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<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFF9FFB75AB6F8DA40D9CEC4" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus nghiasonii V. S. Dang &amp; Naiki 2021" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571822" ID-Zenodo-Dep="7571822" LSID="urn:lsid:plazi:treatment:43284262FFF9FFB75AB6F8DA40D9CEC4" httpUri="http://treatment.plazi.org/id/43284262FFF9FFB75AB6F8DA40D9CEC4" lastPageId="105" pageId="102">
<subSubSection box="[136,1128,1919,1946]" pageId="102" type="nomenclature">
<paragraph blockId="102.[136,1451,1919,2053]" box="[136,1128,1919,1946]" pageId="102">
<emphasis bold="true" box="[136,171,1919,1945]" pageId="102">50.</emphasis>
<taxonomicName authority="V. S. Dang &amp; Naiki (2021: 34)" authorityName="V. S. Dang &amp; Naiki" authorityPageNumber="34" authorityYear="2021" box="[178,773,1919,1946]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="102" phylum="Tracheophyta" rank="species" species="nghiasonii">
<emphasis bold="true" box="[178,437,1919,1946]" italics="true" pageId="102">Lasianthus nghiasonii</emphasis>
V.S.Dang &amp; Naiki (2021: 34)
</taxonomicName>
<figureCitation box="[803,881,1919,1946]" captionStart="FIGURE 77" captionStartId="103.[136,229,1784,1806]" captionTargetBox="[167,1418,194,1758]" captionTargetId="figure-19@103.[165,1418,193,1760]" captionTargetPageId="103" captionText="FIGURE 77. Lasianthus nghiasonii V.S.Dang &amp; Naiki (from V4450, VNM): A. Branch with fruit, B. Stipules, C. Abaxial leaf surfaces, DE. Inflorescence showing side view flower buds, F. Flower, GH. Fruits, I. Seeds." figureDoi="http://doi.org/10.5281/zenodo.7571790" httpUri="https://zenodo.org/record/7571790/files/figure.png" pageId="102">Fig. 77</figureCitation>
(plate),
<figureCitation box="[975,1054,1919,1945]" captionStart="FIGURE 78" captionStartId="104.[136,229,1799,1821]" captionTargetBox="[176,1405,209,1763]" captionTargetId="figure-25@104.[165,1418,193,1775]" captionTargetPageId="104" captionText="FIGURE 78. Distribution maps of Lasianthus nghiasonii, L. nhatrangensis, L. ngoclinhensis, L. nuichuaensis and L. oblongilobus." figureDoi="http://doi.org/10.5281/zenodo.7571792" httpUri="https://zenodo.org/record/7571792/files/figure.png" pageId="102">Fig. 78</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection lastPageId="105" pageId="102" type="materials_examined">
<paragraph blockId="102.[136,1451,1919,2053]" pageId="102">
<emphasis bold="true" box="[136,194,1959,1980]" pageId="102">Type:</emphasis>
<materialsCitation box="[219,1451,1959,1981]" collectingDate="2016-02-26" collectorName="Tagane S. &amp; Dang V. S. &amp; Nguyen" country="Vietnam" elevation="1464" location="Bidoup Nui Ba National Park" pageId="102" specimenCount="1" stateProvince="Lam Dong">
<collectingCountry box="[219,332,1959,1980]" name="Vietnam" pageId="102">VIETNAM</collectingCountry>
.
