id="D2AFD5EC06F354E65866DF785B276BDF" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Rumbang, Sarawak&quot;.
</paragraph>
<paragraph id="5C8E01C8EB7BF47FE26711C02FBABCD4" pageId="0" pageNumber="295">
<emphasis id="FE1EA86D407B30FD48AE20614E22C791" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="EF3DD85984D30C1ACAD9B93BC5179FCE" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="E8CD9931B4887591109026E699933907" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Apr. 2016-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14478, ME 14986, ME 15025, MZU.MOL.22.144, MZU.MOL.16.113.
</paragraph>
<paragraph id="04680AE3ECA219A52EEBA4F41BF60174" pageId="0" pageNumber="295">
<emphasis id="A5DD37E6D20D16E716E773D12BF43382" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak. Endemic to western Sarawak region (
<bibRefCitation id="B74BF605C00F390920B855BD73F3B2D7" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="2A4B7083763C98F4F0E8ED6ADEEC552A" pageId="0" pageNumber="295">
<emphasis id="7D62956568220DCA5B23EE0473F861F6" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails found on wet vertical limestone surfaces and was observed foraging inside the rock crevices and cave walls, avoiding direct exposure to light.
</paragraph>
<paragraph id="1A6F94FFD8EF740FB530ACA2F55AA0C1" pageId="0" pageNumber="295">
<emphasis id="5FE15D3D69152046B0AF0CECB4DB1C0F" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="394260120831D45BB38020C79035F030" baseAuthorityName="E. A. Smith" baseAuthorityYear="1894" class="Gastropoda" family="Diplommatinidae" genus="Plectostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectostoma pumilio" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="pumilio">
<emphasis id="A0889F3B5CDE517A9C355FBE1A07028A" italics="true" pageId="0" pageNumber="295">Plectostoma pumilio</emphasis>
</taxonomicName>
(E. A Smith, 1894a)
</emphasis>
</paragraph>
<paragraph id="9138C26B8D44CCD52D24F83E1DC38AC6" pageId="0" pageNumber="295">
Figs 
<figureCitation id="B7A4819A751F34C5D25F2F2F9C313A24" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A Opisthostoma brachyacrum lambii Vermeulen, 1991 ME 14480 B Opisthostoma tridens Vermeulen, 1991 ME 14481 C Plectostoma anisopterum (Vermeulen, 1994) ME 14479 D Plectostoma austeni (E. A. Smith, 1894 a) ME 14478 Gua Rumbang E Plectostoma pumilio (E. A. Smith, 1894 a) ME 14477." figureDoi="10.3897/zookeys.1198.116265.figure9" httpUri="https://binary.pensoft.net/fig/1034248" pageId="0" pageNumber="295">9E</figureCitation>
, 
<figureCitation id="BBD5199C6BD6452365185F83EF23F28E" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Living snails from Gua Rumbang. A Plectostoma pumilio (E. A. Smith, 1894 a) ME 14477 B Plectostoma austeni (E. A. Smith, 1894 a) ME 14477 C Rahula raricostulata (E. A. Smith, 1893) MZU. MOL. 22.139 D Semi adult Macrochlamys infans (Reeve, 1854) MZU. MOL. 23.144. Images not to scale." figureDoi="10.3897/zookeys.1198.116265.figure21" httpUri="https://binary.pensoft.net/fig/1034260" pageId="0" pageNumber="295">21A</figureCitation>
</paragraph>
<paragraph id="3D61325FBBCBD5C07B8C59DB5E27A74C" pageId="0" pageNumber="295">
<emphasis id="94C369F583F9A96A22C0A21B621E37B2" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Rumbang, Sarawak&quot;.
</paragraph>
<paragraph id="8F7D3CDB0B4C7CD1361037474A094824" pageId="0" pageNumber="295">
<emphasis id="44C1BE9D9B7C9B87FA4460B244776EBA" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="5D24D0E4C2791EF2DDC7496AEFF0012E" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="CCF87AF24184AB632D50F87A8356B7ED" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Apr. 2016-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14477, ME 14985, ME 15024, MZU.MOL.16.114.
</paragraph>
<paragraph id="9C845C7043B86A18739935B50689FFE6" pageId="0" pageNumber="295">
<emphasis id="FF8CBA083A9DC3952DE26C726F89DCDA" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities in Serian-Padawan limestone hills in western Sarawak. Endemic to western Sarawak region (
<bibRefCitation id="FB78A357F72CCEFE1BE3FCE8C88FE058" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" pagination="75 - 191" publicationUrl="http://natuurtijdschriften.nl/record/597042" refId="B53" refString="Vermeulen, JJ, 1994. Notes on the non-marine mollusks of the island of Borneo 6. The genus Opisthostoma (Gastropoda Prosobranchia: Diplommatinidae), part 2. Basteria 58: 75 - 191, http://natuurtijdschriften.nl/record/597042" title="Notes on the non-marine mollusks of the island of Borneo 6. The genus Opisthostoma (Gastropoda Prosobranchia: Diplommatinidae), part 2." url="http://natuurtijdschriften.nl/record/597042" volume="58" year="1994">Vermeulen 1994</bibRefCitation>
).
</paragraph>
<paragraph id="779082377203608A5AE94B426A31C3F8" pageId="0" pageNumber="295">
<emphasis id="A4F52F020CA795E8F88D6C9F82BFAF89" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails found on wet vertical limestone surfaces covered with mosses.
</paragraph>
<paragraph id="FBBEF8C92AFD8D68142AC9AC384A2665" pageId="0" pageNumber="295">
<emphasis id="08E3D8010D0833DA3116B1A0A53151F9" bold="true" pageId="0" pageNumber="295">
Subclass 
<taxonomicName id="BEB92D106BB5EE3B9BF1B31963794FE4" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" pageId="0" pageNumber="295" phylum="Mollusca" rank="superOrder" superOrder="Heterobranchia">Heterobranchia</taxonomicName>
Burmeister, 1837
</emphasis>
</paragraph>
<paragraph id="379B7B5A25CE50D055520CD26CC96061" pageId="0" pageNumber="295">
<emphasis id="62A7861B48D5C4005FD9406E691410DC" bold="true" pageId="0" pageNumber="295">
Family 
<taxonomicName id="451C63623F6E4A292E9542D6C309212D" authorityName="Swainson" authorityYear="1840" class="Gastropoda" family="Achatinidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="family">Achatinidae</taxonomicName>
Swainson, 1840
</emphasis>
</paragraph>
<paragraph id="ADB22AB2446B5C77C3DC7ADB4E81E700" pageId="0" pageNumber="295">
<emphasis id="15D276CB37B41121984FF9798D3E503F" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="0F630717CD26D69B29CCD7172DE8DD80" baseAuthorityName="Potiez &amp; Michaud" baseAuthorityYear="1838" class="Gastropoda" family="Achatinidae" genus="Allopeas" kingdom="Animalia" lsidName="Allopeas clavulinum" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="clavulinum">
<emphasis id="21A00BDDEC80CE9AAEF9DE6A8EF739C4" italics="true" pageId="0" pageNumber="295">Allopeas clavulinum</emphasis>
</taxonomicName>
(Potiez &amp; Michaud, 1838)
</emphasis>
</paragraph>
<paragraph id="7FB10CFA814C34C3E5D02828FFDE04AA" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="89728D182DAAD5FC177DC732627720AC" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A Allopeas clavulinum (Potiez &amp; Michaud, 1838) ME 14485 B Allopeas gracile (T. Hutton, 1834) ME 14486 C Opeas hannense (Rang, 1831) ME 14484 D Paropeas achatinaceum (L. Pfeiffer, 1846) ME 14487." figureDoi="10.3897/zookeys.1198.116265.figure10" httpUri="https://binary.pensoft.net/fig/1034249" pageId="0" pageNumber="295">10A</figureCitation>
</paragraph>
<caption id="BC91F82AE3505420985A11F401236672" doi="10.3897/zookeys.1198.116265.figure10" httpUri="https://binary.pensoft.net/fig/1034249" pageId="0" pageNumber="295" start="Figure 10" startId="F10">
<paragraph id="2D559CF85C1CCA8984F5A7A7B828773B" pageId="0" pageNumber="295">
<emphasis id="CF26EFA3E6379D3ACCDE6B89C17A3381" bold="true" pageId="0" pageNumber="295">Figure 10.</emphasis>
<emphasis id="631591919C573D859FC3BEB61DCE8667" bold="true" pageId="0" pageNumber="295">A</emphasis>
<taxonomicName id="753DD0E648A7F6058E90373C6C97CDA1" baseAuthorityName="Potiez &amp; Michaud" baseAuthorityYear="1838" class="Gastropoda" family="Achatinidae" genus="Allopeas" kingdom="Animalia" lsidName="Allopeas clavulinum" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="clavulinum">
<emphasis id="27CD56A07FFA667BA570784902F4B5BD" italics="true" pageId="0" pageNumber="295">Allopeas clavulinum</emphasis>
</taxonomicName>
(Potiez &amp; Michaud, 1838) ME 14485 
<emphasis id="B82C81C65D30309D52409F124BE9E7D2" bold="true" pageId="0" pageNumber="295">B</emphasis>
<taxonomicName id="A244C82B690395F17E5E1A4E7BFCED10" baseAuthorityName="T. Hutton" baseAuthorityYear="1834" class="Gastropoda" family="Achatinidae" genus="Allopeas" kingdom="Animalia" lsidName="Allopeas gracile" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="gracile">
<emphasis id="71D4F81D7CB518547D4C067BED7A13B5" italics="true" pageId="0" pageNumber="295">Allopeas gracile</emphasis>
</taxonomicName>
(T. Hutton, 1834) ME 14486 
<emphasis id="8B5E5FFFFA4E179357C7D16D9F9D4EBC" bold="true" pageId="0" pageNumber="295">C</emphasis>
<taxonomicName id="4546F427BCD6734FD8831D2116C9F8A9" baseAuthorityName="Rang" baseAuthorityYear="1831" class="Gastropoda" family="Achatinidae" genus="Opeas" kingdom="Animalia" lsidName="Opeas hannense" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="hannense">
<emphasis id="60FD51B1029E863F109E2927F764B44F" italics="true" pageId="0" pageNumber="295">Opeas hannense</emphasis>
</taxonomicName>
(Rang, 1831) ME 14484 
<emphasis id="121078559CBFB0DAC6C78D0F1CB8EAAF" bold="true" pageId="0" pageNumber="295">D</emphasis>
<taxonomicName id="4C260184DA72C87E2CE252622DCC36E3" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1846" class="Gastropoda" family="Achatinidae" genus="Paropeas" kingdom="Animalia" lsidName="Paropeas achatinaceum" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="achatinaceum">
<emphasis id="EA04E22F01965FADE4D5C501473EFB34" italics="true" pageId="0" pageNumber="295">Paropeas achatinaceum</emphasis>
</taxonomicName>
(L. Pfeiffer, 1846) ME 14487.
</paragraph>
</caption>
<paragraph id="2ACA3EF14FF0AAB03124667837AB10D5" pageId="0" pageNumber="295">
<emphasis id="60C36C17849DF88C0B1EF63217E18BE7" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
<normalizedToken id="90D6B0A330D6FB86A5FCD142BB2FF2CA" originalValue="“L’ile">&quot;L'ile</normalizedToken>
Bourbon&quot; [= La 
<normalizedToken id="F0B8669C893044BED92CA8BADAC0D56C" originalValue="Réunion">Reunion</normalizedToken>
].
</paragraph>
<paragraph id="ECBFC75F90D4FD816FF16B6FCDEF18C4" pageId="0" pageNumber="295">
<emphasis id="DEFB1A5F502E2093653790DF2269A725" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="4FF621BCED4C896E9078E11B16ECAB34" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="B046393A2DF5BDD4E8A7D13D89C31DE6" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14485, ME 14990, ME 15035, MZU.MOL.22.159.
</paragraph>
<paragraph id="74B8037924977703FDBF97A3B9256ED2" pageId="0" pageNumber="295">
<emphasis id="CF1219DBD233B4994FAB8FEC6AB65F58" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed in Borneo. Circumtropical (
<bibRefCitation id="DBE13429E27D9091B98439AA0C595B95" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
; 
<bibRefCitation id="69EFFEF9FE70A63BA492DA7BE5DFD939" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
).
</paragraph>
<paragraph id="4C8833243B8C46EA8E872912E2A33678" pageId="0" pageNumber="295">
<emphasis id="BD92E70B3F99E364AB506D841903E65B" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="0011841EC845107F55DAA5D0AF8EF5FF" pageId="0" pageNumber="295">
<emphasis id="9BEE783932A82C830D575E6A4185954C" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="EC92679EA110B00851BB78677A675C33" baseAuthorityName="T. Hutton" baseAuthorityYear="1834" class="Gastropoda" family="Achatinidae" genus="Allopeas" kingdom="Animalia" lsidName="Allopeas gracile" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="gracile">
<emphasis id="FF474CC163D926D623C164D0BD804E04" italics="true" pageId="0" pageNumber="295">Allopeas gracile</emphasis>
</taxonomicName>
(T. Hutton, 1834)
</emphasis>
</paragraph>
<paragraph id="5A07184A3B8A02D1F04FFD66A31A940E" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="FDC1DE85D2BA80CE996A2F7480F52B1A" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A Allopeas clavulinum (Potiez &amp; Michaud, 1838) ME 14485 B Allopeas gracile (T. Hutton, 1834) ME 14486 C Opeas hannense (Rang, 1831) ME 14484 D Paropeas achatinaceum (L. Pfeiffer, 1846) ME 14487." figureDoi="10.3897/zookeys.1198.116265.figure10" httpUri="https://binary.pensoft.net/fig/1034249" pageId="0" pageNumber="295">10B</figureCitation>
</paragraph>
<paragraph id="D03A0B43CF63A306932CC7334DA5A5D4" pageId="0" pageNumber="295">
<emphasis id="52B7FCDC3552EFB51888E0E5E872AEAE" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Mirzapoor, India&quot;.
</paragraph>
<paragraph id="40411D2C13D44758B0AC16D40A1EAAAB" pageId="0" pageNumber="295">
<emphasis id="43D624562986FB50F4C1BBB925A4F975" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="6FBE7B0BFFD57D4CAB4CDC5BF7B98BB5" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="35E124C30B4CA5DAB7EA2612FD0DF40C" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14486, ME 14991, ME 15036, MZU.MOL.22.161.
</paragraph>
<paragraph id="CB1C3080660F45DF961131B1667AAB1C" pageId="0" pageNumber="295">
<emphasis id="893E2468E58C9545B4FE88D708A7B880" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed in Borneo. Circumtropical (
<bibRefCitation id="23408BAABCEB84AB735778FB53A1BB26" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
; 
<bibRefCitation id="7B77899904952FD2173773D6F06D6283" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
).
</paragraph>
<paragraph id="27512DEA633B2218CE4CA03C2FC7C26E" pageId="0" pageNumber="295">
<emphasis id="DA8273B60AA456C2B81E6A27EB521AD2" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="0FC69E8115A3C2B3B040E73088ADD3DC" pageId="0" pageNumber="295">
<emphasis id="8DBF3FA0FF02BDCA77C55DEC57CAF601" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="FF3741A63CA3B53E039E64BC615C889C" baseAuthorityName="Rang" baseAuthorityYear="1831" class="Gastropoda" family="Achatinidae" genus="Opeas" kingdom="Animalia" lsidName="Opeas hannense" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="hannense">
<emphasis id="D1FD58A466497651BC78CCF997153D6A" italics="true" pageId="0" pageNumber="295">Opeas hannense</emphasis>
</taxonomicName>
(Rang, 1831)
</emphasis>
</paragraph>
<paragraph id="8005B690F560D0DC911DC3B73D9E5FFE" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="3C92E2C55820705D60D8530FBDBD269C" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A Allopeas clavulinum (Potiez &amp; Michaud, 1838) ME 14485 B Allopeas gracile (T. Hutton, 1834) ME 14486 C Opeas hannense (Rang, 1831) ME 14484 D Paropeas achatinaceum (L. Pfeiffer, 1846) ME 14487." figureDoi="10.3897/zookeys.1198.116265.figure10" httpUri="https://binary.pensoft.net/fig/1034249" pageId="0" pageNumber="295">10C</figureCitation>
</paragraph>
<paragraph id="691A630F4B9945D01B82137090FA0D06" pageId="0" pageNumber="295">
<emphasis id="C6A95FCA640F0956891864640F52E04F" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Village of Hann, Cap Vert peninsula, Dakar, Senegal&quot;.
</paragraph>
<paragraph id="26B17BCF7E9F901EA0DA239AE7EDE612" pageId="0" pageNumber="295">
<emphasis id="23452FB8653D15E956F6A65CCD3E0C6A" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="211D7380464ECCAC3750FDFE53E1FE14" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="98E93A456E4381BA12425E550BE83820" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14484, ME 14989, ME 15034, MZU.MOL.22.158.
</paragraph>
<paragraph id="784541043850D6BDA0F1ED0DF9D711D1" pageId="0" pageNumber="295">
<emphasis id="4EDD2F9E1850D73F802D7025C7BD515E" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak. Also found in Sabah. Distributed from Central America to Africa and Pacific (
<bibRefCitation id="5DE2A467651E1AC835FA15E1C196BA85" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
; 
<bibRefCitation id="6BA299A0B4FE2DE026DBC4B4F87CC5B3" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
).
</paragraph>
<paragraph id="43F94AF686DC1D638C06F60CA29DB384" pageId="0" pageNumber="295">
<emphasis id="451CEA3AE625C3E0CC3D0B3CBAD28835" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="FC48B81A646654E86ECD1728B182016A" pageId="0" pageNumber="295">
<emphasis id="1EBB45FDE421553011C7D215A7FA725F" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="A6E82BC4A37164CA20A5581EFED2455E" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1846" class="Gastropoda" family="Achatinidae" genus="Paropeas" kingdom="Animalia" lsidName="Paropeas achatinaceum" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="achatinaceum">
<emphasis id="8EA26873E5945AC823C312F6F87F22D8" italics="true" pageId="0" pageNumber="295">Paropeas achatinaceum</emphasis>
</taxonomicName>
(L. Pfeiffer, 1846)
</emphasis>
</paragraph>
<paragraph id="D5EA06C38AED92436EE89AE63EE6D3B3" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="CCC6CC1FD262F19FBA3D8770DA1FD8E3" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A Allopeas clavulinum (Potiez &amp; Michaud, 1838) ME 14485 B Allopeas gracile (T. Hutton, 1834) ME 14486 C Opeas hannense (Rang, 1831) ME 14484 D Paropeas achatinaceum (L. Pfeiffer, 1846) ME 14487." figureDoi="10.3897/zookeys.1198.116265.figure10" httpUri="https://binary.pensoft.net/fig/1034249" pageId="0" pageNumber="295">10D</figureCitation>
</paragraph>
<paragraph id="CDEC22521F07288E8128AD046BCFACD2" pageId="0" pageNumber="295">
<emphasis id="03D8A28FBCAF7A80EFBCCB44F0C8BDE5" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
<normalizedToken id="BDFC7BD5605DE686DF5BB0C934B16EA8" originalValue="“Java”">&quot;Java&quot;</normalizedToken>
.
</paragraph>
<paragraph id="A42F2C437CE8E971AA124747F2640205" pageId="0" pageNumber="295">
<emphasis id="917783D6103CC39146E58C3BB9B4B9E2" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="89C0FAAE06509E894D1FCD088DEAED20" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="670479E04B32F1DAFD4C156C75F30AE0" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Apr. 2016-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14487, ME 14992, ME 15037, MZU.MOL.16.147, MZU.MOL.22.160.
</paragraph>
<paragraph id="8A63EA5044F1A74EB3B6E2A36EC1D1B0" pageId="0" pageNumber="295">
<emphasis id="C1718809C51265AC5D455B90024B9458" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed in Borneo. Distributed from South to East Asia, South-east Asia, and Pacific Islands (
<bibRefCitation id="F61B8DFB0E938F0F624EB9EE43F9695F" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
; 
<bibRefCitation id="F5E6E8349927CE27AB8E4280457CEDD3" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
).
</paragraph>
<paragraph id="75129BE8B29671640746B7DFF88DE2BB" pageId="0" pageNumber="295">
<emphasis id="2137FA8429794A78FE77D607DF4CAF62" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="235FE30AB1825E783FC58B60E0A42619" pageId="0" pageNumber="295">
<emphasis id="73CD99A1E88BF64EBAD9D33621812434" bold="true" pageId="0" pageNumber="295">
Family 
<taxonomicName id="BCD8176DD0E6236E5641D3F345062E5F" authorityName="Gulick" authorityYear="1873" class="Gastropoda" family="Achatinellidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Orthurethra" pageId="0" pageNumber="295" phylum="Mollusca" rank="family">Achatinellidae</taxonomicName>
Gulick, 1873
</emphasis>
</paragraph>
<paragraph id="47A78CCCD2EACDCE00FB12D2358DF159" pageId="0" pageNumber="295">
<emphasis id="0937AA1A7528B830A4428F1AA6D28FD5" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="0EA01EDD06A630623296003974BF95BE" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" class="Gastropoda" family="Achatinellidae" genus="Elasmias" kingdom="Animalia" lsidName="Elasmias sundanum" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="sundanum">
<emphasis id="CA530993686917F1EF2856A82B793D4A" italics="true" pageId="0" pageNumber="295">Elasmias sundanum</emphasis>
</taxonomicName>
(
<normalizedToken id="9692C92B6D2C643F6FEF2F9E844D7483" originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1897)
</emphasis>
</paragraph>
<paragraph id="CF78550BA99E46830D70D411C001D1CE" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="103DF40B80E00F69C3FD8763BEC7BF7E" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A Elasmias sundanum (Moellendorff, 1897) ME 15058 B Rahula raricostulata (E. A. Smith, 1893) MZU. MOL. 22.139 C Macrochlamys sanctijohni (Godwin-Austen, 1891) ME 14500 D Semi adult Macrochlamys infans (Reeve, 1854) ME 14501 E Vitrinula glutinosa (Metcalfe, 1852) ME 14502." figureDoi="10.3897/zookeys.1198.116265.figure11" httpUri="https://binary.pensoft.net/fig/1034250" pageId="0" pageNumber="295">11A</figureCitation>
</paragraph>
<caption id="316B632AB30D7C9BAB895F7F8A4BDCB9" doi="10.3897/zookeys.1198.116265.figure11" httpUri="https://binary.pensoft.net/fig/1034250" pageId="0" pageNumber="295" start="Figure 11" startId="F11">
<paragraph id="8B5F0255DBD001AA2127C0E07652D934" pageId="0" pageNumber="295">
<emphasis id="3E3954EA7CBD51F7D0CD1B14A4D96739" bold="true" pageId="0" pageNumber="295">Figure 11.</emphasis>
<emphasis id="432B86D12F3736B86067C420E9462CF2" bold="true" pageId="0" pageNumber="295">A</emphasis>
<taxonomicName id="2398837D6A8EAF54EE8353A534A11858" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" class="Gastropoda" family="Achatinellidae" genus="Elasmias" kingdom="Animalia" lsidName="Elasmias sundanum" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="sundanum">
<emphasis id="099E891B26E77DA4EE962F26AE405E7A" italics="true" pageId="0" pageNumber="295">Elasmias sundanum</emphasis>
</taxonomicName>
(
<normalizedToken id="2769FAE76F2BAC08A95916170AFAC210" originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1897) ME 15058 
<emphasis id="03D53157C12046DAE54C61C2E3460442" bold="true" pageId="0" pageNumber="295">B</emphasis>
<taxonomicName id="F351355C316FBF4481866D930A5011B0" baseAuthorityName="E. A. Smith" baseAuthorityYear="1893" class="Gastropoda" family="Ariophantidae" genus="Rahula" kingdom="Animalia" lsidName="Rahula raricostulata" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="raricostulata">
<emphasis id="C9DCDD94AFB2030DBCE60DC92FD36F17" italics="true" pageId="0" pageNumber="295">Rahula raricostulata</emphasis>
</taxonomicName>
(E. A. Smith, 1893) MZU.MOL.22.139 
<emphasis id="25B69CA366F8C79729F73F8166A071E3" bold="true" pageId="0" pageNumber="295">C</emphasis>
<taxonomicName id="2E2A07CF7E58B1A2939B845A96139A78" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1891" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" lsidName="Macrochlamys sanctijohni" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="sanctijohni">
<emphasis id="1F85596FCD7E4CC5CB1BF5765851220E" italics="true" pageId="0" pageNumber="295">Macrochlamys sanctijohni</emphasis>
</taxonomicName>
(Godwin-Austen, 1891) ME 14500 
<emphasis id="3960ACAA52E1B1E1DBCC2BC6A71C1F98" bold="true" pageId="0" pageNumber="295">D</emphasis>
Semi adult 
<taxonomicName id="CE0B78D0884856D0DCA7017DB16427D3" baseAuthorityName="Reeve" baseAuthorityYear="1854" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" lsidName="Macrochlamys infans" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="infans">
<emphasis id="B1755A56E654556181FE35CC22341BED" italics="true" pageId="0" pageNumber="295">Macrochlamys infans</emphasis>
</taxonomicName>
(Reeve, 1854) ME 14501 
<emphasis id="F917DBA268335DA03D486881F30A3A6F" bold="true" pageId="0" pageNumber="295">E</emphasis>
<taxonomicName id="2147C3EDE48A62AA7BA9C1220F740FBD" baseAuthorityName="Metcalfe" baseAuthorityYear="1852" class="Gastropoda" family="Ariophantidae" genus="Vitrinula" kingdom="Animalia" lsidName="Vitrinula glutinosa" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="glutinosa">
<emphasis id="F803BD63954005B550DAFEA724EF956F" italics="true" pageId="0" pageNumber="295">Vitrinula glutinosa</emphasis>
</taxonomicName>
(Metcalfe, 1852) ME 14502.
</paragraph>
</caption>
<paragraph id="BF99A0C35C4712F5B3166C1FC54989EE" pageId="0" pageNumber="295">
<emphasis id="48084D3843A5567C15001159D39E27A5" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
<normalizedToken id="B869A2989334C96380982D6B1E6F1578" originalValue="“Java”">&quot;Java&quot;</normalizedToken>
.
</paragraph>
<paragraph id="8861BA3D895CED248C3E74AEA35EE872" pageId="0" pageNumber="295">
<emphasis id="B4B63FCC665FC9A6C27F772A24481150" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="EA5465D8BB215E259C754EAC10485C2B" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="4AE8417D658A8D1534E32982EE78EA27" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14514, ME 15058, MZU.MOL.23.149.
</paragraph>
<paragraph id="972A65AE200BC3E50393F7CEF1DBCB10" pageId="0" pageNumber="295">
<emphasis id="F8EC83F2EDA46FE0E8256D6A5326479D" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak to Niah at further northern Sarawak (
<bibRefCitation id="AEC267B51D2A3492489B5F7A0DF5153C" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
). Also found in Sumatra and Java (
<bibRefCitation id="ED41432ABF9F1B4490FAB4D81A63E07A" DOI="https://doi.org/10.14203/treubia.v21i2.2661" author="van Benthem-Jutting, WSS" journalOrPublisher="Treubia" pageId="0" pageNumber="295" pagination="291 - 435" refId="B50" refString="van Benthem-Jutting, WSS, 1952. Systematic studies on the non-marine mollusca of the Indo- Australian Archipelago, III. Critical Revision of the Javanese Pulmonate Land-snails of the Familes Ellobiidae to Limacidae, with an Appendix on Helicarionidae. Treubia 21 (2): 291 - 435, DOI: https://doi.org/10.14203/treubia.v21i2.2661" title="Systematic studies on the non-marine mollusca of the Indo- Australian Archipelago, III. Critical Revision of the Javanese Pulmonate Land-snails of the Familes Ellobiidae to Limacidae, with an Appendix on Helicarionidae." url="https://doi.org/10.14203/treubia.v21i2.2661" volume="21" year="1952">van Benthem-Jutting 1952</bibRefCitation>
).
</paragraph>
<paragraph id="8FFE235401A47D357A66BD31C2EB6CEE" pageId="0" pageNumber="295">
<emphasis id="230CBEFD064EB8644B1DC5066E0C6FC9" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="4A1AAABD51C9082A9E11A449B9143D71" pageId="0" pageNumber="295">
<emphasis id="4D194CBAE6DB0216CEC36A4BFCBE1FB8" bold="true" pageId="0" pageNumber="295">
Family 
<taxonomicName id="E598DCF1DF6913CC6D17201A0D6B8A72" authorityName="Godwin-Austen" authorityYear="1883" class="Gastropoda" family="Ariophantidae" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="family">Ariophantidae</taxonomicName>
Godwin-Austen, 1883
</emphasis>
</paragraph>
<paragraph id="E0C656ACE7FCDB21569C2ACD9687CBF7" pageId="0" pageNumber="295">
<emphasis id="0F6F22EA01726F33916E2575CDD312DC" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="E918055C8C59146E20D158F874685D03" baseAuthorityName="E. A. Smith" baseAuthorityYear="1893" class="Gastropoda" family="Ariophantidae" genus="Rahula" kingdom="Animalia" lsidName="Rahula raricostulata" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="raricostulata">
<emphasis id="0F33BD3650E1850C9EE3B85094E9F51D" italics="true" pageId="0" pageNumber="295">Rahula raricostulata</emphasis>
</taxonomicName>
(E. A. Smith, 1893)
</emphasis>
</paragraph>
<paragraph id="57A18EDA0495484BFEB523083E068DD5" pageId="0" pageNumber="295">
Figs 
<figureCitation id="8A77E8B295F956753030BED7BE5A7F12" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A Elasmias sundanum (Moellendorff, 1897) ME 15058 B Rahula raricostulata (E. A. Smith, 1893) MZU. MOL. 22.139 C Macrochlamys sanctijohni (Godwin-Austen, 1891) ME 14500 D Semi adult Macrochlamys infans (Reeve, 1854) ME 14501 E Vitrinula glutinosa (Metcalfe, 1852) ME 14502." figureDoi="10.3897/zookeys.1198.116265.figure11" httpUri="https://binary.pensoft.net/fig/1034250" pageId="0" pageNumber="295">11B</figureCitation>
, 
<figureCitation id="AB71CF3368E682AC6E90597E107A124B" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Living snails from Gua Rumbang. A Plectostoma pumilio (E. A. Smith, 1894 a) ME 14477 B Plectostoma austeni (E. A. Smith, 1894 a) ME 14477 C Rahula raricostulata (E. A. Smith, 1893) MZU. MOL. 22.139 D Semi adult Macrochlamys infans (Reeve, 1854) MZU. MOL. 23.144. Images not to scale." figureDoi="10.3897/zookeys.1198.116265.figure21" httpUri="https://binary.pensoft.net/fig/1034260" pageId="0" pageNumber="295">21C</figureCitation>
</paragraph>
<paragraph id="9410374C043B61079203F80B682453C7" pageId="0" pageNumber="295">
<emphasis id="E9B341BFFB923DC1D02E78284789F0F7" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Busau or Busan, Sarawak&quot;.
</paragraph>
<paragraph id="32A5A32508EFF0B0137BA4B18019CBC4" pageId="0" pageNumber="295">
<emphasis id="4D6B291ADBD299B188BC5F0664115814" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="92E678153388AE83417A8DFF9B2AB070" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="CBDB2B01BF55559223242862EC7DEFD3" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14495, ME 15043, MZU.MOL.22.139.
</paragraph>
<paragraph id="59A1A91509DDFE8021AC8889187F9364" pageId="0" pageNumber="295">
<emphasis id="EF430F497C96E5F043282C136FDA4496" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak. Endemic to western Sarawak (
<bibRefCitation id="7983A2C661B0F4DA0798FD82945F7A49" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="F3D87838B508DC109C03A73488898B7F" pageId="0" pageNumber="295">
<emphasis id="2B77F2791BED0D640F5C6538234FD09A" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were observed foraging on leaf surface of trees at the base of limestone cliffs.
</paragraph>
<paragraph id="99E91BD443E63138B1438AF86E4011CC" pageId="0" pageNumber="295">
<emphasis id="5C28508A81CF7F92B66682A19C36D34A" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="D67E561BC12A6901533AAFBB5DDA69C1" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1891" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" lsidName="Macrochlamys sanctijohni" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="sanctijohni">
<emphasis id="FC3EEB3C7233C3240CFC1F242F4BFAB8" italics="true" pageId="0" pageNumber="295">Macrochlamys sanctijohni</emphasis>
</taxonomicName>
(Godwin-Austen, 1891)
</emphasis>
</paragraph>
<paragraph id="477BEA7CC2BFEAE98DAD7E50D0210218" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="C5A47FDB7B63FE9CA223334FCE90F5BE" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A Elasmias sundanum (Moellendorff, 1897) ME 15058 B Rahula raricostulata (E. A. Smith, 1893) MZU. MOL. 22.139 C Macrochlamys sanctijohni (Godwin-Austen, 1891) ME 14500 D Semi adult Macrochlamys infans (Reeve, 1854) ME 14501 E Vitrinula glutinosa (Metcalfe, 1852) ME 14502." figureDoi="10.3897/zookeys.1198.116265.figure11" httpUri="https://binary.pensoft.net/fig/1034250" pageId="0" pageNumber="295">11C</figureCitation>
</paragraph>
<paragraph id="606BFCD9E6BB44EF899B7D6A6AB6793F" pageId="0" pageNumber="295">
<emphasis id="E0CB2F4A42455A2BFA0D812A024D01A9" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Busan Hills&quot;.
</paragraph>
<paragraph id="A8A38D19ED62CC1E4568B0EDAA938E08" pageId="0" pageNumber="295">
<emphasis id="D4EA316E53467E1655DB472CAE8EC246" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="F6BD06DD1E426AE6BCBAA23DD61553B1" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="27242F7035D157291DA5EB74E52ECE64" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14500, ME 14998, ME 15048, MZU.MOL.22.451.
</paragraph>
<paragraph id="4F63787A705CE50FF631F1D4268B74F5" pageId="0" pageNumber="295">
<emphasis id="1CF2EC0695587DBC037969C60D3149B6" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak to Niah in northern Sarawak (
<bibRefCitation id="CBCA9D8231E10EE3E41879C8A0B3A5F1" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
). Also found in Palawan, Philippines (
<bibRefCitation id="1447B9D5C3CDFBDBE677DCB86B4E3ADE" author="Smith, EA" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="295" pagination="97 - 127" publicationUrl="https://www.biodiversitylibrary.org/page/30982902" refId="B48" refString="Smith, EA, 1895. On a collection of land-shells from Sarawak, British North Borneo, Palawan, and other neighboring islands. Proceedings of the Zoological Society of London 63 (1): 97 - 127, https://www.biodiversitylibrary.org/page/30982902" title="On a collection of land-shells from Sarawak, British North Borneo, Palawan, and other neighboring islands." url="https://www.biodiversitylibrary.org/page/30982902" volume="63" year="1895">Smith 1895</bibRefCitation>
).
</paragraph>
<paragraph id="A290FC6EA7269F687EF0AE29BC821F10" pageId="0" pageNumber="295">
<emphasis id="27F51BF32B9D06ADF3EB983A606A19E9" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="54A2FBDA7F1565FAA28C05167DC04347" pageId="0" pageNumber="295">
<emphasis id="B1EBA40674D3123EB0FD05695FB73F80" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="8CB8C0278523707C81550743A66968CE" baseAuthorityName="Reeve" baseAuthorityYear="1854" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" lsidName="Macrochlamys infans" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="infans">
<emphasis id="38FA41D3E5EF7679DA8FC346B9FB8FF2" italics="true" pageId="0" pageNumber="295">Macrochlamys infans</emphasis>
</taxonomicName>
(Reeve, 1854)
</emphasis>
</paragraph>
<paragraph id="49AE758451EEB3D49DBCA907EC253406" pageId="0" pageNumber="295">
Figs 
<figureCitation id="E3EAB66616141B06E3A5A574EFECC2CF" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A Elasmias sundanum (Moellendorff, 1897) ME 15058 B Rahula raricostulata (E. A. Smith, 1893) MZU. MOL. 22.139 C Macrochlamys sanctijohni (Godwin-Austen, 1891) ME 14500 D Semi adult Macrochlamys infans (Reeve, 1854) ME 14501 E Vitrinula glutinosa (Metcalfe, 1852) ME 14502." figureDoi="10.3897/zookeys.1198.116265.figure11" httpUri="https://binary.pensoft.net/fig/1034250" pageId="0" pageNumber="295">11D</figureCitation>
, 
<figureCitation id="6BAE8FB718DB291F1A099FA34488DA4F" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Living snails from Gua Rumbang. A Plectostoma pumilio (E. A. Smith, 1894 a) ME 14477 B Plectostoma austeni (E. A. Smith, 1894 a) ME 14477 C Rahula raricostulata (E. A. Smith, 1893) MZU. MOL. 22.139 D Semi adult Macrochlamys infans (Reeve, 1854) MZU. MOL. 23.144. Images not to scale." figureDoi="10.3897/zookeys.1198.116265.figure21" httpUri="https://binary.pensoft.net/fig/1034260" pageId="0" pageNumber="295">21D</figureCitation>
</paragraph>
<paragraph id="DEAF1E9582C4E6F30D7DCA6E54CF41E5" pageId="0" pageNumber="295">
<emphasis id="8408BD27F14135CB201BDCD1282F61B5" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
<normalizedToken id="674DA7B469CE10A81707676B2AA3DAE0" originalValue="“Borneo”">&quot;Borneo&quot;</normalizedToken>
.
</paragraph>
<paragraph id="8F0D5BD25203D12BDBE7F5FC46224464" pageId="0" pageNumber="295">
<emphasis id="45D2A17207A7D62DF8BC909740099E78" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="A4A31BFDD207FD4C22DA24C2C2548926" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="A86B0FEFB81ED83B9CDE73E52D92846D" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Apr. 2016-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14501, ME 15061, MZU.MOL.16.118, MZU.MOL.22.449, MZU.MOL.23.144.
</paragraph>
<paragraph id="EC9A82D46AE7C8E2FDCE054BD0E297FB" pageId="0" pageNumber="295">
<emphasis id="889BEACDB12E1E861D122AA8898058E1" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed in Borneo. Distributed from Sumatra to Philippines (
<bibRefCitation id="A3A21524C9769E6AF60D052BF5314E61" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
).
</paragraph>
<paragraph id="B30AC02D36E41F492AB1C932308D841E" pageId="0" pageNumber="295">
<emphasis id="771739B34BBB224226021F24313E6DF8" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Empty shells were found among leaf-litter and plant debris at the base of limestone cliffs while living snails were found foraging on leaf surfaces.
</paragraph>
<paragraph id="913371DCDAB26616E1CD66E8C59F4A4E" pageId="0" pageNumber="295">
<emphasis id="342CB50DF87D868BCB30DBAD9EA0E963" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="DD3F1BA1506AFB29F67432D72C0CB385" baseAuthorityName="Metcalfe" baseAuthorityYear="1852" class="Gastropoda" family="Ariophantidae" genus="Vitrinula" kingdom="Animalia" lsidName="Vitrinula glutinosa" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="glutinosa">
<emphasis id="57EA2B0A2DED57F91C05365011F956FF" italics="true" pageId="0" pageNumber="295">Vitrinula glutinosa</emphasis>
</taxonomicName>
(Metcalfe, 1852)
</emphasis>
</paragraph>
<paragraph id="CA7244966FCEB2619E40792FE24650D5" pageId="0" pageNumber="295">
Figs 
<figureCitation id="32C95FCF3472CD5FE3FF136AC08142C1" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A Elasmias sundanum (Moellendorff, 1897) ME 15058 B Rahula raricostulata (E. A. Smith, 1893) MZU. MOL. 22.139 C Macrochlamys sanctijohni (Godwin-Austen, 1891) ME 14500 D Semi adult Macrochlamys infans (Reeve, 1854) ME 14501 E Vitrinula glutinosa (Metcalfe, 1852) ME 14502." figureDoi="10.3897/zookeys.1198.116265.figure11" httpUri="https://binary.pensoft.net/fig/1034250" pageId="0" pageNumber="295">11E</figureCitation>
, 
<figureCitation id="4AAFBF3C13FE741644DAED08584A5B70" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Living snails from Gua Rumbang. A Juvenile Vitrinula glutinosa (Metcalfe, 1852) MZU. MOL. 23.146 B Kaliella barrakporensis (Reeve, 1852) MZU. MOL. 23.143 C Kaliella busauensis (E. A. Smith, 1895) MZU. MOL. 22.144 D Kaliella rumbangensis (E. A. Smith, 1895) MZU. MOL. 22.163. Images not to scale." figureDoi="10.3897/zookeys.1198.116265.figure22" httpUri="https://binary.pensoft.net/fig/1034261" pageId="0" pageNumber="295">22A</figureCitation>
</paragraph>
<paragraph id="BDED9C11E1FCA52CAEDE8525AD5D690B" pageId="0" pageNumber="295">
<emphasis id="E181F4D81B689CE354F1E6845E8E942B" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
<normalizedToken id="C333D10C2D89FC3791C94A7A7F1EFE49" originalValue="“Borneo”">&quot;Borneo&quot;</normalizedToken>
.
</paragraph>
<paragraph id="B6486A68861BB0A33027A630E3D202A6" pageId="0" pageNumber="295">
<emphasis id="0800D333DE718FD6B91ACEF9F65FE50A" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="28A4B5E95E1D93F2D8F9479AEB3D8DA6" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="261B45ACC501C62162C857F000A2FFC2" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14502, ME 15049, MZU.MOL.22.130, MZU.MOL.23.146.
</paragraph>
<paragraph id="AC9104EDEDA35498B1FB4A11C5E6DF76" pageId="0" pageNumber="295">
<emphasis id="5F1D752EFF6CD9252B9E338C2FD29A9B" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak to Mulu at northern Sarawak. Endemic to Borneo (
<bibRefCitation id="D28A974D400AC4D0C35A8DA2F6F9F601" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="F97B89B2794543706F7641E0D9BCC7F8" pageId="0" pageNumber="295">
<emphasis id="B35BF6730233108445E3921EABBEF973" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Empty shells were found among leaf-litter and plant debris at the base of limestone cliffs while living snails were found foraging on leaf surfaces. Individuals of this species show variability in the height of the spire and in the colour (pale to dark brown) (
<bibRefCitation id="A8593120E42D36EF85D340B5E6885705" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="A617A0F652B26281329D8CB22624321D" pageId="0" pageNumber="295">
<emphasis id="DC1AB836227F343027C02F0787695C41" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="18AEE11BE4D9B65D297A7273CACDBEDA" authorityName="Marzuki, T. S. Liew &amp; Mohd-Azlan" authorityYear="2021" class="Gastropoda" family="Ariophantidae" genus="Microcystina" kingdom="Animalia" lsidName="Microcystina arabii" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="arabii">
<emphasis id="528433DF7948ADC7FD1863FFFF33AE6D" italics="true" pageId="0" pageNumber="295">Microcystina arabii</emphasis>
</taxonomicName>
Marzuki, Liew &amp; Mohd-Azlan, 2021
</emphasis>
</paragraph>
<paragraph id="B54619EA824F5CCCCA3E1996C772DE9A" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="07526D7A069EA18F2613F8E74DE25A32" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A Microcystina arabii Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 14505 B Microcystina kilat Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 15050 C Microcystina paripari Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 15052 D Microcystina physotrochus Vermeulen, Liew &amp; Schilthuizen, 2015 ME 15060 E Microcystina vitreiformis (Moellendorff, 1897) MZU. MOL. 16.117." figureDoi="10.3897/zookeys.1198.116265.figure12" httpUri="https://binary.pensoft.net/fig/1034251" pageId="0" pageNumber="295">12A</figureCitation>
</paragraph>
<caption id="BE11CDBE89398ADAC6DC2444900606C2" doi="10.3897/zookeys.1198.116265.figure12" httpUri="https://binary.pensoft.net/fig/1034251" pageId="0" pageNumber="295" start="Figure 12" startId="F12">
<paragraph id="5E2CD4B42A457A8BF620A01D6F6DEED0" pageId="0" pageNumber="295">
<emphasis id="EC1C59EB4BA8CF823D77A7D3150B1D65" bold="true" pageId="0" pageNumber="295">Figure 12.</emphasis>
<emphasis id="1DF0F0BC1056A2D5267B0446920E29DB" bold="true" pageId="0" pageNumber="295">A</emphasis>
<taxonomicName id="F679492656E77C457A45AA39CB6A96B9" authorityName="Marzuki, T. S. Liew &amp; Mohd-Azlan" authorityYear="2021" class="Gastropoda" family="Ariophantidae" genus="Microcystina" kingdom="Animalia" lsidName="Microcystina arabii" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="arabii">
<emphasis id="C6912E520720D28A01B7903C5A439054" italics="true" pageId="0" pageNumber="295">Microcystina arabii</emphasis>
</taxonomicName>
Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 14505 
<emphasis id="4708C3550E5BCCEF8991374901CD1F7B" bold="true" pageId="0" pageNumber="295">B</emphasis>
<taxonomicName id="E51C11A4B4F3EB5B70DCEF5ECCA7478B" authorityName="Marzuki, T. S. Liew &amp; Mohd-Azlan" authorityYear="2021" class="Gastropoda" family="Ariophantidae" genus="Microcystina" kingdom="Animalia" lsidName="Microcystina kilat" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="kilat">
<emphasis id="75BBD1166E1EF1B29DD907A89995AE2B" italics="true" pageId="0" pageNumber="295">Microcystina kilat</emphasis>
</taxonomicName>
Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 15050 
<emphasis id="334FC1B23675DC3A13BBE9E46F887B57" bold="true" pageId="0" pageNumber="295">C</emphasis>
<taxonomicName id="1B2201FF372244CD2AA0B7611DB71843" authorityName="Marzuki, T. S. Liew &amp; Mohd-Azlan" authorityYear="2021" class="Gastropoda" family="Ariophantidae" genus="Microcystina" kingdom="Animalia" lsidName="Microcystina paripari" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="paripari">
<emphasis id="A470E6A39FF62D99AC5E726DE8F261B8" italics="true" pageId="0" pageNumber="295">Microcystina paripari</emphasis>
</taxonomicName>
Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 15052 
<emphasis id="2A7C03ED03DF49202E338DA944D91DCB" bold="true" pageId="0" pageNumber="295">D</emphasis>
<taxonomicName id="834F22E63F39E9E46811AFE2CDDA5AB9" authorityName="Vermeulen, Liew &amp; Schilthuizen" authorityYear="2015" class="Gastropoda" family="Ariophantidae" genus="Microcystina" kingdom="Animalia" lsidName="Microcystina physotrochus" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="physotrochus">
<emphasis id="FE29067CC49D574F16A1A8FB845B403D" italics="true" pageId="0" pageNumber="295">Microcystina physotrochus</emphasis>
</taxonomicName>
Vermeulen, Liew &amp; Schilthuizen, 2015 ME 15060 
<emphasis id="9F21C66FB976D22839CD7E804FC07211" bold="true" pageId="0" pageNumber="295">E</emphasis>
<taxonomicName id="FACD6D371505259E4D10CD884638493D" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" class="Gastropoda" family="Ariophantidae" genus="Microcystina" kingdom="Animalia" lsidName="Microcystina vitreiformis" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="vitreiformis">
<emphasis id="04EAEF67F9A078558967C918B3540640" italics="true" pageId="0" pageNumber="295">Microcystina vitreiformis</emphasis>
</taxonomicName>
(
<normalizedToken id="815191231B0EAB66CBBF52152FE3FF0A" originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1897) MZU.MOL.16.117.
</paragraph>
</caption>
<paragraph id="CAFBFB9A374AB666120CB5130DF6990D" pageId="0" pageNumber="295">
<emphasis id="6AD3957671399A6F8B699D8521D7445B" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Malaysia, Sarawak, Kuching, Division, Bukit Sokwang (Site 2), northern site of Gunung Doya, limestone hill along Skio road, 2.05 miles E Bau&quot;.
</paragraph>
<paragraph id="A67556E45FC7F0767E76CA07CB57B3A8" pageId="0" pageNumber="295">
<emphasis id="08D7E18D872B69CDEBCFF57680097D01" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="1C4487763087FEDF72CAFDC1714D5C18" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="86A7F9A91CB7460EC5AD524466A42968" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14505, MZU.MOL.23.152.
</paragraph>
<paragraph id="F085F51C6DFD494B1FB5E769B7C318FA" pageId="0" pageNumber="295">
<emphasis id="33B62CDDB189DE4F77B43A2615BE1C8D" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak. Endemic to western Sarawak (
<bibRefCitation id="1012F62C813D66CEE911553CEB365ACA" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="66FFD7299CF14246D909622656B3FBA5" pageId="0" pageNumber="295">
<emphasis id="D2AB2CA431D35B4C535A611F3E7F64DC" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="A3FC796303D549DB1D23F9E4AB73D290" pageId="0" pageNumber="295">
<emphasis id="630DDB6E5405C275C46B181A71EC7BED" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="070FF189318E46DB41910EBCEDA80242" authorityName="Marzuki, T. S. Liew &amp; Mohd-Azlan" authorityYear="2021" class="Gastropoda" family="Ariophantidae" genus="Microcystina" kingdom="Animalia" lsidName="Microcystina kilat" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="kilat">
<emphasis id="3A44221646AFF28A68DF9609E18F3961" italics="true" pageId="0" pageNumber="295">Microcystina kilat</emphasis>
</taxonomicName>
Marzuki, Liew &amp; Mohd-Azlan, 2021
</emphasis>
</paragraph>
<paragraph id="5DFA7C9BDAA6A7BE5595A34CC1C8ECEA" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="64E2C16F01CF9CA18678C5C32D137201" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A Microcystina arabii Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 14505 B Microcystina kilat Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 15050 C Microcystina paripari Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 15052 D Microcystina physotrochus Vermeulen, Liew &amp; Schilthuizen, 2015 ME 15060 E Microcystina vitreiformis (Moellendorff, 1897) MZU. MOL. 16.117." figureDoi="10.3897/zookeys.1198.116265.figure12" httpUri="https://binary.pensoft.net/fig/1034251" pageId="0" pageNumber="295">12B</figureCitation>
</paragraph>
<paragraph id="C40171B4C3848053342112C11E83BC93" pageId="0" pageNumber="295">
<emphasis id="6B4491627313CE3DA29628E027A5D717" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Malaysia, Sarawak, Kuching Division, Lobang Angin (Site 2), limestone outcrop near Sungai Sarawak Kanan, 1.75 miles W Bau&quot;.
</paragraph>
<paragraph id="769B51828BBACBC76210D9BFCC54F745" pageId="0" pageNumber="295">
<emphasis id="542ECE436D7B357C3B17796658DFB68D" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="EA2657F9DEC43E6D8760E5FE4D0FAF27" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="73D1C568D3E393829435C49AFAE6134A" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 15050, MZU.MOL.23.151.
</paragraph>
<paragraph id="D1AC6D34B201A1D59FBC607BCB329710" pageId="0" pageNumber="295">
<emphasis id="C00FAB5D6238B7D876B558FD8853C691" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak. Endemic to western Sarawak (
<bibRefCitation id="7A39A35C5138416FBE6DCFC79A992014" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="F5DC6EC798993B1020059033EBC37D57" pageId="0" pageNumber="295">
<emphasis id="DEFD9A3005EF58E6A1B6191AC58F5E69" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="5BD3386DEF0D2029682A306B9BEA8ADA" pageId="0" pageNumber="295">
<emphasis id="E2F3039DBBE6B793CB79803C574E8988" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="78124F1B6F98115FE0D617D6E86702AF" authorityName="Marzuki, T. S. Liew &amp; Mohd-Azlan" authorityYear="2021" class="Gastropoda" family="Ariophantidae" genus="Microcystina" kingdom="Animalia" lsidName="Microcystina paripari" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="paripari">
<emphasis id="4E38D705AFA493D8E6C5D8A4869330FB" italics="true" pageId="0" pageNumber="295">Microcystina paripari</emphasis>
</taxonomicName>
Marzuki, Liew &amp; Mohd-Azlan, 2021
</emphasis>
</paragraph>
<paragraph id="BEF4D13CCB3F79E4596C6062A1E7FEA3" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="08F0EA5C6C05E6143ADE313BB56ACBB5" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A Microcystina arabii Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 14505 B Microcystina kilat Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 15050 C Microcystina paripari Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 15052 D Microcystina physotrochus Vermeulen, Liew &amp; Schilthuizen, 2015 ME 15060 E Microcystina vitreiformis (Moellendorff, 1897) MZU. MOL. 16.117." figureDoi="10.3897/zookeys.1198.116265.figure12" httpUri="https://binary.pensoft.net/fig/1034251" pageId="0" pageNumber="295">12C</figureCitation>
</paragraph>
<paragraph id="6C2B9DCDD10C4AA3353D33663DA6F008" pageId="0" pageNumber="295">
<emphasis id="10F84AB4028EE8F9D9E9D740A70E035A" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Malaysia, Sarawak, Kuching Division, Fairy Cave (Site 2), south part of Gunung Kapor, 4 miles SW Bau&quot;.
</paragraph>
<paragraph id="26CC4CE853DD71ADAF52F2620C7780CC" pageId="0" pageNumber="295">
<emphasis id="D6AA0E9CB18D012A79884E0FB0F3FB1E" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="0A23E4C5DE240BD791ADE96466E2193D" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="A0DC3D833486953647983E68CCC940BB" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14503, ME 15000, ME 15052, MZU.MOL.22.132.
</paragraph>
<paragraph id="3A27BA41E1B3B36319A590CC32ADD326" pageId="0" pageNumber="295">
<emphasis id="A006C7A6F9442B7DEE3A044FD25DC29C" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak. Endemic to western Sarawak (
<bibRefCitation id="8E52B865F1CACF90A612F912099584F6" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="A087779C9B920492F8F070EFC26593A7" pageId="0" pageNumber="295">
<emphasis id="2BA211AC720F8D38A6D905E519A7B2E2" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="211B71993C4043A3AB977F03686C9136" pageId="0" pageNumber="295">
<emphasis id="90DECCC92096D56AE9C6F5BD14BE7942" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="76D9F89513CB2332DA1B44BEBD1E3A73" authorityName="Vermeulen, Liew &amp; Schilthuizen" authorityYear="2015" class="Gastropoda" family="Ariophantidae" genus="Microcystina" kingdom="Animalia" lsidName="Microcystina physotrochus" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="physotrochus">
<emphasis id="5738F8B91C5B4B6E7F089F64DF57C3FD" italics="true" pageId="0" pageNumber="295">Microcystina physotrochus</emphasis>
</taxonomicName>
Vermeulen, Liew &amp; Schilthuizen, 2015
</emphasis>
</paragraph>
<paragraph id="DDAB571559304C01E1C35C6E29CF5DC1" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="4D622FF6C5237DD23D391F4887553FF5" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A Microcystina arabii Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 14505 B Microcystina kilat Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 15050 C Microcystina paripari Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 15052 D Microcystina physotrochus Vermeulen, Liew &amp; Schilthuizen, 2015 ME 15060 E Microcystina vitreiformis (Moellendorff, 1897) MZU. MOL. 16.117." figureDoi="10.3897/zookeys.1198.116265.figure12" httpUri="https://binary.pensoft.net/fig/1034251" pageId="0" pageNumber="295">12D</figureCitation>
</paragraph>
<paragraph id="F0B6D98D30ACA46BC8BAA43F5B0EB226" pageId="0" pageNumber="295">
<emphasis id="7074FB10B1566DD71A51D5351932BD8A" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Malaysia, Sabah, Sandakan Province, Kinabatangan Valley, Batu Keruak 2, near Sukau&quot;.
</paragraph>
<paragraph id="AD125B5D5DCF530DEDC53A306CE83963" pageId="0" pageNumber="295">
<emphasis id="1562FF4325E2184BBC4C08746CC72BC9" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="BADF651438AEE4D131B8BE7B608C4BBD" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="48B987201A17D198791158B5D91D42AC" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 15060.
</paragraph>
<paragraph id="94220B640B01C2CF94D7257486FB1D8C" pageId="0" pageNumber="295">
<emphasis id="A98314F88C6B89E7E64B7EAD6D94FBCC" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak to Niah in northern Sarawak. Also found in Sabah. Endemic to Borneo (
<bibRefCitation id="7AF6DA8EBFDF587B23B9BA780243BDA5" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
; 
<bibRefCitation id="79A045CD58C72D2FF049C34633D3358D" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
).
</paragraph>
<paragraph id="52CEDF78739DE3ECDDFA0EC85C79B22E" pageId="0" pageNumber="295">
<emphasis id="B5253EED9DE32B2AE0EFB4BE7473A435" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="7E9DC1A41473E7D66B40260BA805C2AB" pageId="0" pageNumber="295">
<emphasis id="E28FD6D02EA195CD0D613057F50AE8F8" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="205ABDC346FDF867CE76E61FCCAD6AF3" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" class="Gastropoda" family="Ariophantidae" genus="Microcystina" kingdom="Animalia" lsidName="Microcystina vitreiformis" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="vitreiformis">
<emphasis id="DAA1CD7881E23F1A2E5B80B61D5B79DF" italics="true" pageId="0" pageNumber="295">Microcystina vitreiformis</emphasis>
</taxonomicName>
(
<normalizedToken id="312DF40FEF0D2F745F14521726C50EC9" originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1897)
</emphasis>
</paragraph>
<paragraph id="562758E2A1A68929B2E3E90AD8BF8D6E" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="76E7B66202A370D7B098FE6BB7CE714E" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A Microcystina arabii Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 14505 B Microcystina kilat Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 15050 C Microcystina paripari Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 15052 D Microcystina physotrochus Vermeulen, Liew &amp; Schilthuizen, 2015 ME 15060 E Microcystina vitreiformis (Moellendorff, 1897) MZU. MOL. 16.117." figureDoi="10.3897/zookeys.1198.116265.figure12" httpUri="https://binary.pensoft.net/fig/1034251" pageId="0" pageNumber="295">12E</figureCitation>
</paragraph>
<paragraph id="498EB6F97C3A288DAF9E97ED34067934" pageId="0" pageNumber="295">
<emphasis id="A522627722EC9930D8020EB90D472B68" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
<normalizedToken id="C4D71C498FE28593F343A0EA4A948456" originalValue="“Java”">&quot;Java&quot;</normalizedToken>
.
</paragraph>
<paragraph id="63351026FDD2F9424850099A78D360A3" pageId="0" pageNumber="295">
<emphasis id="2A20BDF98117490F7CC9D71E569EC24F" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="7DC8114A8B9D631C6386C2082DA2E686" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="C91E794DB5A6ABC0CF80781AD8D720B5" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Apr. 2016-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 15051, MZU.MOL.16.117.
</paragraph>
<paragraph id="193837AD6CF2194A4A69DE45835392F6" pageId="0" pageNumber="295">
<emphasis id="46D7E34AEF5E2553C3BCD708E76FB3D0" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak to Niah in northern Sarawak. Also found in Java and its adjacent islands, Indonesia (Nurinsiyah, 2021).
</paragraph>
<paragraph id="9811A2D93C32A3B12C076C9A06FA2496" pageId="0" pageNumber="295">
<emphasis id="78A56EAF396E010040EBA11388091A5A" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Only empty shells were found.
</paragraph>
<paragraph id="14FB58E2AA14C261166BC92739941FA8" pageId="0" pageNumber="295">
<emphasis id="5E39738D714BFF874827E08687AAB65E" bold="true" pageId="0" pageNumber="295">
Family 
<taxonomicName id="EFA94E5EB09D8B1D77A0BC27B6B23D28" class="Gastropoda" family="Camaenidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Sigmurethra" pageId="0" pageNumber="295" phylum="Mollusca" rank="family">Camaenidae</taxonomicName>
Pilsbry, 1895
</emphasis>
</paragraph>
<paragraph id="196CBC7A68680FA7215A082C1875C718" pageId="0" pageNumber="295">
<emphasis id="4C91DC07199C74B32903C0928478DA65" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="F6FF649C4704B5D3D6CDA545609EF427" authorityName="Fulton" authorityYear="1896" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphidromus angulatus" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="angulatus">
<emphasis id="71E13FCEBC092C23C2FECAF6AB5F8346" italics="true" pageId="0" pageNumber="295">Amphidromus angulatus</emphasis>
</taxonomicName>
Fulton, 1896
</emphasis>
</paragraph>
<paragraph id="8FEA203BA06A51B6B26E56213F3F8539" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="74805B6CA6E92AFF15D05DD1FFEE3E56" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A Amphidromus angulatus Fulton, 1896 MZU. MOL. 22.147 B Amphidromus epidemiae Wang, 2021 ME 14507 C Ganesella acris (Benson, 1859) MZU. MOL. 22.150 D Philalanka kusana (Aldrich, 1889) MZU. MOL. 16.111 E Sundacharopa argos Vermeulen &amp; Liew, 2022 ME 14511." figureDoi="10.3897/zookeys.1198.116265.figure13" httpUri="https://binary.pensoft.net/fig/1034252" pageId="0" pageNumber="295">13A</figureCitation>
</paragraph>
<caption id="D7A30608B563E6546A478A72D18B6905" doi="10.3897/zookeys.1198.116265.figure13" httpUri="https://binary.pensoft.net/fig/1034252" pageId="0" pageNumber="295" start="Figure 13" startId="F13">
<paragraph id="70837C6BC4D2808A3DF8C83B4709866D" pageId="0" pageNumber="295">
<emphasis id="6656EF632C33A1A1EEF1ABAF130FFE29" bold="true" pageId="0" pageNumber="295">Figure 13.</emphasis>
<emphasis id="55A31542DDEE77FE7A47415892E84A00" bold="true" pageId="0" pageNumber="295">A</emphasis>
<taxonomicName id="0C330AB1E2BA01D510A256A8C86D234E" authorityName="Fulton" authorityYear="1896" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphidromus angulatus" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="angulatus">
<emphasis id="8854E7BCD91CFD24D3D6A3D567EC3C55" italics="true" pageId="0" pageNumber="295">Amphidromus angulatus</emphasis>
</taxonomicName>
Fulton, 1896 MZU.MOL.22.147 
<emphasis id="837F5F249571EB4C3295ACB2418136A4" bold="true" pageId="0" pageNumber="295">B</emphasis>
<taxonomicName id="75734699269550EEDC50F7BBBC055BD8" authorityName="Y. -C. Wang" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphidromus epidemiae" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="epidemiae">
<emphasis id="CE364E173F279A493F66DEB4D9561C91" italics="true" pageId="0" pageNumber="295">Amphidromus epidemiae</emphasis>
</taxonomicName>
Wang, 2021 ME 14507 
<emphasis id="A6CA99122D65CD91C951CC03D08BACD3" bold="true" pageId="0" pageNumber="295">C</emphasis>
<taxonomicName id="29F0EC50A4F2C3FB76D7DC8C6B83D486" baseAuthorityName="Benson" baseAuthorityYear="1859" class="Gastropoda" family="Camaenidae" genus="Ganesella" kingdom="Animalia" lsidName="Ganesella acris" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="acris">
<emphasis id="AAA1717E608DF24B4EB03DF859C8C99A" italics="true" pageId="0" pageNumber="295">Ganesella acris</emphasis>
</taxonomicName>
(Benson, 1859) MZU.MOL.22.150 
<emphasis id="127F59942C23025853BFBF185F66A2FE" bold="true" pageId="0" pageNumber="295">D</emphasis>
<taxonomicName id="747DF53B4B7063A023F5F318A568FB3E" baseAuthorityName="Aldrich" baseAuthorityYear="1889" class="Gastropoda" family="Charopidae" genus="Philalanka" kingdom="Animalia" lsidName="Philalanka kusana" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="kusana">
<emphasis id="64FAD58B3CF91B5F4C71233F8774AFA1" italics="true" pageId="0" pageNumber="295">Philalanka kusana</emphasis>
</taxonomicName>
(Aldrich, 1889) MZU.MOL.16.111 
<emphasis id="8E10C64C9CDB167BCAAAF877CDAE2315" bold="true" pageId="0" pageNumber="295">E</emphasis>
<taxonomicName id="B34F1651AF107407BDB71A3B24601C97" authorityName="Vermeulen &amp; Liew" authorityYear="2022" genus="Sundacharopa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sundacharopa argos" pageId="0" pageNumber="295" rank="species" species="argos">
<emphasis id="863A56187FE1360369DA8CE43AE7F83E" italics="true" pageId="0" pageNumber="295">Sundacharopa argos</emphasis>
</taxonomicName>
Vermeulen &amp; Liew, 2022 ME 14511.
</paragraph>
</caption>
<paragraph id="330397B28C934220B5B956532E8AF3A2" pageId="0" pageNumber="295">
<emphasis id="7763256386C7D3FF2633BE6B0EF3E374" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
<normalizedToken id="28B9CFAC5DFA1636EECE6CA990942E89" originalValue="“Sarawak”">&quot;Sarawak&quot;</normalizedToken>
.
</paragraph>
<paragraph id="5439F7A86EFC8000127A3F4AB11B4382" pageId="0" pageNumber="295">
<emphasis id="4C7BF9DBD43F060BA0FDBA1C7A671CA1" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="7FA00B3CE71411EFDA3F45E0621547D6" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="2F1A605D1186EC9721C53B020BB31BAD" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; MZU.MOL.22.147.
