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<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFB1FFF05AB6FDEB43FCC980" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus chrysoneurus Miq. 1857" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571622" ID-Zenodo-Dep="7571622" LSID="urn:lsid:plazi:treatment:43284262FFB1FFF05AB6FDEB43FCC980" httpUri="http://treatment.plazi.org/id/43284262FFB1FFF05AB6FDEB43FCC980" pageId="46">
<subSubSection box="[136,1123,590,617]" pageId="46" type="nomenclature">
<paragraph blockId="46.[136,1452,590,904]" box="[136,1123,590,617]" pageId="46">
<emphasis bold="true" box="[136,171,590,616]" pageId="46">14.</emphasis>
<taxonomicName ID-CoL="3SCJF" ID-ENA="358834" authority="(Korth.) Miq. (1857: 322)" authorityName="Miq." authorityPageNumber="322" authorityYear="1857" baseAuthorityName="Korth." box="[178,769,590,617]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="46" phylum="Tracheophyta" rank="species" species="chrysoneurus">
<emphasis bold="true" box="[178,471,590,617]" italics="true" pageId="46">Lasianthus chrysoneurus</emphasis>
(Korth.) Miq. (1857: 322)
</taxonomicName>
<figureCitation box="[799,876,590,616]" captionStart="FIGURE 33" captionStartId="47.[136,229,1795,1817]" captionTargetBox="[171,1413,212,1767]" captionTargetId="figure-18@47.[166,1418,209,1770]" captionTargetPageId="47" captionText="FIGURE 33. Lasianthus chrysoneurus (Korth.) Miq. (AF from Dang 301 and G from V10916, VNM and HN): A. Branch with flower buds, B. Close-up of young branch and leaves, C. Stipules, D. Abaxial leaf surfaces, EF. Inflorescence showing side view flower buds and bracts, G. Young and mature fruits (G: photos by Shuichiro Tagane)." figureDoi="http://doi.org/10.5281/zenodo.7571650" httpUri="https://zenodo.org/record/7571650/files/figure.png" pageId="46">Fig. 33</figureCitation>
(plate),
<figureCitation box="[970,1049,590,616]" captionStart="FIGURE 32" captionStartId="45.[136,229,1781,1803]" captionTargetBox="[176,1410,204,1747]" captionTargetId="figure-17@45.[165,1418,193,1757]" captionTargetPageId="45" captionText="FIGURE 32. Distribution maps of Lasianthus chinensis, L. chrysoneurus, L. curtisii and L. dakglaiensis." figureDoi="http://doi.org/10.5281/zenodo.7571646" httpUri="https://zenodo.org/record/7571646/files/figure.png" pageId="46">Fig. 32</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection pageId="46" type="materials_examined">
<paragraph blockId="46.[136,1452,590,904]" box="[136,1225,630,652]" pageId="46">
<emphasis bold="true" box="[136,194,630,651]" pageId="46">Type:</emphasis>
<materialsCitation box="[219,1221,630,652]" country="Indonesia" location="Backer" pageId="46" specimenCount="1" stateProvince="Jawa Timur" typeStatus="neotype">
<collectingCountry box="[219,352,630,651]" name="Indonesia" pageId="46">INDONESIA</collectingCountry>
.
<collectingRegion box="[361,406,630,651]" country="Indonesia" name="Jawa Timur" pageId="46">Java</collectingRegion>
,
<collectingRegion box="[416,524,630,651]" country="Indonesia" name="Jawa Barat" pageId="46">Jawa Barat</collectingRegion>
, 1917,
<emphasis box="[594,727,630,651]" italics="true" pageId="46">
<location LSID="urn:lsid:plazi:treatment:43284262FFB1FFF05AB6FDEB43FCC980:CE5EA5AFFFB1FFF0586CFDD342B0CEC3" box="[594,662,630,651]" country="Indonesia" name="Backer" pageId="46" stateProvince="Jawa Timur">Backer</location>
22515
</emphasis>
(
<typeStatus box="[740,818,630,651]" pageId="46">neotype</typeStatus>
L! [L0306249], designated by Zhu 2002)
</materialsCitation>
.
</paragraph>
<paragraph blockId="46.[136,1452,590,904]" pageId="46">
<taxonomicName authority="Korth. (1851: 222)" authorityName="Korth." authorityPageNumber="222" authorityYear="1851" box="[136,572,666,688]" class="Magnoliopsida" family="Rubiaceae" genus="Mephitidia" kingdom="Plantae" order="Gentianales" pageId="46" phylum="Tracheophyta" rank="species" species="chrysoneura">
<emphasis box="[136,371,666,687]" italics="true" pageId="46">Mephitidia chrysoneura</emphasis>
<materialsCitation box="[381,572,666,688]" country="Indonesia" location="Backer" municipality="Type" pageId="46" specimenCount="1" stateProvince="Jawa Timur" typeStatus="neotype">Korth. (1851: 222)</materialsCitation>
</taxonomicName>
.
<collectingMunicipality box="[585,635,666,687]" pageId="46">Type</collectingMunicipality>
:—
<collectingCountry box="[662,797,666,687]" name="Indonesia" pageId="46">INDONESIA</collectingCountry>
.
<collectingRegion box="[809,854,666,687]" country="Indonesia" name="Jawa Timur" pageId="46">Java</collectingRegion>
,
<collectingRegion box="[868,979,666,687]" country="Indonesia" name="Jawa Barat" pageId="46">Jawa Barat</collectingRegion>
, 1917,
<emphasis box="[1057,1195,666,687]" italics="true" pageId="46">
<location LSID="urn:lsid:plazi:treatment:43284262FFB1FFF05AB6FDEB43FCC980:CE5EA5AFFFB1FFF05E1FFD3F4443CEE7" box="[1057,1125,666,687]" country="Indonesia" municipality="Type" name="Backer" pageId="46" stateProvince="Jawa Timur">Backer</location>
22515
</emphasis>
(
<typeStatus box="[1211,1289,666,687]" pageId="46">neotype</typeStatus>
L! [L0306249], designated by Zhu 2002).
</paragraph>
<paragraph blockId="46.[136,1452,590,904]" box="[136,1342,738,760]" pageId="46">
<taxonomicName authority="Hook. f. (1880: 183)" authorityName="Hook. f." authorityPageNumber="183" authorityYear="1880" box="[136,503,738,760]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="46" phylum="Tracheophyta" rank="species" species="kurzii">
<emphasis box="[136,302,738,759]" italics="true" pageId="46">Lasianthus kurzii</emphasis>
<materialsCitation box="[308,503,738,760]" country="Myanmar" location="Type" municipality="Chopedang" pageId="46" specimenCount="1" typeStatus="holotype">Hook.f. (1880: 183)</materialsCitation>
</taxonomicName>
.
<location LSID="urn:lsid:plazi:treatment:43284262FFB1FFF05AB6FDEB43FCC980:CE5EA5AFFFB1FFF0583EFD474217CEBF" box="[512,561,738,759]" country="Myanmar" municipality="Chopedang" name="Type" pageId="46">Type</location>
:—
<collectingCountry box="[588,718,738,759]" name="Myanmar" pageId="46">MYANMAR</collectingCountry>
.
<collectingMunicipality box="[727,839,738,760]" pageId="46">Chopedang</collectingMunicipality>
, 1827,
<emphasis box="[909,1058,738,759]" italics="true" pageId="46">Wall. Cat. 8310</emphasis>
(
<typeStatus box="[1071,1157,738,760]" pageId="46">holotype</typeStatus>
K! [K000031671].
</paragraph>
<paragraph blockId="46.[136,1452,590,904]" pageId="46">
<taxonomicName authority="Pierre ex Pit. (1924: 383)" authorityName="Pit." authorityPageNumber="383" authorityYear="1924" box="[136,584,774,796]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="46" phylum="Tracheophyta" rank="species" species="hoaensis">
<emphasis box="[136,332,774,795]" italics="true" pageId="46">Lasianthus hoaensis</emphasis>
<materialsCitation box="[338,584,774,796]" country="Vietnam" county="Type" location="Pierre" municipality="Bien Hoa" pageId="46" specimenCount="1" stateProvince="Dong Nai" typeStatus="syntype">Pierre ex Pit. (1924: 383)</materialsCitation>
</taxonomicName>
.
<collectingCounty box="[593,643,774,795]" pageId="46">Type</collectingCounty>
:—
<collectingCountry box="[671,785,774,795]" name="Vietnam" pageId="46">VIETNAM</collectingCountry>
.
<collectingRegion box="[794,983,774,796]" country="Vietnam" name="Dong Nai" pageId="46">Dong Nai Province</collectingRegion>
,
<collectingMunicipality box="[992,1086,774,796]" pageId="46">Bien Hoa</collectingMunicipality>
,
<emphasis box="[1096,1210,774,795]" italics="true" pageId="46">
<location LSID="urn:lsid:plazi:treatment:43284262FFB1FFF05AB6FDEB43FCC980:CE5EA5AFFFB1FFF05E76FCA344A2CF53" box="[1096,1156,774,795]" country="Vietnam" county="Type" municipality="Bien Hoa" name="Pierre" pageId="46" stateProvince="Dong Nai">Pierre</location>
1316
</emphasis>
(
<typeStatus box="[1223,1299,774,795]" pageId="46">syntype</typeStatus>
P!; isosyntypes BM!, E!, K!).
</paragraph>
<paragraph blockId="46.[136,1452,590,904]" pageId="46">
<taxonomicName authority="Pit. (1924: 383)" authorityName="Pit." authorityPageNumber="383" authorityYear="1924" box="[136,660,846,868]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="46" phylum="Tracheophyta" rank="variety" species="hoaensis" variety="microphyllus">
<emphasis box="[136,331,846,867]" italics="true" pageId="46">Lasianthus hoaensis</emphasis>
<materialsCitation box="[336,660,846,868]" collectingDate="1919-08-06" country="Cambodia" location="Type" municipality="Mont. de l'Lephant" pageId="46" specimenCount="1" typeStatus="lectotype">
var.
<emphasis box="[377,501,846,867]" italics="true" pageId="46">microphyllus</emphasis>
Pit. (1924: 383)
</materialsCitation>
</taxonomicName>
.
<location LSID="urn:lsid:plazi:treatment:43284262FFB1FFF05AB6FDEB43FCC980:CE5EA5AFFFB1FFF058A3FCEB42E9CF2B" box="[669,719,846,867]" country="Cambodia" municipality="Mont. de l'Lephant" name="Type" pageId="46">Type</location>
:—
<collectingCountry box="[747,881,846,867]" name="Cambodia" pageId="46">CAMBODIA</collectingCountry>
.
<collectingMunicipality box="[889,1077,846,868]" pageId="46">Mont. de lLephant</collectingMunicipality>
,
<date box="[1086,1227,846,868]" pageId="46" value="1919-08-06">
<collectingDate box="[1086,1227,846,868]" pageId="46" value="1919-08-06">6 August 1919</collectingDate>
</date>
,
<emphasis box="[1236,1350,846,867]" italics="true" pageId="46">Poilane s.n.</emphasis>
(
<typeStatus box="[1362,1452,846,868]" pageId="46">lectotype</typeStatus>
P! [P02285244], designated by
<bibRefCitation author="Napiroon, T. &amp; Chayamarit, K. &amp; Dawson, S. &amp; Till, W. &amp; Balslev, H." box="[493,695,882,904]" pageId="46" pagination="1 - 38" refId="ref54811" refString="Napiroon, T., Chayamarit, K., Dawson, S., Till, W. &amp; Balslev, H. (2020) A synopsis of Lasianthus (Lasiantheae, Rubiaceae) in Thailand and two additional new species. Phytotaxa 439 (1): 1 - 38. https: // doi. org / 10.11646 / phytotaxa. 439.1.1" type="journal article" year="2020">
Napiroon
<emphasis box="[589,638,882,903]" italics="true" pageId="46">et al.</emphasis>
2020
</bibRefCitation>
).
</paragraph>
<paragraph blockId="46.[136,1452,950,1481]" pageId="46">
<emphasis bold="true" box="[136,396,950,976]" pageId="46">
<collectingRegion box="[136,262,950,976]" pageId="46">Specimens</collectingRegion>
examined:
</emphasis>
<materialsCitation collectingDate="2019-05-09" collectionCode="VNM" collectorName="Dang V. S. &amp; Hoang N. S. &amp; Dang" country="Vietnam" elevation="810" location="Gia Nghia District" pageId="46" specimenCount="1" stateProvince="Dak Nong">
<collectingCountry box="[425,557,950,976]" name="Vietnam" pageId="46">VIETNAM</collectingCountry>
.
<emphasis bold="true" box="[570,671,950,976]" pageId="46">Central:</emphasis>
<collectingRegion box="[680,910,950,977]" country="Vietnam" name="Dak Nong" pageId="46">Dak Nong Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFB1FFF05AB6FDEB43FCC980:CE5EA5AFFFB1FFF059A2FC134455CF98" box="[924,1139,950,976]" country="Vietnam" name="Gia Nghia District" pageId="46" stateProvince="Dak Nong">Gia Nghia District</location>
,
<quantity box="[1153,1226,950,977]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="46" unit="m" value="810.0">
<elevation box="[1153,1226,950,977]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="46" unit="m" value="810.0">810 m</elevation>
</quantity>
elev.,
<date box="[1304,1447,950,977]" pageId="46" value="2019-05-09">
<collectingDate box="[1304,1447,950,977]" pageId="46" value="2019-05-09">9 May 2019</collectingDate>
</date>
,
<emphasis box="[136,537,986,1013]" italics="true" pageId="46">
<collectorName box="[136,247,987,1013]" pageId="46">Dang V.S.</collectorName>
&amp;
<collectorName box="[282,415,987,1013]" pageId="46">Hoang N.S.</collectorName>
,
<collectorName box="[426,488,987,1012]" pageId="46">Dang</collectorName>
301
</emphasis>
(
<collectionCode box="[552,623,986,1012]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="46" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2019-06-11" collectionCode="VNM" collectorName="Dang V. S. &amp; Hoang N. S. &amp; Dang" country="Vietnam" elevation="680" location="Ta Dung National Park" pageId="46" specimenCount="1" stateProvince="Central">
in the same location,
<location LSID="urn:lsid:plazi:treatment:43284262FFB1FFF05AB6FDEB43FCC980:CE5EA5AFFFB1FFF0594FFC7E4450CFBC" box="[881,1142,986,1013]" country="Vietnam" name="Ta Dung National Park" pageId="46" stateProvince="Central">Ta Dung National Park</location>
,
<quantity box="[1153,1224,986,1013]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="46" unit="m" value="680.0">
<elevation box="[1153,1224,986,1013]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="46" unit="m" value="680.0">680 m</elevation>
</quantity>
elev.,
<date box="[1296,1447,986,1013]" pageId="46" value="2019-06-11">
<collectingDate box="[1296,1447,986,1013]" pageId="46" value="2019-06-11">11 June 2019</collectingDate>
</date>
,
<emphasis box="[136,533,1022,1049]" italics="true" pageId="46">
<collectorName box="[136,246,1023,1049]" pageId="46">Dang V.S.</collectorName>
&amp;
<collectorName box="[280,412,1023,1049]" pageId="46">Hoang N.S.</collectorName>
,
<collectorName box="[422,484,1023,1048]" pageId="46">Dang</collectorName>
310
</emphasis>
(
<collectionCode box="[547,618,1022,1048]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="46" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation accessionNumber="V10916 &amp; V10994" collectingDate="2019-08" collectionCode="HN, KAG, MAK, VNM" collectorName="Oguri E. &amp; Tagane S. &amp; Sugai K. &amp; Suyama &amp; Bui" country="Vietnam" elevation="887" location="Kon Ka Kinh National Park" pageId="46" specimenCount="1" stateProvince="Gia Lai">
<collectingRegion box="[637,829,1022,1048]" country="Vietnam" name="Gia Lai" pageId="46">Gia Lai Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFB1FFF05AB6FDEB43FCC980:CE5EA5AFFFB1FFF05979FC5A4459C850" box="[839,1151,1022,1049]" country="Vietnam" name="Kon Ka Kinh National Park" pageId="46" stateProvince="Gia Lai">Kon Ka Kinh National Park</location>
,
<quantity box="[1162,1232,1022,1049]" metricMagnitude="2" metricUnit="m" metricValue="8.87" pageId="46" unit="m" value="887.0">
<elevation box="[1162,1232,1022,1049]" metricMagnitude="2" metricUnit="m" metricValue="8.87" pageId="46" unit="m" value="887.0">887 m</elevation>
</quantity>
elev.,
<date box="[1301,1447,1022,1048]" pageId="46" value="2019-08">
<collectingDate box="[1301,1447,1022,1048]" pageId="46" value="2019-08">August 2019</collectingDate>
</date>
,
<emphasis box="[136,950,1058,1085]" italics="true" pageId="46">
<collectorName box="[136,236,1059,1085]" pageId="46">Oguri E.</collectorName>
,
<collectorName box="[248,359,1059,1085]" pageId="46">Tagane S.</collectorName>
,
<collectorName box="[371,470,1059,1085]" pageId="46">Sugai K.</collectorName>
,
<collectorName box="[481,570,1059,1085]" pageId="46">Suyama</collectorName>
Y. &amp;
<collectorName box="[633,672,1059,1084]" pageId="46">Bui</collectorName>
H.Q.
<accessionNumber box="[740,950,1058,1085]" isEnumeration="true" pageId="46">V10916 &amp; V10994</accessionNumber>
</emphasis>
(
<collectionCode box="[965,1009,1058,1084]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="46" type="Herbarium">HN</collectionCode>
,
<collectionCode box="[1020,1083,1058,1084]" country="Japan" lsid="urn:lsid:biocol.org:col:15116" name="Kagoshima University Museum - Herbarium" pageId="46" type="Herbarium">KAG</collectionCode>
,
<collectionCode box="[1095,1163,1058,1084]" country="Japan" lsid="urn:lsid:biocol.org:col:14216" name="Tokyo Metropolitan University" pageId="46" type="Herbarium">MAK</collectionCode>
,
<collectionCode box="[1174,1244,1058,1084]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="46" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation collectingDate="1931-08-20" collectionCode="VNM" collectorName="Poilane" country="Vietnam" location="Nui Chua Chan" municipality="Xuan Loc District" pageId="46" specimenCount="1" stateProvince="Dong Nai">
<emphasis bold="true" box="[1262,1382,1058,1084]" pageId="46">Southern:</emphasis>
<collectingRegion country="Vietnam" name="Dong Nai" pageId="46">Dong Nai Province</collectingRegion>
,
<collectingMunicipality box="[298,504,1094,1121]" pageId="46">Xuan Loc District</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:43284262FFB1FFF05AB6FDEB43FCC980:CE5EA5AFFFB1FFF0583DFBE34293C828" box="[515,693,1094,1120]" country="Vietnam" municipality="Xuan Loc District" name="Nui Chua Chan" pageId="46" stateProvince="Dong Nai">Nui Chua Chan</location>
,
<date box="[705,885,1094,1120]" pageId="46" value="1931-08-20">
<collectingDate box="[705,885,1094,1120]" pageId="46" value="1931-08-20">20 August 1931</collectingDate>
</date>
,
<emphasis box="[898,1062,1094,1120]" italics="true" pageId="46">
<collectorName box="[898,985,1094,1120]" pageId="46">Poilane</collectorName>
19336
</emphasis>
(
<collectionCode box="[1078,1149,1094,1120]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="46" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="1931-08-24" collectionCode="VNM" collectorName="Poilane" country="Vietnam" elevation="750" location="Nui Chua Chan" municipality="Xuan Loc District" pageId="46" specimenCount="1" stateProvince="Southern">
in the same location,
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="46" unit="m" value="750.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="46" unit="m" value="750.0">750 m</elevation>
</quantity>
elev.,
<date box="[231,410,1130,1156]" pageId="46" value="1931-08-24">
<collectingDate box="[231,410,1130,1156]" pageId="46" value="1931-08-24">24 August 1931</collectingDate>
</date>
,
<emphasis box="[422,586,1130,1156]" italics="true" pageId="46">
<collectorName box="[422,509,1130,1156]" pageId="46">Poilane</collectorName>
18381
</emphasis>
(
<collectionCode box="[601,672,1130,1156]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="46" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="46" type="distribution">
<paragraph blockId="46.[136,1452,950,1481]" pageId="46">
<emphasis bold="true" box="[189,345,1166,1192]" pageId="46">Distribution:</emphasis>
—Widespread in tropical and subtropical regions in Asia (
<collectingCountry box="[988,1108,1166,1192]" name="Cambodia" pageId="46">Cambodia</collectingCountry>
,
<collectingCountry box="[1118,1188,1166,1192]" name="China" pageId="46">China</collectingCountry>
,
<collectingCountry box="[1197,1257,1166,1192]" name="India" pageId="46">India</collectingCountry>
,
<collectingCountry box="[1268,1380,1166,1192]" name="Indonesia" pageId="46">Indonesia</collectingCountry>
,
<collectingCountry box="[1390,1447,1167,1193]" name="Laos" pageId="46">Laos</collectingCountry>
,
<collectingCountry box="[136,247,1203,1229]" name="Myanmar" pageId="46">Myanmar</collectingCountry>
,
<collectingCountry box="[259,479,1202,1229]" name="Papua New Guinea" pageId="46">Papua New Guinea</collectingCountry>
,
<collectingCountry box="[491,596,1202,1228]" name="Thailand" pageId="46">Thailand</collectingCountry>
). In
<collectingCountry box="[645,743,1202,1228]" name="Vietnam" pageId="46">Vietnam</collectingCountry>
,
<taxonomicName box="[754,1036,1202,1228]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="46" phylum="Tracheophyta" rank="species" species="chrysoneurus">
<emphasis box="[754,1036,1202,1228]" italics="true" pageId="46">Lasianthus chrysoneurus</emphasis>
</taxonomicName>
occurs in
<collectingRegion box="[1154,1274,1202,1229]" country="Vietnam" name="Dak Nong" pageId="46">Dak Nong</collectingRegion>
,
<collectingRegion box="[1286,1399,1202,1229]" country="Vietnam" name="Dong Nai" pageId="46">Dong Nai</collectingRegion>
,
<collectingRegion country="Vietnam" name="Gia Lai" pageId="46">Gia Lai</collectingRegion>
, and
<collectingRegion box="[235,359,1239,1265]" country="Vietnam" name="Lam Dong" pageId="46">Lam Dong</collectingRegion>
.
</paragraph>
<paragraph blockId="46.[136,1452,950,1481]" box="[189,1418,1274,1301]" pageId="46">
<emphasis bold="true" box="[189,423,1274,1300]" pageId="46">Habitat &amp; Ecology:</emphasis>
—Growing in primary and secondary montane forests or open areas at
<quantity box="[1218,1359,1274,1301]" metricMagnitude="2" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="46" unit="m" value="800.0" valueMax="1000.0" valueMin="600.0">6001000 m</quantity>
elev.
</paragraph>
<paragraph blockId="46.[136,1452,950,1481]" box="[189,857,1310,1337]" pageId="46">
<emphasis bold="true" box="[189,323,1310,1336]" pageId="46">Phenology:</emphasis>
—Flowering and fruiting from May to October.
</paragraph>
<paragraph blockId="46.[136,1452,950,1481]" box="[189,656,1346,1375]" pageId="46">
<emphasis bold="true" box="[189,405,1346,1372]" pageId="46">Vernacular name:</emphasis>
—Xú hưƠng biên hòa.
</paragraph>
<paragraph blockId="46.[136,1452,950,1481]" pageId="46">
<emphasis bold="true" box="[189,265,1382,1408]" pageId="46">Notes:</emphasis>
<taxonomicName box="[294,575,1382,1408]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="46" phylum="Tracheophyta" rank="species" species="chrysoneurus">
<emphasis box="[294,575,1382,1408]" italics="true" pageId="46">Lasianthus chrysoneurus</emphasis>
</taxonomicName>
was clarified in the revisions by
<bibRefCitation author="Zhu, H. &amp; Taylor, C. M." box="[952,1191,1382,1408]" pageId="46" pagination="185 - 198" refId="ref55538" refString="Zhu, H. &amp; Taylor, C. M. (2011) Lasianthus. In: Wu, Z. Y., Raven, P. H. &amp; Hong, D. Y. (Eds.) Flora of China, vol. 19. Science Press, Beijing &amp; Missouri Botanical Garden Press, St. Louis, pp. 185 - 198." type="book chapter" year="2011">Zhu &amp; Taylor (2011)</bibRefCitation>
,
<bibRefCitation author="Zhu, H. &amp; Roos, M. C. &amp; Ridsdale, C. E." box="[1204,1399,1382,1408]" pageId="46" pagination="1 - 102" refId="ref55608" refString="Zhu, H., Roos, M. C. &amp; Ridsdale, C. E. (2012) A taxonomic revision of the Malesian species of Lasianthus (Rubiaceae). Blumea 57: 1 - 102. https: // doi. org / 10.3767 / 000651912 X 652012" type="journal article" year="2012">
Zhu
<emphasis box="[1257,1314,1382,1408]" italics="true" pageId="46">et al.</emphasis>
(2012)
</bibRefCitation>
, and
<bibRefCitation author="Napiroon, T. &amp; Chayamarit, K. &amp; Dawson, S. &amp; Till, W. &amp; Balslev, H." box="[136,387,1418,1444]" pageId="46" pagination="1 - 38" refId="ref54811" refString="Napiroon, T., Chayamarit, K., Dawson, S., Till, W. &amp; Balslev, H. (2020) A synopsis of Lasianthus (Lasiantheae, Rubiaceae) in Thailand and two additional new species. Phytotaxa 439 (1): 1 - 38. https: // doi. org / 10.11646 / phytotaxa. 439.1.1" type="journal article" year="2020">
Napiroon
<emphasis box="[248,303,1418,1444]" italics="true" pageId="46">et al.</emphasis>
(2020)
</bibRefCitation>
. This species is easily recognizable in the field in having appressed pubescent branches, stipules up to
<quantity box="[200,265,1454,1481]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="46" unit="mm" value="8.0">8 mm</quantity>
long, bracts
<quantity box="[409,474,1454,1481]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="46" unit="mm" value="5.0">5 mm</quantity>
long, and fruits subglobose, white when ripe.
</paragraph>
</subSubSection>
</treatment>
</document>