<collectingRegion box="[341,539,1959,1981]" country="Vietnam" name="Lam Dong" pageId="102">Lam Dong Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFF9FFB75AB6F8DA40D9CEC4:CE5EA5AFFFF9FFB8581AF8024365CBF5" box="[548,835,1959,1981]" country="Vietnam" name="Bidoup Nui Ba National Park" pageId="102" stateProvince="Lam Dong">Bidoup Nui Ba National Park</location>
,
<quantity box="[845,919,1959,1981]" metricMagnitude="3" metricUnit="m" metricValue="1.464" pageId="102" unit="m" value="1464.0">
<elevation box="[845,919,1959,1981]" metricMagnitude="3" metricUnit="m" metricValue="1.464" pageId="102" unit="m" value="1464.0">1464 m</elevation>
</quantity>
elev.,
<date box="[981,1154,1959,1981]" pageId="102" value="2016-02-26">
<collectingDate box="[981,1154,1959,1981]" pageId="102" value="2016-02-26">26 February 2016</collectingDate>
</date>
,
<emphasis box="[1164,1451,1959,1980]" italics="true" pageId="102">
<collectorName box="[1164,1260,1959,1980]" pageId="102">Tagane S.</collectorName>
,
<collectorName box="[1271,1369,1959,1980]" pageId="102">Dang V.S.</collectorName>
,
<collectorName box="[1378,1451,1959,1980]" pageId="102">Nguyen</collectorName>
</emphasis>
</materialsCitation>
<materialsCitation collectionCode="VNM, FU" collectorName="Wai J. V" country="Japan" location="Iriomote Station" pageId="102" specimenCode="VNM00042919, VNM00042920, VNM00042921" specimenCount="1" typeStatus="holotype">
<emphasis box="[189,375,1995,2016]" italics="true" pageId="102">
<taxonomicNameLabel box="[189,232,1995,2016]" pageId="102" rank="variety">V.N.</taxonomicNameLabel>
,
<collectorName box="[243,327,1995,2016]" pageId="102">Wai J. V</collectorName>
4450
</emphasis>
(
<typeStatus box="[390,476,1995,2017]" pageId="102">holotype</typeStatus>
<collectionCode box="[483,544,1995,2016]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="102" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[560,720,1995,2016]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="102" type="Herbarium">VNM00042919</specimenCode>
]; isotype
<collectionCode box="[815,852,1995,2016]" country="China" name="Fudan University, Department of Biology" pageId="102">FU</collectionCode>
!,
<collectionCode box="[867,928,1995,2016]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="102" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[945,1102,1995,2016]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="102" type="Herbarium">VNM00042920</specimenCode>
,
<specimenCode box="[1113,1270,1995,2016]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="102" type="Herbarium">VNM00042921</specimenCode>
], the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFF9FFB75AB6F8DA40D9CEC4:CE5EA5AFFFF9FFB85A83F84A417CC44D" box="[189,346,2031,2053]" country="Japan" name="Iriomote Station" pageId="102">Iriomote Station</location>
(
<collectingCountry box="[359,418,2031,2052]" name="Japan" pageId="102">Japan</collectingCountry>
))
</materialsCitation>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7571790" ID-Zenodo-Dep="7571790" httpUri="https://zenodo.org/record/7571790/files/figure.png" pageId="103" startId="103.[136,229,1784,1806]" targetBox="[167,1418,194,1758]" targetPageId="103">
<paragraph blockId="103.[136,1452,1784,1842]" pageId="103">
<emphasis bold="true" box="[136,266,1784,1806]" pageId="103">FIGURE 77.</emphasis>
