214 lines
27 KiB
XML
214 lines
27 KiB
XML
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<mods:title id="5DCD7604171654C4FFDC9B2F43C81203">A new cryptic species and review of the east-Andean leaf chafer genus Mesomerodon Ohaus, 1905 (Coleoptera, Scarabaeidae, Rutelinae)</mods:title>
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<mods:namePart id="F40695E44405D9AB61FE30FFB0E0E935">Seidel, Matthias</mods:namePart>
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<mods:namePart id="A67AA161F2963AEA9D71B8C6DD986101">Jameson, Mary L.</mods:namePart>
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<mods:namePart id="BA49B3A79D4285D362779EC023BB725D">Stone, Rachel L.</mods:namePart>
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<mods:title id="E8BEED6494E4B25CD66B947A78861BBF">ZooKeys</mods:title>
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<mods:date id="B50724919AA79886204C231639019F6B">2017</mods:date>
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<mods:number id="CA4369D45204883C7BD78C7A500A3DE0">671</mods:number>
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<mods:url id="FEBE3D5E5B91C394E1BEC140A515C032">http://dx.doi.org/10.3897/zookeys.671.11815</mods:url>
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<mods:classification id="6E09FB30343BBF6FFD3F04C63A803171">journal article</mods:classification>
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<mods:identifier id="060E46E2761A1FBF768DFF644EFDAE17" type="DOI">http://dx.doi.org/10.3897/zookeys.671.11815</mods:identifier>
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<treatment id="42FE50612D310F32E319CE700DD0C0AF" ID-GBIF-Taxon="128810346" LSID="urn:lsid:zoobank.org:act:4B70B243-1A6A-445D-9C33-5E859DE8B086" httpUri="http://treatment.plazi.org/id/42FE50612D310F32E319CE700DD0C0AF" lastPageId="8" lastPageNumber="69" pageId="5" pageNumber="66">
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<subSubSection id="0364F0C703EF286E0827EA01A87CE5B7" pageId="5" pageNumber="66" type="nomenclature">
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<paragraph id="CA0FE4CA4614448BF91731C0A64DB09D" pageId="5" pageNumber="66">
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<taxonomicName id="E94A722E83DC53103BEF0FB737E5D1C7" ID-CoL="424HY" LSID="http://zoobank.org/4B70B243-1A6A-445D-9C33-5E859DE8B086" authority="Seidel, Jameson, & Stone" class="Insecta" family="Scarabaeidae" genus="Mesomerodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesomerodon barclayi" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="barclayi">
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<pageBreakToken id="3B7E62EEC41C101B2C168DF69B468620" pageId="5" pageNumber="66" start="start">Mesomerodon</pageBreakToken>
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barclayi Seidel, Jameson, & Stone
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</taxonomicName>
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<taxonomicNameLabel id="C9C907BDF377FE18FACB3AC114F609DC" pageId="5" pageNumber="66">sp. n.</taxonomicNameLabel>
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Figs 1-3, 8, 15, 18, 22, 25
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</paragraph>
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</subSubSection>
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<subSubSection id="2F3B46CBF8D2BB3BD56662F02E29BC8E" lastPageId="8" lastPageNumber="69" pageId="6" pageNumber="67" type="type material">
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<paragraph id="E1FDDA6609140659ADD46AAA17D59E18" pageId="6" pageNumber="67">
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<pageBreakToken id="67141D9360D67FBAD59E806A145F430D" pageId="6" pageNumber="67" start="start">Type</pageBreakToken>
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material.
