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<document id="53CE13F2A80FB9ACC53A7F74F4F91F4A" ID-DOI="10.3897/zookeys.1213.131283" ID-ZooBank="8FF3F5AB-3DF7-4DE9-AAC4-DADA05435A8E" ID-publisher-id="131283" URI-arpha="C564F4B3-64FD-5C10-A07E-5CECD7D2453C" URI-zoobank="http://zoobank.org/8FF3F5AB-3DF7-4DE9-AAC4-DADA05435A8E" article-type="research-article" checkinTime="1727362523881" checkinUser="pensoft" docAuthor="Kundrata, Robin &amp; Packova, Gabriela" docDate="2024" docId="E51A1E16E0A75E26ACC812A3043A6A0D" docLanguage="en" docName="ZooKeys 1213: 183-197" docOrigin="ZooKeys 1213" docSource="https://zookeys.pensoft.net/article/131283/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Namibdrilus Kundrata &amp; Packova, 2024, gen. nov." docType="treatment" docUuid="81EF4D52-F9D3-4BF0-B7FD-8BDBD216032E" docUuidSource="ZooBank" docVersion="2" dtd-version="3.0" lastPageNumber="197" masterDocId="C564F4B364FD5C10A07E5CECD7D2453C" masterDocTitle="Discovery of a new soft-bodied click-beetle genus from Namibia with a unique morphology leads to a modified diagnosis of Drilini (Coleoptera, Elateridae)" masterLastPageNumber="197" masterPageNumber="183" pageNumber="183" updateTime="1727362885818" updateUser="ExternalLinkService">
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<mods:title id="1AB7FB3F4907416604434508703BA6D3">Discovery of a new soft-bodied click-beetle genus from Namibia with a unique morphology leads to a modified diagnosis of Drilini (Coleoptera, Elateridae)</mods:title>
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<mods:namePart id="9D0CEB14E205D6065C97A117B87670E5">Kundrata, Robin</mods:namePart>
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<mods:affiliation id="E049D38DF4D8F9899097B2801EC7E1B0">Department of Zoology, Faculty of Science, Palacky University, 17. listopadu 50, 77146 Olomouc, Czech Republic</mods:affiliation>
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<mods:namePart id="2D0AA07B7F35BA40E024512D7DFED26E">Packova, Gabriela</mods:namePart>
<mods:nameIdentifier id="FDBADC42D083C214515564234C848F6D" type="ORCID">0000-0001-7949-619X</mods:nameIdentifier>
<mods:affiliation id="603540BFA8FFF4A1C6EE4E0CBC6D93DB">Department of Zoology, Faculty of Science, Palacky University, 17. listopadu 50, 77146 Olomouc, Czech Republic</mods:affiliation>
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<mods:date id="977990A1E8B327860FB4F730086A9908">2024</mods:date>
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<treatment id="E51A1E16E0A75E26ACC812A3043A6A0D" ID-DOI="http://doi.org/10.5281/zenodo.13844115" ID-Zenodo-Dep="13844115" ID-arpha="E51A1E16-E0A7-5E26-ACC8-12A3043A6A0D" ID-zoobank="https://zoobank.org/81EF4D52-F9D3-4BF0-B7FD-8BDBD216032E" LSID="urn:lsid:zoobank.org:act:81EF4D52-F9D3-4BF0-B7FD-8BDBD216032E" LSID-ZBK="urn:lsid:zoobank.org:act:81EF4D52-F9D3-4BF0-B7FD-8BDBD216032E" httpUri="http://treatment.plazi.org/id/E51A1E16E0A75E26ACC812A3043A6A0D">
<subSubSection id="EE3BACF7D14AF1934F65BDF5074025AC" type="nomenclature">
<paragraph id="D7FA69A7E15AD7077219666AEA0ED1BB">
<heading id="329647F76E82B17C76F2B38B91FE15A1" reason="label">Genus</heading>
<taxonomicName id="61DAA77FE8491E5F769BCEA40A855140" authorityName="Kundrata &amp; Packova" authorityYear="2024" class="Insecta" family="Elateridae" genus="Namibdrilus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="8AB10FF5D79AB6C86A1EEBA3BB32967F" italics="true">Namibdrilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3379730ADB9A44AF6DBFBCA6D5CD9C98" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="65D631511987A022C6A698A150209294" type="description">
<paragraph id="859DDA0D755B319FA01105BA06E622CC">
<figureCitation id="1843710372752BD9FA322A180054B8F5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Namibdrilus albertalleni gen. et sp. nov., holotype, male A habitus, dorsal view B habitus, ventral view. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1213.131283.figure1" httpUri="https://binary.pensoft.net/fig/1142208">Figs 1</figureCitation>
,
<figureCitation id="F31929404FDE1D964735D91C8C9F4A62" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Namibdrilus albertalleni gen. et sp. nov., holotype, male A head with basal antennomeres, dorsal view B head, frontolateral view C head, frontal view D head, ventral view E left antenna, dorsal view. Scale bars: 1.0 mm (A D); 2.0 mm (E)." figureDoi="10.3897/zookeys.1213.131283.figure2" httpUri="https://binary.pensoft.net/fig/1142209">2</figureCitation>
,
<figureCitation id="50DD88D19D0C8118F396871AF564A51B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Namibdrilus albertalleni gen. et sp. nov., holotype, male A head and prothorax, dorsal view B head and prothorax, lateral view C prothorax and mesothorax, ventral view D metathorax, ventral view E apical portion of elytra, dorsal view F right protarsus, dorsal view G left mesotarsus, dorsal view H left metatarsus, dorsal view. Scale bars: 1.0 mm (A, D); 0.5 mm (B, C, E, F H)." figureDoi="10.3897/zookeys.1213.131283.figure3" httpUri="https://binary.pensoft.net/fig/1142210">3</figureCitation>
,
<figureCitation id="0DA95DFB0701880B8BC6B3289B6852B3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Namibdrilus albertalleni gen. et sp. nov., holotype, male A abdomen, ventral view B male genitalia (undissected), dorsal view C abdominal tergites IX and X, dorsal view D abdominal sternite IX, ventral view E male genitalia, dorsal view F male genitalia, lateral view G male genitalia, ventral view. Scale bars: 3.0 mm (A); 0.2 mm (B); 0.5 mm (C G)." figureDoi="10.3897/zookeys.1213.131283.figure4" httpUri="https://binary.pensoft.net/fig/1142211">4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENLAC" type="type species">
<paragraph id="A55F62E02372B1E5B4167CD512641DC2">
<heading id="25829874C0BE8A1EC6FC2ED5CC5E563C" reason="title">
<typeStatus id="41ED6FD15F46570FA37B415896F52589">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="4FA3B104E8650E2E11ED99FC61343A05">
<taxonomicName id="64FA43C805A965AAFC30DBEA700F79E5" authorityName="Kundrata &amp; Packova" authorityYear="2024" class="Insecta" family="Elateridae" genus="Namibdrilus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="albertalleni" status="sp. nov.">
<emphasis id="41455C841CCC42E8E94F717841868078" italics="true">Namibdrilus albertalleni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7715F39D9E46EDB6B5A277A9A3F1A5E8" rank="species">sp. nov.</taxonomicNameLabel>
; here designated.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5LAC" type="etymology">
<paragraph id="68832460B3D8F03F42526781CB07D457">
<heading id="66DDCF8BFBA3152C69B676F8E52047CB" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="BD7DD2F67FA17E159DC6C32E052BE1CE">
The generic name is derived from the
<collectingCountry id="007AAA0C9CEE490E9C38D18ADAFCFC38" name="Namibia">Republic of Namibia</collectingCountry>
, plus
<taxonomicName id="6DEA3FEA7A72522B6172409B1EC63110" authorityName="Olivier" authorityYear="1790" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="8E7881E404682660C5B3A69E21523F17" italics="true">Drilus</emphasis>
</taxonomicName>
, a genus name in
<taxonomicName id="32D721DA8E1A9C2D760691FE350A9756" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Elateridae</taxonomicName>
:
<taxonomicName id="D4B4A6CD982DF635C890FBB6C1DDCF00" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="tribe" tribe="Drilini">Drilini</taxonomicName>
. Gender: masculine.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVMAC" type="diagnosis">
<paragraph id="AF95C9BC7143CA18CB49AFF431907BEF">
<heading id="026412B033DCC714A256C9581CF0E92A" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="72267851C382C229D18AA4B1E91291D1">
<taxonomicName id="552403D92DC78FEB5E207CDBE762526E" authorityName="Kundrata &amp; Packova" authorityYear="2024" class="Insecta" family="Elateridae" genus="Namibdrilus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="9EBC80D6428F31D514CA921DB864ECC2" italics="true">Namibdrilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="16958307CB9FAB147EFDADB4BB947FFB" rank="genus">gen. nov.</taxonomicNameLabel>
can be unequivocally distinguished from its congeners by the robust unidentate mandibles (Fig.
