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205 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.147.1898" ID-GBIF-Dataset="5207e87b-8044-498d-a80d-7275387282d2" ID-PMC="PMC3286249" ID-Pensoft-Pub="1313-2970-147-277" ID-PubMed="22371667" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-147-277" ModsDocOrigin="ZooKeys 147" ModsDocTitle="Cladistic assessment of subtribal affinities within the tribe Moriomorphini with description of Rossjoycea glacialis, gen. n. and sp. n. from the South Island, and revision of Meonochilus Liebherr and Marris from the North Island, New Zealand (Coleoptera, Carabidae)" checkinTime="1451249665881" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2011" docId="43ACA789D6F9E0C352E02CFE577D7066" docLanguage="en" docName="ZooKeys 147: 277-335" docOrigin="ZooKeys 147" docSource="http://dx.doi.org/10.3897/zookeys.147.1898" docTitle="Meonochilus eplicatus" docType="treatment" docVersion="3" lastPageNumber="316" masterDocId="554A3015B71F903E5450EF01FFFDC71C" masterDocTitle="Cladistic assessment of subtribal affinities within the tribe Moriomorphini with description of Rossjoycea glacialis, gen. n. and sp. n. from the South Island, and revision of Meonochilus Liebherr and Marris from the North Island, New Zealand (Coleoptera, Carabidae)" masterLastPageNumber="335" masterPageNumber="277" pageNumber="312" updateTime="1668152665929" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Cladistic assessment of subtribal affinities within the tribe Moriomorphini with description of Rossjoycea glacialis, gen. n. and sp. n. from the South Island, and revision of Meonochilus Liebherr and Marris from the North Island, New Zealand (Coleoptera, Carabidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liebherr, James K.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>147</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>277</mods:start>
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<mods:end>335</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.147.1898</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.147.1898</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-147-277</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152032015" LSID="urn:lsid:plazi:treatment:43ACA789D6F9E0C352E02CFE577D7066" httpUri="http://treatment.plazi.org/id/43ACA789D6F9E0C352E02CFE577D7066" lastPageId="39" lastPageNumber="316" pageId="35" pageNumber="312">
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<subSubSection pageId="35" pageNumber="312" type="nomenclature">
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<paragraph pageId="35" pageNumber="312">
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5.
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<taxonomicName LSID="urn:lsid:zoobank.org:act:" authority="Broun" class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus eplicatus" order="Coleoptera" pageId="35" pageNumber="312" phylum="Arthropoda" rank="species" species="eplicatus">Meonochilus eplicatus (Broun)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="35" pageNumber="312" type="reference_group">
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<paragraph pageId="35" pageNumber="312">
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<taxonomicName class="Insecta" family="Carabidae" genus="Tarastethus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tarastethus eplicatus" order="Coleoptera" pageId="35" pageNumber="312" phylum="Arthropoda" rank="species" species="eplicatus">Tarastethus eplicatus</taxonomicName>
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<bibRefCitation author="Broun, T" journalOrPublisher="New Zealand Institute Bulletin" pageId="41" pageNumber="318" pagination="591 - 666" title="Descriptions of new genera and species of Coleoptera." volume="1" year="1923">Broun, 1923</bibRefCitation>
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: 675.
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</paragraph>
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<paragraph pageId="35" pageNumber="312">
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<taxonomicName class="Insecta" family="Carabidae" genus="Molopsida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Molopsida eplicatus" order="Coleoptera" pageId="35" pageNumber="312" phylum="Arthropoda" rank="species" species="eplicatus">Molopsida eplicatus</taxonomicName>
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,
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<bibRefCitation pageId="35" pageNumber="312">Britton, 1940</bibRefCitation>
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: 477.
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</paragraph>
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<paragraph pageId="35" pageNumber="312">
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax eplicatus" order="Coleoptera" pageId="35" pageNumber="312" phylum="Arthropoda" rank="species" species="eplicatus">Mecyclothorax eplicatus</taxonomicName>
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,
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<bibRefCitation pageId="35" pageNumber="312">
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Larochelle and
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<normalizedToken originalValue="Larivière">Lariviere</normalizedToken>
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, 2001
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</bibRefCitation>
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: 62.
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</paragraph>
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<paragraph pageId="35" pageNumber="312">
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<taxonomicName class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus eplicatus" order="Coleoptera" pageId="35" pageNumber="312" phylum="Arthropoda" rank="species" species="eplicatus">Meonochilus eplicatus</taxonomicName>
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,
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<bibRefCitation author="Liebherr, JK" journalOrPublisher="Tijdschrift voor Entomologie" pageId="41" pageNumber="318" title="Taxonomic revision of the Mecyclothorax beetles (Coleoptera: Carabidae) of Oahu: epithets as epitaphs for an endangered fauna? Systematic Entomology 34: 649 - 687." url="doi: 10.1111/j.1365-3113.2009.00477.x" year="2009 a">Liebherr and Marris, 2009</bibRefCitation>
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: 10.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="36" lastPageNumber="313" pageId="35" pageNumber="312" type="diagnosis">
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<paragraph pageId="35" pageNumber="312">Diagnosis.</paragraph>
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<paragraph lastPageId="36" lastPageNumber="313" pageId="35" pageNumber="312">
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Individuals of this species can be told by the broadly and moderately convex elytral intervals, and distinctly punctate striae (Fig. 10B)-though the punctures are less distinct than in
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<taxonomicName class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus rectus" order="Coleoptera" pageId="35" pageNumber="312" phylum="Arthropoda" rank="species" species="rectus">Meonochilus rectus</taxonomicName>
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or
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<taxonomicName class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus spiculatus" order="Coleoptera" pageId="35" pageNumber="312" phylum="Arthropoda" rank="species" species="spiculatus">Meonochilus spiculatus</taxonomicName>
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(Fig. 9B, 9C)-with the intervals covered with elongate transverse microsculpture that varies based on sex. Males exhibit glossy elytral intervals with remnants of microsculpture in the striae, whereas females exhibit more broadly distributed sculpticells over portions, or the entire surfaces of the intervals. The pronotal hind angles are denticulate with the basolateral margins on each side converging slightly immediately anterad the angles. The basolateral marginal bead is broadly convex inside the hind angles. The body is narrower than in
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<taxonomicName class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus placens" order="Coleoptera" pageId="35" pageNumber="312" phylum="Arthropoda" rank="species" species="placens">Meonochilus placens</taxonomicName>
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, with the pronotal base more constricted, and the elytral humeri less expanded laterally (Fig. 10B, 10C). The male aedeagal median lobe apex is broadly rounded and slightly asym
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<pageBreakToken pageId="36" pageNumber="313" start="start">metrically</pageBreakToken>
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expanded ventrally (Fig. 16A), but not nearly to the degree as observed in males of
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<taxonomicName class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus placens" order="Coleoptera" pageId="36" pageNumber="313" phylum="Arthropoda" rank="species" species="placens">Meonochilus placens</taxonomicName>
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(Fig. 16D). As in
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<taxonomicName class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus amplipennis" order="Coleoptera" pageId="36" pageNumber="313" phylum="Arthropoda" rank="species" species="amplipennis">Meonochilus amplipennis</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus placens" order="Coleoptera" pageId="36" pageNumber="313" phylum="Arthropoda" rank="species" species="placens">Meonochilus placens</taxonomicName>
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, the male aedeagal internal sac is membranous. Standardized body length 4.9-5.9 mm.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="37" lastPageNumber="314" pageId="36" pageNumber="313" type="male genitalia">
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<paragraph pageId="36" pageNumber="313">Male Genitalia.</paragraph>
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<paragraph lastPageId="37" lastPageNumber="314" pageId="36" pageNumber="313">
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(n = 4). Aedeagal median lobe with variable sagittal crest, from broad and deep to shallow and short, median lobe shaft with straight ventral margin, apex slightly downturned and broadly rounded (Fig. 16A); internal sac membranous, densely covered with fine microspicules, more heavily pigmented roll of membrane situated to right of gonopore; ventral right paramere stalked, length
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<pageBreakToken pageId="37" pageNumber="314" start="start">0.60</pageBreakToken>
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<normalizedToken originalValue="–0.66×">-0.66x</normalizedToken>
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distance from parameral articulation to outer face of apex, base thinner and apical half inflated, apex broadly rounded, apical half of ventral margin to tip lined with 7-15 setae, dorsal portion of tip with only several shorter setae accompanying ventral series, and dorsal edge glabrous or with 1 seta; dorsal left paramere broad, conchoid, length equal to 0.75
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<normalizedToken originalValue="–0.80×">-0.80x</normalizedToken>
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distance from parameral articulation to external face of apex, rounded apex with 2-3 larger setae interspersed with 2-3 small setae (Fig. 16B).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="37" pageNumber="314" type="female reproductive tract">
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<paragraph pageId="37" pageNumber="314">Female Reproductive Tract.</paragraph>
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<paragraph pageId="37" pageNumber="314">
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(n = 2) Bursa copulatrix elongate, narrowed apically to rounded apex, distance from between gonocoxal bases to apex 1.55
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<normalizedToken originalValue="×">x</normalizedToken>
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maximal bursal breadth (Fig. 17A); spermathecal gland duct entering bursa on left side of apex, the spermatheca not discernible among the membranous rolls of cuticle comprising bursal apex; spermathecal gland duct a short connective between stem of ductile reservoir and elongate, expanded basal tube that enters bursal wall; common oviduct ventrally joined to bursa, expanded to greatest breadth immediately distad juncture; basal gonocoxite 1 with lateroapical series of 1-2 setae, and mesal series of 8-10 setae in apical half of mesal surface (Fig. 17B); apical gonocoxite 2 broadly subtriangular, with 2 broad lateral ensiform setae and 1 dorsal ensiform seta on dorsomedial surface; apical sensory furrow bearing 2 nematiform setae at 0.80
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<normalizedToken originalValue="×">x</normalizedToken>
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coxite length.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="37" pageNumber="314" type="types">
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<paragraph pageId="37" pageNumber="314">Types.</paragraph>
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<paragraph pageId="37" pageNumber="314">
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Lectotype female (BMNH), mounted on unmarked white platen, labeled: Type (round, red-bordered label) // Pakarau. / 19.5.18. // New Zealand / Broun Coll. / Brit. Mus. / 1922 - 482. // Tarastethus / eplicatus // Lectotype ♀ / Tarastethus / eplicatus / Broun, 1923 / det. J.K.
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<bibRefCitation author="Liebherr, JK" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="41" pageNumber="318" pagination="15 - 76" title="The Mecyclothorax beetles (Coleoptera: Carabidae: Moriomorphini) of West Maui, Hawaii: taxonomy, biogeography, and conservation." url="doi: 10.1002/mmnd.201100005" volume="58" year="2011">Liebherr 2011</bibRefCitation>
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(black-bordered red label). Paralectotype female (BMNH), identically mounted and labeled, except for: Paratype (round yellow-bordered label) [first label] // Kaiangaroa / 12.5.1918 [second label] //
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<normalizedToken originalValue="…">...</normalizedToken>
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Paralectotype ♀
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<normalizedToken originalValue="…">...</normalizedToken>
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[fifth label].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="37" pageNumber="314" type="non-type material">
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<paragraph pageId="37" pageNumber="314">Non-type Material.</paragraph>
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<paragraph pageId="37" pageNumber="314">
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ND: Kara, Whangarei,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-35.720333">35°43.22'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="174.30017">174°18.01'E</geoCoordinate>
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, ix-1942, Fairburn (NZAC, 1); Kawakawa, Whangae Stream,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-35.377335">35°22.64'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="174.06467">174°03.88'E</geoCoordinate>
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, 14-xi-1994, Townsend (JITC, 1); Mangamuka Gorge Walkwy., pittrap/wet broadleaf for.,
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-35.183334">35°11'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="173.45">173°27'E</geoCoordinate>
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, 350 m el., 27-
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<normalizedToken originalValue="v–">v-</normalizedToken>
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29-vi-1999,
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<normalizedToken originalValue="Larivière">Lariviere</normalizedToken>
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/Larochelle (NZAC, 1); Mangawhai Gorge, under stone at night,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-36.068165">36°04.09'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="174.58366">174°35.02'E</geoCoordinate>
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, 25-i-1948, Pritchard (NZAC, 2); Ngaiotonga Sce. Res., litter,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-35.3175">35°19.05'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="174.256">174°15.36'E</geoCoordinate>
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, 20-i-1972, Ramsay, (NZAC, 1); Puketi St. For., litter,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-35.225334">35°13.52'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="173.74184">173°44.51'E</geoCoordinate>
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, 21-i-1972, Ramsay (NZAC, 2); Rangiahua,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-35.267166">35°16.03'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="173.61966">173°37.18'E</geoCoordinate>
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, 01-xi-1966, Cumber (NZAC, 1); Waimatenui,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-35.612">35°36.72'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="173.72366">173°43.42'E</geoCoordinate>
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, 06-i-1942, Clarke (AMNZ, 1), 09-iii-1961, Cumber (NZAC, 1); Waipoua St. For., litter,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-35.6095">35°36.57'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="173.53433">173°32.06'E</geoCoordinate>
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, 07-xii-1961, Townsend (NZAC, 1), sifted litter, 80/121,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-35.6095">35°36.57'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="173.53433">173°32.06'E</geoCoordinate>
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, 25-xi-1980, Kuschel (NZAC, 1); Waitangi St. For., litter and decayed wood, 81/117,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-35.253166">35°15.19'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="174.00934">174°00.56'E</geoCoordinate>
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, 02-xi-1981, Kuschel (NZAC, 1).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="39" lastPageNumber="316" pageId="37" pageNumber="314" type="distribution and habitat">
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<paragraph pageId="37" pageNumber="314">Distribution and Habitat.</paragraph>
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<paragraph lastPageId="38" lastPageNumber="315" pageId="37" pageNumber="314">
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Of the three Northland species,
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<taxonomicName class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus eplicatus" order="Coleoptera" pageId="37" pageNumber="314" phylum="Arthropoda" rank="species" species="eplicatus">Meonochilus eplicatus</taxonomicName>
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as currently circumscribed exhibits the broadest geographic range (Fig. 15). Though genitalic characters do not support dismemberment of this species, the variation in elytra microsculpture should be studied further with additional material from across the range.
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<pageBreakToken pageId="38" pageNumber="315" start="start">Specimens</pageBreakToken>
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collected to date have been from ground-level microhabitats, collected via pitfall trapping in wet broadleaf forest, from leaf litter, or under stones.
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</paragraph>
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<caption pageId="39" pageNumber="316">
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<paragraph pageId="39" pageNumber="316">
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<pageBreakToken pageId="39" pageNumber="316" start="start">Figure</pageBreakToken>
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17. Female reproductive tract structures,
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<taxonomicName class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus" order="Coleoptera" pageId="39" pageNumber="316" phylum="Arthropoda" rank="genus">Meonochilus</taxonomicName>
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spp., ventral view A female reproductive tract,
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<taxonomicName class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus eplicatus" order="Coleoptera" pageId="39" pageNumber="316" phylum="Arthropoda" rank="species" species="eplicatus">Meonochilus eplicatus</taxonomicName>
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B left gonocoxite,
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<taxonomicName class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus eplicatus" order="Coleoptera" pageId="39" pageNumber="316" phylum="Arthropoda" rank="species" species="eplicatus">Meonochilus eplicatus</taxonomicName>
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C female reproductive tract,
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<taxonomicName class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus placens" order="Coleoptera" pageId="39" pageNumber="316" phylum="Arthropoda" rank="species" species="placens">Meonochilus placens</taxonomicName>
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D left gonocoxite,
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<taxonomicName class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meonochilus placens" order="Coleoptera" pageId="39" pageNumber="316" phylum="Arthropoda" rank="species" species="placens">Meonochilus placens</taxonomicName>
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(see Table 1 for abbreviations).
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |