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<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFC9FF895AB6FB734402CC94" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus lacduongensis V. S. Dang &amp; Naiki 2021" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571768" ID-Zenodo-Dep="7571768" LSID="urn:lsid:plazi:treatment:43284262FFC9FF895AB6FB734402CC94" httpUri="http://treatment.plazi.org/id/43284262FFC9FF895AB6FB734402CC94" lastPageId="87" pageId="86">
<subSubSection box="[136,1170,1238,1265]" pageId="86" type="nomenclature">
<paragraph blockId="86.[136,1452,1238,1372]" box="[136,1170,1238,1265]" pageId="86">
<emphasis bold="true" box="[136,171,1238,1264]" pageId="86">40.</emphasis>
<taxonomicName authority="V. S. Dang &amp; Naiki (2021: 42)" authorityName="V. S. Dang &amp; Naiki" authorityPageNumber="42" authorityYear="2021" box="[178,815,1238,1265]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="86" phylum="Tracheophyta" rank="species" species="lacduongensis">
<emphasis bold="true" box="[178,479,1238,1265]" italics="true" pageId="86">Lasianthus lacduongensis</emphasis>
V.S.Dang &amp; Naiki (2021: 42)
</taxonomicName>
<figureCitation box="[845,923,1238,1265]" captionStart="FIGURE 65" captionStartId="88.[136,229,1794,1816]" captionTargetBox="[167,1418,194,1768]" captionTargetId="figure-25@88.[165,1418,193,1769]" captionTargetPageId="88" captionText="FIGURE 65. Lasianthus lacduongensis V.S.Dang &amp; Naiki (from V1840, VNM): A. Branch, B. Close-up of young branch and leaves, C. Stipules, D. Abaxial leaf surfaces, EF. Young fruits, HI. Mature fruit, J. Seeds." figureDoi="http://doi.org/10.5281/zenodo.7571752" httpUri="https://zenodo.org/record/7571752/files/figure.png" pageId="86">Fig. 65</figureCitation>
(plate),
<figureCitation box="[1017,1096,1238,1264]" captionStart="FIGURE 62" captionStartId="84.[136,229,1796,1818]" captionTargetBox="[176,1405,208,1763]" captionTargetId="figure-24@84.[165,1418,193,1773]" captionTargetPageId="84" captionText="FIGURE 62. Distribution maps of Lasianthus khanhvinhensis, L. konchurangensis, L. kontumensis and L. lacduongensis." figureDoi="http://doi.org/10.5281/zenodo.7571744" httpUri="https://zenodo.org/record/7571744/files/figure.png" pageId="86">Fig. 62</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection pageId="86" type="materials_examined">
<paragraph blockId="86.[136,1452,1238,1372]" pageId="86">
<emphasis bold="true" box="[136,194,1278,1299]" pageId="86">Type:</emphasis>
<materialsCitation collectingDate="1840-05" collectingDateMax="2014-02-19" collectingDateMin="1840-05" collectionCode="VNM, FU" collectorName="Toyama H. &amp; Tagane S. &amp; Dang V. S. &amp; Nagamasu H. &amp; Naiki A. &amp; Tran H. &amp; Yang C. J. &amp; Ngoc N. V. &amp; Cuong N. Q. &amp; Hieu H. N. P. &amp; Loi X. N." country="Vietnam" elevation="1490" location="Iriomote Station" municipality="Bidoup Nui Ba National Park" pageId="86" specimenCode="VNM00042910, VNM00042911, VNM00042912" specimenCount="1" stateProvince="Lam Dong" typeStatus="holotype">
<collectingCountry box="[219,332,1278,1299]" name="Vietnam" pageId="86">VIETNAM</collectingCountry>
.
<collectingRegion box="[342,541,1278,1300]" country="Vietnam" name="Lam Dong" pageId="86">Lam Dong Province</collectingRegion>
,
<collectingMunicipality box="[551,840,1278,1300]" pageId="86">Bidoup Nui Ba National Park</collectingMunicipality>
,
<quantity box="[850,924,1278,1299]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="86" unit="m" value="1490.0">
<elevation box="[850,924,1278,1299]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="86" unit="m" value="1490.0">1490 m</elevation>
</quantity>
elev.,
<date box="[988,1162,1278,1300]" pageId="86" value="2014-02-19">
<collectingDate box="[988,1162,1278,1300]" pageId="86" value="2014-02-19">19 February 2014</collectingDate>
</date>
,
<emphasis italics="true" pageId="86">
<collectorName box="[1173,1281,1279,1300]" pageId="86">Toyama H.</collectorName>
,
<collectorName box="[1291,1387,1278,1300]" pageId="86">Tagane S.</collectorName>
,
<collectorName pageId="86">Dang V.S.</collectorName>
,
<collectorName box="[240,379,1315,1336]" pageId="86">Nagamasu H.</collectorName>
,
<collectorName box="[391,475,1314,1335]" pageId="86">Naiki A.</collectorName>
,
<collectorName box="[488,569,1315,1336]" pageId="86">Tran H.</collectorName>
,
<collectorName box="[582,678,1314,1336]" pageId="86">Yang C.J.</collectorName>
with
<collectorName box="[737,840,1315,1336]" pageId="86">
Ngoc
<taxonomicNameLabel box="[797,840,1315,1336]" pageId="86" rank="variety">N.V.</taxonomicNameLabel>
</collectorName>
,
<collectorName box="[852,974,1314,1335]" pageId="86">Cuong N.Q.</collectorName>
,
<collectorName box="[986,1109,1314,1336]" pageId="86">Hieu H.N.P.</collectorName>
,
<collectorName box="[1121,1206,1314,1336]" pageId="86">Loi X.N.</collectorName>
<date box="[1214,1277,1314,1335]" pageId="86" value="1840-05">
<collectingDate box="[1214,1277,1314,1335]" pageId="86" value="1840-05">V1840</collectingDate>
</date>
</emphasis>
(
<typeStatus box="[1293,1379,1314,1336]" pageId="86">holotype</typeStatus>
<collectionCode box="[1387,1448,1314,1335]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="86" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[195,355,1350,1371]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="86" type="Herbarium">VNM00042910</specimenCode>
]; isotypes
<collectionCode box="[455,492,1350,1371]" country="China" name="Fudan University, Department of Biology" pageId="86">FU</collectionCode>
!,
<collectionCode box="[505,566,1350,1371]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="86" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[580,736,1350,1371]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="86" type="Herbarium">VNM00042911</specimenCode>
,
<specimenCode box="[746,903,1350,1371]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="86" type="Herbarium">VNM00042912</specimenCode>
], the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFC9FF895AB6FB734402CC94:CE5EA5AFFFC9FF885E02FAE344FEC914" box="[1084,1240,1350,1372]" country="Vietnam" municipality="Bidoup Nui Ba National Park" name="Iriomote Station" pageId="86" stateProvince="Lam Dong">Iriomote Station</location>
(Japan))
</materialsCitation>
.
</paragraph>
<paragraph blockId="86.[136,1452,1418,2093]" pageId="86">
<emphasis bold="true" box="[136,397,1418,1444]" pageId="86">
<collectingRegion box="[136,262,1418,1444]" pageId="86">Specimens</collectingRegion>
examined:
</emphasis>
<materialsCitation collectingDate="1840-05" collectingDateMax="2160-05" collectingDateMin="1840-05" collectionCode="FU, VNM" collectorName="Toyama H. &amp; Tagane S. &amp; Dang V. S. &amp; Nagamasu H. &amp; Naiki A. &amp; Tran H. &amp; Yang C. J. &amp; Ngoc N. V. &amp; Cuong N. Q. &amp; Hieu H. N. P. &amp; Loi X. N. &amp; Japan" country="Vietnam" county="Bidoup Nui Ba National Park" elevation="1490" latitude="12.117172" location="Iriomote Station" longLatPrecision="1" longitude="108.94546" municipality="Iriomote Station" pageId="86" specimenCount="1" stateProvince="Lam Dong">
<collectingCountry box="[426,558,1418,1444]" name="Vietnam" pageId="86">VIETNAM</collectingCountry>
.
<emphasis bold="true" box="[571,672,1418,1444]" pageId="86">Central:</emphasis>
<collectingRegion box="[682,918,1418,1445]" country="Vietnam" name="Lam Dong" pageId="86">Lam Dong Province</collectingRegion>
,
<collectingCounty box="[933,1280,1418,1445]" pageId="86">Bidoup Nui Ba National Park</collectingCounty>
,
<quantity box="[1294,1383,1418,1445]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="86" unit="m" value="1490.0">
<elevation box="[1294,1383,1418,1445]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="86" unit="m" value="1490.0">1490 m</elevation>
</quantity>
elev.,
<geoCoordinate box="[136,314,1454,1480]" degrees="12" direction="north" minutes="10" orientation="latitude" pageId="86" precision="1" seconds="26.91" value="12.174142">12°1026.91N</geoCoordinate>
,
<geoCoordinate box="[324,514,1454,1481]" degrees="108" direction="east" minutes="41" orientation="longitude" pageId="86" precision="1" seconds="53.94" value="108.69832">108°4153.94E</geoCoordinate>
, in evergreen forests,
<date box="[752,950,1454,1480]" pageId="86" value="2014-02-19">
<collectingDate box="[752,950,1454,1480]" pageId="86" value="2014-02-19">19 February 2014</collectingDate>
</date>
,
<emphasis italics="true" pageId="86">
<collectorName box="[959,1082,1455,1480]" pageId="86">Toyama H.</collectorName>
,
<collectorName box="[1091,1200,1455,1481]" pageId="86">Tagane S.</collectorName>
,
<collectorName box="[1211,1322,1455,1481]" pageId="86">Dang V.S.</collectorName>
,
<collectorName pageId="86">Nagamasu H.</collectorName>
,
<collectorName box="[176,268,1490,1517]" pageId="86">Naiki A.</collectorName>
,
<collectorName box="[278,366,1491,1516]" pageId="86">Tran H.</collectorName>
,
<collectorName box="[376,482,1491,1517]" pageId="86">Yang C.J.</collectorName>
with
<collectorName box="[541,655,1491,1516]" pageId="86">
Ngoc
<taxonomicNameLabel box="[606,655,1491,1516]" pageId="86" rank="variety">N.V.</taxonomicNameLabel>
</collectorName>
,
<collectorName box="[664,800,1491,1517]" pageId="86">Cuong N.Q.</collectorName>
,
<collectorName box="[810,947,1491,1516]" pageId="86">Hieu H.N.P.</collectorName>
,
<collectorName box="[956,1049,1491,1516]" pageId="86">Loi X.N.</collectorName>
<date box="[1054,1128,1490,1516]" pageId="86" value="1840-05">
<collectingDate box="[1054,1128,1490,1516]" pageId="86" value="1840-05">V1840</collectingDate>
</date>
</emphasis>
(
<collectionCode box="[1141,1181,1490,1516]" country="China" name="Fudan University, Department of Biology" pageId="86">FU</collectionCode>
,
<collectionCode box="[1190,1258,1491,1517]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="86" type="Herbarium">VNM</collectionCode>
, the herbarium of
<collectingMunicipality box="[136,319,1526,1552]" pageId="86">Iriomote Station</collectingMunicipality>
(Japan)); in the same location,
<quantity box="[673,758,1526,1553]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="86" unit="m" value="1490.0">
<elevation box="[673,758,1526,1553]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="86" unit="m" value="1490.0">1490 m</elevation>
</quantity>
elev.,
<geoCoordinate box="[831,1010,1526,1552]" degrees="12" direction="north" minutes="10" orientation="latitude" pageId="86" precision="1" seconds="26.91" value="12.174142">12°1026.91N</geoCoordinate>
,
<geoCoordinate box="[1022,1211,1526,1553]" degrees="108" direction="east" minutes="41" orientation="longitude" pageId="86" precision="1" seconds="53.94" value="108.69832">108°4153.94E</geoCoordinate>
, in evergreen forests,
<date box="[136,335,1562,1588]" pageId="86" value="2014-02-19">
<collectingDate box="[136,335,1562,1588]" pageId="86" value="2014-02-19">19 February 2014</collectingDate>
</date>
,
<emphasis italics="true" pageId="86">
<collectorName box="[345,468,1563,1588]" pageId="86">Toyama H.</collectorName>
,
<collectorName box="[478,588,1563,1589]" pageId="86">Tagane S.</collectorName>
,
<collectorName box="[599,711,1563,1589]" pageId="86">Dang V.S.</collectorName>
,
<collectorName box="[721,877,1563,1588]" pageId="86">Nagamasu H.</collectorName>
,
<collectorName box="[887,979,1562,1589]" pageId="86">Naiki A.</collectorName>
,
<collectorName box="[989,1078,1563,1588]" pageId="86">Tran H.</collectorName>
,
<collectorName box="[1088,1194,1563,1589]" pageId="86">Yang C.J.</collectorName>
with
<collectorName box="[1253,1367,1563,1588]" pageId="86">
Ngoc
<taxonomicNameLabel box="[1318,1367,1563,1588]" pageId="86" rank="variety">N.V.</taxonomicNameLabel>
</collectorName>
,
<collectorName pageId="86">Cuong N.Q.</collectorName>
,
<collectorName box="[203,340,1599,1624]" pageId="86">Hieu H.N.P.</collectorName>
,
<collectorName box="[351,445,1599,1624]" pageId="86">Loi X.N.</collectorName>
<date box="[451,525,1598,1624]" pageId="86" value="1930-05">
<collectingDate box="[451,525,1598,1624]" pageId="86" value="1930-05">V1930</collectingDate>
</date>
</emphasis>
(
<collectionCode box="[539,579,1598,1624]" country="China" name="Fudan University, Department of Biology" pageId="86">FU</collectionCode>
,
<collectionCode box="[589,657,1599,1625]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="86" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFC9FF895AB6FB734402CC94:CE5EA5AFFFC9FF885963F99B4435CA10" box="[861,1043,1598,1624]" country="Vietnam" county="Bidoup Nui Ba National Park" latitude="12.117172" longLatPrecision="1" longitude="108.94546" municipality="Iriomote Station" name="Iriomote Station" pageId="86" stateProvince="Lam Dong">Iriomote Station</location>
(Japan)).
<collectingRegion box="[1155,1392,1598,1625]" country="Vietnam" name="Khanh Hoa" pageId="86">Khanh Hoa Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFC9FF895AB6FB734402CC94:CE5EA5AFFFC9FF885F45F99A417BCA35" country="Vietnam" county="Bidoup Nui Ba National Park" latitude="12.117172" longLatPrecision="1" longitude="108.94546" municipality="Iriomote Station" name="Hon Ba Nature Reserve" pageId="86" stateProvince="Lam Dong">Hon Ba Nature Reserve</location>
,
<quantity box="[359,443,1634,1661]" metricMagnitude="3" metricUnit="m" metricValue="1.533" pageId="86" unit="m" value="1533.0">
<elevation box="[359,443,1634,1661]" metricMagnitude="3" metricUnit="m" metricValue="1.533" pageId="86" unit="m" value="1533.0">1533 m</elevation>
</quantity>
elev.,
<geoCoordinate box="[514,693,1634,1660]" degrees="12" direction="north" minutes="07" orientation="latitude" pageId="86" precision="1" seconds="01.82" value="12.117172">12°0701.82N</geoCoordinate>
,
<geoCoordinate box="[704,893,1634,1661]" degrees="108" direction="east" minutes="56" orientation="longitude" pageId="86" precision="1" seconds="43.63" value="108.94546">108°5643.63E</geoCoordinate>
, in evergreen forests,
<date box="[1136,1354,1634,1661]" pageId="86" value="2014-11-25">
<collectingDate box="[1136,1354,1634,1661]" pageId="86" value="2014-11-25">25 November 2014</collectingDate>
</date>
,
<emphasis italics="true" pageId="86">
<collectorName pageId="86">Toyama H.</collectorName>
,
<collectorName box="[176,286,1671,1697]" pageId="86">Tagane S.</collectorName>
,
<collectorName box="[298,410,1671,1697]" pageId="86">Dang V.S.</collectorName>
,
<collectorName box="[421,576,1671,1696]" pageId="86">Nagamasu H.</collectorName>
,
<collectorName box="[587,679,1670,1697]" pageId="86">Naiki A.</collectorName>
,
<collectorName box="[690,779,1671,1696]" pageId="86">Tran H.</collectorName>
,
<collectorName box="[789,896,1671,1697]" pageId="86">Yang C.J.</collectorName>
with
<collectorName box="[956,1092,1671,1697]" pageId="86">Cuong N.Q.</collectorName>
,
<collectorName box="[1102,1237,1671,1696]" pageId="86">Hieu H.N.P.</collectorName>
<date box="[1242,1315,1670,1696]" pageId="86" value="2160-05">
<collectingDate box="[1242,1315,1670,1696]" pageId="86" value="2160-05">V2160</collectingDate>
</date>
</emphasis>
(
<collectionCode box="[1329,1369,1670,1696]" country="China" name="Fudan University, Department of Biology" pageId="86">FU</collectionCode>
,
<collectionCode box="[1379,1447,1671,1697]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="86" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFC9FF895AB6FB734402CC94:CE5EA5AFFFC9FF885B74F90F4226CA8C" box="[330,512,1706,1732]" country="Vietnam" county="Bidoup Nui Ba National Park" latitude="12.117172" longLatPrecision="1" longitude="108.94546" municipality="Iriomote Station" name="Iriomote Station" pageId="86" stateProvince="Lam Dong">Iriomote Station</location>
(
<collectorName box="[527,596,1706,1732]" pageId="86">Japan</collectorName>
))
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="87" pageId="86" type="distribution">
<paragraph blockId="86.[136,1452,1418,2093]" pageId="86">
<emphasis bold="true" box="[189,345,1742,1768]" pageId="86">Distribution:</emphasis>
—Endemic to
<collectingCountry box="[504,599,1742,1768]" name="Vietnam" pageId="86">Vietnam</collectingCountry>
(known from the Hon Ba Nature Reserve in
<collectingRegion box="[1087,1319,1742,1769]" country="Vietnam" name="Khanh Hoa" pageId="86">Khanh Hoa Province</collectingRegion>
and Bidoup Nui Ba National Park in
<collectingRegion box="[414,646,1778,1805]" country="Vietnam" name="Lam Dong" pageId="86">Lam Dong Province</collectingRegion>
).
</paragraph>
<paragraph blockId="86.[136,1452,1418,2093]" box="[189,921,1814,1841]" pageId="86">
<emphasis bold="true" box="[189,423,1814,1840]" pageId="86">Habitat &amp; Ecology:</emphasis>
—In evergreen forests at
<quantity box="[707,862,1814,1841]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="86" unit="m" value="1400.0" valueMax="1600.0" valueMin="1200.0">12001600 m</quantity>
elev.
</paragraph>
<paragraph blockId="86.[136,1452,1418,2093]" box="[189,934,1850,1877]" pageId="86">
<emphasis bold="true" box="[189,323,1850,1876]" pageId="86">Phenology:</emphasis>
—Flowering and fruiting from February to November.
</paragraph>
<paragraph blockId="86.[136,1452,1418,2093]" box="[189,673,1886,1915]" pageId="86">
<emphasis bold="true" box="[189,405,1886,1912]" pageId="86">Vernacular name:</emphasis>
—Xú hưƠng lạc dưƠng.
</paragraph>
<paragraph blockId="86.[136,1452,1418,2093]" pageId="86">
<emphasis bold="true" box="[189,265,1922,1948]" pageId="86">Notes:</emphasis>
<taxonomicName box="[294,586,1922,1948]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="86" phylum="Tracheophyta" rank="species" species="lacduongensis">
<emphasis box="[294,586,1922,1948]" italics="true" pageId="86">Lasianthus lacduongensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName authority="H. Zhu (2012: 62)" authorityName="H. Zhu" authorityPageNumber="62" authorityYear="2012" box="[745,1208,1922,1948]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="86" phylum="Tracheophyta" rank="species" species="membranaceoideus">
<emphasis box="[745,995,1922,1948]" italics="true" pageId="86">L. membranaceoideus</emphasis>
H.
<bibRefCitation author="Zhu, H. &amp; Roos, M. C. &amp; Ridsdale, C. E." box="[1029,1208,1922,1948]" pageId="86" pagination="1 - 102" refId="ref55608" refString="Zhu, H., Roos, M. C. &amp; Ridsdale, C. E. (2012) A taxonomic revision of the Malesian species of Lasianthus (Rubiaceae). Blumea 57: 1 - 102. https: // doi. org / 10.3767 / 000651912 X 652012" type="journal article" year="2012">Zhu (2012: 62)</bibRefCitation>
</taxonomicName>
which is endemic to Borneo in having the shape of leaves and bracts inconspicuous, but differs from the latter in having sparsely tomentose on young branches (vs. glabrous), more secondary veins (810 on each side of midrib vs. 68 on each side of midrib), shorter petioles (
<quantity box="[323,417,2030,2057]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="86" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">57 mm</quantity>
long vs.
<quantity box="[519,626,2030,2057]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="86" unit="mm" value="8.0" valueMax="10.0" valueMin="6.0">610 mm</quantity>
long), and fruits with 2 (vs. 4) pyrenes.
</paragraph>
<paragraph blockId="86.[136,1452,1418,2093]" lastBlockId="87.[136,1451,158,220]" lastPageId="87" pageId="86">
<taxonomicName box="[189,482,2066,2092]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="86" phylum="Tracheophyta" rank="species" species="lacduongensis">
<emphasis box="[189,482,2066,2092]" italics="true" pageId="86">Lasianthus lacduongensis</emphasis>
</taxonomicName>
was previously recognized by Dang
<emphasis box="[914,973,2066,2092]" italics="true" pageId="86">et al.</emphasis>
(2019) based on the specimen
<emphasis box="[1337,1451,2066,2092]" italics="true" pageId="86">Dang 193</emphasis>
(VNM) as a new record of “
<taxonomicName authority="H. Zhu" authorityName="H. Zhu" authorityYear="2012" box="[444,769,158,184]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="87" phylum="Tracheophyta" rank="species" species="membranaceoideus">
<emphasis box="[444,690,158,184]" italics="true" pageId="87">L. membranaceoideus</emphasis>
H.Zhu
</taxonomicName>
” for the flora of
<collectingCountry box="[951,1050,158,184]" name="Vietnam" pageId="87">Vietnam</collectingCountry>
. However, after careful examination of morphological and molecular evidence, it is
<taxonomicName authority="V. S. Dang &amp; Naiki." authorityName="V. S. Dang &amp; Naiki." box="[665,1060,194,220]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="87" phylum="Tracheophyta" rank="species" species="lacduogensis">
<emphasis box="[665,840,194,220]" italics="true" pageId="87">L. lacduogensis</emphasis>
V.S.Dang &amp; Naiki.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>