<figureCitation id="05FF84F74DB29CEF167073DA82677147" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Namibdrilus albertalleni gen. et sp. nov., holotype, male A head with basal antennomeres, dorsal view B head, frontolateral view C head, frontal view D head, ventral view E left antenna, dorsal view. Scale bars: 1.0 mm (A D); 2.0 mm (E)." figureDoi="10.3897/zookeys.1213.131283.figure2" httpUri="https://binary.pensoft.net/fig/1142209">2 A D</figureCitation>
) and the aedeagal median lobe dorsally without a subapical hook (Fig.
<figureCitation id="DBBE52144B23D047EE3BCBD766CD2B99" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Namibdrilus albertalleni gen. et sp. nov., holotype, male A abdomen, ventral view B male genitalia (undissected), dorsal view C abdominal tergites IX and X, dorsal view D abdominal sternite IX, ventral view E male genitalia, dorsal view F male genitalia, lateral view G male genitalia, ventral view. Scale bars: 3.0 mm (A); 0.2 mm (B); 0.5 mm (C G)." figureDoi="10.3897/zookeys.1213.131283.figure4" httpUri="https://binary.pensoft.net/fig/1142211">4 B, E G</figureCitation>
). Additionally, the following combination of characters can be used to recognize the genus: body (Fig.
<figureCitation id="94FFAE1A80A21F3BEA0E9D271345FC6C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Namibdrilus albertalleni gen. et sp. nov., holotype, male A habitus, dorsal view B habitus, ventral view. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1213.131283.figure1" httpUri="https://binary.pensoft.net/fig/1142208">1 A, B</figureCitation>
) elongate; frontoclypeal region (Fig.
<figureCitation id="5B590B82F9D09CAC406C2CA82DD71D98" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Namibdrilus albertalleni gen. et sp. nov., holotype, male A head with basal antennomeres, dorsal view B head, frontolateral view C head, frontal view D head, ventral view E left antenna, dorsal view. Scale bars: 1.0 mm (A D); 2.0 mm (E)." figureDoi="10.3897/zookeys.1213.131283.figure2" httpUri="https://binary.pensoft.net/fig/1142209">2 A D</figureCitation>
) strongly produced forwards, gradually narrowed toward apex, apically carinate and widely rounded; eyes (Fig.
<figureCitation id="90AF30DA3C4A828E9F7C8DC985E17046" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Namibdrilus albertalleni gen. et sp. nov., holotype, male A head with basal antennomeres, dorsal view B head, frontolateral view C head, frontal view D head, ventral view E left antenna, dorsal view. Scale bars: 1.0 mm (A D); 2.0 mm (E)." figureDoi="10.3897/zookeys.1213.131283.figure2" httpUri="https://binary.pensoft.net/fig/1142209">2 A D</figureCitation>
) large, their frontal separation 1.15 times eye diameter; antenna (Fig.
<figureCitation id="BB246C846AAC409174E235831B8CEF73" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Namibdrilus albertalleni gen. et sp. nov., holotype, male A head with basal antennomeres, dorsal view B head, frontolateral view C head, frontal view D head, ventral view E left antenna, dorsal view. Scale bars: 1.0 mm (A D); 2.0 mm (E)." figureDoi="10.3897/zookeys.1213.131283.figure2" httpUri="https://binary.pensoft.net/fig/1142209">2 E</figureCitation>
) robust, strongly serrate; pronotum (Fig.
