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<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFC4FF855AB6FDEB4262C914" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus lancifolius Hook. f. 1880" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571778" ID-Zenodo-Dep="7571778" LSID="urn:lsid:plazi:treatment:43284262FFC4FF855AB6FDEB4262C914" httpUri="http://treatment.plazi.org/id/43284262FFC4FF855AB6FDEB4262C914" pageId="91">
<subSubSection box="[136,1023,590,617]" pageId="91" type="nomenclature">
<paragraph blockId="91.[136,1138,590,652]" box="[136,1023,590,617]" pageId="91">
<emphasis bold="true" box="[136,171,590,616]" pageId="91">42.</emphasis>
<taxonomicName ID-CoL="3SCN3" ID-ENA="1603731" authority="Hook. f. (1880: 187)" authorityName="Hook. f." authorityPageNumber="187" authorityYear="1880" box="[178,666,590,617]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="91" phylum="Tracheophyta" rank="species" species="lancifolius">
<emphasis bold="true" box="[178,437,590,617]" italics="true" pageId="91">Lasianthus lancifolius</emphasis>
Hook.f. (1880: 187)
</taxonomicName>
<figureCitation box="[697,775,590,616]" captionStart="FIGURE 68" captionStartId="92.[136,229,1768,1790]" captionTargetBox="[170,1415,194,1738]" captionTargetId="figure-24@92.[167,1418,193,1744]" captionTargetPageId="92" captionText="FIGURE 68. Lasianthus lancifolius Hook.f. (from Dang 272, VNM and the herbarium of Iriomote Station (Japan)): A. Branch, B. Closeup of expanding young leaves, C. Stipules, D. Abaxial leaf surfaces, E. Adaxial leaf surfaces." figureDoi="http://doi.org/10.5281/zenodo.7571762" httpUri="https://zenodo.org/record/7571762/files/figure.png" pageId="91">Fig. 68</figureCitation>
(plate),
<figureCitation box="[870,949,590,616]" captionStart="FIGURE 67" captionStartId="90.[136,229,1794,1816]" captionTargetBox="[180,1405,206,1757]" captionTargetId="figure-24@90.[165,1418,193,1769]" captionTargetPageId="90" captionText="FIGURE 67. Distribution maps of Lasianthus lamdongensis, L. lancifolius, L. larsenii, L. latifolius and L. lecomtei." figureDoi="http://doi.org/10.5281/zenodo.7571758" httpUri="https://zenodo.org/record/7571758/files/figure.png" pageId="91">Fig. 67</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection pageId="91" type="materials_examined">
<paragraph blockId="91.[136,1138,590,652]" box="[136,1138,630,652]" pageId="91">
<emphasis bold="true" box="[136,194,630,651]" pageId="91">Type:</emphasis>
<materialsCitation box="[219,1133,630,652]" collectorName="Hooker &amp; Thomson" country="India" location="INDIA" pageId="91" specimenCount="1" typeStatus="lectotype">
<collectingCountry box="[219,289,630,651]" name="India" pageId="91">INDIA</collectingCountry>
. Khasia,
<emphasis box="[377,598,630,651]" italics="true" pageId="91">
<collectorName box="[377,449,630,651]" pageId="91">Hooker</collectorName>
&amp;
<collectorName box="[480,568,630,651]" pageId="91">Thomson</collectorName>
12
</emphasis>
(
<typeStatus box="[610,700,630,652]" pageId="91">lectotype</typeStatus>
K! [K000031624], designated by
<bibRefCitation author="Deb, D. B. &amp; Gangopadhyay, M." box="[1029,1126,630,652]" pageId="91" pagination="265 - 308" refId="ref54255" refString="Deb, D. B. &amp; Gangopadhyay, M. (1991) Taxonomic study of the genus Lasianthus Jack (Rubiaceae) in India. Journal of Economic and Taxonomic Botany 15 (2): 265 - 308." type="journal article" year="1991">Deb 1991</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="91.[136,1452,698,1373]" pageId="91">
<emphasis bold="true" box="[136,393,698,724]" pageId="91">
<collectingRegion box="[136,262,698,724]" pageId="91">Specimens</collectingRegion>
examined:
</emphasis>
<materialsCitation collectingDate="2018-11-17" collectionCode="VNM" collectorName="Dang V. S. &amp; Naiki A. &amp; Dang &amp; Japan" country="Vietnam" county="City" elevation="1267" location="Iriomote Station" municipality="Ba Vi National Park" pageId="91" specimenCount="1" stateProvince="Ha Noi">
<collectingCountry box="[422,554,698,724]" name="Vietnam" pageId="91">VIETNAM</collectingCountry>
.
<emphasis bold="true" box="[563,684,698,724]" pageId="91">Northern:</emphasis>
<collectingRegion box="[691,772,698,725]" country="Vietnam" name="Ha Noi" pageId="91">Ha Noi</collectingRegion>
<collectingCounty box="[778,827,698,724]" pageId="91">City</collectingCounty>
,
<collectingMunicipality box="[838,1064,698,725]" pageId="91">Ba Vi National Park</collectingMunicipality>
,
<quantity box="[1074,1159,698,725]" metricMagnitude="3" metricUnit="m" metricValue="1.267" pageId="91" unit="m" value="1267.0">
<elevation box="[1074,1159,698,725]" metricMagnitude="3" metricUnit="m" metricValue="1.267" pageId="91" unit="m" value="1267.0">1267 m</elevation>
</quantity>
elev.,
<date box="[1230,1447,698,724]" pageId="91" value="2018-11-17">
<collectingDate box="[1230,1447,698,724]" pageId="91" value="2018-11-17">17 November 2018</collectingDate>
</date>
,
<emphasis box="[136,492,734,761]" italics="true" pageId="91">
<collectorName box="[136,246,735,761]" pageId="91">Dang V.S.</collectorName>
&amp;
<collectorName box="[279,371,734,761]" pageId="91">Naiki A.</collectorName>
,
<collectorName box="[382,444,735,760]" pageId="91">Dang</collectorName>
272
</emphasis>
(
<collectionCode box="[506,576,734,760]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="91" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFC4FF855AB6FDEB4262C914:CE5EA5AFFFC4FF855936FD7B439BCEB0" box="[776,957,734,760]" country="Vietnam" county="City" municipality="Ba Vi National Park" name="Iriomote Station" pageId="91" stateProvince="Ha Noi">Iriomote Station</location>
(Japan)); in the same location,
<quantity box="[1303,1387,734,761]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="91" unit="m" value="1210.0">
<elevation box="[1303,1387,734,761]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="91" unit="m" value="1210.0">1210 m</elevation>
</quantity>
elev.,
<date box="[136,355,770,796]" pageId="91" value="2018-11-17">
<collectingDate box="[136,355,770,796]" pageId="91" value="2018-11-17">17 November 2018</collectingDate>
</date>
,
<emphasis box="[366,744,770,797]" italics="true" pageId="91">
<collectorName box="[366,477,771,797]" pageId="91">Dang V.S.</collectorName>
&amp;
<collectorName box="[513,606,770,797]" pageId="91">Naiki A.</collectorName>
,
<collectorName box="[618,680,771,796]" pageId="91">Dang</collectorName>
272a
</emphasis>
(
<collectionCode box="[758,828,770,796]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="91" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFC4FF855AB6FDEB4262C914:CE5EA5AFFFC4FF855E36FCA74499CF54" box="[1032,1215,770,796]" country="Vietnam" county="City" municipality="Ba Vi National Park" name="Iriomote Station" pageId="91" stateProvince="Ha Noi">Iriomote Station</location>
(
<collectorName box="[1230,1299,770,796]" pageId="91">Japan</collectorName>
))
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="91" type="distribution">
<paragraph blockId="91.[136,1452,698,1373]" pageId="91">
<emphasis bold="true" box="[189,345,806,832]" pageId="91">Distribution:</emphasis>
—Widespread in tropical and subtropical regions in Asia (
<collectingCountry box="[1046,1181,806,832]" name="Bangladesh" pageId="91">Bangladesh</collectingCountry>
,
<collectingCountry box="[1198,1282,806,832]" name="Bhutan" pageId="91">Bhutan</collectingCountry>
,
<collectingCountry box="[1300,1370,806,832]" name="China" pageId="91">China</collectingCountry>
,
<collectingCountry box="[1387,1447,806,832]" name="India" pageId="91">India</collectingCountry>
,
<collectingCountry box="[136,241,842,868]" name="Thailand" pageId="91">Thailand</collectingCountry>
). In
<collectingCountry box="[289,387,842,868]" name="Vietnam" pageId="91">Vietnam</collectingCountry>
,
<taxonomicName box="[397,645,842,868]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="91" phylum="Tracheophyta" rank="species" species="lancifolius">
<emphasis box="[397,645,842,868]" italics="true" pageId="91">Lasianthus lancifolius</emphasis>
</taxonomicName>
occurs in
<collectingRegion box="[761,846,842,869]" country="Vietnam" name="Ha Noi" pageId="91">Ha Noi</collectingRegion>
,
<collectingRegion box="[857,955,842,868]" country="Vietnam" name="Phu Tho" pageId="91">Phu Tho</collectingRegion>
, and
<collectingRegion box="[1013,1135,842,868]" country="Vietnam" name="Vinh Phuc" pageId="91">Vinh Phuc</collectingRegion>
.
</paragraph>
<paragraph blockId="91.[136,1452,698,1373]" pageId="91">
<emphasis bold="true" box="[189,420,878,904]" pageId="91">Habitat &amp; Ecology:</emphasis>
—Growing under the shade of secondary forest at
<quantity box="[976,1130,878,905]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="91" unit="m" value="1150.0" valueMax="1300.0" valueMin="1000.0">10001300 m</quantity>
elev. This species is found in the forest where dominated by species
<taxonomicName authority="V. S. Dang &amp; Naiki (2020: 49)" authorityName="V. S. Dang &amp; Naiki" authorityPageNumber="49" authorityYear="2020" box="[566,1131,914,940]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="91" phylum="Tracheophyta" rank="species" species="thuyanae">
<emphasis box="[566,798,914,940]" italics="true" pageId="91">Lasianthus thuyanae</emphasis>
V.S.
<bibRefCitation author="Dang, V. S. &amp; Naiki, A." box="[845,1131,914,940]" pageId="91" pagination="49 - 54" refId="ref54210" refString="Dang, V. S. &amp; Naiki, A. (2020) Two new species of Lasianthus (Rubiaceae) from northern Vietnam. Annales Botanici Fennici 57: 49 - 54. https: // doi. org / 10.5735 / 085.057.0106" type="journal article" year="2020">Dang &amp; Naiki (2020: 49)</bibRefCitation>
</taxonomicName>
,
<taxonomicName authority="V. S. Dang &amp; Naiki (2020: 51)" authorityName="V. S. Dang &amp; Naiki" authorityPageNumber="51" authorityYear="2020" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="91" phylum="Tracheophyta" rank="species" species="tamdaoensis">
<emphasis box="[1142,1311,914,940]" italics="true" pageId="91">L. tamdaoensis</emphasis>
V.S.
<bibRefCitation author="Dang, V. S. &amp; Naiki, A." pageId="91" pagination="49 - 54" refId="ref54210" refString="Dang, V. S. &amp; Naiki, A. (2020) Two new species of Lasianthus (Rubiaceae) from northern Vietnam. Annales Botanici Fennici 57: 49 - 54. https: // doi. org / 10.5735 / 085.057.0106" type="journal article" year="2020">Dang &amp; Naiki (2020: 51)</bibRefCitation>
</taxonomicName>
,
<taxonomicName authority="Hance (1885: 324)" authorityName="Hance" authorityPageNumber="324" authorityYear="1885" box="[334,641,950,977]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="91" phylum="Tracheophyta" rank="species" species="fordii">
<emphasis box="[334,424,950,976]" italics="true" pageId="91">L. fordii</emphasis>
Hance (1885: 324)
</taxonomicName>
,
<taxonomicName box="[652,825,950,977]" class="Magnoliopsida" family="Phyllanthaceae" genus="Phyllanthus" kingdom="Plantae" order="Malpighiales" pageId="91" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[652,786,950,976]" italics="true" pageId="91">Phyllanthus</emphasis>
sp.
</taxonomicName>
,
<taxonomicName box="[837,1001,950,977]" class="Magnoliopsida" family="Melastomataceae" genus="Melastoma" kingdom="Plantae" order="Myrtales" pageId="91" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[837,961,950,976]" italics="true" pageId="91">Melastoma</emphasis>
sp.
</taxonomicName>
,
<taxonomicName authority="(Finet &amp; Gagnep.) Figlar (2000: 21)" authorityName="Figlar" authorityPageNumber="21" authorityYear="2000" baseAuthorityName="Finet &amp; Gagnep." class="Magnoliopsida" family="Magnoliaceae" genus="Magnolia" kingdom="Plantae" order="Magnoliales" pageId="91" phylum="Tracheophyta" rank="species" species="cavaleriei">
<emphasis box="[1012,1240,950,976]" italics="true" pageId="91">Magnolia cavaleriei</emphasis>
(Finet &amp; Gagnep.) Figlar (2000: 21)
</taxonomicName>
,
<taxonomicName authority="Hemsl. (1890: 422)" authorityName="Hemsl." authorityPageNumber="422" authorityYear="1890" box="[359,803,986,1013]" class="Magnoliopsida" family="Aristolochiaceae" genus="Asarum" kingdom="Plantae" order="Piperales" pageId="91" phylum="Tracheophyta" rank="species" species="maximum">
<emphasis box="[359,563,987,1013]" italics="true" pageId="91">Asarum maximum</emphasis>
Hemsl. (1890: 422)
</taxonomicName>
,
<taxonomicName authority="Blume (1851: 305)" authorityName="Blume" authorityPageNumber="305" authorityYear="1851" box="[820,1297,986,1013]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="91" phylum="Tracheophyta" rank="species" species="myrsinifolia">
<emphasis box="[820,1062,986,1013]" italics="true" pageId="91">Quercus myrsinifolia</emphasis>
Blume (1851: 305)
</taxonomicName>
,
<taxonomicName authority="Rehder &amp; E. H. Wilson (1916: 203)" authorityName="Rehder &amp; E. H. Wilson" authorityPageNumber="203" authorityYear="1916" class="Magnoliopsida" family="Fagaceae" genus="Castanopsis" kingdom="Plantae" order="Fagales" pageId="91" phylum="Tracheophyta" rank="species" species="fissa">
<emphasis italics="true" pageId="91">Castanopsis fissa</emphasis>
Rehder&amp;E.H.Wilson(1916:203)
</taxonomicName>
,
<taxonomicName authority="Lecomte (1913: 75)" authorityName="Lecomte" authorityPageNumber="75" authorityYear="1913" box="[574,1059,1022,1049]" class="Magnoliopsida" family="Lauraceae" genus="Cinnamomum" kingdom="Plantae" order="Laurales" pageId="91" phylum="Tracheophyta" rank="species" species="balansae">
<emphasis box="[574,835,1022,1049]" italics="true" pageId="91">Cinnamomum balansae</emphasis>
Lecomte(1913:75)
</taxonomicName>
,
<taxonomicName authority="Hu (1924: 228)" authorityName="Hu" authorityPageNumber="228" authorityYear="1924" box="[1066,1447,1022,1049]" class="Magnoliopsida" family="Magnoliaceae" genus="Magnolia" kingdom="Plantae" order="Magnoliales" pageId="91" phylum="Tracheophyta" rank="species" species="fordiana">
<emphasis box="[1066,1273,1022,1048]" italics="true" pageId="91">Magnolia fordiana</emphasis>
Hu(1924:228)
</taxonomicName>
,
<taxonomicName box="[136,355,1058,1084]" class="Magnoliopsida" family="Sapotaceae" genus="Madhuca" kingdom="Plantae" order="Ericales" pageId="91" phylum="Tracheophyta" rank="species" species="pasquieri">
<emphasis box="[136,355,1058,1084]" italics="true" pageId="91">Madhuca pasquieri</emphasis>
</taxonomicName>
H.J.Lam (1925: 182),
<taxonomicName authority="Seem. (1863: 226)" authorityName="Seem." authorityPageNumber="226" authorityYear="1863" box="[612,1066,1058,1084]" class="Magnoliopsida" family="Bignoniaceae" genus="Markhamia" kingdom="Plantae" order="Lamiales" pageId="91" phylum="Tracheophyta" rank="species" species="stipulata">
<emphasis box="[612,848,1058,1084]" italics="true" pageId="91">Markhamia stipulata</emphasis>
Seem. (1863: 226)
</taxonomicName>
,
<taxonomicName authority="Lecomte (1920: 346)" authorityName="Lecomte" authorityPageNumber="346" authorityYear="1920" class="Magnoliopsida" family="Sapotaceae" genus="Eberhardtia" kingdom="Plantae" order="Ericales" pageId="91" phylum="Tracheophyta" rank="species" species="tonkinensis">
<emphasis box="[1078,1346,1058,1084]" italics="true" pageId="91">Eberhardtia tonkinensis</emphasis>
Lecomte (1920: 346)
</taxonomicName>
.
</paragraph>
<paragraph blockId="91.[136,1452,698,1373]" box="[189,950,1130,1157]" pageId="91">
<emphasis bold="true" box="[189,323,1130,1156]" pageId="91">Phenology:</emphasis>
—Flowering and fruiting from September to December.
</paragraph>
<paragraph blockId="91.[136,1452,698,1373]" box="[189,699,1166,1195]" pageId="91">
<emphasis bold="true" box="[189,405,1166,1192]" pageId="91">Vernacular name:</emphasis>
—Xú hưƠng lá ngọn giáo.
</paragraph>
<paragraph blockId="91.[136,1452,698,1373]" pageId="91">
<emphasis bold="true" box="[189,265,1202,1228]" pageId="91">Notes:</emphasis>
<taxonomicName box="[294,545,1202,1228]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="91" phylum="Tracheophyta" rank="species" species="lancifolius">
<emphasis box="[294,545,1202,1228]" italics="true" pageId="91">Lasianthus lancifolius</emphasis>
</taxonomicName>
was described in 1880 by Hooker. It is characterized by having lanceolate to elliptic-lanceolate leaves, secondary veins 89 on each side of midrib, corolla puberulous outside and pubescent inside, lobes 5, and fruits globose,
<quantity box="[445,538,1274,1301]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="91" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">35 mm</quantity>
diam.
</paragraph>
<paragraph blockId="91.[136,1452,698,1373]" pageId="91">
In
<collectingCountry box="[223,321,1310,1336]" name="Vietnam" pageId="91">Vietnam</collectingCountry>
, we collected this species on the way to the highest peak in the montane natural forest of Ba Vi National Park at 1210 and
<quantity box="[435,520,1346,1373]" metricMagnitude="3" metricUnit="m" metricValue="1.267" pageId="91" unit="m" value="1267.0">1267 m</quantity>
elev.
</paragraph>
</subSubSection>
</treatment>
</document>