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</paragraph>
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<paragraph id="9ABC5B670931349C446E57404F8C47C2" lastPageId="8" lastPageNumber="69" pageId="6" pageNumber="67">
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Holotype male and 19 paratypes (10 males, 9 females). Holotype male at ZMHB with label data: a) "Rutelide, Pacayacu, 23.8.37" (handwritten), b) "
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<taxonomicName id="84EB50A3623A7DF59F1973C001A41D86" class="Insecta" family="Scarabaeidae" genus="Mesomerodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesomerodon spinipenne" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="spinipenne">Mesomerodon spinipenne</taxonomicName>
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Ohs., MNHUB Berlin" (typeset), c) male genitalia card mounted, d) our red holotype label. Paratype female at MSPC with label data: a) "Ecuador, Pacayacu, 3.X.37, Dr. Schultze - Rhonhof S.G." (typeset and handwritten), b) "Pacayacu 3.10.37" (handwritten), c) "loan from Zool.Mus. Berlin" (typeset), d) "
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<taxonomicName id="7D159105327D0C640169C701FA465845" class="Insecta" family="Scarabaeidae" genus="Mesomerodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesomerodon spinipenne" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="spinipenne">Mesomerodon spinipenne</taxonomicName>
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Ohs., MNHUB Berlin" (typeset), e) our yellow paratype label. Paratype male at ZMHB with label data: a) "O.ECUADOR, Sarayacu Feyer" (typeset), b) "
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<taxonomicName id="75DFA0BF5817318C6E3418633FDAD3C3" class="Insecta" family="Scarabaeidae" genus="Mesomerodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesomerodon spinipenne" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="spinipenne">Mesomerodon spinipenne</taxonomicName>
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, Cotype ♂ Ohs." (handwritten on red label), c) "kein typus" (handwritten), d) male genitalia card mounted e) our yellow paratype label. Paratype female at ZMHB with label data: a) "O.ECUADOR, Sarayacu Feyer" (typeset), b) "
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<taxonomicName id="0E2AD8EB528F8C746F50290351CEF184" class="Insecta" family="Scarabaeidae" genus="Mesomerodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesomerodon spinipenne" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="spinipenne">Mesomerodon spinipenne</taxonomicName>
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, Cotype ♀ Ohs." (handwritten on red label), c) "kein Typus" (handwritten), d) our yellow paratype label. Paratype male at ZMHB with label data: a) "O.ECUADOR, Sarayacu Feyer" (typeset), b)
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<normalizedToken id="0F590EBD6B2DAB33326A33834F409D5C" originalValue="“88882”">"88882"</normalizedToken>
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(typeset), c)
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<normalizedToken id="F4ABAC5AE302761FA0A0A0DF06D6F37B" originalValue="“♂”">"♂"</normalizedToken>
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(typeset on black bordered label), d) "
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<taxonomicName id="1CC689415BC1BF953B4E93395A2F7C27" class="Insecta" family="Scarabaeidae" genus="Mesomerodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesomerodon spinipenne" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="spinipenne">Mesomerodon spinipenne</taxonomicName>
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Ohs., MNHUB Berlin" (typeset), e) male genitalia card mounted, f) our yellow paratype label. Paratype female at ZMHB with label data: a) "Rutelide u. Cyclocephala, Pacayacu 24.9.37" (handwritten), b) "
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<taxonomicName id="39902A16EBDA14AC26FB4A68FA67E070" class="Insecta" family="Scarabaeidae" genus="Mesomerodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesomerodon spinipenne" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="spinipenne">Mesomerodon spinipenne</taxonomicName>
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Ohs., MNHUB Berlin" (typeset), c) our yellow paratype label. Paratype female at ZMHB with label data: a) "O. Ecuador, Puyo, A. Schulze" (typeset), b) "Ohaus determ.,
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<taxonomicName id="9E29E8A8EB295B123F04F9A11E5BED53" class="Insecta" family="Scarabaeidae" genus="Mesomerodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesomerodon spinipenne" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="spinipenne">Mesomerodon spinipenne</taxonomicName>
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♀ Ohs." (typeset and handwritten), c) our yellow paratype label. Paratype female at ZMHB with label data: a) "O. Ecuador, Puyo, A. Schulze" (typeset), b) "
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<taxonomicName id="B15A989D14DFBB96BCBBABF9B8E83609" class="Insecta" family="Scarabaeidae" genus="Mesomerodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesomerodon spinipenne" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="spinipenne">Mesomerodon spinipenne</taxonomicName>
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Ohs." (handwritten), c)
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<normalizedToken id="416146419DA9EBA0DCB9529A28397F4E" originalValue="“♀”">"♀"</normalizedToken>
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(typeset), d) our yellow paratype label. Paratype male at CCECL with label data: a) ECUADOR (Orellana), Payamino Research Station, 0°29'967"S, 77°17'083"W, 300m Tropical Rainforest, At M.V. light at night, 30.
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<normalizedToken id="58EDC2C726EA48C97A8A4B61BFE0476D" originalValue="vii–">vii-</normalizedToken>
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12 viii.2007" (typeset), b) male genitalia card mounted, c) our yellow paratype label. Paratype male at MSPC with label data: a) ECUADOR (Orellana), Payamino Research Station, 0°29'967"S, 77°17'083"W, 300m Tropical Rainforest, At M.V. light at night, 30.vii -12 viii.2007" (typeset), b) "coll. CPDT Gillett, BMNH(E) 2007-65" (typeset), c) "DNA extract No: Meso1, deposited at MSPC" (handwritten), d) male genitalia card mounted, e) "Matthias Seidel Collection 2016", f) our yellow paratype label. Paratypes (2 males) at BMNH with label data: a) ECUADOR (Orellana), Payamino Research Station, 0°29'967"S, 77°17'083"W, 300m Tropical Rainforest, At M.V. light at night, 30.vii -12 viii.2007" (typeset), b) "coll. CPDT Gillett, BMNH(E) 2007-65" (typeset), c) "DNA extract No: Meso2, deposited at MSPC; no PCR amplification" (handwritten), d) male genitalia card mounted, e) our yellow paratype label. Paratype male at SLCC with label data: a) "PARAGUAY: Alto Parana, 1.XI.1990" (typeset), b) "Collection S. Le Tirant" (typeset), c) "supposedly mislabeled locality, des. Seidel 2016" (typeset), d) "MESOMERODON SPINIPENNE OHAUS, det. M.E. Jameson 2003" (typeset and handwritten), d) male genitalia card mounted, e) our yellow paratype
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<pageBreakToken id="27316177F144F3444060B51964F86F9B" pageId="7" pageNumber="68" start="start">label</pageBreakToken>
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. Paratype male at IRSNB with label data: a) "ECUADOR (Orellana), Payamino Territory, 300m Tropical Rainforest, July 2007" (typeset), b) "DNA extract: Rut105, det. M.Seidel 2016" (typeset and handwritten), c) male genitalia card mounted, d) our yellow paratype label. Paratype male at ZSM with label data: a) "O.Ecuador, Puyo, A. Schulze, 6.V.35" (typeset and handwritten), b) "
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<taxonomicName id="07139C8567CACD2C9237F5FF4CEF672E" class="Insecta" family="Scarabaeidae" genus="Mesomerodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesomerodon spinipenne" order="Coleoptera" pageId="7" pageNumber="68" phylum="Arthropoda" rank="species" species="spinipenne">Mesomerodon spinipenne</taxonomicName>
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Ohs." (handwritten), c) "Staatssammlung
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<normalizedToken id="7BDE3930848D7EAB7A966B81CB92F583" originalValue="München">Muenchen</normalizedToken>
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, 1975, Erwerb Coll. Machatschke", d) male genitalia card mounted, e) our yellow paratype label. Paratype female at NMPC with label data: a) "O.Ecuador, Puyo, A. Schulze, 6.V.35" (typeset and handwritten), b) "Ohaus determ.,
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<taxonomicName id="79AA95471BA5BB8FB4A0CC8B505E3A20" class="Insecta" family="Scarabaeidae" genus="Mesomerodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesomerodon spinipenne" order="Coleoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="species" species="spinipenne">
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<pageBreakToken id="A728884324097D7CDCC4044A2C0789A5" pageId="8" pageNumber="69" start="start">Mesomerodon</pageBreakToken>
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spinipenne
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</taxonomicName>
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♀ Ohs." (typeset and handwritten), c) "Staatssammlung
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<normalizedToken id="17F5D2D01D73CF1927A11113337A2361" originalValue="München">Muenchen</normalizedToken>
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, 1975, Erwerb Coll. Machatschke", d) our yellow paratype label. Paratype female at ZSM with label data: a) "O.Ecuador, Sarayacu Feyer" (typeset), b) "Paratypus,