</paragraph>
<paragraph id="D2B0A7931C7BBF471370BA0D6746C03B" pageId="0" pageNumber="295">
<emphasis id="C34E089A6B935BFB47FA645565478629" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak, to Niah, further northern Sarawak. Also found in West Kalimantan (
<bibRefCitation id="FCC2625E596B46F72A7EFC12159246C3" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="BA8DB661FF1C851F6D8D3B0B96F562A7" pageId="0" pageNumber="295">
<emphasis id="E72A5C16F5C21F7D5B4DC412A876CC11" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were observed foraging on leaf surface of trees at the base of limestone cliffs.
</paragraph>
<paragraph id="C0D8874CFBFF7B79428930DC40C86662" pageId="0" pageNumber="295">
<emphasis id="0AD1351501BAF774427B3F26CF954E75" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="D9C7C19464EF54499893A5D7AAB43B9F" authorityName="Y. -C. Wang" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphidromus epidemiae" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="epidemiae">
<emphasis id="27441460A59A4AC7EE4DA27431B747E9" italics="true" pageId="0" pageNumber="295">Amphidromus epidemiae</emphasis>
</taxonomicName>
Wang, 2021
</emphasis>
</paragraph>
<paragraph id="2D40C2B67A19151C4A2DD7C99E46AE75" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="196A83C8C3077EC4B680A0DF42E0AFA6" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A Amphidromus angulatus Fulton, 1896 MZU. MOL. 22.147 B Amphidromus epidemiae Wang, 2021 ME 14507 C Ganesella acris (Benson, 1859) MZU. MOL. 22.150 D Philalanka kusana (Aldrich, 1889) MZU. MOL. 16.111 E Sundacharopa argos Vermeulen &amp; Liew, 2022 ME 14511." figureDoi="10.3897/zookeys.1198.116265.figure13" httpUri="https://binary.pensoft.net/fig/1034252" pageId="0" pageNumber="295">13B</figureCitation>
</paragraph>
<paragraph id="18DA90BF98861ECDE02A812190B01E09" pageId="0" pageNumber="295">
<emphasis id="3D80CE1B9DC9A236200C9BB42590A792" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Sarawak, Kuching Division, Bau&quot;.
</paragraph>
<paragraph id="370138D8C426035E1A15F10E38528727" pageId="0" pageNumber="295">
<emphasis id="62C862232EC16DF4094BDD9BB721368E" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="4DAF106D9863068F8E2DF144987CF806" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="90D1BEF31B6A681B34308B404C2F1169" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; ME 14507, ME 15004, ME 15054, MZU.MOL.22.183.
</paragraph>
<paragraph id="21C98933E5F1ABDFED7321FE463EF073" pageId="0" pageNumber="295">
<emphasis id="8CD2AB52487F2AB3C7DA592FC9110AD7" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak. Endemic to western Sarawak.
</paragraph>
<paragraph id="01590F6C5079AC3573B5632A88807EF9" pageId="0" pageNumber="295">
<emphasis id="092DE7D8ED20C79BDD7494C9BF693842" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="04FF775722027EC54416928EC2628AF3" pageId="0" pageNumber="295">
<emphasis id="6774BA2D94CFF05BE183C27D949545AE" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="314344F0F2C5D7B4331EB863834DEDEB" baseAuthorityName="Benson" baseAuthorityYear="1859" class="Gastropoda" family="Camaenidae" genus="Ganesella" kingdom="Animalia" lsidName="Ganesella acris" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="acris">
<emphasis id="CD3E8A66A3322DB1198F181A1184BB58" italics="true" pageId="0" pageNumber="295">Ganesella acris</emphasis>
</taxonomicName>
(Benson, 1859)
</emphasis>
</paragraph>
<paragraph id="662B6F6FC2891FD31F454C2B79C8B678" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="DEE03AF1B150896DE393A8EBEC934348" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A Amphidromus angulatus Fulton, 1896 MZU. MOL. 22.147 B Amphidromus epidemiae Wang, 2021 ME 14507 C Ganesella acris (Benson, 1859) MZU. MOL. 22.150 D Philalanka kusana (Aldrich, 1889) MZU. MOL. 16.111 E Sundacharopa argos Vermeulen &amp; Liew, 2022 ME 14511." figureDoi="10.3897/zookeys.1198.116265.figure13" httpUri="https://binary.pensoft.net/fig/1034252" pageId="0" pageNumber="295">13C</figureCitation>
</paragraph>
<paragraph id="4B1233C1885D74C803C08D6E0D0621C8" pageId="0" pageNumber="295">
<emphasis id="7B86CBC2B1F7203FB47CD4FC15A36CEE" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Teria 
<normalizedToken id="31FBA1B08089574379A379D66A38A299" originalValue="Ghát">Ghat</normalizedToken>
montium 
<normalizedToken id="6192ED27B74DE46E2D5814ED0D931B63" originalValue="Khasiæ”">Khasiae&quot;</normalizedToken>
[= Khasi Hills, Teria Ghat, India].
</paragraph>
<paragraph id="2B52C3213DB259B1F6684B117E989918" pageId="0" pageNumber="295">
<emphasis id="C2E7BC8C0DDB9E6B5618BC3D6FB188E0" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="689783AEC819ACBE59EC89DFE9E46982" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="048577331B893CE3DA0E58A3E7B38F06" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Apr. 2016-2 Sep. 2022; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14508, MZU.MOL.22.150, MZU.MOL.16.146.
</paragraph>
<paragraph id="16C9275ABA4D1F3122888C6817D0661B" pageId="0" pageNumber="295">
<emphasis id="704DB5F86F73ED88B73978369089E079" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed with scattered localities in Borneo. Distributed from Sumatra to Java Indonesia, and South to Southeast Asian mainland (
<bibRefCitation id="7C68FF23C2D293F85C0D434584161808" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
; 
<bibRefCitation id="20078B170A6BC95BF031D53900A9EDBD" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
).
</paragraph>
<paragraph id="EAD69CCFD4092145878AA57C7BBA3D9A" pageId="0" pageNumber="295">
<emphasis id="6F08E0F0DB1C5B27072F737CC61505F1" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="1DC3E9ABA236F1BA9EB3C50090039ABA" pageId="0" pageNumber="295">
<emphasis id="8E5C3362BB78CE54FD06828EC26911C8" bold="true" pageId="0" pageNumber="295">
Family 
<taxonomicName id="F1BB48423858081D6EF2482CF30D1CF1" authorityName="F. W. Hutton" authorityYear="1884" class="Gastropoda" family="Charopidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pulmonata" pageId="0" pageNumber="295" phylum="Mollusca" rank="family">Charopidae</taxonomicName>
Hutton, 1884
</emphasis>
</paragraph>
<paragraph id="8EDB9C8D7CD3D95FB9FCE27BC964A45E" pageId="0" pageNumber="295">
<emphasis id="57F9FE25209C4D5007CD0A46DFE79B35" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="D77D088A08F97C88EEC99DD6B5FA814D" baseAuthorityName="Aldrich" baseAuthorityYear="1889" class="Gastropoda" family="Charopidae" genus="Philalanka" kingdom="Animalia" lsidName="Philalanka kusana" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="kusana">
<emphasis id="E6F7149A78B1B7A3ED22AFE3536F29E4" italics="true" pageId="0" pageNumber="295">Philalanka kusana</emphasis>
</taxonomicName>
(Aldrich, 1889)
</emphasis>
</paragraph>
<paragraph id="2526D8E60D8EF7316DBA7CFD16D908C2" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="0BE1015E274EE7D1645603D708244865" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A Amphidromus angulatus Fulton, 1896 MZU. MOL. 22.147 B Amphidromus epidemiae Wang, 2021 ME 14507 C Ganesella acris (Benson, 1859) MZU. MOL. 22.150 D Philalanka kusana (Aldrich, 1889) MZU. MOL. 16.111 E Sundacharopa argos Vermeulen &amp; Liew, 2022 ME 14511." figureDoi="10.3897/zookeys.1198.116265.figure13" httpUri="https://binary.pensoft.net/fig/1034252" pageId="0" pageNumber="295">13D</figureCitation>
</paragraph>
<paragraph id="0333DA8E4F37F4700354EE57A2B8F556" pageId="0" pageNumber="295">
<emphasis id="680BBAFEC17ECB692779DC962810138A" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Kusan and Penggiron districts in South-eastern Borneo&quot;.
</paragraph>
<paragraph id="A0ADCEBECA25D3CD6EDFEF5010F1EFBE" pageId="0" pageNumber="295">
<emphasis id="9E2CF2FF94C167D2CCEE2E5765C1F61F" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="5A69E20A2FB2C14E06560433945B22DA" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="91AE315A159C8D63E88B0E670E047C9B" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Apr. 2016-2 Sep. 2022; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14509, MZU.MOL.16.111.
</paragraph>
<paragraph id="01E80C2B573B738C5BC2E44A17BECA4D" pageId="0" pageNumber="295">
<emphasis id="7F5FE1A4B51F0355A98789AAD967CB39" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed in Borneo. Distributed from West Malaysia to Papua (
<bibRefCitation id="4B97996E40FD97EC593D4DEE78C8F4C5" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
; 
<bibRefCitation id="BB8F8C0895D94EE59A1FE808A457816D" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
).
</paragraph>
<paragraph id="83AF9141E0FCB6A1BBD8041CC463CF4F" pageId="0" pageNumber="295">
<emphasis id="CD2F5767C65BC8B326C39E566B4BF31B" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="E42A71FB5B17E4C3BF636937B9AB1D23" pageId="0" pageNumber="295">
<emphasis id="9F395F72DA23395FBA98973160236335" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="CDD15188099ED90FC207449B7966F120" authorityName="Vermeulen &amp; Liew" authorityYear="2022" genus="Sundacharopa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Sundacharopa argos" pageId="0" pageNumber="295" rank="species" species="argos">
<emphasis id="14D658037C3784664F5BD582C17D8201" italics="true" pageId="0" pageNumber="295">Sundacharopa argos</emphasis>
</taxonomicName>
Vermeulen &amp; Liew, 2022
</emphasis>
</paragraph>
<paragraph id="F9E026572F9D1864B28E78F781526891" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="6B6BA795A652C2019877373FF79CD82E" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A Amphidromus angulatus Fulton, 1896 MZU. MOL. 22.147 B Amphidromus epidemiae Wang, 2021 ME 14507 C Ganesella acris (Benson, 1859) MZU. MOL. 22.150 D Philalanka kusana (Aldrich, 1889) MZU. MOL. 16.111 E Sundacharopa argos Vermeulen &amp; Liew, 2022 ME 14511." figureDoi="10.3897/zookeys.1198.116265.figure13" httpUri="https://binary.pensoft.net/fig/1034252" pageId="0" pageNumber="295">13E</figureCitation>
</paragraph>
<paragraph id="DF663403570CD721A33C3AEB25F61F1C" pageId="0" pageNumber="295">
<emphasis id="6A18A3DC4D779D312C596D6A76D5B0C3" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Malaysia, Sabah, upper Padas River valley, Long Pa Sia&quot;.
</paragraph>
<paragraph id="BCC7EB63A299B4E11B60B743CA0867F8" pageId="0" pageNumber="295">
<emphasis id="42399A924C4250C3E52E1C0C593CACA3" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="58DD5696B32EA95ED81DB959F27E38CE" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="F626C1BEF89C5A011A051136A28DB352" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14511.
</paragraph>
<paragraph id="49F497FA568319087C2C0A2BDB33B19B" pageId="0" pageNumber="295">
<emphasis id="D26469B319C7BED3AB7BEBB6BFC06D12" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed in Borneo. Endemic to Borneo (
<bibRefCitation id="C93DE157854F7A60CA8BB44C04615DAA" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
).
</paragraph>
<paragraph id="47EEC8B8F15F7CF3161D4067B2237716" pageId="0" pageNumber="295">
<emphasis id="BFB2FD310F0E2E880FC1BDB5FB380FB7" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="3CDF9A5A386DBAD18F4A376D03B006A9" pageId="0" pageNumber="295">
<emphasis id="17E68DF56C2B44ED887876517A48CF8D" bold="true" pageId="0" pageNumber="295">
Family 
<taxonomicName id="7AD7077EB4557FFB6C192AD6C2C79364" authorityName="Thiele" authorityYear="1931" class="Gastropoda" family="Chronidae" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="family">Chronidae</taxonomicName>
Thiele, 1931
</emphasis>
</paragraph>
<paragraph id="AB0F491306FBAB056A037D94683F708E" pageId="0" pageNumber="295">
<emphasis id="C8B94C12481731B49FB70FF2B4AE7703" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="7B77CF14FFB23492C6025A1FC74ACC4D" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1853" class="Gastropoda" family="Chronidae" genus="Kaliella" kingdom="Animalia" lsidName="Kaliella barrakporensis" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="barrakporensis">
<emphasis id="0527E5138FBBECC6B7790F5C3C3414A4" italics="true" pageId="0" pageNumber="295">Kaliella barrakporensis</emphasis>
</taxonomicName>
(Reeve, 1852)
</emphasis>
</paragraph>
<paragraph id="BC1CE59F85CCE93257A7AFE77136691D" pageId="0" pageNumber="295">
Figs 
<figureCitation id="19368F1FC0461D9241C437E31453EB12" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A Kaliella barrakporensis (Reeve, 1852) ME 14489 B Kaliella busauensis (E. A. Smith, 1895) ME 14488 C Kaliella calculosa (Gould, 1852) ME 14493 D Kaliella microconus (Mousson, 1865) ME 14490." figureDoi="10.3897/zookeys.1198.116265.figure14" httpUri="https://binary.pensoft.net/fig/1034253" pageId="0" pageNumber="295">14A</figureCitation>
, 
<figureCitation id="D4FCA4BED919BF428645B8BC1E555EFA" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Living snails from Gua Rumbang. A Juvenile Vitrinula glutinosa (Metcalfe, 1852) MZU. MOL. 23.146 B Kaliella barrakporensis (Reeve, 1852) MZU. MOL. 23.143 C Kaliella busauensis (E. A. Smith, 1895) MZU. MOL. 22.144 D Kaliella rumbangensis (E. A. Smith, 1895) MZU. MOL. 22.163. Images not to scale." figureDoi="10.3897/zookeys.1198.116265.figure22" httpUri="https://binary.pensoft.net/fig/1034261" pageId="0" pageNumber="295">22B</figureCitation>
</paragraph>
<caption id="1B13E7A863F9D3EBA9D3BE7AFBECBE24" doi="10.3897/zookeys.1198.116265.figure14" httpUri="https://binary.pensoft.net/fig/1034253" pageId="0" pageNumber="295" start="Figure 14" startId="F14">
<paragraph id="656490BBC43BE35E701A05205B70A6AC" pageId="0" pageNumber="295">
<emphasis id="D59C67C8ACC489894E08DF3C8DB18254" bold="true" pageId="0" pageNumber="295">Figure 14.</emphasis>
<emphasis id="3EAFA1A4E78CF70262911AB1857E4DD3" bold="true" pageId="0" pageNumber="295">A</emphasis>
<taxonomicName id="487377B3EAE6E5E1327D88EC2B5C06B3" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1853" class="Gastropoda" family="Chronidae" genus="Kaliella" kingdom="Animalia" lsidName="Kaliella barrakporensis" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="barrakporensis">
<emphasis id="A3B57E07D139DCAFA28C269343E24E63" italics="true" pageId="0" pageNumber="295">Kaliella barrakporensis</emphasis>
</taxonomicName>
(Reeve, 1852) ME 14489 
<emphasis id="79753619274CBC7AE27C0F7616AF985A" bold="true" pageId="0" pageNumber="295">B</emphasis>
<taxonomicName id="F8570C74077A20F3EDB44468C4FBC76B" baseAuthorityName="E. A. Smith" baseAuthorityYear="1895" class="Gastropoda" family="Chronidae" genus="Kaliella" kingdom="Animalia" lsidName="Kaliella busauensis" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="busauensis">
<emphasis id="D7919CE67B6074B753FDADAF4611B2F9" italics="true" pageId="0" pageNumber="295">Kaliella busauensis</emphasis>
</taxonomicName>
(E. A. Smith, 1895) ME 14488 
<emphasis id="C7A09EACC8595637120E598C21B498E9" bold="true" pageId="0" pageNumber="295">C</emphasis>
<taxonomicName id="510E8849D2E4248B47B6F1BADB92130B" baseAuthorityName="A. Gould" baseAuthorityYear="1852" class="Gastropoda" family="Chronidae" genus="Kaliella" kingdom="Animalia" lsidName="Kaliella calculosa" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="calculosa">
<emphasis id="AAB6085863E9C112DDFE0692A6A90B01" italics="true" pageId="0" pageNumber="295">Kaliella calculosa</emphasis>
</taxonomicName>
(Gould, 1852) ME 14493 
<emphasis id="D52345605C21CDE31B21AC04D2E2153E" bold="true" pageId="0" pageNumber="295">D</emphasis>
<taxonomicName id="52BF8CBC9B212BFC9234F339582A5A03" baseAuthorityName="Mousson" baseAuthorityYear="1865" class="Gastropoda" family="Chronidae" genus="Kaliella" kingdom="Animalia" lsidName="Kaliella microconus" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="microconus">
<emphasis id="FF4A0331633951A84DED42D6EE5881A4" italics="true" pageId="0" pageNumber="295">Kaliella microconus</emphasis>
</taxonomicName>
(Mousson, 1865) ME 14490.
</paragraph>
</caption>
<paragraph id="4D25A3AB1B62407100224ACD1DB8CDF2" pageId="0" pageNumber="295">
<emphasis id="29AFBCB03C1C17ADA914D73C9DBC9C53" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Barrakpore Indiae (Bacon)&quot; [= Barrackpore, West Bengal, India].
</paragraph>
<paragraph id="BC7DCFDA0D608B2D0812ABD1B13666DF" pageId="0" pageNumber="295">
<emphasis id="39FD6A3789594937A5DE73B57786F3CD" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="5728B82606AC7F60A7AD72D8CB2270E4" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="FF5B63D30C48E8C248E34B6CA89942C2" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14489, ME 14994, ME 15039, MZU.MOL.22.148, MZU.MOL.23.143.
</paragraph>
<paragraph id="8721D327D1F5BB408774970EC3778D42" pageId="0" pageNumber="295">
<emphasis id="C770908421DBE4CB8D4EBC856997067D" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed in Borneo. Distributed from Africa and South Asia mainland to Indo-Australian archipelago and Europe (
<bibRefCitation id="CA06EAA94989AEDB7D65B9FF01B75D9A" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
; 
<bibRefCitation id="E8E40ADE48ED36935F44B42209F5C2CC" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
).
</paragraph>
<paragraph id="4308DED3225C94EBF75BD1C09256C088" pageId="0" pageNumber="295">
<emphasis id="092E71F1F4E8DED7806D73116A073622" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were observed foraging on leaf surface of trees at the base of limestone cliffs.
</paragraph>
<paragraph id="26A86E46775777D9203484AA78B363A6" pageId="0" pageNumber="295">
<emphasis id="AA6A26F48CEEB0B854144C4316C5A9C2" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="4941EB6BD7D7B05C8FCD5B42EA83FA65" baseAuthorityName="E. A. Smith" baseAuthorityYear="1895" class="Gastropoda" family="Chronidae" genus="Kaliella" kingdom="Animalia" lsidName="Kaliella busauensis" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="busauensis">
<emphasis id="EFBE2836DA8D7043A1D80B7FF8AC68D1" italics="true" pageId="0" pageNumber="295">Kaliella busauensis</emphasis>
</taxonomicName>
(E. A. Smith, 1895)
</emphasis>
</paragraph>
<paragraph id="18AA9C01E709DC6B4E421C18E385BF13" pageId="0" pageNumber="295">
Figs 
<figureCitation id="5F7657F0D7F564A7CFF63B902362759F" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A Kaliella barrakporensis (Reeve, 1852) ME 14489 B Kaliella busauensis (E. A. Smith, 1895) ME 14488 C Kaliella calculosa (Gould, 1852) ME 14493 D Kaliella microconus (Mousson, 1865) ME 14490." figureDoi="10.3897/zookeys.1198.116265.figure14" httpUri="https://binary.pensoft.net/fig/1034253" pageId="0" pageNumber="295">14B</figureCitation>
, 
<figureCitation id="61E6ECBDCF27F1028861E3D68D1C8829" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Living snails from Gua Rumbang. A Juvenile Vitrinula glutinosa (Metcalfe, 1852) MZU. MOL. 23.146 B Kaliella barrakporensis (Reeve, 1852) MZU. MOL. 23.143 C Kaliella busauensis (E. A. Smith, 1895) MZU. MOL. 22.144 D Kaliella rumbangensis (E. A. Smith, 1895) MZU. MOL. 22.163. Images not to scale." figureDoi="10.3897/zookeys.1198.116265.figure22" httpUri="https://binary.pensoft.net/fig/1034261" pageId="0" pageNumber="295">22C</figureCitation>
</paragraph>
<paragraph id="F94223373FBE221D2853A89CF17F0A1C" pageId="0" pageNumber="295">
<emphasis id="1DEF0E7B304EA3498E8F067ACCF97ADE" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Busau, Sarawak&quot;.
</paragraph>
<paragraph id="1AC583CE988346BCE7249ADC8CE45882" pageId="0" pageNumber="295">
<emphasis id="34F0F539D895A105ADF94C541595E612" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="77C799E3371DC7BAE434539B21547FC9" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="CDF30092103FC1084819970666550692" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Apr. 2016-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14488, ME 14993, ME 15038, MZU.MOL.16.115, MZU.MOL.22.144.
</paragraph>
<paragraph id="2D0FD7DD3C92C1B75893B9367F8DDBAB" pageId="0" pageNumber="295">
<emphasis id="E53BB34BB1B0EDC7C5CD4F33F3D303A4" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak. Endemic to western Sarawak.
</paragraph>
<paragraph id="4EC074EDF1EBC47431DC823287A191C5" pageId="0" pageNumber="295">
<emphasis id="5A7ACF8535FB8F6B1DAAB447705EF358" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were observed foraging on leaf surface of trees at the base of limestone cliffs. 
<bibRefCitation id="D69D49BA367E60B30C5038058E88C335" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew (2022)</bibRefCitation>
mentioned that this species is synonymous with 
<taxonomicName id="799F1CE8D12073266C90971B379A6847" lsidName="K. barrakporensis" pageId="0" pageNumber="295" rank="species" species="barrakporensis">
<emphasis id="A4E5A24314956A8E6C309591E31BDA1C" italics="true" pageId="0" pageNumber="295">K. barrakporensis</emphasis>
</taxonomicName>
. In contrast, 
<bibRefCitation id="59BCFCCDF42D15E53CDD329FE2654507" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. (2021)</bibRefCitation>
listed it as separate species. 
<taxonomicName id="503B8CABEB56C22C25AC51605E8FE0C9" baseAuthorityName="E. A. Smith" baseAuthorityYear="1895" class="Gastropoda" family="Chronidae" genus="Kaliella" kingdom="Animalia" lsidName="Kaliella busauensis" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="busauensis">
<emphasis id="01C11689301680C0EE934D00C286E838" italics="true" pageId="0" pageNumber="295">Kaliella busauensis</emphasis>
</taxonomicName>
has a higher, dark brown shell with a cancellated shell surface caused by prominent spiral grooves and oblique radial riblets.
</paragraph>
<paragraph id="436F4757BFD22869872A7620BAEFA6A5" pageId="0" pageNumber="295">
<emphasis id="34926A84E5F478C182576D63E79AA508" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="02D878DD57B101769D232D7669E68766" baseAuthorityName="A. Gould" baseAuthorityYear="1852" class="Gastropoda" family="Chronidae" genus="Kaliella" kingdom="Animalia" lsidName="Kaliella calculosa" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="calculosa">
<emphasis id="5840BB059CC3FE490A9FC2DE157EF9D3" italics="true" pageId="0" pageNumber="295">Kaliella calculosa</emphasis>
</taxonomicName>
(Gould, 1852)
</emphasis>
</paragraph>
<paragraph id="9C7AFDB2D447F64224ECE2CE1270CB99" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="5BD57F6101CB6B947A4A5FD4CA851D7A" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A Kaliella barrakporensis (Reeve, 1852) ME 14489 B Kaliella busauensis (E. A. Smith, 1895) ME 14488 C Kaliella calculosa (Gould, 1852) ME 14493 D Kaliella microconus (Mousson, 1865) ME 14490." figureDoi="10.3897/zookeys.1198.116265.figure14" httpUri="https://binary.pensoft.net/fig/1034253" pageId="0" pageNumber="295">14C</figureCitation>
</paragraph>
<paragraph id="1508D7C33FB9054856C09B86CA11971D" pageId="0" pageNumber="295">
<emphasis id="5A41EC4B237390CE41E54AE92C9FF5CF" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
<normalizedToken id="4069F4BC7438D4A312F5D12AB47F8999" originalValue="“Tahiti”">&quot;Tahiti&quot;</normalizedToken>
[= Tahiti Island, French Polynesia].
</paragraph>
<paragraph id="7FFE7F33FAE029D1EBB4815B01EF4D90" pageId="0" pageNumber="295">
<emphasis id="DA44FBBAE9EB5BEF9CEB12963C25A367" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="C338EFC60309B5D4B6E72F0D41A8DC72" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="88DD214799B4444B2E67B62A9BB6255C" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14493, ME 14506, MZU.MOL.22.450.
</paragraph>
<paragraph id="F2D89244BD68F811721A9BDA49B8C087" pageId="0" pageNumber="295">
<emphasis id="849F822FBB2973F4981780C7A33B0FA2" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed in Borneo. Distributed from India to Australia and Pacific (
<bibRefCitation id="1762E79B6220AF457040D7340024091A" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
; 
<bibRefCitation id="8FB4E870DD5874A9D1E1B6551B05E890" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
).
</paragraph>
<paragraph id="64836FDE7DA08901188BD992D742CC25" pageId="0" pageNumber="295">
<emphasis id="D2E234519F42C829E566770694CBD830" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were observed foraging on leaf surface of trees at the base of limestone cliffs.
</paragraph>
<paragraph id="FAD4EDC05D38960C421048604532DEF9" pageId="0" pageNumber="295">
<emphasis id="B20437F141E3AD8D96A019EC37AC7B14" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="B7ABEBAD3F8F33A77F6D1F4374AB466D" baseAuthorityName="Mousson" baseAuthorityYear="1865" class="Gastropoda" family="Chronidae" genus="Kaliella" kingdom="Animalia" lsidName="Kaliella microconus" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="microconus">
<emphasis id="CC9FAEA36A7652EDCB8829BDEAE30B32" italics="true" pageId="0" pageNumber="295">Kaliella microconus</emphasis>
</taxonomicName>
(Mousson, 1865)
</emphasis>
</paragraph>
<paragraph id="6330B9F566DD5D65F520A64449E592E9" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="3EE4BD1D566F5F895DC2B46035D505AA" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A Kaliella barrakporensis (Reeve, 1852) ME 14489 B Kaliella busauensis (E. A. Smith, 1895) ME 14488 C Kaliella calculosa (Gould, 1852) ME 14493 D Kaliella microconus (Mousson, 1865) ME 14490." figureDoi="10.3897/zookeys.1198.116265.figure14" httpUri="https://binary.pensoft.net/fig/1034253" pageId="0" pageNumber="295">14D</figureCitation>
</paragraph>
<paragraph id="4DB48FA497E3640D233C10F0EE9C0F94" pageId="0" pageNumber="295">
<emphasis id="110A10B352B3618E589EC93B38D22B6A" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Lomma-Lomma (Viti)&quot; [= Loma Loma, Fiji].
</paragraph>
<paragraph id="4F47EC3275D4FF2E44257A922A462F4C" pageId="0" pageNumber="295">
<emphasis id="6B28EBAB38FA14078C08B4782A8911AA" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="48AE07F830692AC1BB7B1F5A09E7E106" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="44E66FC37CB02CD8294ACB5E6A2151D7" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14490, ME 14995, ME 15040, MZU.MOL.22.168.
</paragraph>
<paragraph id="9E22501B26921C2630775A4917AA8415" pageId="0" pageNumber="295">
<emphasis id="F86E74D442E896366A3CC9D588DC6833" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed in Borneo. Distributed from South-east Asia to Australia and the Pacific Islands (
<bibRefCitation id="8A70B0E934B53CA3DF3F8D6143AA8BF4" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
; 
<bibRefCitation id="77AF7EB483647BF6E9291261A2B7FC76" author="Vermeulen, JJ" journalOrPublisher="Backhuys Publishers, Leiden, Netherlands" pageId="0" pageNumber="295" refId="B58" refString="Vermeulen, JJ, Whitten, AJ, 1998. Fauna Malesiana guide to the land snails of Bali. Backhuys Publishers, Leiden, Netherlands" title="Fauna Malesiana guide to the land snails of Bali." year="1998">Vermeulen and Whitten 1998</bibRefCitation>
).
</paragraph>
<paragraph id="6B5D83E724F78FEFD437C81856A6B3C8" pageId="0" pageNumber="295">
<emphasis id="5511F7926FA62082A6E1DE3E94677760" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were observed foraging on leaf surface of trees at the base of limestone cliffs.
</paragraph>
<paragraph id="27B6DE5B5252ED5F180AB2300515A227" pageId="0" pageNumber="295">
<emphasis id="2D354B249F79624087105D2604B3E580" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="B5EB6486261D36249F762148ADADAF91" class="Gastropoda" family="Chronidae" genus="Kaliella" kingdom="Animalia" lsidName="Kaliella rumbangensis" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="rumbangensis">
<emphasis id="6CEA7CE733BE426C2840E43D4E5464DA" italics="true" pageId="0" pageNumber="295">Kaliella rumbangensis</emphasis>
</taxonomicName>
(E. A. Smith, 1895)
</emphasis>
</paragraph>
<paragraph id="BCDB24CC2D644347346642AAE4253F12" pageId="0" pageNumber="295">
Figs 
<figureCitation id="8E6B894C704FE42B044A5CF47C9AD492" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. A Kaliella rumbangensis (E. A. Smith, 1895) ME 14491 B Kaliella scandens (Cox, 1872) ME 14492 C Exrhysota brookei (A. Adams &amp; Reeve, 1850) MZU. MOL. 22.151." figureDoi="10.3897/zookeys.1198.116265.figure15" httpUri="https://binary.pensoft.net/fig/1034254" pageId="0" pageNumber="295">15A</figureCitation>
, 
<figureCitation id="2E4366E0112327D49FDB260C1C4B7212" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Living snails from Gua Rumbang. A Juvenile Vitrinula glutinosa (Metcalfe, 1852) MZU. MOL. 23.146 B Kaliella barrakporensis (Reeve, 1852) MZU. MOL. 23.143 C Kaliella busauensis (E. A. Smith, 1895) MZU. MOL. 22.144 D Kaliella rumbangensis (E. A. Smith, 1895) MZU. MOL. 22.163. Images not to scale." figureDoi="10.3897/zookeys.1198.116265.figure22" httpUri="https://binary.pensoft.net/fig/1034261" pageId="0" pageNumber="295">22D</figureCitation>
</paragraph>
<caption id="8551EE0E9041E40746BB85B6B346287B" doi="10.3897/zookeys.1198.116265.figure15" httpUri="https://binary.pensoft.net/fig/1034254" pageId="0" pageNumber="295" start="Figure 15" startId="F15">
<paragraph id="6150D834B7369B9AD64EA4A47518E253" pageId="0" pageNumber="295">
<emphasis id="BA23DB24D4B5E89582A4C6109A4FE44C" bold="true" pageId="0" pageNumber="295">Figure 15.</emphasis>
<emphasis id="F397E719CA17E0C634F9F506D5E5CA56" bold="true" pageId="0" pageNumber="295">A</emphasis>
<taxonomicName id="47A5DCC852C530BD435AD31CAA785737" class="Gastropoda" family="Chronidae" genus="Kaliella" kingdom="Animalia" lsidName="Kaliella rumbangensis" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="rumbangensis">
<emphasis id="DDA1621CBE299FE2E7F53DC9AB90EE76" italics="true" pageId="0" pageNumber="295">Kaliella rumbangensis</emphasis>
</taxonomicName>
(E. A. Smith, 1895) ME 14491 
<emphasis id="0DB7CA3DBCEF9FA3A2D2E564AA1CBD9C" bold="true" pageId="0" pageNumber="295">B</emphasis>
<taxonomicName id="8D9B6D0F02066FE3391CC64CEC68ABA4" baseAuthorityName="Cox" baseAuthorityYear="1872" class="Gastropoda" family="Chronidae" genus="Kaliella" kingdom="Animalia" lsidName="Kaliella scandens" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="scandens">
<emphasis id="0093A89F5CAAA9BBA12DF1FC9EF1B44C" italics="true" pageId="0" pageNumber="295">Kaliella scandens</emphasis>
</taxonomicName>
(Cox, 1872) ME 14492 
<emphasis id="A96177A02539BE4393BC91E0542F0F7A" bold="true" pageId="0" pageNumber="295">C</emphasis>
<taxonomicName id="D6469238C61E7580D7753365BABB4553" baseAuthorityName="A. Adams &amp; Reeve" baseAuthorityYear="1850" class="Gastropoda" family="Chronidae" genus="Exrhysota" kingdom="Animalia" lsidName="Exrhysota brookei" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="brookei">
<emphasis id="3BDA170C8C624CF174FA37502A26C156" italics="true" pageId="0" pageNumber="295">Exrhysota brookei</emphasis>
</taxonomicName>
(A. Adams &amp; Reeve, 1850) MZU.MOL.22.151.
</paragraph>
</caption>
<paragraph id="4024E532CFC4539BD0F21D0C30932710" pageId="0" pageNumber="295">
<emphasis id="B23C5E5230B6696691D4DC70D4DB3090" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Rumbang, Sarawak and Mount Rabong&quot;.
</paragraph>
<paragraph id="F2931FFF1C4FD2C6DCF2C4CDFCE5281A" pageId="0" pageNumber="295">
<emphasis id="30776FBC557548D5E9E965EF18836EAC" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="0DB7DA124C96D65602C96BD2A8043282" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="25B0E4F14DE87B972C9D825CC9F6E742" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14491, ME 14504, ME 14996, ME 15041, MZU.MOL.22.163, MZU.MOL.22.145.
</paragraph>
<paragraph id="A27CDC4DCDDBF08FEEB395933DC8B829" pageId="0" pageNumber="295">
<emphasis id="25A7A08434D843557A7AB7F7B3A23048" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Padawan limestone hills in western Sarawak only. Endemic to western Sarawak (Smith, 1895).
</paragraph>
<paragraph id="9083FCF90F69265951B9404902B6D42F" pageId="0" pageNumber="295">
<emphasis id="C61C79438D3FCEC0323AEA8C55998680" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were observed foraging on leaf surface of trees at the base of limestone cliffs. 
<bibRefCitation id="8314F04E0E6D9A2FBF7FF5E3C10867EA" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew (2022)</bibRefCitation>
listed this as synonymous with 
<taxonomicName id="7151FA676C05AABEE5389507A699196F" lsidName="K. barrakporensis" pageId="0" pageNumber="295" rank="species" species="barrakporensis">
<emphasis id="FD941B2CC4B3C9C34D140B7B9F33DAF7" italics="true" pageId="0" pageNumber="295">K. barrakporensis</emphasis>
</taxonomicName>
. 
<taxonomicName id="0358946604A4836820CE61DCE3E62C16" class="Gastropoda" family="Chronidae" genus="Kaliella" kingdom="Animalia" lsidName="Kaliella rumbangensis" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="rumbangensis">
<emphasis id="97A3DDF7B70DBBC0B178777F06A3C40B" italics="true" pageId="0" pageNumber="295">Kaliella rumbangensis</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="1B9D540CC08701D6E93793ED50BC3C76" lsidName="K. barrakporensis" pageId="0" pageNumber="295" rank="species" species="barrakporensis">
<emphasis id="86221A09AF7291D8D78FF1860FF8C904" italics="true" pageId="0" pageNumber="295">K. barrakporensis</emphasis>
</taxonomicName>
by its smaller size and by the inconspicuous (or even absent) peripheral keel on the last whorl.
</paragraph>
<paragraph id="69503B4DE5C62CDFBFF49C3121C7F387" pageId="0" pageNumber="295">
<emphasis id="3FD50905760DDD57F0536DD6747DA854" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="31EE9138EDC3CD4446345A75C1CB3770" baseAuthorityName="Cox" baseAuthorityYear="1872" class="Gastropoda" family="Chronidae" genus="Kaliella" kingdom="Animalia" lsidName="Kaliella scandens" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="scandens">
<emphasis id="8400DB7C34454E1C9AEC36245BD5270C" italics="true" pageId="0" pageNumber="295">Kaliella scandens</emphasis>
</taxonomicName>
(Cox, 1872)
</emphasis>
</paragraph>
<paragraph id="ED76A393AA91EFD723826DA6CE1D74E0" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="8059CEA094157BCD67D07BE0E06A345D" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. A Kaliella rumbangensis (E. A. Smith, 1895) ME 14491 B Kaliella scandens (Cox, 1872) ME 14492 C Exrhysota brookei (A. Adams &amp; Reeve, 1850) MZU. MOL. 22.151." figureDoi="10.3897/zookeys.1198.116265.figure15" httpUri="https://binary.pensoft.net/fig/1034254" pageId="0" pageNumber="295">15B</figureCitation>
</paragraph>
<paragraph id="14DDDFD41C67C62E39F24D13D8A1930F" pageId="0" pageNumber="295">
<emphasis id="8A641CEE1D7BD7099B7ED1FDC89DA4EC" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Port Macquarie, east coast of Australia&quot;.
</paragraph>
<paragraph id="E29E8143D9C329DCAFB928DE92D6B4A1" pageId="0" pageNumber="295">
<emphasis id="9B8973591C8C9A3F26385172DA55218D" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="71D4ED49834A23D854B110734D6BA82E" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="FD2A06CB4A2A9DCD13F65A8BAAE53A16" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14492, ME 14997, ME 15042, MZU.MOL.22.444.
</paragraph>
<paragraph id="B31B132418C9BE24A60D60D535AB83E8" pageId="0" pageNumber="295">
<emphasis id="605E963DDC766C57A9FEED5CCD62283A" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed in Borneo. Distributed from South-east Asia to Australia and the Pacific Islands (
<bibRefCitation id="B11B3BE7A550B707A3A9BF2622F0798C" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
; 
<bibRefCitation id="68B10B91931E220C48237348D32643E1" DOI="https://doi.org/10.3897/zookeys.531.6097" author="Vermeulen, JJ" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 139" refId="B60" refString="Vermeulen, JJ, Liew, TS, Schilthuizen, M, 2015. Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species. ZooKeys 531: 1 - 139, DOI: https://doi.org/10.3897/zookeys.531.6097" title="Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species." url="https://doi.org/10.3897/zookeys.531.6097" volume="531" year="2015">Vermeulen et al. 2015</bibRefCitation>
).
</paragraph>
<paragraph id="0FD5AFFCE5FBDE8A40E00BBDFBB3678F" pageId="0" pageNumber="295">
<emphasis id="DC8D1AB62566E89C4B41AC19FCC8A3B0" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were observed foraging on leaf surface of trees at the base of limestone cliffs.
</paragraph>
<paragraph id="8CFE6542D30CFA60D9EA5CD5A79BB772" pageId="0" pageNumber="295">
<emphasis id="34B287452A033F8C480E40AA6C3406AE" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="DE547E8326E37C1F100092B460D5F536" baseAuthorityName="A. Adams &amp; Reeve" baseAuthorityYear="1850" class="Gastropoda" family="Chronidae" genus="Exrhysota" kingdom="Animalia" lsidName="Exrhysota brookei" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="brookei">
<emphasis id="8739435B08732904AE52D5C04A1E664A" italics="true" pageId="0" pageNumber="295">Exrhysota brookei</emphasis>
</taxonomicName>
(A. Adams &amp; Reeve, 1850)
</emphasis>
</paragraph>
<paragraph id="1A9AEC09430FD3548B9E5869BBB3E9E2" pageId="0" pageNumber="295">
Figs 
<figureCitation id="13A1C33F3F51FC627A53A5B9F42CBE41" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. A Kaliella rumbangensis (E. A. Smith, 1895) ME 14491 B Kaliella scandens (Cox, 1872) ME 14492 C Exrhysota brookei (A. Adams &amp; Reeve, 1850) MZU. MOL. 22.151." figureDoi="10.3897/zookeys.1198.116265.figure15" httpUri="https://binary.pensoft.net/fig/1034254" pageId="0" pageNumber="295">15C</figureCitation>
, 
<figureCitation id="E1FAFDBB6204A2F34487C0F4C778F6D8" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Living snail from Gua Rumbang. Exrhysota brookei (A. Adams &amp; Reeve, 1850) MZU. MOL. 22.151." figureDoi="10.3897/zookeys.1198.116265.figure23" httpUri="https://binary.pensoft.net/fig/1034262" pageId="0" pageNumber="295">23</figureCitation>
</paragraph>
<paragraph id="8814B96C3942B6BB403123F11CB8CDC8" pageId="0" pageNumber="295">
<emphasis id="FD243095BC62E0E7F0A5C8CA446DD15A" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Mountains of Borneo&quot;.
</paragraph>
<paragraph id="638A8F6B6AA063E9A0F0A3E07B683D94" pageId="0" pageNumber="295">
<emphasis id="7476246EE8B5AF7771D8FB06C95AA10F" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="EB07567479B6DB21E62D779493A72BCE" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="EE74D6C03060D3EF0AAFC4211C1BB5F9" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; MZU.MOL.22.151.
</paragraph>
<paragraph id="8807B136C7EFEC8B3CB8A57A145BE3B8" pageId="0" pageNumber="295">
<emphasis id="7DBB7A9138C5602A0B3EB25029328A57" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widespread in Borneo. Endemic to Borneo (
<bibRefCitation id="885D9EDD91BAE4848376C85C3A5D8839" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="72BBDA6D9CA78A15AA7AA24F7CC93A02" pageId="0" pageNumber="295">
<emphasis id="8B9C72ABD460B31F10678D98EF7313DD" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were found foraging in limestone crevices and in leaf litter. This is the largest native land snail species in Borneo.
</paragraph>
<paragraph id="66DE12658936C0825A5551E2575DC028" pageId="0" pageNumber="295">
<emphasis id="042072F1753ADB54254A78C9C4F2EAE4" bold="true" pageId="0" pageNumber="295">
Family 
<taxonomicName id="EBAE2678B9FF4B67F375EE503E38BB56" authorityName="Panha &amp; Naggs" authorityYear="2010" class="Gastropoda" family="Diapheridae" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="family">Diapheridae</taxonomicName>
Panha &amp; Naggs, 2010
</emphasis>
</paragraph>
<paragraph id="820618EA8B6046243F111D60D4FE9C2C" pageId="0" pageNumber="295">
<emphasis id="057AB252C93BE11A8784840EBE97190F" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="0422B216CACB960B2CB0C854FCB3C227" authorityName="Laidlaw" authorityYear="1950" class="Gastropoda" family="Diapheridae" genus="Platycochlium" kingdom="Animalia" lsidName="Platycochlium sarawakense" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="sarawakense">
<emphasis id="AE362A51894A7C5069DA9549FD47577D" italics="true" pageId="0" pageNumber="295">Platycochlium sarawakense</emphasis>
</taxonomicName>
Laidlaw, 1950
</emphasis>
</paragraph>
<paragraph id="D51A529CD8C375DA12E7305C4052CE9B" pageId="0" pageNumber="295">
Figs 
<figureCitation id="5E1F30327FF544B73AAAC8ED1709D1AC" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. A Platycochlium sarawakense Laidlaw, 1950 ME 14483 B Dyakia subdebilis E. A. Smith, 1895 ME 14496 C Rhinocochlis nasuta (Metcalfe, 1852) ME 14493 D Juvenile Geotrochus conicoides (Metcalfe, 1852) ME 15057." figureDoi="10.3897/zookeys.1198.116265.figure16" httpUri="https://binary.pensoft.net/fig/1034255" pageId="0" pageNumber="295">16A</figureCitation>
, 
<figureCitation id="56398351D85B05F3BADA19952CE52873" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Living snails from Gua Rumbang. A Platycochlium sarawakense Laidlaw, 1950 MZU. MOL. 22.153 B Dyakia subdebilis E. A. Smith, 1895 MZU. MOL. 23.136 C Helicarion dyakanum (Godwin-Austen, 1891) ME 14499 D Paralaoma sarawakensis Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 14510. All imaged not to scale." figureDoi="10.3897/zookeys.1198.116265.figure24" httpUri="https://binary.pensoft.net/fig/1034263" pageId="0" pageNumber="295">24A</figureCitation>
</paragraph>
<caption id="7BF3BCFD7DAFBB27C391E5A47A16E165" doi="10.3897/zookeys.1198.116265.figure16" httpUri="https://binary.pensoft.net/fig/1034255" pageId="0" pageNumber="295" start="Figure 16" startId="F16">
<paragraph id="3B58259975980277EE25CBC56B8AF613" pageId="0" pageNumber="295">
<emphasis id="283CB432ABD82F5A8991FDEFEF9B605B" bold="true" pageId="0" pageNumber="295">Figure 16.</emphasis>
<emphasis id="D82EBDE8DC35193EDEB5AFF3D48E2E47" bold="true" pageId="0" pageNumber="295">A</emphasis>
<taxonomicName id="2E13ADBFF6588CDD541155F6915269B9" authorityName="Laidlaw" authorityYear="1950" class="Gastropoda" family="Diapheridae" genus="Platycochlium" kingdom="Animalia" lsidName="Platycochlium sarawakense" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="sarawakense">
<emphasis id="C7C5B8F2D8EB482535FD25E2F95A6212" italics="true" pageId="0" pageNumber="295">Platycochlium sarawakense</emphasis>
</taxonomicName>
Laidlaw, 1950 ME 14483 
<emphasis id="6DF02642B125B05C1AB57E308CF4E580" bold="true" pageId="0" pageNumber="295">B</emphasis>
<taxonomicName id="7EB2A77F0CD19A417951645E48C14CAE" authorityName="E. A. Smith" authorityYear="1895" class="Gastropoda" family="Dyakiidae" genus="Dyakia" kingdom="Animalia" lsidName="Dyakia subdebilis" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="subdebilis">
<emphasis id="12767E7091071D9A606E93A23D5A242C" italics="true" pageId="0" pageNumber="295">Dyakia subdebilis</emphasis>
</taxonomicName>
E. A. Smith, 1895 ME 14496 
<emphasis id="54A4858697FEE7691603A238B7B5AA9E" bold="true" pageId="0" pageNumber="295">C</emphasis>
<taxonomicName id="3D6689E1E7F282697A01259616C7D2F7" baseAuthorityName="Metcalfe" baseAuthorityYear="1852" class="Gastropoda" family="Dyakiidae" genus="Rhinocochlis" kingdom="Animalia" lsidName="Rhinocochlis nasuta" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="nasuta">
<emphasis id="1E093C925571672F16F40E5BA0E91149" italics="true" pageId="0" pageNumber="295">Rhinocochlis nasuta</emphasis>
</taxonomicName>
(Metcalfe, 1852) ME 14493 
<emphasis id="68D0AFDC43536DC785AEF2E0F125E9A4" bold="true" pageId="0" pageNumber="295">D</emphasis>
Juvenile 
<taxonomicName id="4F964378E12C24B83F55EB3F4AEAD547" baseAuthorityName="Metcalfe" baseAuthorityYear="1852" class="Gastropoda" family="Trochomorphidae" genus="Geotrochus" kingdom="Animalia" lsidName="Geotrochus conicoides" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="conicoides">
<emphasis id="D614CE5E1E622BD5C34A6690D854B1A9" italics="true" pageId="0" pageNumber="295">Geotrochus conicoides</emphasis>
</taxonomicName>
(Metcalfe, 1852) ME 15057.
</paragraph>
</caption>
<paragraph id="8BC7864BEC5589DCF6B798E68DA22134" pageId="0" pageNumber="295">
<emphasis id="A275EFD44D084C248811022C4565517D" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Gunong Kapor, Bau District, Sarawak&quot;.
</paragraph>
<paragraph id="8CF34FA808382D27C020708BBA93EA0B" pageId="0" pageNumber="295">
<emphasis id="77A5B2CEDC9402FDD91FB238E5710852" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="40548F005AA99F4EF89236B38F574B96" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="D93F516ADB1D226D4FA500E01948A1F7" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14483, ME 14988, ME 15033, MZU.MOL.22.153.
</paragraph>
<paragraph id="FB502CCC5BF1F7DF37BE1CDA6A8CCF6F" pageId="0" pageNumber="295">
<emphasis id="04E97633717D1A762CE12EB2B97BE705" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak. Endemic to western Sarawak (
<bibRefCitation id="D1E25A1D5B17DED85B4D206BFBB7F576" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="9B5A37CBAD299D40D09494E520D52E5D" pageId="0" pageNumber="295">
<emphasis id="C5577279A5A4E7B3D05A762BAEE6E644" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails found foraging among leaf-litter and plant debris at the base of limestone cliffs.
</paragraph>
<paragraph id="FEC3162EDF1586D451A0DFB1CD5750B2" pageId="0" pageNumber="295">
<emphasis id="014263B8549CFB3EE8114CEF2135929D" bold="true" pageId="0" pageNumber="295">
Family 
<taxonomicName id="FF1007BB2A10AF0EA030EC44A7AF1DE6" authorityName="Gude &amp; B. B. Woodward" authorityYear="1921" class="Gastropoda" family="Dyakiidae" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="family">Dyakiidae</taxonomicName>
Gude &amp; B. B. Woodward, 1921
</emphasis>
</paragraph>
<paragraph id="B49DEBC43699D3217816A174667C36AD" pageId="0" pageNumber="295">
<emphasis id="084C7CBA0A85243035CCCA85CA4B9F54" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="F1592E758C7F97A1E008DA38A854325E" authorityName="E. A. Smith" authorityYear="1895" class="Gastropoda" family="Dyakiidae" genus="Dyakia" kingdom="Animalia" lsidName="Dyakia subdebilis" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="subdebilis">
<emphasis id="E14D931B867B853D8360DBEB736AA341" italics="true" pageId="0" pageNumber="295">Dyakia subdebilis</emphasis>
</taxonomicName>
E. A. Smith, 1895
</emphasis>
</paragraph>
<paragraph id="F424A8E6CFE67AF6484B3070DDC05BD9" pageId="0" pageNumber="295">
Figs 
<figureCitation id="985F893FB76555BEFF9C1F4556894A1A" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. A Platycochlium sarawakense Laidlaw, 1950 ME 14483 B Dyakia subdebilis E. A. Smith, 1895 ME 14496 C Rhinocochlis nasuta (Metcalfe, 1852) ME 14493 D Juvenile Geotrochus conicoides (Metcalfe, 1852) ME 15057." figureDoi="10.3897/zookeys.1198.116265.figure16" httpUri="https://binary.pensoft.net/fig/1034255" pageId="0" pageNumber="295">16B</figureCitation>
, 
<figureCitation id="586B9FCF38CB0C243326B074929F98EA" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Living snails from Gua Rumbang. A Platycochlium sarawakense Laidlaw, 1950 MZU. MOL. 22.153 B Dyakia subdebilis E. A. Smith, 1895 MZU. MOL. 23.136 C Helicarion dyakanum (Godwin-Austen, 1891) ME 14499 D Paralaoma sarawakensis Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 14510. All imaged not to scale." figureDoi="10.3897/zookeys.1198.116265.figure24" httpUri="https://binary.pensoft.net/fig/1034263" pageId="0" pageNumber="295">24B</figureCitation>
</paragraph>
<paragraph id="A9FD08BC56F4095B66DCC42C85C58C19" pageId="0" pageNumber="295">
<emphasis id="10DC1340B7740572D665A8FA181D76A1" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
<normalizedToken id="CF932AE6D7040D37729AE5DAA7F074A0" originalValue="“Sarawak”">&quot;Sarawak&quot;</normalizedToken>
.
</paragraph>
<paragraph id="62F759750B09275B32BBB3941BEAE548" pageId="0" pageNumber="295">
<emphasis id="8608F5D42E9CF10ECAE87B4B90B85013" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="A73525BF80D3B3F21711DF6CCE7976E5" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="9296A0BCA6CD5586A9B0F97FD96DA402" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14496, ME 15044, MZU.MOL.22.141, MZU.MOL.23.136.
</paragraph>
<paragraph id="94592C999D6832F3A7787C7E34B3F01D" pageId="0" pageNumber="295">
<emphasis id="A3F8F6C4F4C6C4DE2CA16AA00E2EC023" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak. Endemic to western Sarawak (
<bibRefCitation id="47C8BADB5B93C430FCC97CA19C869176" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="9A868DD7C73A93CCD44AA7718036308B" pageId="0" pageNumber="295">
<emphasis id="388AADCBE19E8AB08DADA806C49B3F7B" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were found foraging on leaf surface of trees at the base of limestone cliffs and on limestone surfaces covered with mosses and lichens.
</paragraph>
<paragraph id="181E12B50378E5C8E1794FF18BBF857C" pageId="0" pageNumber="295">
<emphasis id="ADB589124A197688387693B1C76B50CC" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="EAD13541F905CDCA469785EF8AE6F4A1" baseAuthorityName="Metcalfe" baseAuthorityYear="1852" class="Gastropoda" family="Dyakiidae" genus="Rhinocochlis" kingdom="Animalia" lsidName="Rhinocochlis nasuta" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="nasuta">
<emphasis id="A4FA9DC5D47DC3A83DE93A8C90FA4593" italics="true" pageId="0" pageNumber="295">Rhinocochlis nasuta</emphasis>
</taxonomicName>
(Metcalfe, 1852)
</emphasis>
</paragraph>
<paragraph id="A4DE4C7FAA63DF25963EE10C19F9E61F" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="1F6A0715C81B193D5B3E2E2FC4C1C743" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. A Platycochlium sarawakense Laidlaw, 1950 ME 14483 B Dyakia subdebilis E. A. Smith, 1895 ME 14496 C Rhinocochlis nasuta (Metcalfe, 1852) ME 14493 D Juvenile Geotrochus conicoides (Metcalfe, 1852) ME 15057." figureDoi="10.3897/zookeys.1198.116265.figure16" httpUri="https://binary.pensoft.net/fig/1034255" pageId="0" pageNumber="295">16C</figureCitation>
</paragraph>
<paragraph id="956EDF4798071B8F7BF2C5BCAD020AB8" pageId="0" pageNumber="295">
<emphasis id="43D346EE5AD57111537ED25BEBB0C26E" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
<normalizedToken id="1C642BC0C9925334DA9F63B1DE4D0674" originalValue="“Borneo”">&quot;Borneo&quot;</normalizedToken>
.
</paragraph>
<paragraph id="4274CAF656689674B230485CD252D849" pageId="0" pageNumber="295">
<emphasis id="21DA8635C267E8D2CD281F93868896EF" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="69068AA16C14B1CAE75F3590BF081EC9" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="30286186F169E12021CC78B4BC875CAC" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14497, ME 15002, ME 15045, MZU.MOL.22.146.
</paragraph>
<paragraph id="746626F5233677D0CA657221104E1E1F" pageId="0" pageNumber="295">
<emphasis id="B09E622C6EF1C3704075609A4CF1B53C" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed in Borneo. Endemic to Borneo (
<bibRefCitation id="E256E6834EEF7B131060B204ADE1DC25" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="EB7A664AD237C15021BB34554339B838" pageId="0" pageNumber="295">
<emphasis id="84C17537626DC2C90E471C7D2EB44CAF" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were observed foraging on leaf surface of trees at the base of limestone cliffs.
</paragraph>
<paragraph id="7C04641CAF880A5521573998DE69A30A" pageId="0" pageNumber="295">
<emphasis id="C02E97F7F1B71639D288D1C7E8D90A67" bold="true" pageId="0" pageNumber="295">
Family 
<taxonomicName id="8635B48693F8A84A13FCA0F00A5D54DE" authorityName="Schileyko" authorityYear="2002" class="Gastropoda" family="Geotrochidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="family">Geotrochidae</taxonomicName>
Schileyko, 2002
</emphasis>
</paragraph>
<paragraph id="374135973A18BB686B8488E077CD5F79" pageId="0" pageNumber="295">
<emphasis id="D136EAC3BA39AE210EDA83C251FA6347" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="B93C98190468675609EBF90B8D285CCA" baseAuthorityName="Metcalfe" baseAuthorityYear="1852" class="Gastropoda" family="Trochomorphidae" genus="Geotrochus" kingdom="Animalia" lsidName="Geotrochus conicoides" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="conicoides">
<emphasis id="FC0B6D348A0B75EADB9044533B8836F7" italics="true" pageId="0" pageNumber="295">Geotrochus conicoides</emphasis>
</taxonomicName>
(Metcalfe, 1852)
</emphasis>
</paragraph>
<paragraph id="95C135A2722704EBAB21648D63D7D856" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="253D2B59481D79CC36F284571504810E" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. A Platycochlium sarawakense Laidlaw, 1950 ME 14483 B Dyakia subdebilis E. A. Smith, 1895 ME 14496 C Rhinocochlis nasuta (Metcalfe, 1852) ME 14493 D Juvenile Geotrochus conicoides (Metcalfe, 1852) ME 15057." figureDoi="10.3897/zookeys.1198.116265.figure16" httpUri="https://binary.pensoft.net/fig/1034255" pageId="0" pageNumber="295">16D</figureCitation>
</paragraph>
<paragraph id="F96722FDBF060AAF7E1760076941A9DB" pageId="0" pageNumber="295">
<emphasis id="8C3B7E0E2D74928AFA30C595F2FFAB06" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
<normalizedToken id="E6D6972CF409BC0635DF9D0E7374C9B2" originalValue="“Borneo”">&quot;Borneo&quot;</normalizedToken>
.
</paragraph>
<paragraph id="FF59140C8D50B4B53495E7C607E9EFF3" pageId="0" pageNumber="295">
<emphasis id="239BF659557C3222179018F5527B87C6" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="F560764EEEE113F9C29D5B623AF03338" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="EAFA9D6540CCA5A3D7EF7CE2350FB432" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 15057, MZU.MOL.23.150
</paragraph>
<paragraph id="4B33A33C58B84E770E117EA2215DBD10" pageId="0" pageNumber="295">
<emphasis id="A9B7C0FA52999E8D3B5419C58836883C" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed in Borneo. Widespread. Also found in Sumatra, Indonesia and Palawan, Philippines (
<bibRefCitation id="BF0E788BCD4CBA328323046B2F502468" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
; 
<bibRefCitation id="F413685B56192418B632FFE0AB4EE71A" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
).
</paragraph>
<paragraph id="61240A666ABCD8E4A8E9A3D6B612FD37" pageId="0" pageNumber="295">
<emphasis id="495251A906D54BC813965A703A6EA668" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="2D105D96EACDCC9CF9AE1F6ADC1E2CB0" pageId="0" pageNumber="295">
<emphasis id="9F3AC5355A25D6FA478CBC105F985CD9" bold="true" pageId="0" pageNumber="295">
Family 
<taxonomicName id="BAB0CDD5146C860DFD94F00D6EA8847D" authorityName="Bourguignat" authorityYear="1877" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="family">Helicarionidae</taxonomicName>
Bourguignat, 1877
</emphasis>
</paragraph>
<paragraph id="F4C54AF19DB660330F6A938D4FEA492E" pageId="0" pageNumber="295">
<emphasis id="95EB51B254294FDF727046220E7CC1DC" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="4ED8AFF725B01236D446620C60875213" authorityName="E.A.Smith" authorityYear="1899" class="Gastropoda" family="Helicarionidae" genus="Helicarion" kingdom="Animalia" lsidName="Helicarion dyakanum" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="dyakanum">
<emphasis id="E8796DD9D207307B55C7CD4E56C87167" italics="true" pageId="0" pageNumber="295">Helicarion dyakanum</emphasis>
</taxonomicName>
(Godwin-Austen, 1891)
</emphasis>
</paragraph>
<paragraph id="D18E03E73C9685FD4119580E774FD20A" pageId="0" pageNumber="295">
Figs 
<figureCitation id="797252F4FA57E423A8C52DF813FB9FCA" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. A Helicarion dyakanum (Godwin-Austen, 1891) ME 14499 B Paralaoma sarawakensis Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 14510 C Pupina evansi Godwin-Austen, 1889 ME 15029 D Videna nepiadelphos Vermeulen &amp; Liew, 2022 ME 14513 E Pupisoma dioscoricola (C. B. Adams, 1845) ME 14498." figureDoi="10.3897/zookeys.1198.116265.figure17" httpUri="https://binary.pensoft.net/fig/1034256" pageId="0" pageNumber="295">17A</figureCitation>
, 
<figureCitation id="DAB45668C7BAFEF9AE8C23C2995EF9B1" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Living snails from Gua Rumbang. A Platycochlium sarawakense Laidlaw, 1950 MZU. MOL. 22.153 B Dyakia subdebilis E. A. Smith, 1895 MZU. MOL. 23.136 C Helicarion dyakanum (Godwin-Austen, 1891) ME 14499 D Paralaoma sarawakensis Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 14510. All imaged not to scale." figureDoi="10.3897/zookeys.1198.116265.figure24" httpUri="https://binary.pensoft.net/fig/1034263" pageId="0" pageNumber="295">24C</figureCitation>
</paragraph>
<caption id="E2CFAEBEA069B58F110E2B721A67FF1E" doi="10.3897/zookeys.1198.116265.figure17" httpUri="https://binary.pensoft.net/fig/1034256" pageId="0" pageNumber="295" start="Figure 17" startId="F17">
<paragraph id="3477A544A1C091EEBB42C4AA751E46FA" pageId="0" pageNumber="295">
<emphasis id="EAFD4833047ADE4CA07ECE8A1E383973" bold="true" pageId="0" pageNumber="295">Figure 17.</emphasis>
<emphasis id="53CD794AD453F9E4433A996C5FA3E02C" bold="true" pageId="0" pageNumber="295">A</emphasis>
<taxonomicName id="3B5EB7865890FE58221E768D24A7E811" authorityName="E.A.Smith" authorityYear="1899" class="Gastropoda" family="Helicarionidae" genus="Helicarion" kingdom="Animalia" lsidName="Helicarion dyakanum" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="dyakanum">
<emphasis id="68E9EBDAD9D6AEF53FEE30EFF3860F3E" italics="true" pageId="0" pageNumber="295">Helicarion dyakanum</emphasis>
</taxonomicName>
(Godwin-Austen, 1891) ME 14499 
<emphasis id="C1644FD7925ED01B0C31F1BADE8F2FBB" bold="true" pageId="0" pageNumber="295">B</emphasis>
<taxonomicName id="E44AFF2DD094BBD6097E3A5189634C05" authorityName="Marzuki, T. S. Liew &amp; Mohd-Azlan" authorityYear="2021" class="Gastropoda" family="Punctidae" genus="Paralaoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paralaoma sarawakensis" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="sarawakensis">
<emphasis id="5604EFF76F549F9471C1DAA70326D018" italics="true" pageId="0" pageNumber="295">Paralaoma sarawakensis</emphasis>
</taxonomicName>
Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 14510 
<emphasis id="AF9DC97303B3A986017B927DAE9AFC43" bold="true" pageId="0" pageNumber="295">C</emphasis>
<taxonomicName id="25B27BABEA0E62004FE2720F0EE57FC1" authorityName="Godwin-Austen" authorityYear="1889" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina evansi" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="evansi">
<emphasis id="5DC5E91FD5C8CE3F76DB9DCCDFA1DE34" italics="true" pageId="0" pageNumber="295">Pupina evansi</emphasis>
</taxonomicName>
Godwin-Austen, 1889 ME 15029 
<emphasis id="B1F801863163625BE1300A7EC4FEBC86" bold="true" pageId="0" pageNumber="295">D</emphasis>
<taxonomicName id="4E3C90015DFB5F117F02BA278EED9C06" authorityName="Vermeulen &amp; Liew" authorityYear="2022" class="Gastropoda" family="Trochomorphidae" genus="Videna" kingdom="Animalia" lsidName="Videna nepiadelphos" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="nepiadelphos">
<emphasis id="F7A8C2BE761C707443F5E6E826DD0DCF" italics="true" pageId="0" pageNumber="295">Videna nepiadelphos</emphasis>
</taxonomicName>
Vermeulen &amp; Liew, 2022 ME 14513 
<emphasis id="B84FBD9814DF15CE7ACFAF8907B5893A" bold="true" pageId="0" pageNumber="295">E</emphasis>
<taxonomicName id="8881071FFDDEDDC208B22DFEF1178B6B" baseAuthorityName="C. B. Adams" baseAuthorityYear="1845" class="Gastropoda" family="Valloniidae" genus="Pupisoma" kingdom="Animalia" lsidName="Pupisoma dioscoricola" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="dioscoricola">
<emphasis id="7944E71D8CFEE7436B8388024BF9ECF5" italics="true" pageId="0" pageNumber="295">Pupisoma dioscoricola</emphasis>
</taxonomicName>
(C. B. Adams, 1845) ME 14498.
</paragraph>
</caption>
<paragraph id="E855DFB38574A9942FB425F464633AD6" pageId="0" pageNumber="295">
<emphasis id="0076EE0054454B145B06D34374412430" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Busan Hills, Borneo&quot; [= Jambusan Hills, Bau, Sarawak].
</paragraph>
<paragraph id="20431D4815D13BDC809A52EA94AE6829" pageId="0" pageNumber="295">
<emphasis id="30C291599CCA1DFF0C85FD56C95A4ACB" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="866CAEA7C0BFC6C18B6CC4F43655B620" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="5E965A12DC96E2DB079F91694A45B1BE" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14499, ME 15001.
</paragraph>
<paragraph id="3541C7C4B771C918C9F143D0F8AC2898" pageId="0" pageNumber="295">
<emphasis id="7A63AB77DC4BC89E3F0E238EDE6D3012" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed in Sarawak. In Sabah on Mount Trusmadi only. Endemic to Borneo (
<bibRefCitation id="A0888F1F5C1DA1D5BE38BB9931057CAE" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
; 
<bibRefCitation id="FC8CEAB425AC7D818C47A6F66CFDA933" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
).
</paragraph>
<paragraph id="AEE121B12D97AA256704838995DCE847" pageId="0" pageNumber="295">
<emphasis id="43BDB490ECEBD68BB7C1ADC3B9002575" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were observed foraging on leaf surface of trees at the base of limestone cliffs.
</paragraph>
<paragraph id="37C784B081704C7562771A5411833B8E" pageId="0" pageNumber="295">
<emphasis id="252DD158EEB5D7FA09FE7C8E01CE020D" bold="true" pageId="0" pageNumber="295">
Family 
<taxonomicName id="304E06AB23F86439691F3E9350AAD335" authorityName="Morse" authorityYear="1864" class="Gastropoda" family="Punctidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pulmonata" pageId="0" pageNumber="295" phylum="Mollusca" rank="family">Punctidae</taxonomicName>
Morse, 1864
</emphasis>
</paragraph>
<paragraph id="E38A6EDA4546BBFBE91D29F8E431BFC7" pageId="0" pageNumber="295">
<emphasis id="622432AB9F2DA6ADB4A50054D9F61336" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="93B42C61B9AF5DB58977685EBC3D8C5E" authorityName="Marzuki, T. S. Liew &amp; Mohd-Azlan" authorityYear="2021" class="Gastropoda" family="Punctidae" genus="Paralaoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paralaoma sarawakensis" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="sarawakensis">
<emphasis id="45E0DE51FC5769EBF77C389BC65BAFC6" italics="true" pageId="0" pageNumber="295">Paralaoma sarawakensis</emphasis>
</taxonomicName>
Marzuki, Liew &amp; Mohd-Azlan, 2021
</emphasis>
</paragraph>
<paragraph id="50BF8527DF76645D41BB85136CBD41D9" pageId="0" pageNumber="295">
Figs 
<figureCitation id="D3937903DC45858355E9E25E53C3108B" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. A Helicarion dyakanum (Godwin-Austen, 1891) ME 14499 B Paralaoma sarawakensis Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 14510 C Pupina evansi Godwin-Austen, 1889 ME 15029 D Videna nepiadelphos Vermeulen &amp; Liew, 2022 ME 14513 E Pupisoma dioscoricola (C. B. Adams, 1845) ME 14498." figureDoi="10.3897/zookeys.1198.116265.figure17" httpUri="https://binary.pensoft.net/fig/1034256" pageId="0" pageNumber="295">17B</figureCitation>
, 
<figureCitation id="3C09393ED121B7D70154071DD5686C63" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Living snails from Gua Rumbang. A Platycochlium sarawakense Laidlaw, 1950 MZU. MOL. 22.153 B Dyakia subdebilis E. A. Smith, 1895 MZU. MOL. 23.136 C Helicarion dyakanum (Godwin-Austen, 1891) ME 14499 D Paralaoma sarawakensis Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 14510. All imaged not to scale." figureDoi="10.3897/zookeys.1198.116265.figure24" httpUri="https://binary.pensoft.net/fig/1034263" pageId="0" pageNumber="295">24D</figureCitation>
</paragraph>
<paragraph id="0963CB23B5549D3AA6AF9AFC8F7761CA" pageId="0" pageNumber="295">
<emphasis id="8F481DB2548F4E2F64E60876E73DE2D9" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Limestone hill along Skio road, 2.05 miles E Bau, Northern site of Gua Doya, Bukit Sokwang (Site 3), Kuching Division, Sarawak, Malaysia&quot;.
</paragraph>
<paragraph id="2F824A264A541906CF0D8F25E189671E" pageId="0" pageNumber="295">
<emphasis id="3CDB5F0C84B8C66AAF804E263B5357E3" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="BC282927E2D120CA763303AFCD2D5D2D" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="29A6DF18E8A1E1D99136609F905CF5FC" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14510, ME 15055, MZU.MOL.22.448.
</paragraph>
<paragraph id="A2781A991DCDEF719D82C28AF1058AD2" pageId="0" pageNumber="295">
<emphasis id="5B97DCF15DBC2A14620CDDA66C202F25" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak. Also found in Baram, in northern Sarawak. Endemic to Sarawak (
<bibRefCitation id="A52E38D906E8B63F7D5EEDA99666C119" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="D0D3A5C585934BD5277D55E24FCD24E5" pageId="0" pageNumber="295">
<emphasis id="105901CFD9C73A92469561A35C11CA82" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were found foraging among leaf-litter and plant debris at the base of limestone cliffs.
</paragraph>
<paragraph id="3C147E7F76F17E4A655C329826FE0B8B" pageId="0" pageNumber="295">
<emphasis id="DC89AC38159DB321A3E86C4A1143C933" bold="true" pageId="0" pageNumber="295">
Family 
<taxonomicName id="2979A0DD945ADC0A4995818EF856B1E9" authorityName="L. Pfeiffer" authorityYear="1853" class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="family">Pupinidae</taxonomicName>
L. Pfeiffer, 1853
</emphasis>
</paragraph>
<paragraph id="A8102A4778B4593C1C09CAF6AFD69E69" pageId="0" pageNumber="295">
<emphasis id="C57CA990C6C45C8A1189FCF92B69DD0A" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="86FEA90F1E44C0C78401E3789DB7C2DF" authorityName="Godwin-Austen" authorityYear="1889" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina evansi" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="evansi">
<emphasis id="9A801EA0BEB215F8EC9F964026B3BDD3" italics="true" pageId="0" pageNumber="295">Pupina evansi</emphasis>
</taxonomicName>
Godwin-Austen, 1889
</emphasis>
</paragraph>
<paragraph id="FAB266B2F6B982739A2306C57DEC50F2" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="80C4CF8DA55A47249449B6EDA8609120" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. A Helicarion dyakanum (Godwin-Austen, 1891) ME 14499 B Paralaoma sarawakensis Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 14510 C Pupina evansi Godwin-Austen, 1889 ME 15029 D Videna nepiadelphos Vermeulen &amp; Liew, 2022 ME 14513 E Pupisoma dioscoricola (C. B. Adams, 1845) ME 14498." figureDoi="10.3897/zookeys.1198.116265.figure17" httpUri="https://binary.pensoft.net/fig/1034256" pageId="0" pageNumber="295">17C</figureCitation>
</paragraph>
<paragraph id="981F1B6875B56BA7DD5EA166989046D5" pageId="0" pageNumber="295">
<emphasis id="DC0F3CE4990A92D6313661698E70221D" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;From deposit in Cave A, Borneo&quot; [= Tupak Cave, Jambusan Hills, see Cranbrook 2013)].
</paragraph>
<paragraph id="A13BC865180D1E27055B97D51489C96E" pageId="0" pageNumber="295">
<emphasis id="A70F4283C248EED6A5FC765B0DA1AD64" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="D39ECC2A8379EFE4B8A6BAF92D2D7B55" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="350D12EB4D101EDA9CD6EF68B36C31E2" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 15029.
</paragraph>
<paragraph id="A93A15A524DB81097826ED6F6805458D" pageId="0" pageNumber="295">
<emphasis id="3DCE609A35015745FBF8621A1B68C21E" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities in Serian-Padawan limestone hills in western Sarawak (
<bibRefCitation id="3454A241C6E5B363A541D4B3E3283DED" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
). Also found in Sirhassen, Natuna Island (
<bibRefCitation id="E5989487515017BE0A2A24C5870F262A" DOI="https://doi.org/10.1080/00222939408677737" author="Smith, EA" journalOrPublisher="Journal of the Linnean Society of London, Zoology" pageId="0" pageNumber="295" refId="B47" refString="Smith, EA, 1894b. On the land-shells of the Natuna Islands. The Annals and Magazine of Natural History, including Zoology. Botany, and Geology Series 6 13: 453-463. https://doi.org/10.1080/00222939408677737" title="On the land-shells of the Natuna Islands. The Annals and Magazine of Natural History, including Zoology. Botany, and Geology Series 6 13: 453 - 463." url="https://doi.org/10.1080/00222939408677737" year="1894 b">Smith 1894b</bibRefCitation>
).
</paragraph>
<paragraph id="EDE3DE2FBD7A8244D0101C94727CD743" pageId="0" pageNumber="295">
<emphasis id="0DD678A508D8CC3683F0FD2AF99BCA5E" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Among leaf-litter and plant debris at the base of limestone cliffs. Only empty shells were found.
</paragraph>
<paragraph id="178E6227D3525D5C67FAA4000912A31D" pageId="0" pageNumber="295">
<emphasis id="5976225982568A971439D079B64BC2EC" bold="true" pageId="0" pageNumber="295">
Family 
<taxonomicName id="FB7F89AA7688440BBFA9D5B60FA7D707" authorityName="Mollendorff" authorityYear="1890" class="Gastropoda" family="Trochomorphidae" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="family">Trochomorphidae</taxonomicName>
<normalizedToken id="44F627E43DCD8CA928C15016AF3489B7" originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1890
</emphasis>
</paragraph>
<paragraph id="61A89ACDDA71065247D567A7BC0A0882" pageId="0" pageNumber="295">
<emphasis id="2A425FEA635827E72A7925D89E7029A7" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="CC699414BF5A938466D39FCAC15D135E" authorityName="Vermeulen &amp; Liew" authorityYear="2022" class="Gastropoda" family="Trochomorphidae" genus="Videna" kingdom="Animalia" lsidName="Videna nepiadelphos" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="nepiadelphos">
<emphasis id="330AB6E57A85D33A63550D9146CF0AB6" italics="true" pageId="0" pageNumber="295">Videna nepiadelphos</emphasis>
</taxonomicName>
Vermeulen &amp; Liew, 2022
</emphasis>
</paragraph>
<paragraph id="DDF147F426CC2175AAAEF67E664E28D9" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="EEC5BC4D6F4E56116915124DE8B71DB9" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. A Helicarion dyakanum (Godwin-Austen, 1891) ME 14499 B Paralaoma sarawakensis Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 14510 C Pupina evansi Godwin-Austen, 1889 ME 15029 D Videna nepiadelphos Vermeulen &amp; Liew, 2022 ME 14513 E Pupisoma dioscoricola (C. B. Adams, 1845) ME 14498." figureDoi="10.3897/zookeys.1198.116265.figure17" httpUri="https://binary.pensoft.net/fig/1034256" pageId="0" pageNumber="295">17D</figureCitation>
</paragraph>
<paragraph id="135A8E84BFF10F240FC13CE90A1BF787" pageId="0" pageNumber="295">
<emphasis id="251A2A29462770C3437A029BDF68484D" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Danum valley Conservation Area, Tawau Prov., Sabah&quot;.
</paragraph>
<paragraph id="C3F672D6AC882BA0231139377A2BB435" pageId="0" pageNumber="295">
<emphasis id="A18D9ACA582FF939C1F1A088CDAEF179" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="5F15501B4E10825255704F001910AB32" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="266B906BF8EEFC352C1C3377F24332C9" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14513, ME 15056, MZU.MOL.22.447.
</paragraph>
<paragraph id="7E6CF3B14B5F17BDE5BDF0D96C32A890" pageId="0" pageNumber="295">
<emphasis id="FE25CB16B6B2414ACF23B69717B4085F" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed in Borneo. Widespread. Also found in Panaitan island, Indonesia (
<bibRefCitation id="BB24D158425D9BF076857CE187173CCC" DOI="https://doi.org/10.6084/m9.figshare.17156255.v4" author="Vermeulen, JJ" journalOrPublisher="Basteria" pageId="0" pageNumber="295" refId="B57" refString="Vermeulen, JJ, Liew, TS, 2022. Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu. https://doi.org/10.6084/m9.figshare.17156255.v4" title="Land snails and slugs of Sabah and Labuan (Malaysia). Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Kota Kinabalu." url="https://doi.org/10.6084/m9.figshare.17156255.v4" year="2022">Vermeulen and Liew 2022</bibRefCitation>
)
</paragraph>
<paragraph id="785062432AB6633FE67E778A48629EDE" pageId="0" pageNumber="295">
<emphasis id="3F8CBE6FFDDC077170C6667F8386DD1E" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Only empty shells were found among leaf-litter and plant debris at the base of limestone cliffs.
</paragraph>
<paragraph id="86B2C46B950BED258B5F525CFE16174B" pageId="0" pageNumber="295">
<emphasis id="7F9FF4805589DA8E27EDB7BEA214164A" bold="true" pageId="0" pageNumber="295">
Family 
<taxonomicName id="D77D80F552686BBCA221DF593AACA329" authorityName="Morse" authorityYear="1864" class="Gastropoda" family="Valloniidae" kingdom="Animalia" lsidName="" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="family">Valloniidae</taxonomicName>
Morse, 1864
</emphasis>
</paragraph>
<paragraph id="05EC2E5E6C13FE737FA146EC0BF757DF" pageId="0" pageNumber="295">
<emphasis id="60EC196F8DE7D4836C0F898070F26BCC" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="52BB82B85F9AB7C9BD9D4AAC98724159" baseAuthorityName="C. B. Adams" baseAuthorityYear="1845" class="Gastropoda" family="Valloniidae" genus="Pupisoma" kingdom="Animalia" lsidName="Pupisoma dioscoricola" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="dioscoricola">
<emphasis id="40FD3EB060AE4C7EFC84C8C60915B1DF" italics="true" pageId="0" pageNumber="295">Pupisoma dioscoricola</emphasis>
</taxonomicName>
(C. B. Adams, 1845)
</emphasis>
</paragraph>
<paragraph id="AEE9E8312D0F988A06E183EA56ED76D5" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="D33943965A98FACA030EC4FAC0DEED0E" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. A Helicarion dyakanum (Godwin-Austen, 1891) ME 14499 B Paralaoma sarawakensis Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 14510 C Pupina evansi Godwin-Austen, 1889 ME 15029 D Videna nepiadelphos Vermeulen &amp; Liew, 2022 ME 14513 E Pupisoma dioscoricola (C. B. Adams, 1845) ME 14498." figureDoi="10.3897/zookeys.1198.116265.figure17" httpUri="https://binary.pensoft.net/fig/1034256" pageId="0" pageNumber="295">17E</figureCitation>
</paragraph>
<paragraph id="6892121500D604FA2CB48AC9CAD5D0C7" pageId="0" pageNumber="295">
<emphasis id="B2F842B7C816F0CA9BC0F645D9664665" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
<normalizedToken id="CB7D86252E4ADF2EDD16C5CBB5DADDDC" originalValue="“Jamaica”">&quot;Jamaica&quot;</normalizedToken>
.
</paragraph>
<paragraph id="3C9C047D39A3F771FA8CE849A8DED59D" pageId="0" pageNumber="295">
<emphasis id="ACBB3414B8CABAA23E243641DB2EA6C0" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="A42F017F53AAD59779201EAC022CF310" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="C31555BE4C61474AAF7AF52A26045CC8" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022-28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14498, ME 15003, ME 15046, MZU.MOL.22.445.
</paragraph>
<paragraph id="C4514F7A0E01A2AF1025A2161D092EC9" pageId="0" pageNumber="295">
<emphasis id="7918A7FEA490924A07FA7816B407F39F" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Widely distributed in Borneo. Widespread. Distributed in Africa, Asia, Australia, and America (
<bibRefCitation id="38A52CD0469A0BDAA16B9C9D48E1FACA" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="701EBFBE922BB052061440141EEE05EC" pageId="0" pageNumber="295">
<emphasis id="BF0539986EC801739580396FB95A6E71" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were observed foraging on leaf surface of trees at the base of limestone cliffs.
</paragraph>
<paragraph id="367600A65E7FE19C72CD4C6F4215B322" pageId="0" pageNumber="295">
<emphasis id="55D4E2E3B65CAC1068A9BE92D35331D5" bold="true" pageId="0" pageNumber="295">
Subclass 
<taxonomicName id="78876F6812E614F627F18A3C49E8BD69" class="Gastropoda" kingdom="Animalia" lsidName="" pageId="0" pageNumber="295" phylum="Mollusca" rank="subClass" subClass="Neritimorpha">Neritimorpha</taxonomicName>
Golikov &amp; Starobogatov, 1975
</emphasis>
</paragraph>
<paragraph id="564D9B13261A5C537ADB4AA62A621F66" pageId="0" pageNumber="295">
<emphasis id="9845EF40D10BDD301921068EEE186D2F" bold="true" pageId="0" pageNumber="295">
Family 
<taxonomicName id="8A938E56CFD5A98EDBEB6796F901A5CD" authorityName="Troschel" authorityYear="1857" class="Gastropoda" family="Hydrocenidae" kingdom="Animalia" lsidName="" order="Cycloneritida" pageId="0" pageNumber="295" phylum="Mollusca" rank="family">Hydrocenidae</taxonomicName>
Troschel, 1857
</emphasis>
</paragraph>
<paragraph id="131EE0AFC88687B5923D227FE00749B0" pageId="0" pageNumber="295">
<emphasis id="95C12F198C02C1A1AEE2476A2E555419" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="28280CF75D79E412FA69A1991B2E4A76" authorityName="E. A. Smith" authorityYear="1895" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa everetti" order="Cycloneritida" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="everetti">
<emphasis id="7ECFA876A219F9D5088D8E4AA80F0499" italics="true" pageId="0" pageNumber="295">Georissa everetti</emphasis>
</taxonomicName>
E. A. Smith, 1895
</emphasis>
</paragraph>
<paragraph id="D72F86956B61C5A84011ED349331FCCF" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="E205C2886B3D06805374163B23C44AC3" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. A Georissa everetti E. A. Smith, 1895 ME 14482 B Georissa hungerfordi Godwin-Austen, 1889 ME 15032." figureDoi="10.3897/zookeys.1198.116265.figure18" httpUri="https://binary.pensoft.net/fig/1034257" pageId="0" pageNumber="295">18A</figureCitation>
</paragraph>
<caption id="B8F1C47C5B76409F572A034A5AC1664E" doi="10.3897/zookeys.1198.116265.figure18" httpUri="https://binary.pensoft.net/fig/1034257" pageId="0" pageNumber="295" start="Figure 18" startId="F18">
<paragraph id="2DB2E6E9809B53E911D386EFF22E8101" pageId="0" pageNumber="295">
<emphasis id="30F7EB00F2E6B37826EA50A6296CB688" bold="true" pageId="0" pageNumber="295">Figure 18.</emphasis>
<emphasis id="77851E31F145E4E8C8E51F3C7A89861D" bold="true" pageId="0" pageNumber="295">A</emphasis>
<taxonomicName id="5B286B0B8215CE26415DD4AA313C83C1" authorityName="E. A. Smith" authorityYear="1895" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa everetti" order="Cycloneritida" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="everetti">
<emphasis id="27BC578D8338247A75D055A0649FC25C" italics="true" pageId="0" pageNumber="295">Georissa everetti</emphasis>
</taxonomicName>
E. A. Smith, 1895 ME 14482 
<emphasis id="F51D5EAE9E7FCC1867C4FDA6FF79FB88" bold="true" pageId="0" pageNumber="295">B</emphasis>
<taxonomicName id="BCEDEA4DA625A56631BDC55A944CECB3" authorityName="Godwin-Austen" authorityYear="1889" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa hungerfordi" order="Cycloneritida" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="hungerfordi">
<emphasis id="15C516E3DBB6345395C633A8637D82B7" italics="true" pageId="0" pageNumber="295">Georissa hungerfordi</emphasis>
</taxonomicName>
Godwin-Austen, 1889 ME 15032.
</paragraph>
</caption>
<paragraph id="FC7FCB25D07C01E3B0BC5BD36BC6A302" pageId="0" pageNumber="295">
<emphasis id="09782906E6D48B4D4A950B6DF2D7E4DD" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
&quot;Rumbang, W. Sarawak&quot;.
</paragraph>
<paragraph id="F917C61F3FE8D00A196B9E67D69BD0E2" pageId="0" pageNumber="295">
<emphasis id="C95450A3880F001838C42A36998628B0" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="050264EBF9FC53C71D3E59B4F55A5BCF" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="488E2D98B7721873F44E2E18339CAC49" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 2 Sep. 2022; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 14482, MZU.MOL.22.157.
</paragraph>
<paragraph id="9AFBFDFE6C9D22AAA48F482FC87ABC1C" pageId="0" pageNumber="295">
<emphasis id="765323BE83F320DF23BD6C4025CC587A" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak to Niah in northern Sarawak. Also found in Sabah, Sepulut valley. Widespread. Endemic to Borneo (
<bibRefCitation id="32449531C0296AC2FD28ECBF16D1AA0B" DOI="https://doi.org/10.3897/zookeys.840.33326" author="Khalik, MZ" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="35 - 86" refId="B24" refString="Khalik, MZ, Hendriks, KP, Vermeulen, JJ, Schilthuizen, M, 2019. Conchological and molecular analysis of the &quot;non-scaly&quot; Bornean Georissa with descriptions of three new species (Gastropoda, Neritimorpha, Hydrocenidae). ZooKeys 840: 35 - 86, DOI: https://doi.org/10.3897/zookeys.840.33326" title="Conchological and molecular analysis of the &quot; non-scaly &quot; Bornean Georissa with descriptions of three new species (Gastropoda, Neritimorpha, Hydrocenidae)." url="https://doi.org/10.3897/zookeys.840.33326" volume="840" year="2019">Khalik et al. 2019</bibRefCitation>
; 
<bibRefCitation id="9EB04A76DB690AA629A0BBC8333D5196" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="1B0AA38A9C190A8188507DAA58B6387D" pageId="0" pageNumber="295">
<emphasis id="11CE378B4E866AD6EFD65E1ABF881E24" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were observed foraging on wet limestone wall surfaces covered with mosses.
</paragraph>
<paragraph id="189676F6708537F00947C8218FE6971C" pageId="0" pageNumber="295">
<emphasis id="E722D3FC606D8A64A409840F26BDE82C" bold="true" pageId="0" pageNumber="295">
<taxonomicName id="08EF8A044AF7CF25921C9D3041ACC6DA" authorityName="Godwin-Austen" authorityYear="1889" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa hungerfordi" order="Cycloneritida" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="hungerfordi">
<emphasis id="7B06A64DD6432C1548F324EC4287978E" italics="true" pageId="0" pageNumber="295">Georissa hungerfordi</emphasis>
</taxonomicName>
Godwin-Austen, 1889
</emphasis>
</paragraph>
<paragraph id="9109F0D9F879D460859211C6C6068CEC" pageId="0" pageNumber="295">
Fig. 
<figureCitation id="31994A66F4C1D536885BE2DC24DCC3F4" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. A Georissa everetti E. A. Smith, 1895 ME 14482 B Georissa hungerfordi Godwin-Austen, 1889 ME 15032." figureDoi="10.3897/zookeys.1198.116265.figure18" httpUri="https://binary.pensoft.net/fig/1034257" pageId="0" pageNumber="295">18B</figureCitation>
</paragraph>
<paragraph id="BCB071A017B9E21E2EBFE316635986E7" pageId="0" pageNumber="295">
<emphasis id="E96E2CF81E7AC97DE4FC34143A0B597F" bold="true" pageId="0" pageNumber="295">Type locality.</emphasis>
<normalizedToken id="3DFFCD4197260C6966BA04D4978782D3" originalValue="“Borneo”">&quot;Borneo&quot;</normalizedToken>
.
</paragraph>
<paragraph id="71CE505261766E7BF7CF5F95DF00F1DC" pageId="0" pageNumber="295">
<emphasis id="E751059081C0415F0F947B96D060A2FB" bold="true" pageId="0" pageNumber="295">Material examined.</emphasis>
Malaysia • Sarawak, Padawan, Gua Rumbang; 
<geoCoordinate id="191BFDDB9D49C85FE69DC3D2D83B337C" degrees="1" direction="north" minutes="16.77" orientation="latitude" precision="9" value="1.2795">1°16.77'N</geoCoordinate>
, 
<geoCoordinate id="FF5E33E6DA69D70100AB5534040D97F1" degrees="110" direction="east" minutes="15.69" orientation="longitude" precision="9" value="110.2615">110°15.69'E</geoCoordinate>
; 28 Jun. 2023; N.S. Nasir, M.E Marzuki, J.Y. Lee, and M.Z Khalik leg.; ME 15032.
</paragraph>
<paragraph id="82DFC826CD0CC56E3C39BDC7709F0583" pageId="0" pageNumber="295">
<emphasis id="079CF20DFB465F194F5FE6FDB490A5D8" bold="true" pageId="0" pageNumber="295">Distribution.</emphasis>
Scattered localities between Bau and Serian-Padawan limestone hills in western Sarawak. Endemic to western Sarawak (
<bibRefCitation id="D56B75E6716C2C8F26C3F2FAA2FCAE7D" DOI="https://doi.org/10.3897/zookeys.840.33326" author="Khalik, MZ" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="35 - 86" refId="B24" refString="Khalik, MZ, Hendriks, KP, Vermeulen, JJ, Schilthuizen, M, 2019. Conchological and molecular analysis of the &quot;non-scaly&quot; Bornean Georissa with descriptions of three new species (Gastropoda, Neritimorpha, Hydrocenidae). ZooKeys 840: 35 - 86, DOI: https://doi.org/10.3897/zookeys.840.33326" title="Conchological and molecular analysis of the &quot; non-scaly &quot; Bornean Georissa with descriptions of three new species (Gastropoda, Neritimorpha, Hydrocenidae)." url="https://doi.org/10.3897/zookeys.840.33326" volume="840" year="2019">Khalik et al. 2019</bibRefCitation>
; 
<bibRefCitation id="98373AA5C960C1D200DE4F33F913BBE5" DOI="https://doi.org/10.3897/zookeys.1035.60843" author="Marzuki, ME" journalOrPublisher="ZooKeys" pageId="0" pageNumber="295" pagination="1 - 113" refId="B31" refString="Marzuki, ME, Liew, TS, Mohd Azlan, J, 2021. Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1 - 113, DOI: https://doi.org/10.3897/zookeys.1035.60843" title="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species." url="https://doi.org/10.3897/zookeys.1035.60843" volume="1035" year="2021">Marzuki et al. 2021</bibRefCitation>
).
</paragraph>
<paragraph id="0B6558100F0163449158B91451BF9A10" pageId="0" pageNumber="295">
<emphasis id="66305A3ACA4680F39A96BCAFDDB72023" bold="true" pageId="0" pageNumber="295">Remarks.</emphasis>
Living snails were observed foraging on wet limestone wall surfaces covered with mosses.
</paragraph>
<caption id="885E9FD46E06CABA2881AA50BB2D7FBC" doi="10.3897/zookeys.1198.116265.figure19" httpUri="https://binary.pensoft.net/fig/1034258" pageId="0" pageNumber="295" start="Figure 19" startId="F19">
<paragraph id="1E35EC324113DD00BEAEE8C870CB5FDC" pageId="0" pageNumber="295">
<emphasis id="FB89A2D7D6B738753CFE3F80C6F1DEF3" bold="true" pageId="0" pageNumber="295">Figure 19.</emphasis>
Living snails from Gua Rumbang. 
<emphasis id="5BA5537B811BC23FBBEC392AFAFC325A" bold="true" pageId="0" pageNumber="295">A</emphasis>
<taxonomicName id="73CFDA519C4C5D334AD761458DF68FE3" baseAuthorityName="H. Adams" baseAuthorityYear="1871" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" lsidName="Pincerna globosa" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="globosa">
<emphasis id="5F1CD185C510DFEE96AD8EE7F618221F" italics="true" pageId="0" pageNumber="295">Pincerna globosa</emphasis>
</taxonomicName>
(H. Adams, 1871) MZU.MOL.22.135 
<emphasis id="11DDC21CA70BE44446705065BF3AE5C9" bold="true" pageId="0" pageNumber="295">B</emphasis>
<taxonomicName id="6CE94DE2B157EF02C7EC59E7A17C2462" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1889" class="Gastropoda" family="Alycaeidae" genus="Stomacosmethis" kingdom="Animalia" lsidName="Stomacosmethis hosei" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="hosei">
<emphasis id="3C5DCB727BA9F711A868178069262081" italics="true" pageId="0" pageNumber="295">Stomacosmethis hosei</emphasis>
</taxonomicName>
(Godwin-Austen, 1889) MZU.MOL.22.136 
<emphasis id="1B645B7CF1AEAB9BFC99E705E9D844E4" bold="true" pageId="0" pageNumber="295">C</emphasis>
<taxonomicName id="4D349579F4D63B547C03A43AC4E9C5F2" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1889" class="Gastropoda" family="Cyclophoridae" genus="Craspedotropis" kingdom="Animalia" lsidName="Craspedotropis borneensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="borneensis">
<emphasis id="735225EF00A14E2743D5529453C055FE" italics="true" pageId="0" pageNumber="295">Craspedotropis borneensis</emphasis>
</taxonomicName>
(Godwin-Austen, 1889) MZU.MOL.23.141 
<emphasis id="C206622E7EE2B99CB71CB1CA77A325AE" bold="true" pageId="0" pageNumber="295">D</emphasis>
<taxonomicName id="A368E1CC0775D4DEB83CF0382E776C90" baseAuthorityName="E. A. Smith" baseAuthorityYear="1893" class="Gastropoda" family="Cyclophoridae" genus="Japonia" kingdom="Animalia" lsidName="Japonia borneensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="borneensis">
<emphasis id="62C179FC36558C904B7E84654E09C905" italics="true" pageId="0" pageNumber="295">Japonia borneensis</emphasis>
</taxonomicName>
(E. A. Smith, 1893) MZU. MOL.23.138. Images not to scale.
</paragraph>
</caption>
<caption id="B58A3DF80589742BC9CC92360DE7F610" doi="10.3897/zookeys.1198.116265.figure20" httpUri="https://binary.pensoft.net/fig/1034259" pageId="0" pageNumber="295" start="Figure 20" startId="F20">
<paragraph id="7F57DEF3AC5655026B9D4469BA8CC75A" pageId="0" pageNumber="295">
<emphasis id="5F8FA4665CDD26F030E68B66A505CB86" bold="true" pageId="0" pageNumber="295">Figure 20.</emphasis>
Living snails from Gua Rumbang. 
<emphasis id="AF3AFC9534F8FDA727F194CF2181D111" bold="true" pageId="0" pageNumber="295">A</emphasis>
<taxonomicName id="396134F09112252BFB95CB81D047614C" baseAuthorityName="Mousson" baseAuthorityYear="1849" class="Gastropoda" family="Cyclophoridae" genus="Opisthoporus" kingdom="Animalia" lsidName="Opisthoporus biciliatus" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="biciliatus">
<emphasis id="0A81561B5AB61A1634166E8115A35EA8" italics="true" pageId="0" pageNumber="295">Opisthoporus biciliatus</emphasis>
</taxonomicName>
(Mousson, 1849) ME 14979 
<emphasis id="C89181178B705ECF738B0305F33C34FD" bold="true" pageId="0" pageNumber="295">B</emphasis>
<taxonomicName id="9A99D82527928F0D702C50D85EABACC0" baseAuthorityName="H. Adams &amp; A. Adams" baseAuthorityYear="1851" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina adversa" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="adversa">
<emphasis id="22E24279C62F89869D0109457F0C97D7" italics="true" pageId="0" pageNumber="295">Diplommatina adversa</emphasis>
</taxonomicName>
(H. Adams &amp; A. Adams, 1851) MZU. MOL.23.140 
<emphasis id="16768DAC15E17B770659FEE316B0EF78" bold="true" pageId="0" pageNumber="295">C</emphasis>
<taxonomicName id="8A2FE19B974C5CF59847430FC1DE58D7" authorityName="E. A. Smith" authorityYear="1893" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplommatina baritensis" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="baritensis">
<emphasis id="6901C3C398225FC99175AADFCE2C98AF" italics="true" pageId="0" pageNumber="295">Diplommatina baritensis</emphasis>
</taxonomicName>
E. A. Smith, 1893 MZU. MOL.23.137 
<emphasis id="531BCF32B32D38A15EF35BF7605183C3" bold="true" pageId="0" pageNumber="295">D</emphasis>
<taxonomicName id="E528570FB9DA9E4C8D58781619A6A0E5" baseAuthorityName="Vermeulen" baseAuthorityYear="1994" class="Gastropoda" family="Diplommatinidae" genus="Plectostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectostoma anisopterum" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="anisopterum">
<emphasis id="316D34503EF1B569EBF5008957EF1471" italics="true" pageId="0" pageNumber="295">Plectostoma anisopterum</emphasis>
</taxonomicName>
ME 14479 (Vermeulen, 1994). All images not to scale.
</paragraph>
</caption>
<caption id="E27967C26E6323406072186E67E2CDCC" doi="10.3897/zookeys.1198.116265.figure21" httpUri="https://binary.pensoft.net/fig/1034260" pageId="0" pageNumber="295" start="Figure 21" startId="F21">
<paragraph id="6D37B0073763F5966920DF61F5A1DA72" pageId="0" pageNumber="295">
<emphasis id="7BD73E08AD3F58F67A9D08046C6C6D32" bold="true" pageId="0" pageNumber="295">Figure 21.</emphasis>
Living snails from Gua Rumbang. 
<emphasis id="B1CEA26D5A3818CBC8022EC481415567" bold="true" pageId="0" pageNumber="295">A</emphasis>
<taxonomicName id="36DE690C9529B471BC71B312B60FDA2A" baseAuthorityName="E. A. Smith" baseAuthorityYear="1894" class="Gastropoda" family="Diplommatinidae" genus="Plectostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectostoma pumilio" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="pumilio">
<emphasis id="2D019D526189A4738D8C12F90D94AD08" italics="true" pageId="0" pageNumber="295">Plectostoma pumilio</emphasis>
</taxonomicName>
(E. A. Smith, 1894a) ME 14477 
<emphasis id="EE4D59F4FA8D6586DA50A34EFE1B7DAA" bold="true" pageId="0" pageNumber="295">B</emphasis>
<taxonomicName id="AF9BB4B1CF89B54C112B60F5D9C72CD5" baseAuthorityName="E. A. Smith" baseAuthorityYear="1894" class="Gastropoda" family="Diplommatinidae" genus="Plectostoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectostoma austeni" order="Architaenioglossa" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="austeni">
<emphasis id="157F773D3764DBCC2CD952EBCD0B6F1B" italics="true" pageId="0" pageNumber="295">Plectostoma austeni</emphasis>
</taxonomicName>
(E. A. Smith, 1894a) ME 14477 
<emphasis id="5DAD30F6313F5BFAD3DDE44CC2A04D12" bold="true" pageId="0" pageNumber="295">C</emphasis>
<taxonomicName id="4A9D703044D90BE83875EBF070622BC7" baseAuthorityName="E. A. Smith" baseAuthorityYear="1893" class="Gastropoda" family="Ariophantidae" genus="Rahula" kingdom="Animalia" lsidName="Rahula raricostulata" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="raricostulata">
<emphasis id="BDD51B9E9B0AD03980E5899427E58FDF" italics="true" pageId="0" pageNumber="295">Rahula raricostulata</emphasis>
</taxonomicName>
(E. A. Smith, 1893) MZU.MOL.22.139 
<emphasis id="69414FBEA18F004D97849A52A3C1416A" bold="true" pageId="0" pageNumber="295">D</emphasis>
Semi adult 
<taxonomicName id="45234984E8B073C08C1F52BA827E2D72" baseAuthorityName="Reeve" baseAuthorityYear="1854" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" lsidName="Macrochlamys infans" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="infans">
<emphasis id="6EBD36B3DB4165753F65EB3AC1842536" italics="true" pageId="0" pageNumber="295">Macrochlamys infans</emphasis>
</taxonomicName>
(Reeve, 1854) MZU.MOL.23.144. Images not to scale.
</paragraph>
</caption>
<caption id="75F618706C93FCF5C64CD639BE206EC3" doi="10.3897/zookeys.1198.116265.figure22" httpUri="https://binary.pensoft.net/fig/1034261" pageId="0" pageNumber="295" start="Figure 22" startId="F22">
<paragraph id="6744FD4C15BAE2F3F44DEE8B3A306B48" pageId="0" pageNumber="295">
<emphasis id="9C7D92CB1012CAF0B97427013A8C979D" bold="true" pageId="0" pageNumber="295">Figure 22.</emphasis>
Living snails from Gua Rumbang. 
<emphasis id="C627BF7231164397A566BAEB66F22EC1" bold="true" pageId="0" pageNumber="295">A</emphasis>
Juvenile 
<taxonomicName id="6DF72AD015936540D4EEA58D2EEFB0E1" baseAuthorityName="Metcalfe" baseAuthorityYear="1852" class="Gastropoda" family="Ariophantidae" genus="Vitrinula" kingdom="Animalia" lsidName="Vitrinula glutinosa" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="glutinosa">
<emphasis id="80411C0EFE87FB16A26199700C816D0C" italics="true" pageId="0" pageNumber="295">Vitrinula glutinosa</emphasis>
</taxonomicName>
(Metcalfe, 1852) MZU.MOL.23.146 
<emphasis id="5BE91C160DA8412B5858B229199351D8" bold="true" pageId="0" pageNumber="295">B</emphasis>
<taxonomicName id="4331292E60B723C844310C01A55E3565" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1853" class="Gastropoda" family="Chronidae" genus="Kaliella" kingdom="Animalia" lsidName="Kaliella barrakporensis" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="barrakporensis">
<emphasis id="FAF50F9B14B8659ED6024D1DA5E132AF" italics="true" pageId="0" pageNumber="295">Kaliella barrakporensis</emphasis>
</taxonomicName>
(Reeve, 1852) MZU.MOL.23.143 
<emphasis id="DB748FA3E962AAFD11DA2F7C1D1B5194" bold="true" pageId="0" pageNumber="295">C</emphasis>
<taxonomicName id="ACFB4D77C6B01523F2FB8FAA326E9214" baseAuthorityName="E. A. Smith" baseAuthorityYear="1895" class="Gastropoda" family="Chronidae" genus="Kaliella" kingdom="Animalia" lsidName="Kaliella busauensis" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="busauensis">
<emphasis id="B3253F5BD26E76773247AA0C6A2B51AB" italics="true" pageId="0" pageNumber="295">Kaliella busauensis</emphasis>
</taxonomicName>
(E. A. Smith, 1895) MZU.MOL.22.144 
<emphasis id="4B7553C853C1FBD8B5449D24A8B9A976" bold="true" pageId="0" pageNumber="295">D</emphasis>
<taxonomicName id="B31BD39434D61FD2ABF8CDBAC1C2C678" class="Gastropoda" family="Chronidae" genus="Kaliella" kingdom="Animalia" lsidName="Kaliella rumbangensis" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="rumbangensis">
<emphasis id="2629B94887EA0903768E2BF043E34504" italics="true" pageId="0" pageNumber="295">Kaliella rumbangensis</emphasis>
</taxonomicName>
(E. A. Smith, 1895) MZU.MOL.22.163. Images not to scale.
</paragraph>
</caption>
<caption id="EC3D9F8EE868E09975F02D4A16C677D2" doi="10.3897/zookeys.1198.116265.figure23" httpUri="https://binary.pensoft.net/fig/1034262" pageId="0" pageNumber="295" start="Figure 23" startId="F23">
<paragraph id="3DD7D7C30C27AD8110766C1530E3E3F7" pageId="0" pageNumber="295">
<emphasis id="74B4009B36CB865F2830757E42C5E566" bold="true" pageId="0" pageNumber="295">Figure 23.</emphasis>
Living snail from Gua Rumbang. 
<taxonomicName id="AACCE41D98ACA0A364E4D99B911E5660" baseAuthorityName="A. Adams &amp; Reeve" baseAuthorityYear="1850" class="Gastropoda" family="Chronidae" genus="Exrhysota" kingdom="Animalia" lsidName="Exrhysota brookei" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="brookei">
<emphasis id="BEEA0982AEDCDCEB1E57285928FDD3D0" italics="true" pageId="0" pageNumber="295">Exrhysota brookei</emphasis>
</taxonomicName>
(A. Adams &amp; Reeve, 1850) MZU.MOL.22.151.
</paragraph>
</caption>
<caption id="FEE4AB04833E5C1D55896C5AC4E7A507" doi="10.3897/zookeys.1198.116265.figure24" httpUri="https://binary.pensoft.net/fig/1034263" pageId="0" pageNumber="295" start="Figure 24" startId="F24">
<paragraph id="105A609093CF8CF707069CC1F91F8CB7" pageId="0" pageNumber="295">
<emphasis id="B83441824C7F360B5A4E420CE34BA822" bold="true" pageId="0" pageNumber="295">Figure 24.</emphasis>
Living snails from Gua Rumbang. 
<emphasis id="8BD370BB6BE2D8CA3F3E5FE8F8759C7E" bold="true" pageId="0" pageNumber="295">A</emphasis>
<taxonomicName id="C038FD7E285A280543620981D50825DE" authorityName="Laidlaw" authorityYear="1950" class="Gastropoda" family="Diapheridae" genus="Platycochlium" kingdom="Animalia" lsidName="Platycochlium sarawakense" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="sarawakense">
<emphasis id="C58F29A33BA5245071E5E74D580CB3FA" italics="true" pageId="0" pageNumber="295">Platycochlium sarawakense</emphasis>
</taxonomicName>
Laidlaw, 1950 MZU.MOL.22.153 
<emphasis id="6732D2A36C2E36E2D1979263B3A39A94" bold="true" pageId="0" pageNumber="295">B</emphasis>
<taxonomicName id="B735983298AF57A55F0282D3494CC5CB" authorityName="E. A. Smith" authorityYear="1895" class="Gastropoda" family="Dyakiidae" genus="Dyakia" kingdom="Animalia" lsidName="Dyakia subdebilis" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="subdebilis">
<emphasis id="5F463CD086734E84FD81AB9C47A2F8D3" italics="true" pageId="0" pageNumber="295">Dyakia subdebilis</emphasis>
</taxonomicName>
E. A. Smith, 1895 MZU.MOL.23.136 
<emphasis id="8A1A7F5BFE261087DCD9B3916A0455A0" bold="true" pageId="0" pageNumber="295">C</emphasis>
<taxonomicName id="7F2B3D02FE5A0FEFA4467103AE2242C0" authorityName="E.A.Smith" authorityYear="1899" class="Gastropoda" family="Helicarionidae" genus="Helicarion" kingdom="Animalia" lsidName="Helicarion dyakanum" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="dyakanum">
<emphasis id="B5A90E01402669316BB22F427D8342FC" italics="true" pageId="0" pageNumber="295">Helicarion dyakanum</emphasis>
</taxonomicName>
(Godwin-Austen, 1891) ME 14499 
<emphasis id="05BF623AF0A4AD271374E031169EDA6D" bold="true" pageId="0" pageNumber="295">D</emphasis>
<taxonomicName id="93A7096B3FC932644DE4F35D6112BF02" authorityName="Marzuki, T. S. Liew &amp; Mohd-Azlan" authorityYear="2021" class="Gastropoda" family="Punctidae" genus="Paralaoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paralaoma sarawakensis" order="Stylommatophora" pageId="0" pageNumber="295" phylum="Mollusca" rank="species" species="sarawakensis">
<emphasis id="EBFB0FDD2978FAC61FE501123E32D685" italics="true" pageId="0" pageNumber="295">Paralaoma sarawakensis</emphasis>
</taxonomicName>
Marzuki, Liew &amp; Mohd-Azlan, 2021 ME 14510. All imaged not to scale.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>