<taxonomicName authority="V. S. Dang &amp; Naiki" authorityName="V. S. Dang &amp; Naiki" box="[272,667,1784,1806]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="103" phylum="Tracheophyta" rank="species" species="nghiasonii">
<emphasis box="[272,485,1784,1805]" italics="true" pageId="103">Lasianthus nghiasonii</emphasis>
V.S.Dang &amp; Naiki
</taxonomicName>
(from
<emphasis box="[735,798,1784,1805]" italics="true" pageId="103">V4450</emphasis>
, VNM):
<emphasis bold="true" box="[887,911,1784,1805]" pageId="103">A.</emphasis>
Branch with fruit,
<emphasis bold="true" box="[1095,1118,1784,1805]" pageId="103">B.</emphasis>
Stipules,
<emphasis bold="true" box="[1214,1237,1784,1806]" pageId="103">C.</emphasis>
Abaxial leaf surfaces,
<emphasis bold="true" box="[136,188,1820,1841]" pageId="103">DE.</emphasis>
Inflorescence showing side view flower buds,
<emphasis bold="true" box="[639,658,1820,1841]" pageId="103">F.</emphasis>
Flower,
<emphasis bold="true" box="[743,799,1820,1842]" pageId="103">GH.</emphasis>
Fruits,
<emphasis bold="true" box="[872,887,1820,1841]" pageId="103">I.</emphasis>
Seeds.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7571792" ID-Zenodo-Dep="7571792" box="[136,1399,1799,1821]" httpUri="https://zenodo.org/record/7571792/files/figure.png" pageId="104" startId="104.[136,229,1799,1821]" targetBox="[176,1405,209,1763]" targetPageId="104">
<paragraph blockId="104.[136,1399,1799,1821]" box="[136,1399,1799,1821]" pageId="104">
<emphasis bold="true" box="[136,265,1799,1821]" pageId="104">FIGURE 78.</emphasis>
Distribution maps of
<taxonomicName box="[476,688,1799,1820]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="104" phylum="Tracheophyta" rank="species" species="nghiasonii">
<emphasis box="[476,688,1799,1820]" italics="true" pageId="104">Lasianthus nghiasonii</emphasis>
</taxonomicName>
,
<taxonomicName box="[701,862,1799,1821]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="104" phylum="Tracheophyta" rank="species" species="nhatrangensis">
<emphasis box="[701,862,1799,1821]" italics="true" pageId="104">L. nhatrangensis</emphasis>
</taxonomicName>
,
<emphasis box="[874,1194,1799,1821]" italics="true" pageId="104">
<taxonomicName box="[874,1033,1799,1821]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="104" phylum="Tracheophyta" rank="species" species="ngoclinhensis">L. ngoclinhensis</taxonomicName>
,
<taxonomicName box="[1043,1194,1799,1821]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="104" phylum="Tracheophyta" rank="species" species="nuichuaensis">L. nuichuaensis</taxonomicName>
</emphasis>
and
<taxonomicName box="[1241,1394,1799,1821]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="104" phylum="Tracheophyta" rank="species" species="oblongilobus">L. oblongilobus</taxonomicName>
.
</paragraph>
</caption>
<paragraph blockId="104.[136,1452,1867,2074]" lastBlockId="105.[136,1452,158,652]" lastPageId="105" pageId="104">
<emphasis bold="true" box="[136,397,1867,1893]" pageId="104">Specimens examined:</emphasis>
<materialsCitation collectingDate="2016-02-26" collectorName="Tagane S. &amp; Dang V. S. &amp; Nguyen" country="Vietnam" elevation="1464" latitude="12.171394" location="Bidoup Nui Ba National Park" longLatPrecision="1" longitude="108.69723" municipality="Central" pageId="104" specimenCount="1" stateProvince="Lam Dong">
<collectingCountry box="[426,558,1868,1894]" name="Vietnam" pageId="104">VIETNAM</collectingCountry>
.
<emphasis bold="true" box="[571,672,1867,1893]" pageId="104">
<collectingMunicipality box="[571,667,1867,1893]" pageId="104">Central</collectingMunicipality>
:
</emphasis>
<collectingRegion box="[682,918,1867,1894]" country="Vietnam" name="Lam Dong" pageId="104">Lam Dong Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFF9FFB75AB6F8DA40D9CEC4:CE5EA5AFFFF7FFB6599BF8EE4526CB2D" box="[933,1280,1867,1894]" country="Vietnam" latitude="12.171394" longLatPrecision="1" longitude="108.69723" municipality="Central" name="Bidoup Nui Ba National Park" pageId="104" stateProvince="Lam Dong">Bidoup Nui Ba National Park</location>
,
<quantity box="[1294,1383,1867,1894]" metricMagnitude="3" metricUnit="m" metricValue="1.464" pageId="104" unit="m" value="1464.0">
<elevation box="[1294,1383,1867,1894]" metricMagnitude="3" metricUnit="m" metricValue="1.464" pageId="104" unit="m" value="1464.0">1464 m</elevation>
</quantity>
elev.,
<geoCoordinate box="[136,314,1904,1930]" degrees="12" direction="north" minutes="10" orientation="latitude" pageId="104" precision="1" seconds="17.02" value="12.171394">12°1017.02N</geoCoordinate>
,
<geoCoordinate box="[332,522,1903,1930]" degrees="108" direction="east" minutes="41" orientation="longitude" pageId="104" precision="1" seconds="50.01" value="108.69723">108°4150.01E</geoCoordinate>
, in lower montane evergreen forest,
<date box="[964,1177,1903,1929]" pageId="104" value="2016-02-26">
<collectingDate box="[964,1177,1903,1929]" pageId="104" value="2016-02-26">26 February 2016</collectingDate>
</date>
,
<emphasis italics="true" pageId="104">
<collectorName box="[1194,1310,1904,1930]" pageId="104">Tagane S.</collectorName>
,
<collectorName box="[1328,1447,1904,1930]" pageId="104">Dang V.S.</collectorName>
,
<collectorName box="[136,222,1940,1965]" pageId="104">Nguyen</collectorName>
</emphasis>
</materialsCitation>
<materialsCitation collectingDate="2016-02-21" collectionCode="FU, VNM" collectorName="Wai J. V &amp; Tagane S. &amp; Nagamasu H. &amp; Naiki A. &amp; Dang V. S. &amp; Nguyen" country="Japan" elevation="1533" latitude="12.176131" location="Iriomote Station" longLatPrecision="1" longitude="108.68557" municipality="Central" pageId="104" specimenCount="1" stateProvince="Lam Dong">
<emphasis box="[228,436,1940,1966]" italics="true" pageId="104">
<taxonomicNameLabel box="[228,277,1940,1965]" pageId="104" rank="variety">V.N.</taxonomicNameLabel>
,
<collectorName box="[288,379,1940,1966]" pageId="104">Wai J. V</collectorName>
4450
</emphasis>
(
<collectionCode box="[450,490,1940,1966]" country="China" name="Fudan University, Department of Biology" pageId="104">FU</collectionCode>
,
<collectionCode box="[500,568,1940,1966]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="104" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFF9FFB75AB6F8DA40D9CEC4:CE5EA5AFFFF7FFB6593DF836439FCBE5" box="[771,953,1939,1965]" country="Japan" latitude="12.176131" longLatPrecision="1" longitude="108.68557" name="Iriomote Station" pageId="104">Iriomote Station</location>
(
<collectingCountry box="[967,1036,1940,1966]" name="Japan" pageId="104">Japan</collectingCountry>
)); in the same location,
<quantity box="[1302,1386,1939,1966]" metricMagnitude="3" metricUnit="m" metricValue="1.533" pageId="104" unit="m" value="1533.0">
<elevation box="[1302,1386,1939,1966]" metricMagnitude="3" metricUnit="m" metricValue="1.533" pageId="104" unit="m" value="1533.0">1533 m</elevation>
</quantity>
elev.,
<geoCoordinate box="[136,315,1976,2002]" degrees="12" direction="north" minutes="10" orientation="latitude" pageId="104" precision="1" seconds="34.07" value="12.176131">12°1034.07N</geoCoordinate>
,
<geoCoordinate box="[329,518,1976,2002]" degrees="108" direction="east" minutes="41" orientation="longitude" pageId="104" precision="1" seconds="08.04" value="108.68557">108°4108.04E</geoCoordinate>
, in lower montane evergreen forest,
<date box="[941,1147,1975,2002]" pageId="104" value="2016-02-21">
<collectingDate box="[941,1147,1975,2002]" pageId="104" value="2016-02-21">21 February 2016</collectingDate>
</date>
,
<emphasis italics="true" pageId="104">
<collectorName box="[1160,1273,1976,2002]" pageId="104">Tagane S.</collectorName>
,
<collectorName box="[1288,1447,1976,2001]" pageId="104">Nagamasu H.</collectorName>
,
<collectorName box="[136,229,2011,2038]" pageId="104">Naiki A.</collectorName>
,
<collectorName box="[240,352,2012,2038]" pageId="104">Dang V.S.</collectorName>
,
<collectorName box="[363,449,2012,2037]" pageId="104">Nguyen</collectorName>
</emphasis>
</materialsCitation>
<materialsCitation collectingDate="2014-11-25" collectingDateMax="2167-05" collectingDateMin="2014-11-25" collectionCode="FU, VNM" collectorName="Hoang T. B. &amp; Wai J. V &amp; Naiki A. &amp; Dang V. S. &amp; Tagane S. &amp; Nagamasu H. &amp; Tran H. &amp; Yang C. J. &amp; Hieu H. N. P. &amp; Dang" country="Japan" county="Iriomote Station" elevation="1438" lastPageId="105" latitude="12.118922" location="Khanh Hoa Province" longLatPrecision="1" longitude="108.94167" municipality="Iriomote Station" pageId="104" specimenCount="1" stateProvince="Toyama">
<emphasis box="[455,802,2012,2038]" italics="true" pageId="104">
<taxonomicNameLabel box="[455,504,2012,2037]" pageId="104" rank="variety">V.N.</taxonomicNameLabel>
,
<collectorName box="[514,644,2012,2037]" pageId="104">Hoang T.B.</collectorName>
,
<collectorName box="[654,746,2012,2038]" pageId="104">Wai J. V</collectorName>
3987
</emphasis>
(
<collectionCode box="[816,856,2012,2038]" country="China" name="Fudan University, Department of Biology" pageId="104">FU</collectionCode>
,
<collectionCode box="[866,934,2012,2038]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="104" type="Herbarium">VNM</collectionCode>
, the herbarium of
<collectingCounty box="[1136,1317,2011,2037]" pageId="104">Iriomote Station</collectingCounty>
(
<collectingCountry box="[1332,1401,2012,2038]" name="Japan" pageId="104">Japan</collectingCountry>
)); in the same location,
<quantity box="[342,426,2048,2074]" metricMagnitude="3" metricUnit="m" metricValue="1.438" pageId="104" unit="m" value="1438.0">
<elevation box="[342,426,2048,2074]" metricMagnitude="3" metricUnit="m" metricValue="1.438" pageId="104" unit="m" value="1438.0">1438 m</elevation>
</quantity>
elev., 12˚1023.01”N, 108˚4152.20E,
<date box="[877,1094,2047,2074]" pageId="104" value="2017-11-26">
<collectingDate box="[877,1094,2047,2074]" pageId="104" value="2017-11-26">26 November 2017</collectingDate>
</date>
, in moist evergreen forest,
<emphasis italics="true" lastPageId="105" pageId="104">
<collectorName lastPageId="105" pageId="104">Naiki A.</collectorName>
&amp;
<collectorName box="[194,304,159,185]" pageId="105">Dang V.S.</collectorName>
10101, 10102, 10103
</emphasis>
(
<collectionCode box="[559,629,158,184]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="105" type="Herbarium">VNM</collectionCode>
, the herbarium of
<collectingMunicipality box="[829,1010,158,184]" pageId="105">Iriomote Station</collectingMunicipality>
(
<collectingCountry box="[1024,1093,158,184]" name="Japan" pageId="105">Japan</collectingCountry>
)).
<location LSID="urn:lsid:plazi:treatment:43284262FFF9FFB75AB6F8DA40D9CEC4:CE5EA5AFFFF6FFB75E5FFF3B456ACCF0" box="[1121,1356,158,185]" country="Japan" county="Iriomote Station" latitude="12.118922" longLatPrecision="1" longitude="108.94167" municipality="Iriomote Station" name="Khanh Hoa Province" pageId="105" stateProvince="Toyama">Khanh Hoa Province</location>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFF9FFB75AB6F8DA40D9CEC4:CE5EA5AFFFF6FFB75F68FF3A4111CC95" country="Japan" county="Iriomote Station" latitude="12.118922" longLatPrecision="1" longitude="108.94167" municipality="Iriomote Station" name="Hon Ba Nature Reserve" pageId="105" stateProvince="Toyama">Hon Ba Nature Reserve</location>
,
<quantity box="[321,405,194,221]" metricMagnitude="3" metricUnit="m" metricValue="1.521" pageId="105" unit="m" value="1521.0">
<elevation box="[321,405,194,221]" metricMagnitude="3" metricUnit="m" metricValue="1.521" pageId="105" unit="m" value="1521.0">1521 m</elevation>
</quantity>
elev.,
<geoCoordinate box="[476,655,194,220]" degrees="12" direction="north" minutes="07" orientation="latitude" pageId="105" precision="1" seconds="01.82" value="12.117172">12°0701.82N</geoCoordinate>
,
<geoCoordinate box="[665,855,194,221]" degrees="108" direction="east" minutes="56" orientation="longitude" pageId="105" precision="1" seconds="43.63" value="108.94546">108°5643.63E</geoCoordinate>
, in evergreen forests,
<date box="[1097,1314,194,221]" pageId="105" value="2014-11-25">
<collectingDate box="[1097,1314,194,221]" pageId="105" value="2014-11-25">25 November 2014</collectingDate>
</date>
,
<emphasis italics="true" pageId="105">
<collectingRegion box="[1324,1412,195,220]" country="Japan" name="Toyama" pageId="105">Toyama</collectingRegion>
H.,
<collectorName box="[136,246,231,257]" pageId="105">Tagane S.</collectorName>
,
<collectorName box="[257,369,231,257]" pageId="105">Dang V.S.</collectorName>
,
<collectorName box="[380,535,231,256]" pageId="105">Nagamasu H.</collectorName>
,
<collectorName box="[545,637,230,257]" pageId="105">Naiki A.</collectorName>
,
<collectorName box="[648,737,231,256]" pageId="105">Tran H.</collectorName>
,
<collectorName box="[747,853,231,257]" pageId="105">Yang C.J.</collectorName>
with
<collectorName box="[913,1047,231,256]" pageId="105">Hieu H.N.P.</collectorName>
<date box="[1052,1125,230,256]" pageId="105" value="2167-05">
<collectingDate box="[1052,1125,230,256]" pageId="105" value="2167-05">V2167</collectingDate>
</date>
</emphasis>
(
<collectionCode box="[1139,1179,230,256]" country="China" name="Fudan University, Department of Biology" pageId="105">FU</collectionCode>
,
<collectionCode box="[1189,1257,231,257]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="105" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFF9FFB75AB6F8DA40D9CEC4:CE5EA5AFFFF6FFB75AB6FEAF4119CD6C" box="[136,319,266,292]" country="Japan" county="Iriomote Station" latitude="12.118922" longLatPrecision="1" longitude="108.94167" municipality="Iriomote Station" name="Iriomote Station" pageId="105" stateProvince="Toyama">Iriomote Station</location>
(
<collectingCountry box="[334,403,266,292]" name="Japan" pageId="105">Japan</collectingCountry>
)); in the same location,
<quantity box="[673,758,266,293]" metricMagnitude="3" metricUnit="m" metricValue="1.502" pageId="105" unit="m" value="1502.0">
<elevation box="[673,758,266,293]" metricMagnitude="3" metricUnit="m" metricValue="1.502" pageId="105" unit="m" value="1502.0">1502 m</elevation>
</quantity>
elev.,
<geoCoordinate box="[831,1010,266,292]" degrees="12" direction="north" minutes="07" orientation="latitude" pageId="105" precision="1" seconds="08.12" value="12.118922">12°0708.12N</geoCoordinate>
,
<geoCoordinate box="[1022,1211,266,293]" degrees="108" direction="east" minutes="56" orientation="longitude" pageId="105" precision="1" seconds="30.03" value="108.94167">108°5630.03E</geoCoordinate>
, in evergreen forests,
<date box="[136,288,302,329]" pageId="105" value="2019-06-15">
<collectingDate box="[136,288,302,329]" pageId="105" value="2019-06-15">15 June 2019</collectingDate>
</date>
,
<emphasis box="[299,535,302,329]" italics="true" pageId="105">
<collectorName box="[299,413,303,329]" pageId="105">Dang V.S.</collectorName>
,
<collectorName box="[424,486,303,328]" pageId="105">Dang</collectorName>
348
</emphasis>
(
<collectionCode box="[550,621,302,328]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="105" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="105" type="distribution">
<paragraph blockId="105.[136,1452,158,652]" pageId="105">
<emphasis bold="true" box="[189,345,338,364]" pageId="105">Distribution:</emphasis>
—Endemic to
<collectingCountry box="[508,603,338,364]" name="Vietnam" pageId="105">Vietnam</collectingCountry>
(known from the Bidoup Nui Ba National Park in
<collectingRegion box="[1176,1404,338,365]" country="Vietnam" name="Lam Dong" pageId="105">Lam Dong Province</collectingRegion>
and the Hon Ba Nature Reserve in
<collectingRegion box="[480,720,374,401]" country="Vietnam" name="Khanh Hoa" pageId="105">Khanh Hoa Province</collectingRegion>
).
</paragraph>
<paragraph blockId="105.[136,1452,158,652]" box="[189,1037,410,437]" pageId="105">
<emphasis bold="true" box="[189,423,410,436]" pageId="105">Habitat &amp; Ecology:</emphasis>
—Moist evergreen forest at around
<quantity box="[823,978,410,437]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="105" unit="m" value="1500.0" valueMax="1600.0" valueMin="1400.0">14001600 m</quantity>
elev.
</paragraph>
<paragraph blockId="105.[136,1452,158,652]" box="[189,934,446,473]" pageId="105">
<emphasis bold="true" box="[189,323,446,472]" pageId="105">Phenology:</emphasis>
—Flowering and fruiting from February to November.
</paragraph>
<paragraph blockId="105.[136,1452,158,652]" box="[189,745,482,511]" pageId="105">
<emphasis bold="true" box="[189,405,482,508]" pageId="105">Vernacular name:</emphasis>
—Xú hưƠng hoàng nghĩa sƠn.
</paragraph>
<paragraph blockId="105.[136,1452,158,652]" pageId="105">
<emphasis bold="true" box="[189,265,518,544]" pageId="105">Notes:</emphasis>
<taxonomicName box="[294,544,518,544]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="105" phylum="Tracheophyta" rank="species" species="nghiasonii">
<emphasis box="[294,544,518,544]" italics="true" pageId="105">Lasianthus nghiasonii</emphasis>
</taxonomicName>
is similar to
<taxonomicName authority="Blume (1826: 998)" authorityName="Blume" authorityPageNumber="998" authorityYear="1826" box="[699,1058,518,544]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="105" phylum="Tracheophyta" rank="species" species="inodorus">
<emphasis box="[699,831,518,544]" italics="true" pageId="105">L. inodorus</emphasis>
Blume (1826: 998)
</taxonomicName>
in the shape of leaves, number of secondary veins, and inflorescence
<typeStatus box="[532,583,555,581]" pageId="105">type</typeStatus>
, but differs from the latter in having longer petioles (
<quantity box="[1173,1294,554,581]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="105" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">1015 mm</quantity>
long vs.
<quantity metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="105" unit="mm" value="8.0" valueMax="10.0" valueMin="6.0">610 mm</quantity>
long), bracts
<quantity box="[333,425,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="105" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
long (vs.
<quantity box="[536,628,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="105" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">36 mm</quantity>
long) and larger calyx tubes (
<quantity box="[960,1054,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="105" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
long vs.
<quantity box="[1155,1220,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="105" unit="mm" value="2.0">2 mm</quantity>
long) and fruit color when ripe.
</paragraph>
</subSubSection>
</treatment>
</document>