<figureCitation id="A9ACE0DD5A198C33989656D249E3BA6A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Namibdrilus albertalleni gen. et sp. nov., holotype, male A head and prothorax, dorsal view B head and prothorax, lateral view C prothorax and mesothorax, ventral view D metathorax, ventral view E apical portion of elytra, dorsal view F right protarsus, dorsal view G left mesotarsus, dorsal view H left metatarsus, dorsal view. Scale bars: 1.0 mm (A, D); 0.5 mm (B, C, E, F H)." figureDoi="10.3897/zookeys.1213.131283.figure3" httpUri="https://binary.pensoft.net/fig/1142210">3 A, B</figureCitation>
) roughly subrectangular, 1.30 times as wide as long, lateral carina not developed; prosternum (Fig.
<figureCitation id="F38F265C17D3E40E52B68536E4A0BA10" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Namibdrilus albertalleni gen. et sp. nov., holotype, male A head and prothorax, dorsal view B head and prothorax, lateral view C prothorax and mesothorax, ventral view D metathorax, ventral view E apical portion of elytra, dorsal view F right protarsus, dorsal view G left mesotarsus, dorsal view H left metatarsus, dorsal view. Scale bars: 1.0 mm (A, D); 0.5 mm (B, C, E, F H)." figureDoi="10.3897/zookeys.1213.131283.figure3" httpUri="https://binary.pensoft.net/fig/1142210">3 C</figureCitation>
) distinctly transverse, without well-developed prosternal process; scutellar shield (Fig.
<figureCitation id="46E912F152EEC2B7056B8008880EE63C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Namibdrilus albertalleni gen. et sp. nov., holotype, male A head and prothorax, dorsal view B head and prothorax, lateral view C prothorax and mesothorax, ventral view D metathorax, ventral view E apical portion of elytra, dorsal view F right protarsus, dorsal view G left mesotarsus, dorsal view H left metatarsus, dorsal view. Scale bars: 1.0 mm (A, D); 0.5 mm (B, C, E, F H)." figureDoi="10.3897/zookeys.1213.131283.figure3" httpUri="https://binary.pensoft.net/fig/1142210">3 A</figureCitation>
) apically subtruncate and medially emarginate; mesoventrite (Fig.
<figureCitation id="613CCBDEC9BE82043E709356976A39C9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Namibdrilus albertalleni gen. et sp. nov., holotype, male A head and prothorax, dorsal view B head and prothorax, lateral view C prothorax and mesothorax, ventral view D metathorax, ventral view E apical portion of elytra, dorsal view F right protarsus, dorsal view G left mesotarsus, dorsal view H left metatarsus, dorsal view. Scale bars: 1.0 mm (A, D); 0.5 mm (B, C, E, F H)." figureDoi="10.3897/zookeys.1213.131283.figure3" httpUri="https://binary.pensoft.net/fig/1142210">3 C</figureCitation>
) narrow, v-shaped; elytra (Figs
<figureCitation id="68C5AF5EE76722739AA198ACF473C89E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Namibdrilus albertalleni gen. et sp. nov., holotype, male A habitus, dorsal view B habitus, ventral view. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1213.131283.figure1" httpUri="https://binary.pensoft.net/fig/1142208">1 A</figureCitation>
,
<figureCitation id="6634992E226168B6ACC72F817AAE238E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Namibdrilus albertalleni gen. et sp. nov., holotype, male A head and prothorax, dorsal view B head and prothorax, lateral view C prothorax and mesothorax, ventral view D metathorax, ventral view E apical portion of elytra, dorsal view F right protarsus, dorsal view G left mesotarsus, dorsal view H left metatarsus, dorsal view. Scale bars: 1.0 mm (A, D); 0.5 mm (B, C, E, F H)." figureDoi="10.3897/zookeys.1213.131283.figure3" httpUri="https://binary.pensoft.net/fig/1142210">3 E</figureCitation>
) elongate, with relatively rough surface and without any apparent striae or rows of punctures; abdomen (Figs
<figureCitation id="84B5318A406465695D55695FD0291244" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Namibdrilus albertalleni gen. et sp. nov., holotype, male A habitus, dorsal view B habitus, ventral view. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1213.131283.figure1" httpUri="https://binary.pensoft.net/fig/1142208">1 B</figureCitation>
,
<figureCitation id="DFDAB0D83D599E4F947E4E6FE8CC75A0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Namibdrilus albertalleni gen. et sp. nov., holotype, male A abdomen, ventral view B male genitalia (undissected), dorsal view C abdominal tergites IX and X, dorsal view D abdominal sternite IX, ventral view E male genitalia, dorsal view F male genitalia, lateral view G male genitalia, ventral view. Scale bars: 3.0 mm (A); 0.2 mm (B); 0.5 mm (C G)." figureDoi="10.3897/zookeys.1213.131283.figure4" httpUri="https://binary.pensoft.net/fig/1142211">4 A</figureCitation>
) with eight free visible sternites, the first of which is partly membranous medially; abdominal sternite IX and tergite X (Figs
<figureCitation id="14B117C9CFFAA4C7D777C60D6D9928D5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Namibdrilus albertalleni gen. et sp. nov., holotype, male A abdomen, ventral view B male genitalia (undissected), dorsal view C abdominal tergites IX and X, dorsal view D abdominal sternite IX, ventral view E male genitalia, dorsal view F male genitalia, lateral view G male genitalia, ventral view. Scale bars: 3.0 mm (A); 0.2 mm (B); 0.5 mm (C G)." figureDoi="10.3897/zookeys.1213.131283.figure4" httpUri="https://binary.pensoft.net/fig/1142211">4 C, D</figureCitation>
) not apparently elongate, about 1.5 times as long as wide. Based on its morphology,
<taxonomicName id="C765F896D1649D032854FAB6FAD033FA" authorityName="Kundrata &amp; Packova" authorityYear="2024" class="Insecta" family="Elateridae" genus="Namibdrilus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="1C7D937E3C16A12D2AC54490CDD954A3" italics="true">Namibdrilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8975F5AAE3BD52823515CDF4CAFE064C" rank="genus">gen. nov.</taxonomicNameLabel>
is similar to genera of the clade D (
<taxonomicName id="9D72733531D60787D858DB6B0E3B2403" authorityName="Olivier" authorityYear="1790" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="82484B6E8E82E6227C1983B662329214" italics="true">Drilus</emphasis>
</taxonomicName>
clade) defined by
<bibRefCitation id="C781A8C78D73C60986CAB144DB0B254E" DOI="10.1093/zoolinnean/zlz033" author="Kundrata R &amp; Bocak L" firstAuthor="Kundrata" journalOrPublisher="Zoological Journal of the Linnean Society" pagination="413-452" refId="B16" refString="Kundrata R, Bocak L (2019) Molecular phylogeny reveals the gradual evolutionary transition to soft-bodiedness in click-beetles and identifies sub-Saharan Africa as a cradle of diversity for Drilini (Coleoptera: Elateridae). Zoological Journal of the Linnean Society 187: 413452. https://doi.org/10.1093/zoolinnean/zlz033" title="Molecular phylogeny reveals the gradual evolutionary transition to soft-bodiedness in click-beetles and identifies sub-Saharan Africa as a cradle of diversity for Drilini (Coleoptera: Elateridae)." volume="187" year="2019">Kundrata and Bocak (2019)</bibRefCitation>
. For more information and the comparison of
<taxonomicName id="F84306878D7E8D8F4D04A7D3D41BB165" authorityName="Kundrata &amp; Packova" authorityYear="2024" class="Insecta" family="Elateridae" genus="Namibdrilus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="0C04556367423344886399EB8FA920CD" italics="true">Namibdrilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="93920889C55C6515BDE0B4BECC2A10F5" rank="genus">gen. nov.</taxonomicNameLabel>
with presumably related genera see the identification key and discussion below.
</paragraph>
<caption id="A90E3D1424D20429210EFBD0ADEBAC6E" ID-DOI="10.3897/zookeys.1213.131283.figure1" ID-arpha="AB02C106-5D88-501D-A13C-B02218E824BB" httpUri="https://binary.pensoft.net/fig/1142208" startId="F1">
<paragraph id="11921EEF6A00EE10DCAF196FCE63553A">
<label id="4A11A726254097900BD8D74D664EAC19">Figure 1.</label>
</paragraph>
<paragraph id="31F076F81EC5A0DE6981F30198F65D3C">
<taxonomicName id="89DA344A2A274F28C35E8B4F5B4BD4EB" authorityName="Kundrata &amp; Packova" authorityYear="2024" class="Insecta" family="Elateridae" genus="Namibdrilus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="albertalleni" status="gen. et sp. nov.">
<emphasis id="8C22F55421FF85C330E196AF65FD9F40" italics="true">Namibdrilus albertalleni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9D440A6A1B45255F97C1CBF1712C9156" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype, male
<emphasis id="D5C58D881AAF829DCF88BB497D9A675F" bold="true">A</emphasis>
habitus, dorsal view
<emphasis id="1A731CF86D2FD72DD085AA55BF19DC6C" bold="true">B</emphasis>
habitus, ventral view. Scale bar: 4.0 mm.
</paragraph>
</caption>
<caption id="353C6E57076352C853A8726399B75E7A" ID-DOI="10.3897/zookeys.1213.131283.figure2" ID-arpha="1A937B85-DC28-5B30-8FE2-E051E5F6D4F7" httpUri="https://binary.pensoft.net/fig/1142209" startId="F2">
<paragraph id="8D6CA17496A5105F09A93F1194E181D1">
<label id="5CE8BBA06124BB5DD0B9F19588F8C66C">Figure 2.</label>
</paragraph>
<paragraph id="73820FD541DC4B75EA6BB28441CEBDE1">
<taxonomicName id="9E61546B47A91E68F87B7A09B2042D73" authorityName="Kundrata &amp; Packova" authorityYear="2024" class="Insecta" family="Elateridae" genus="Namibdrilus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="albertalleni" status="gen. et sp. nov.">
<emphasis id="A6449AE67C48E14FACE593225FA45528" italics="true">Namibdrilus albertalleni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F5D503AE8656C7CCAA88B7BA7C1EA9C5" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype, male
<emphasis id="1B7046CDA85D4D376FE5F425C6AE4FA5" bold="true">A</emphasis>
head with basal antennomeres, dorsal view
<emphasis id="4CC05088665F8D9BC2F6FA279EAB37BF" bold="true">B</emphasis>
head, frontolateral view
<emphasis id="9E6060F8FE5B8AE2E26C2B0E72D42D41" bold="true">C</emphasis>
head, frontal view
<emphasis id="4894951B94DE29C9DF2F1613F13A3E43" bold="true">D</emphasis>
head, ventral view
<emphasis id="7A7A54D8D04C0D57C26B091F0B2282C4" bold="true">E</emphasis>
left antenna, dorsal view. Scale bars: 1.0 mm (
<emphasis id="2766156F24F3E0A6686CEC797D42879A" bold="true">A D</emphasis>
); 2.0 mm (
<emphasis id="C9C2749B554279988E5D7484C1414459" bold="true">E</emphasis>
).
</paragraph>
</caption>
<caption id="58A0641E659D067AF0ED438F0DCCFE15" ID-DOI="10.3897/zookeys.1213.131283.figure3" ID-arpha="05CAECC2-2ECE-57E9-8548-6D0F3CBB6AA2" httpUri="https://binary.pensoft.net/fig/1142210" startId="F3">
<paragraph id="E77DE1918399335452C5CC62829E68E4">
<label id="037C23D5F22CAE2675C863DA11851BB7">Figure 3.</label>
</paragraph>
<paragraph id="79FA36128796AE18A8BEE062282356BC">
<taxonomicName id="1F87F935B6C4D830D080EC0230A4DD9E" authorityName="Kundrata &amp; Packova" authorityYear="2024" class="Insecta" family="Elateridae" genus="Namibdrilus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="albertalleni" status="gen. et sp. nov.">
<emphasis id="CDCF53A76739202352EB64036E1D0146" italics="true">Namibdrilus albertalleni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4E4054B01E5F21A3209CF020CDF117B" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype, male
<emphasis id="9FEFAB514CC420C9332BDA9BAF2455B3" bold="true">A</emphasis>
head and prothorax, dorsal view
<emphasis id="D70C26B7B3976E723F62FA2CB147957B" bold="true">B</emphasis>
head and prothorax, lateral view
<emphasis id="F044BC920ED6F2134E535389FD3C1486" bold="true">C</emphasis>
prothorax and mesothorax, ventral view
<emphasis id="AA2594B3E5D624CDE344EF4495E5E6B9" bold="true">D</emphasis>
metathorax, ventral view
<emphasis id="3E16C9C9F36D68242F6168AE3D4FB30A" bold="true">E</emphasis>
apical portion of elytra, dorsal view
<emphasis id="CA3C31915D04ECDCAC031903FE6A98C5" bold="true">F</emphasis>
right protarsus, dorsal view
<emphasis id="F6370AD2728D643EA693D93CE2AE00F9" bold="true">G</emphasis>
left mesotarsus, dorsal view
<emphasis id="4963E4A04566B688CE90A96B450FFC5D" bold="true">H</emphasis>
left metatarsus, dorsal view. Scale bars: 1.0 mm (
<emphasis id="11E42D70CF0FBB764C3D3D3E944149E1" bold="true">A, D</emphasis>
); 0.5 mm (
<emphasis id="A20FED69E1F76CA1914F7E985D3111C4" bold="true">B, C, E, F H</emphasis>
).
</paragraph>
</caption>
<caption id="5BC3AE56E2401B8FC74C2DF1540C5223" ID-DOI="10.3897/zookeys.1213.131283.figure4" ID-arpha="F90985DD-B7EE-512C-B4D6-26676F4CEB8E" httpUri="https://binary.pensoft.net/fig/1142211" startId="F4">
<paragraph id="C33BCF82A556AA930F2B431E3AAFB103">
<label id="BA3F8417C5E58FCE631147C2B568E69F">Figure 4.</label>
</paragraph>
<paragraph id="29D106A93FB4AE0B99CD9AE4891FB314">
<taxonomicName id="B70CB7B10A9295F5AB5B71179F9058B5" authorityName="Kundrata &amp; Packova" authorityYear="2024" class="Insecta" family="Elateridae" genus="Namibdrilus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="albertalleni" status="gen. et sp. nov.">
<emphasis id="27BE82B4E860095DC5290AA72BE2CFBE" italics="true">Namibdrilus albertalleni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CB8596B55B9387DE820EEF91C0F1B39D" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype, male
<emphasis id="9CFFB310C88C830CA0C13F10D2A13F18" bold="true">A</emphasis>
abdomen, ventral view
<emphasis id="7B3CECCC30DAE30AD65490E871A5BFBA" bold="true">B</emphasis>
male genitalia (undissected), dorsal view
<emphasis id="E3B057A7B05D60755C406FA993B9C011" bold="true">C</emphasis>
abdominal tergites IX and X, dorsal view
<emphasis id="9463CF253D27F6C7EAC42DF2238FCBF2" bold="true">D</emphasis>
abdominal sternite IX, ventral view
<emphasis id="E287371DB2B86D83F00D0030581CFF65" bold="true">E</emphasis>
male genitalia, dorsal view
<emphasis id="334298DBCF5A9DF4AB1D625810B5877D" bold="true">F</emphasis>
male genitalia, lateral view
<emphasis id="9A02706600A9D75D5E388B9E7E1FED6A" bold="true">G</emphasis>
male genitalia, ventral view. Scale bars: 3.0 mm (
<emphasis id="9E2794FD54FEAAB816E0B0B8AE94A26D" bold="true">A</emphasis>
); 0.2 mm (
<emphasis id="7BFF491F375B904500479FAF6511722B" bold="true">B</emphasis>
); 0.5 mm (
<emphasis id="0FC2B06AD8DAF996C9E674D1A71FA0E2" bold="true">C G</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ESEAE" type="distribution">
<paragraph id="21CEEB99D3E7E4659BB4CB990A8477C0">
<heading id="1774B5BEEAA6355E57703D4044B46090" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="B647AC3ED68C2EB1588B9BB6FD032549">
<collectingCountry id="49F55536FEADB7C964C00DF68DCAD7A7" name="Namibia">Namibia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>