196 lines
25 KiB
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196 lines
25 KiB
XML
<document id="488B511C2E96E3232A156DB725087A10" ID-CLB-Dataset="21869" ID-DOI="10.5852/ejt.2016.236" ID-GBIF-Dataset="2dfee902-a7fc-472b-bf1d-27029b3504f3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3847840" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590436812908" checkinUser="valdenar" docAuthor="Herbert, David G." docDate="2016" docId="C146B323FFB0FFE6FDDD9A68FD6FF13A" docLanguage="en" docName="ejt-2016-236.pdf.imf" docOrigin="European Journal of Taxonomy 236" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Sheldonia Ancey 1887" docType="treatment" docVersion="9" lastPageNumber="16" masterDocId="3D7FCB5BFFBEFFE9FFC0994BFF82F57F" masterDocTitle="New narrow-range endemic land snails from the sky islands of northern South Africa (Gastropoda: Streptaxidae and Urocyclidae)" masterLastPageNumber="29" masterPageNumber="1" pageNumber="15" updateTime="1698841246942" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="789902F4D0D36F1301E407B14979615F">
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<mods:title id="CE25CA6D981BDE317CEB5154D8F1F14B">New narrow-range endemic land snails from the sky islands of northern South Africa (Gastropoda: Streptaxidae and Urocyclidae)</mods:title>
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<mods:name id="DA529002035F887F9819DA4A22A66724" type="personal">
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<mods:namePart id="011ED16B5CE97FBEE7216DE9965B40DA">Herbert, David G.</mods:namePart>
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<mods:typeOfResource id="C3CBC2672A8B8F5C86A2A9D4C73C460C">text</mods:typeOfResource>
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<mods:title id="9EF122F938F7BF85D558EA478FD732D0">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="085DC2A4F5B3DCEED07F8AF3A56BF041">
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<mods:date id="D09F3363921F1F13CF7EEB809935F29D">2016</mods:date>
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<mods:number id="61850B63EA550821F4C25D7817275F9D">2016-10-12</mods:number>
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<mods:number id="91E591DF13EE15B0C392E22FD982652E">236</mods:number>
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<mods:start id="5F9FA11D7C0EA8A0C9E80AA6D813BA46">1</mods:start>
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<mods:end id="DCB3276B3033676FF45F60594BC11E87">29</mods:end>
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<mods:classification id="0F70CBD70DF9D8D51E6497A8C6537826">journal article</mods:classification>
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<mods:identifier id="E4CF1B0F6BF4090AC191A69BFB895266" type="CLB-Dataset">21869</mods:identifier>
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<mods:identifier id="CB6DE183D0BE34168DD8C7FD32A6171C" type="DOI">10.5852/ejt.2016.236</mods:identifier>
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<treatment id="C146B323FFB0FFE6FDDD9A68FD6FF13A" ID-DOI="http://doi.org/10.5281/zenodo.3854752" ID-GBIF-Taxon="164282655" ID-Zenodo-Dep="3854752" LSID="urn:lsid:plazi:treatment:C146B323FFB0FFE6FDDD9A68FD6FF13A" httpUri="http://treatment.plazi.org/id/C146B323FFB0FFE6FDDD9A68FD6FF13A" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
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<subSubSection id="01F551BEFFB0FFE7FDDD9A68FB94F641" box="[541,1046,803,830]" pageId="14" pageNumber="15" type="nomenclature">
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<paragraph id="49500235FFB0FFE7FDDD9A68FB94F641" blockId="14.[541,1046,803,830]" box="[541,1046,803,830]" pageId="14" pageNumber="15">
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<heading id="1218B559FFB0FFE7FDDD9A68FB94F641" box="[541,1046,803,830]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="15" reason="2">
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Genus
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<taxonomicName id="8EEF79B6FFB0FFE7FDB09A68FC07F642" ID-CoL="7PGPT" authority="Ancey, 1887" authorityName="Ancey" authorityYear="1887" box="[624,901,803,830]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="genus">
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<emphasis id="7B9BDE27FFB0FFE7FDB09A68FD68F641" bold="true" box="[624,746,803,830]" italics="true" pageId="14" pageNumber="15">Sheldonia</emphasis>
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<bibRefCitation id="2D7E7FC4FFB0FFE7FD319A68FC07F642" author="Ancey C. F." box="[753,901,803,829]" pageId="14" pageNumber="15" pagination="53 - 54" refId="ref13486" refString="Ancey C. F. 1887. Descriptions of new genera or subgenera of Helicidae. Conchologist's Exchange 1 (9 - 10): 53 - 54." type="journal article" year="1887">Ancey, 1887</bibRefCitation>
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</taxonomicName>
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(
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<emphasis id="7B9BDE27FFB0FFE7FC569A6FFB8FF642" box="[918,1037,803,829]" italics="true" pageId="14" pageNumber="15">sensu lato</emphasis>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="01F551BEFFB0FFE6FF7D9A2EFDA9F740" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="reference_group">
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<paragraph id="49500235FFB0FFE7FF7D9A2EFE41F6DC" blockId="14.[189,1398,868,1036]" pageId="14" pageNumber="15">
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<treatmentCitationGroup id="69FF251BFFB0FFE7FF7D9A2EFE41F6DC" pageId="14" pageNumber="15">
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<treatmentCitation id="C84E2424FFB0FFE7FF7D9A2EFD80F600" author="Ancey C. F." box="[189,514,868,895]" page="53" pageId="14" pageNumber="15" year="1887">
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<taxonomicName id="8EEF79B6FFB0FFE7FF7D9A2EFD80F600" ID-CoL="7PGPT" authority="Ancey, 1887: 53" authorityName="Ancey" authorityPageNumber="53" authorityYear="1887" box="[189,514,868,895]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="genus">
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<emphasis id="7B9BDE27FFB0FFE7FF7D9A2EFEB0F600" box="[189,306,869,895]" italics="true" pageId="14" pageNumber="15">Sheldonia</emphasis>
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<bibRefCitation id="2D7E7FC4FFB0FFE7FEFA9A2EFD80F600" author="Ancey C. F." box="[314,514,868,895]" pageId="14" pageNumber="15" pagination="53 - 54" refId="ref13486" refString="Ancey C. F. 1887. Descriptions of new genera or subgenera of Helicidae. Conchologist's Exchange 1 (9 - 10): 53 - 54." type="journal article" year="1887">Ancey, 1887: 53</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<typeStatus id="9654BC97FFB0FFE7FDCF9A2EFDCBF600" box="[527,585,869,895]" pageId="14" pageNumber="15">Type</typeStatus>
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species:
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<taxonomicName id="8EEF79B6FFB0FFE7FD799A2EFBADF600" authority="Benson, 1848" authorityName="Benson" authorityYear="1848" box="[697,1071,869,895]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="trotteriana">
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<emphasis id="7B9BDE27FFB0FFE7FD799A2EFC02F601" box="[697,896,869,895]" italics="true" pageId="14" pageNumber="15">Helix trotteriana</emphasis>
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Benson, 1848
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</taxonomicName>
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, by subsequent designation (
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<treatmentCitation id="C84E2424FFB0FFE7FF059AC3FE34F6DC" author="Connolly M." box="[197,438,904,931]" page="467" pageId="14" pageNumber="15" year="1925">
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<bibRefCitation id="2D7E7FC4FFB0FFE7FF059AC3FE34F6DC" author="Connolly M." box="[197,438,904,931]" pageId="14" pageNumber="15" pagination="457 - 479" refId="ref13787" refString="Connolly M. 1925. Notes on African non-marine Mollusca, with descriptions of many new species (continued). Annals and Magazine of Natural History Series 9, 15: 457 - 479, pl. 28." type="journal article" year="1925">Connolly 1925: 467</bibRefCitation>
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</treatmentCitation>
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).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="49500235FFB0FFE7FF7D9AE0FEE1F173" blockId="14.[189,1398,868,1036]" pageId="14" pageNumber="15">
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<treatmentCitationGroup id="69FF251BFFB0FFE7FF7D9AE0FEE1F173" pageId="14" pageNumber="15">
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<treatmentCitation id="C84E2424FFB0FFE7FF7D9AE0FDE5F6BA" author="Godwin-Austen H. H." box="[189,615,939,965]" page="131" pageId="14" pageNumber="15" year="1908">
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<taxonomicName id="8EEF79B6FFB0FFE7FF7D9AE0FDE5F6BA" ID-CoL="830fc108-2ed0-492f-bb03-ae7ed4b10d7b" authority="Godwin-Austen, 1908: 131" authorityName="Godwin-Austen" authorityPageNumber="131" authorityYear="1908" box="[189,615,939,965]" class="Gastropoda" family="Urocyclidae" genus="Peltatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="genus">
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<emphasis id="7B9BDE27FFB0FFE7FF7D9AE0FE9FF6BA" box="[189,285,939,965]" italics="true" pageId="14" pageNumber="15">Peltatus</emphasis>
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<bibRefCitation id="2D7E7FC4FFB0FFE7FEE49AE0FDE5F6BA" author="Godwin-Austen H. H." box="[292,615,939,965]" pageId="14" pageNumber="15" pagination="131 - 133" refId="ref13925" refString="Godwin-Austen H. H. 1908. On Peltatus, gen. nov. Zonitidarum: Type P. hudsoniae (Benson). Annals and Magazine of Natural History Series 8, 1: 131 - 133. Available from http: // biodiversitylibrary. org / page / 24295840 [accessed 10 Aug. 2016]" type="journal article" year="1908">Godwin-Austen, 1908: 131</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<typeStatus id="9654BC97FFB0FFE7FDB29AE7FD2EF6B9" box="[626,684,940,966]" pageId="14" pageNumber="15">Type</typeStatus>
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species (original designation)
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<taxonomicName id="8EEF79B6FFB0FFE7FBD49AE0FB51F6BA" authorityName="Benson" authorityYear="1864" box="[1044,1235,939,965]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="hudsoniae">
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<emphasis id="7B9BDE27FFB0FFE7FBD49AE0FB51F6BA" box="[1044,1235,939,965]" italics="true" pageId="14" pageNumber="15">Helix hudsoniae</emphasis>
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</taxonomicName>
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(
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<emphasis id="7B9BDE27FFB0FFE7FB249AE7FA92F6BA" box="[1252,1296,940,965]" italics="true" pageId="14" pageNumber="15">non</emphasis>
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Benson, 1864)
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<bibRefCitation id="2D7E7FC4FFB0FFE7FEC99A85FD8CF697" author="Godwin-Austen H. H." box="[265,526,974,1000]" pageId="14" pageNumber="15" pagination="131 - 133" refId="ref13925" refString="Godwin-Austen H. H. 1908. On Peltatus, gen. nov. Zonitidarum: Type P. hudsoniae (Benson). Annals and Magazine of Natural History Series 8, 1: 131 - 133. Available from http: // biodiversitylibrary. org / page / 24295840 [accessed 10 Aug. 2016]" type="journal article" year="1908">Godwin-Austen, 1908</bibRefCitation>
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[=
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<taxonomicName id="8EEF79B6FFB0FFE7FDF89A84FB5EF697" authority="Melvill & Ponsonby, 1890" authorityName="Melvill & Ponsonby" authorityYear="1890" box="[568,1244,974,1000]" class="Gastropoda" family="Vitrinidae" genus="Vitrina" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="variety" species="huttoniae" variety="aloicola">
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<emphasis id="7B9BDE27FFB0FFE7FDF89A84FC18F697" box="[568,922,974,1000]" italics="true" pageId="14" pageNumber="15">Vitrina huttoniae var. aloicola</emphasis>
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Melvill & Ponsonby, 1890
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</taxonomicName>
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, cf. Godwin- Austen 1912].
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</treatmentCitationGroup>
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</paragraph>
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<caption id="1D9052BDFFB0FFE7FF7D9E09FCD9F295" ID-DOI="http://doi.org/10.5281/zenodo.3847872" ID-Zenodo-Dep="3847872" httpUri="https://zenodo.org/record/3847872/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1858,1884]" targetBox="[189,1398,1066,1830]" targetPageId="14">
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<paragraph id="49500235FFB0FFE7FF7D9E09FCD9F295" blockId="14.[189,1399,1858,2026]" pageId="14" pageNumber="15">
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<emphasis id="7B9BDE27FFB0FFE7FF7D9E09FE89F223" bold="true" box="[189,267,1858,1884]" pageId="14" pageNumber="15">Fig. 8.</emphasis>
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Map showing the distribution of new urocyclid species in relation to centres of plant endemism (
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<bibRefCitation id="2D7E7FC4FFB0FFE7FF059E2DFE7DF2FF" author="Van Wyk A. E. & Smith G. F." box="[197,511,1894,1920]" pageId="14" pageNumber="15" refId="ref14747" refString="Van Wyk A. E. & Smith G. F. 2001. Regions of Floristic Endemism in Southern Africa. Umdaus Press, Pretoria." type="book" year="2001">Van Wyk & Smith 2001</bibRefCitation>
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). Green circles =
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<taxonomicName id="8EEF79B6FFB0FFE7FD289E2DFBDFF2FF" ID-CoL="4Q8K5" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[744,1117,1894,1920]" class="Gastropoda" family="Urocyclidae" genus="Ptilototheca" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="soutpansbergensis" status="gen. et sp. nov.">
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<emphasis id="7B9BDE27FFB0FFE7FD289E2DFBDFF2FF" box="[744,1117,1894,1920]" italics="true" pageId="14" pageNumber="15">Ptilototheca soutpansbergensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="60A8635CFFB0FFE7FBAB9E2DFAB0F2FF" box="[1131,1330,1894,1920]" pageId="14" pageNumber="15" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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; blue triangles =
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<taxonomicName id="8EEF79B6FFB0FFE7FE869EC2FDEEF2DC" ID-CoL="4X4BN" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[326,620,1929,1955]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="wolkbergensis" status="sp. nov.">
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<emphasis id="7B9BDE27FFB0FFE7FE869EC2FDEEF2DC" box="[326,620,1929,1955]" italics="true" pageId="14" pageNumber="15">Sheldonia wolkbergensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="60A8635CFFB0FFE7FDB59EC2FD57F2DC" box="[629,725,1929,1955]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
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; maroon squares =
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<taxonomicName id="8EEF79B6FFB0FFE7FC039EC2FB4CF2DC" ID-CoL="4X4B4" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[963,1230,1929,1955]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="monsmaripi" status="sp. nov.">
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<emphasis id="7B9BDE27FFB0FFE7FC039EC2FB4CF2DC" box="[963,1230,1929,1955]" italics="true" pageId="14" pageNumber="15">Sheldonia monsmaripi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="60A8635CFFB0FFE7FB179EC2FAB5F2DC" box="[1239,1335,1929,1955]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
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; pale blue = Soutpansberg Centre; pale orange = Wolkberg Centre (Serala Subcentre); pale green = Wolkberg Centre (Blyde Subcentre). Contours at 1000 and 2000 m.
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</paragraph>
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</caption>
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<paragraph id="49500235FFB1FFE6FF7D9840FDA9F740" blockId="15.[189,1399,267,575]" pageId="15" pageNumber="16">
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<taxonomicName id="8EEF79B6FFB1FFE6FF7D9840FEB0F45A" ID-CoL="7PGPT" authorityName="Ancey" authorityYear="1887" box="[189,306,267,293]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="genus">
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<emphasis id="7B9BDE27FFB1FFE6FF7D9840FEB0F45A" box="[189,306,267,293]" italics="true" pageId="15" pageNumber="16">Sheldonia</emphasis>
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</taxonomicName>
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is currently a poorly understood and ill-defined genus that has been used
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<emphasis id="7B9BDE27FFB1FFE6FB0E9847FACBF45A" box="[1230,1353,267,293]" italics="true" pageId="15" pageNumber="16">sensu lato</emphasis>
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for an assemblage of heliciform urocyclids occurring in southern Africa. Unpublished anatomical and molecular studies clearly show that it is a composite entity comprising several distinct, monophyletic lineages that warrant recognition as distinct genus-level entities. In
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<taxonomicName id="8EEF79B6FFB1FFE6FC25983EFBD8F4F0" ID-CoL="7PGPT" authorityName="Ancey" authorityYear="1887" box="[997,1114,373,399]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="genus" sensu="stricto">
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<emphasis id="7B9BDE27FFB1FFE6FC25983EFBD8F4F0" box="[997,1114,373,399]" italics="true" pageId="15" pageNumber="16">Sheldonia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="60A8635CFFB1FFE6FBA4983DFB0BF4F0" box="[1124,1161,373,399]" pageId="15" pageNumber="16" sensu="stricto">
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<emphasis id="7B9BDE27FFB1FFE6FBA4983DFB00F4F0" box="[1124,1154,374,399]" italics="true" pageId="15" pageNumber="16">s.s</emphasis>
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.
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</taxonomicNameLabel>
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the epiphallus is of moderate length, the epiphallic caecum is elongate and arises close to or just distal to the mid-region of the epiphallus, the flagellum is long and divided into distinct F1 and F2 components, the atrium frequently possesses a stimulator and the free oviduct gland is conspicuously enlarged. There is no penial verge or atrial diverticulum and the vas deferens is not thickened and convoluted. In addition, the protoconch is usually punctate.
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</paragraph>
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</subSubSection>
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<subSubSection id="01F551BEFFB1FFE6FF7D9B2DFD6FF13A" pageId="15" pageNumber="16" type="description">
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<paragraph id="49500235FFB1FFE6FF7D9B2DFB5CF6E5" blockId="15.[189,1399,614,923]" pageId="15" pageNumber="16">
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The above is largely in accordance with the description of the genus given by
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<bibRefCitation id="2D7E7FC4FFB1FFE6FB909B2DFA9FF7FE" author="Schileyko A. A." box="[1104,1309,614,641]" pageId="15" pageNumber="16" pagination="1167 - 1307" refId="ref14510" refString="Schileyko A. A. 2002. Treatise on Recent terrestrial pulmonate molluscs. Part 9. Helicarionidae, Gymnarionidae, Rhysotinidae, Ariophantidae. Ruthenica Supplement 2: 1167 - 1307." type="journal article" year="2002">Schileyko (2002)</bibRefCitation>
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, except that he stated that the penial retractor muscle was attached to the summit of the caecum when in reality it is attached to the penis apex. This discrepancy stems from the figure of the reproductive tract of
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<taxonomicName id="8EEF79B6FFB1FFE6FF7D9B9BFED4F795" authorityName="Benson" authorityYear="1864" box="[189,342,720,746]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="hudsoniae">
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<emphasis id="7B9BDE27FFB1FFE6FF7D9B9BFED4F795" box="[189,342,720,746]" italics="true" pageId="15" pageNumber="16">S. hudsoniae</emphasis>
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</taxonomicName>
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provided by
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<bibRefCitation id="2D7E7FC4FFB1FFE6FE389B9BFC89F795" author="Godwin-Austen H. H." box="[504,779,720,746]" pageId="15" pageNumber="16" pagination="131 - 133" refId="ref13925" refString="Godwin-Austen H. H. 1908. On Peltatus, gen. nov. Zonitidarum: Type P. hudsoniae (Benson). Annals and Magazine of Natural History Series 8, 1: 131 - 133. Available from http: // biodiversitylibrary. org / page / 24295840 [accessed 10 Aug. 2016]" type="journal article" year="1908">Godwin-Austen (1908)</bibRefCitation>
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and was corrected in a subsequent paper (Godwin- Austen 1912) in relation to this species and several others, including the
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<typeStatus id="9654BC97FFB1FFE6FC349BBFFBA5F671" box="[1012,1063,756,782]" pageId="15" pageNumber="16">type</typeStatus>
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species. Following
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<bibRefCitation id="2D7E7FC4FFB1FFE6FAC99BB8FE8CF64E" author="Connolly M." pageId="15" pageNumber="16" pagination="1 - 660" refId="ref13827" refString="Connolly M. 1939. A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: 1 - 660." type="journal article" year="1939">Connolly (1939)</bibRefCitation>
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,
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<bibRefCitation id="2D7E7FC4FFB1FFE6FED99A5DFE61F64E" author="Schileyko A. A." box="[281,483,790,817]" pageId="15" pageNumber="16" pagination="1167 - 1307" refId="ref14510" refString="Schileyko A. A. 2002. Treatise on Recent terrestrial pulmonate molluscs. Part 9. Helicarionidae, Gymnarionidae, Rhysotinidae, Ariophantidae. Ruthenica Supplement 2: 1167 - 1307." type="journal article" year="2002">Schileyko (2002)</bibRefCitation>
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treated
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<emphasis id="7B9BDE27FFB1FFE6FDFF9A5DFD5AF64F" box="[575,728,790,816]" italics="true" pageId="15" pageNumber="16">Kerkophorus</emphasis>
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<bibRefCitation id="2D7E7FC4FFB1FFE6FD1F9A5DFC61F64E" author="Godwin-Austen H. H." box="[735,995,790,817]" pageId="15" pageNumber="16" pagination="122 - 139" refId="ref13983" refString="Godwin-Austen H. H. 1912. A review of the South-African land-Mollusca belonging to the family Zonitidae [Part I]. Annals and Magazine of Natural History Series 8, 9: 122 - 139. Available from http: // biodiversitylibrary. org / page / 18638879 [accessed 10 Aug. 2016]" type="journal article" year="1912">Godwin-Austen, 1912</bibRefCitation>
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and
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<taxonomicName id="8EEF79B6FFB1FFE6FBDA9A5DFF7AF62B" authority="Godwin-Austen, 1912" authorityName="Godwin-Austen" authorityYear="1912" class="Gastropoda" family="Urocyclidae" genus="Microkerkus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="genus">
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<emphasis id="7B9BDE27FFB1FFE6FBDA9A5DFB2FF64F" box="[1050,1197,790,816]" italics="true" pageId="15" pageNumber="16">Microkerkus</emphasis>
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<bibRefCitation id="2D7E7FC4FFB1FFE6FB749A5DFF7AF62B" author="Godwin-Austen H. H." pageId="15" pageNumber="16" pagination="122 - 139" refId="ref13983" refString="Godwin-Austen H. H. 1912. A review of the South-African land-Mollusca belonging to the family Zonitidae [Part I]. Annals and Magazine of Natural History Series 8, 9: 122 - 139. Available from http: // biodiversitylibrary. org / page / 18638879 [accessed 10 Aug. 2016]" type="journal article" year="1912">Godwin-Austen, 1912</bibRefCitation>
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</taxonomicName>
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as synonyms of
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<taxonomicName id="8EEF79B6FFB1FFE6FE7F9A71FDB6F62B" authorityName="Ancey" authorityYear="1887" box="[447,564,826,852]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="genus">
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<emphasis id="7B9BDE27FFB1FFE6FE7F9A71FDB6F62B" box="[447,564,826,852]" italics="true" pageId="15" pageNumber="16">Sheldonia</emphasis>
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</taxonomicName>
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, but ongoing studies indicate that this is an oversimplification. Further discussion of this is beyond the scope of the present contribution, but the morphological characters diagnostic of these taxa will be provided, together with a key, in a subsequent publication.
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</paragraph>
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<paragraph id="49500235FFB1FFE6FF7D9A8AFD6FF13A" blockId="15.[189,1399,961,1094]" pageId="15" pageNumber="16">
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The following two species clearly do not belong to
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<taxonomicName id="8EEF79B6FFB1FFE6FCFA9A8AFC2DF6A4" authorityName="Ancey" authorityYear="1887" box="[826,943,961,987]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="genus" sensu="stricto">
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<emphasis id="7B9BDE27FFB1FFE6FCFA9A8AFC2DF6A4" box="[826,943,961,987]" italics="true" pageId="15" pageNumber="16">Sheldonia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="60A8635CFFB1FFE6FC7B9A89FC62F6A3" box="[955,992,962,988]" pageId="15" pageNumber="16" sensu="stricto">
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<emphasis id="7B9BDE27FFB1FFE6FC7B9A89FC5BF6A4" box="[955,985,962,987]" italics="true" pageId="15" pageNumber="16">s.s</emphasis>
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.
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</taxonomicNameLabel>
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, nor are they clearly referable to either
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<emphasis id="7B9BDE27FFB1FFE6FEC69AAEFE1DF680" box="[262,415,997,1023]" italics="true" pageId="15" pageNumber="16">Kerkophorus</emphasis>
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or
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<taxonomicName id="8EEF79B6FFB1FFE6FE049AAEFDD5F680" authorityName="Godwin-Austen" authorityYear="1912" box="[452,599,997,1023]" class="Gastropoda" family="Urocyclidae" genus="Microkerkus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="genus">
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<emphasis id="7B9BDE27FFB1FFE6FE049AAEFDD5F680" box="[452,599,997,1023]" italics="true" pageId="15" pageNumber="16">Microkerkus</emphasis>
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</taxonomicName>
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. Their position relative to other southern African helicoid urocyclids is unclear and for the time being I refer them to
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<taxonomicName id="8EEF79B6FFB1FFE6FCC49D43FCFBF15D" authorityName="Ancey" authorityYear="1887" box="[772,889,1032,1058]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="16" phylum="Mollusca" rank="genus">
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<emphasis id="7B9BDE27FFB1FFE6FCC49D43FCFBF15D" box="[772,889,1032,1058]" italics="true" pageId="15" pageNumber="16">Sheldonia</emphasis>
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</taxonomicName>
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in its broadest sense. Clarification of their affinities must await analysis of molecular data.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |