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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.983.52630" ID-GBIF-Dataset="d3b857f7-25c1-4b21-9d39-4510ab675fca" ID-PMC="PMC7655787" ID-Pensoft-Pub="1313-2970-983-1" ID-Pensoft-UUID="C890C7FD4B2D57A8B1A062305ED42D53" ID-PubMed="33244289" ID-ZooBank="7DED4CE2934C4539945F758930C927F9" ModsDocID="1313-2970-983-1" checkinTime="1604475085656" checkinUser="pensoft" docAuthor="Smith, Sarah M., Beaver, Roger A. &amp; Cognato, Anthony I." docDate="2020" docId="20212819FD6FD7566F3C7399C60B4732" docLanguage="en" docName="ZooKeys 983: 1-442" docOrigin="ZooKeys 983" docSource="http://dx.doi.org/10.3897/zookeys.983.52630" docTitle="Hadrodemius Wood 1980" docType="treatment" docVersion="4" id="C890C7FD4B2D57A8B1A062305ED42D53" lastPageNumber="1" masterDocId="C890C7FD4B2D57A8B1A062305ED42D53" masterDocTitle="A monograph of the Xyleborini (Coleoptera, Curculionidae, Scolytinae) of the Indochinese Peninsula (except Malaysia) and China" masterLastPageNumber="442" masterPageNumber="1" pageNumber="1" updateTime="1668169715782" updateUser="ExternalLinkService">
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<mods:title>A monograph of the Xyleborini (Coleoptera, Curculionidae, Scolytinae) of the Indochinese Peninsula (except Malaysia) and China</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Smith, Sarah M.</mods:namePart>
<mods:affiliation>Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-5173-3736</mods:nameIdentifier>
<mods:nameIdentifier type="email">camptocerus@gmail.com</mods:nameIdentifier>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Beaver, Roger A.</mods:namePart>
<mods:affiliation>161 / 2 Mu 5, Soi Wat Pranon, T. Donkaew, A. Maerim, Chiangmai 50180, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cognato, Anthony I.</mods:namePart>
<mods:affiliation>Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>983</mods:number>
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<mods:start>1</mods:start>
<mods:end>442</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.983.52630</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.983.52630</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-983-1</mods:identifier>
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<treatment ID-GBIF-Taxon="168754629" LSID="urn:lsid:plazi:treatment:20212819FD6FD7566F3C7399C60B4732" httpUri="http://treatment.plazi.org/id/20212819FD6FD7566F3C7399C60B4732" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<taxonomicName authority="Wood, 1980" authorityName="Wood" authorityYear="1980" class="Insecta" family="Curculionidae" genus="Hadrodemius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hadrodemius" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Hadrodemius Wood, 1980</taxonomicName>
</paragraph>
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<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Wood" authorityYear="1980" class="Insecta" family="Curculionidae" genus="Hadrodemius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hadrodemius" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Hadrodemius</taxonomicName>
Wood, 1980: 94.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="">
<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Blandford" authorityYear="1896" class="Insecta" family="Curculionidae" genus="Xyleborus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xyleborus globus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="globus">
<emphasis italics="true" pageId="0" pageNumber="1">Xyleborus globus</emphasis>
</taxonomicName>
Blandford, 1896b; original designation.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Wood" authorityYear="1980" class="Insecta" family="Curculionidae" genus="Hadrodemius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hadrodemius" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hadrodemius</emphasis>
</taxonomicName>
species are distinguished by their large size, 4.9-7.2 mm and stout (less than 2
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide) and hairy appearance; pronotal base and scutellar area ornamented with a dense tuft of hair associated with mesonotal mycangium; scutellum visible only on the basal slope of elytral bases; procoxae contiguous; and the pronotal lateral margins rounded.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="">
<paragraph pageId="0" pageNumber="1">Similar genera.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Wood" authorityYear="1980" class="Insecta" family="Curculionidae" genus="Hadrodemius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hadrodemius" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hadrodemius</emphasis>
</taxonomicName>
is closely related to
<taxonomicName authorityName="Ferrari" authorityYear="1867" class="Insecta" family="Curculionidae" genus="Anisandrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anisandrus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Anisandrus</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Sampson" authorityYear="1911" class="Insecta" family="Curculionidae" genus="Cnestus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cnestus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Cnestus</emphasis>
</taxonomicName>
, and
<taxonomicName authorityName="Reitter" authorityYear="1913" class="Insecta" family="Curculionidae" genus="Xylosandrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xylosandrus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Xylosandrus</emphasis>
</taxonomicName>
, all of which possess a mesonotal mycangium and the associated dense tuft of hair-like setae at the scutellar area and pronotal base (
<bibRefCitation DOI="https://doi.org/10.1111/evo.13219" author="Gohli, J" journalOrPublisher="Evolution" pageId="0" pageNumber="1" pagination="1258 - 1272" refId="B172" refString="Gohli, J, Kirkendall, LR, Smith, SM, Cognato, AI, Hulcr, J, Jordal, BH, 2017. Biological factors contributing to bark and ambrosia beetle species diversification. Evolution 71: 1258 - 1272, DOI: https://doi.org/10.1111/evo.13219" title="Biological factors contributing to bark and ambrosia beetle species diversification." url="https://doi.org/10.1111/evo.13219" volume="71" year="2017">Gohli et al. 2017</bibRefCitation>
; Johnson et al. 2018). These three genera is distinguished from
<taxonomicName authorityName="Wood" authorityYear="1980" class="Insecta" family="Curculionidae" genus="Hadrodemius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hadrodemius" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hadrodemius</emphasis>
</taxonomicName>
by their normal scutellum that is flush with the dorsal surface of the elytra rather than just visible only on the basal slope of the elytral bases.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Wood" authorityYear="1980" class="Insecta" family="Curculionidae" genus="Hadrodemius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hadrodemius" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hadrodemius</emphasis>
</taxonomicName>
occurs in tropical areas from India in the West, through the Oriental region to New Guinea and the Solomon Islands in the East.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="">
<paragraph pageId="0" pageNumber="1">Gallery system.</paragraph>
<paragraph pageId="0" pageNumber="1">
Usually constructed in small stems from 1.5 - 5.0 cm diameter, it comprises a circumferential entrance gallery leading to one to several longitudinal galleries (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2444.1.3" author="Beaver, RA" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="51 - 57" refId="B17" refString="Beaver, RA, 2010. A review of the genus Hadrodemius Wood, with new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 2444: 51 - 57, DOI: https://doi.org/10.11646/zootaxa.2444.1.3" title="A review of the genus Hadrodemius Wood, with new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae)." url="https://doi.org/10.11646/zootaxa.2444.1.3" volume="2444" year="2010">Beaver 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
Further details of the biology are given by
<bibRefCitation author="Browne, FG" journalOrPublisher="Malayan Forest Records" pageId="0" pageNumber="1" pagination="1 - 255" refId="B70" refString="Browne, FG, 1961b. The biology of Malayan Scolytidae and Platypodidae. Malayan Forest Records 22: 1 - 255" title="The biology of Malayan Scolytidae and Platypodidae." volume="22" year="1961 b">Browne (1961b)</bibRefCitation>
,
<bibRefCitation author="Kalshoven, LGE" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="1" pagination="135 - 173" refId="B220" refString="Kalshoven, LGE, 1959b. Studies on the biology of Indonesian Scolytoidea 4. Data on the habits of Scolytidae. Second part. Tijdschrift voor Entomologie 102: 135 - 173" title="Studies on the biology of Indonesian Scolytoidea 4. Data on the habits of Scolytidae. Second part." volume="102" year="1959 b">Kalshoven (1959b)</bibRefCitation>
and
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2444.1.3" author="Beaver, RA" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="51 - 57" refId="B17" refString="Beaver, RA, 2010. A review of the genus Hadrodemius Wood, with new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae). Zootaxa 2444: 51 - 57, DOI: https://doi.org/10.11646/zootaxa.2444.1.3" title="A review of the genus Hadrodemius Wood, with new synonymy and a key to species (Coleoptera: Curculionidae: Scolytinae)." url="https://doi.org/10.11646/zootaxa.2444.1.3" volume="2444" year="2010">Beaver (2010)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>