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<taxonomicName id="B9C933B5D5D10637AADC5588B489619D" class="Insecta" family="Scarabaeidae" genus="Mesomerodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesomerodon spinipenne" order="Coleoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="species" species="spinipenne">Mesomerodon spinipenne</taxonomicName>
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Cotype ♀ Ohs." (typeset and handwritten), c) "Staatssammlung
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<normalizedToken id="6B0F2C275FECAD56D08435039768AAA2" originalValue="München">Muenchen</normalizedToken>
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, 1975, Erwerb Coll. Machatschke", d) "NOT a type specimen of
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<taxonomicName id="7DEA4D4A973EAA0B9A778C53EC73FD63" class="Insecta" family="Scarabaeidae" genus="Mesomerodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesomerodon spinipenne" order="Coleoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="species" species="spinipenne">Mesomerodon spinipenne</taxonomicName>
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, Ohaus, 1905, des. Seidel 2016" (typeset), e) our yellow paratype label. Paratype male at FMNH with label data: a) "ECUADOR: Pastaza; 300m confluence R. Macuma & R. Morona VII:17:1971, leg. B. Malkin" (typeset), b) "at light" (typeset), c) "on sand river bank" (typeset), d) male genitalia card mounted, f) our yellow paratype label. Paratype female at FMNH with label data: a) "ECUADOR: Pastaza; 300m confluence R. Macuma & R. Morona VII:17:1971, leg. B. Malkin" (typeset), b) "at light" (typeset), c) "on sand river bank" (typeset), d) "
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<taxonomicName id="EF7CAFFD46C2FAA5E0C27EA28B796108" class="Insecta" family="Scarabaeidae" genus="Mesomerodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesomerodon" order="Coleoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="genus">Mesomerodon</taxonomicName>
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spp??? DET. A.R.Hardy 1980" (typeset and handwritten), f) our yellow paratype label. Paratype female at FMNH with label data: a) "ECUADOR: Pastaza; 300m confluence R. Macuma & R. Morona VII:17:1971, leg. B. Malkin" (typeset), b) our yellow paratype label.
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</paragraph>
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</subSubSection>
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<subSubSection id="B39AECB7E5B71F8E6AF1C6D482E53E23" pageId="8" pageNumber="69" type="description">
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<paragraph id="48AB02A3EC7EC0BCC64D1CC29FBDAEF2" pageId="8" pageNumber="69">Description</paragraph>
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<paragraph id="B9E5091611CABEF50F143CABAE90056D" pageId="8" pageNumber="69">
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(based on 11 males and 9 females). The holotype does not differ significantly from the generic description and suffices for the description of this cryptic species (see
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<normalizedToken id="D455DCC94F387CB6AF7388A353C0F4D7" originalValue="“Remarks”">"Remarks"</normalizedToken>
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). Descriptive details specific to the holotype specimen are indicated. Length from apex of clypeus to apex of pygidium 18-22 mm (♂) (holotype: 19 mm) and 22-25 mm (♀); width at mid-elytra 11-12 mm (♂) (holotype: 11 mm) and 12-15 mm (♀). Color: Cream colored (holotype), tan, or ochre; ventral surface castaneous with weak metallic green reflections. Form: Elytral apices with one short spine (holotype) or swelling (females). Legs: Protarsomere 2 of male ventrally lacking well-defined striae at ventral apex (Fig. 25). Male genitalia (Fig. 22
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<normalizedToken id="93743D36EBE6836AC83B5E686F588456" originalValue="a–c">a-c</normalizedToken>
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): Parameres with elongate, narrow projection (=stem) and with longitudinal, impressed fissure (Fig. 22a); stem gradually and weakly broadened toward apex, subapex lacking paired, spinose projections; ventral sclerite of phallobase with surface concave, apex quadrate (Fig. 22c); lateral view diagnostic (Fig. 22b).
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</paragraph>
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</subSubSection>
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<subSubSection id="F640B438EF209CCCE1C3306E4A8DAD4D" pageId="8" pageNumber="69" type="diagnosis">
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<paragraph id="112023707071652C9A96151130911ECF" pageId="8" pageNumber="69">Diagnosis.</paragraph>
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<paragraph id="DBA495C8AF5728E0F94E32700B0462AE" pageId="8" pageNumber="69">
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Males of
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<taxonomicName id="A1313D3561DF25378CC1AA5C3DC4BFBC" class="Insecta" family="Scarabaeidae" genus="Mesomerodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesomerodon barclayi" order="Coleoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="species" species="barclayi">Mesomerodon barclayi</taxonomicName>
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sp. n. are differentiated from other
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<taxonomicName id="06EFBAF5B300BEA22A3818D7EACEC519" class="Insecta" family="Scarabaeidae" genus="Mesomerodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesomerodon" order="Coleoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="genus">Mesomerodon</taxonomicName>
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species by the following combination of characters: Protarsomere 2 ventrally lacking a striated region at the ventral apex (striated in
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<taxonomicName id="A4E9BCD7711BE2D9B4F4B09FFEBBB801" lsidName="M. spinipenne" pageId="8" pageNumber="69" rank="species" species="spinipenne">M. spinipenne</taxonomicName>
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; apical region lacking striae in
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<taxonomicName id="AD54577B245624A22F119A8877274A53" lsidName="M. gilletti" pageId="8" pageNumber="69" rank="species" species="gilletti">M. gilletti</taxonomicName>
|
||
[Figs 25-27]) and form of the male genitalia (Fig. 22 versus Fig. 24 in
|
||
<taxonomicName id="CC56E7E80DB185421BFC4B3F882F1B02" lsidName="M. spinipenne" pageId="8" pageNumber="69" rank="species" species="spinipenne">M. spinipenne</taxonomicName>
|
||
and Fig. 23 in
|
||
<taxonomicName id="F647354AC65B8BAD8E543DA4C6601EB6" lsidName="M. gilletti" pageId="8" pageNumber="69" rank="species" species="gilletti">M. gilletti</taxonomicName>
|
||
). Females can only be confidently determined when associated to male specimens from the same collecting event.
|
||
</paragraph>
|
||
<caption id="EC30841366465006CDE2F60837BA7740" pageId="8" pageNumber="69">
|
||
<paragraph id="0A69236983EE840708301CB00AC51B10" pageId="8" pageNumber="69">
|
||
Figures 25-27. Form of protarsomeres 2 to 4, ventral view, in
|
||
<taxonomicName id="C6C5FB9EF0352E36DBF0424CC8FED319" class="Insecta" family="Scarabaeidae" genus="Mesomerodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesomerodon" order="Coleoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="genus">Mesomerodon</taxonomicName>
|
||
species. 25
|
||
<taxonomicName id="F1BA456EAE637C793C5EC740B13D83B0" lsidName="M. barclayi" pageId="8" pageNumber="69" rank="species" species="barclayi">M. barclayi</taxonomicName>
|
||
sp. n., showing protarsomere 2 without stiate region at apex 26
|
||
<taxonomicName id="3F34B7B0CE85D53964830D2B5A07C569" lsidName="M. gilletti" pageId="8" pageNumber="69" rank="species" species="gilletti">M. gilletti</taxonomicName>
|
||
showing protarsomere 2 without striate region at apex 27
|
||
<taxonomicName id="0D2022DF437413FC2FEF7773D81A092D" lsidName="M. spinipenne" pageId="8" pageNumber="69" rank="species" species="spinipenne">M. spinipenne</taxonomicName>
|
||
showing protarsomere 2 with striate region apically.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="5FB505CCFDC3E77C4D914230896D9052" pageId="8" pageNumber="69" type="etymology">
|
||
<paragraph id="865F0253436CF8ABA782640AF52F4A50" pageId="8" pageNumber="69">Etymology.</paragraph>
|
||
<paragraph id="2520CAF8DF7E82BA40DD4676CDD56701" pageId="8" pageNumber="69">
|
||
It is our honor to dedicate this species to Max Barclay (Curator,
|
||
<taxonomicName id="C8D53CB3A44DB0D3289C16257D147FA5" lsidName="" order="Coleoptera" pageId="8" pageNumber="69" rank="order">Coleoptera</taxonomicName>
|
||
, Department of Entomology), who invited the first and second authors to the 1st Scarab Symposium at the BMNH in 2014 where cooperation on this work was initiated. Max Barclay has led the way in making biodiversity science more accessible to scientists and citizens alike.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E755CD17AD9CD9F5AB59D7508228DA73" pageId="8" pageNumber="69" type="distribution">
|
||
<paragraph id="F66D2DBD29EF4258C04B4C23C2C28C7A" pageId="8" pageNumber="69">Distribution</paragraph>
|
||
<paragraph id="2B908A04ADD09B0F7687B79DE9735E28" pageId="8" pageNumber="69">
|
||
(Fig. 18). Known from western (lowland) Amazonia areas in Ecuador and occurring in apparent sympatry with
|
||
<taxonomicName id="69A7F4A8FE4EDD06880BFA8A772593FF" lsidName="M. gilletti" pageId="8" pageNumber="69" rank="species" species="gilletti">M. gilletti</taxonomicName>
|
||
.
|
||
<bibRefCitation id="111620CCE3B5F6A6A2C9C0654C7E6153" author="Ohaus, F" journalOrPublisher="Genera Insectorum Fasc" pageId="16" pageNumber="77" pagination="1 - 172" title="Coleoptera lamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. T. 1: Tribus Rutelini" volume="199 A" year="1934">Ohaus (1934</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7BB0B0A40AB24523AB5AA23FDCE5FB21" pageId="8" pageNumber="69">1952</bibRefCitation>
|
||
) recorded
|
||
<taxonomicName id="165134555A1C7466F712499EFD916D82" lsidName="M. spinipenne" pageId="8" pageNumber="69" rank="species" species="spinipenne">M. spinipenne</taxonomicName>
|
||
from Sarayacu, Ecuador. We recovered these specimens in ZMHB and ZSM and identified them as belonging to
|
||
<taxonomicName id="C440AA30FB486F409E4600DE3D97905C" lsidName="M. barclayi" pageId="8" pageNumber="69" rank="species" species="barclayi">M. barclayi</taxonomicName>
|
||
sp. n.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="098D9E42BDCED60C575E0D8405CB6AB6" pageId="8" pageNumber="69" type="locality data">
|
||
<paragraph id="E322C36077470C9C36A1774A617DEBF6" pageId="8" pageNumber="69">Locality data</paragraph>
|
||
<paragraph id="1C09E54CC1A8307A19813941B281FA57" pageId="8" pageNumber="69">(Suppl. material 1). 20 specimens from 9 collections.</paragraph>
|
||
<paragraph id="D1E8D40FD5F2B1BA060B7387978D55A3" pageId="8" pageNumber="69">ECUADOR: Morona-Santiago, Orellana, Pastaza</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="49DC77CCC25D1100A184BB5CC91C7B60" pageId="8" pageNumber="69" type="temporal data">
|
||
<paragraph id="226435254BEF90B8644B664CDB2699FB" pageId="8" pageNumber="69">Temporal data.</paragraph>
|
||
<paragraph id="07876777B2A4F684D48643CC2DEDF4A3" pageId="8" pageNumber="69">Based on label data, this species is known to be active in the months of February, August, and November.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="02403205010DADBB008F262FD6BC28C1" pageId="8" pageNumber="69" type="natural history">
|
||
<paragraph id="BC302904D080DC19B226DC0189B80615" pageId="8" pageNumber="69">Natural history.</paragraph>
|
||
<paragraph id="50939388AAC2399113697ACDCDAFEA7C" pageId="8" pageNumber="69">
|
||
Based on label data, adult
|
||
<taxonomicName id="837447557239BC3EBE901C3E02E8D250" lsidName="M. barclayi" pageId="8" pageNumber="69" rank="species" species="barclayi">M. barclayi</taxonomicName>
|
||
sp. n. is usually collected at light, thus suggesting activity and feeding at night. Individuals probably occur throughout the year. Immature stages are unknown. Specimens have been recorded at an elevation of 300 m.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DAFC11ECA7C757E34EA93A18F2FF01D1" pageId="8" pageNumber="69" type="remarks">
|
||
<paragraph id="8068764C084965DC1EC8EF63B9880AD2" pageId="8" pageNumber="69">Remarks.</paragraph>
|
||
<paragraph id="4E3DCD7D284F6E2B3363470FA0FF8F61" pageId="8" pageNumber="69">
|
||
As with other cryptic species, the overall body form of
|
||
<taxonomicName id="B1FA75A5A4F6BCC6FA6D3B7B48CB3D1B" lsidName="M. barclayi" pageId="8" pageNumber="69" rank="species" species="barclayi">M. barclayi</taxonomicName>
|
||
sp. n. is similar to other species in the genus. The form of the male genitalia (Fig. 22 versus Figs 23-24) however, is sufficient for identification of
|
||
<taxonomicName id="5D0D2A0B8D9D7C20EB2F8C35BB8B58B1" lsidName="M. barclayi" pageId="8" pageNumber="69" rank="species" species="barclayi">M. barclayi</taxonomicName>
|
||
sp. n. When developing our species hypotheses, we examined this morphotype in terms of phenotypic, elevational, and seasonal variation within
|
||
<taxonomicName id="89D614AAA948012461642DC09D542753" lsidName="M. spinipenne" pageId="8" pageNumber="69" rank="species" species="spinipenne">M. spinipenne</taxonomicName>
|
||
or
|
||
<taxonomicName id="5D213E98BD3EA1674D3ADCAF2FDA2197" lsidName="M. gilletti" pageId="8" pageNumber="69" rank="species" species="gilletti">M. gilletti</taxonomicName>
|
||
, but the consistency in the form of the male genitalia led us to conclude that it is a justified species. Apparent sympatry (in location and phenology) with
|
||
<taxonomicName id="14B030530065B3C2BB44A0D56BA0E08A" lsidName="M. gilletti" pageId="8" pageNumber="69" rank="species" species="gilletti">M. gilletti</taxonomicName>
|
||
also led us to carefully examine this species pair. Again, we found that consistency in the male genitalia form was sufficient for diagnosis of both species. Females of
|
||
<taxonomicName id="7FAFBED12C1368C0C85433311C8A864C" lsidName="M. barclayi" pageId="8" pageNumber="69" rank="species" species="barclayi">M. barclayi</taxonomicName>
|
||
sp. n. cannot be identified unless associated with males from the same collecting event.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |