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<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FF1AFF585AB6FB5F44ABCD6C" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus verticillatus Merr. 1935" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571896" ID-Zenodo-Dep="7571896" LSID="urn:lsid:plazi:treatment:43284262FF1AFF585AB6FB5F44ABCD6C" httpUri="http://treatment.plazi.org/id/43284262FF1AFF585AB6FB5F44ABCD6C" lastPageId="134" pageId="133">
<subSubSection box="[136,1111,1274,1301]" pageId="133" type="nomenclature">
<paragraph blockId="133.[136,1452,1274,1552]" box="[136,1111,1274,1301]" pageId="133">
<emphasis bold="true" box="[136,171,1274,1300]" pageId="133">70.</emphasis>
<taxonomicName ID-CoL="6NYG2" ID-ENA="358839" authority="(Lour.) Merr. (1935: 372)" authorityName="Merr." authorityPageNumber="372" authorityYear="1935" baseAuthorityName="Lour." box="[178,742,1274,1301]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="133" phylum="Tracheophyta" rank="species" species="verticillatus">
<emphasis bold="true" box="[178,449,1274,1301]" italics="true" pageId="133">Lasianthus verticillatus</emphasis>
(Lour.) Merr. (1935: 372)
</taxonomicName>
<figureCitation box="[772,863,1274,1300]" captionStart="FIGURE 102" captionStartId="135.[136,229,1763,1785]" captionTargetBox="[171,1412,194,1735]" captionTargetId="figure-17@135.[165,1419,193,1739]" captionTargetPageId="135" captionText="FIGURE 102. Lasianthus verticillatus (Lour.) Merr. (AD from Dang 290 &amp; EJ from Dang 284, VNM and the herbarium of Iriomote Station (Japan)): A. Branch, B. Close-up of expanding young leaves, C. Stipules, D. Abaxial leaf surfaces, EG. Inflorescence showing side view calyx, buds and flower, HI. Young fruits, J. Mature fruits." figureDoi="http://doi.org/10.5281/zenodo.7571864" httpUri="https://zenodo.org/record/7571864/files/figure.png" pageId="133">Fig. 102</figureCitation>
(plate),
<figureCitation box="[957,1037,1274,1300]" captionStart="FIGURE 99" captionStartId="131.[136,229,1809,1831]" captionTargetBox="[176,1405,204,1770]" captionTargetId="figure-17@131.[165,1418,193,1785]" captionTargetPageId="131" captionText="FIGURE 99. Distribution maps of Lasianthus thuathienhuensis, L. thuyanae, L. trichophlebus and L. verticillatus." figureDoi="http://doi.org/10.5281/zenodo.7571856" httpUri="https://zenodo.org/record/7571856/files/figure.png" pageId="133">Fig. 99</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection pageId="133" type="materials_examined">
<paragraph blockId="133.[136,1452,1274,1552]" box="[136,636,1314,1336]" pageId="133">
<emphasis bold="true" box="[136,194,1314,1335]" pageId="133">Type:</emphasis>
<materialsCitation box="[219,632,1314,1336]" collectorName="G. Dausus" country="Vietnam" location="VIETNAM" pageId="133" specimenCount="1" typeStatus="holotype">
<collectingCountry box="[219,332,1314,1335]" name="Vietnam" pageId="133">VIETNAM</collectingCountry>
.
<emphasis box="[341,472,1314,1336]" italics="true" pageId="133">
<collectorName box="[341,442,1314,1336]" pageId="133">G. Dausus</collectorName>
48
</emphasis>
(
<typeStatus box="[485,571,1314,1336]" pageId="133">holotype</typeStatus>
BM!)
</materialsCitation>
.
</paragraph>
<paragraph blockId="133.[136,1452,1274,1552]" box="[136,1005,1350,1372]" pageId="133">
<taxonomicName authority="Lour. (1790: 141)" authorityName="Lour." authorityPageNumber="141" authorityYear="1790" box="[136,496,1350,1372]" class="Magnoliopsida" family="Rubiaceae" genus="Dasus" kingdom="Plantae" order="Gentianales" pageId="133" phylum="Tracheophyta" rank="species" species="verticillatus">
<emphasis box="[136,317,1350,1372]" italics="true" pageId="133">Dasus verticillatus</emphasis>
<materialsCitation box="[323,496,1350,1371]" collectorName="G. Dausus" country="Vietnam" location="VIETNAM" pageId="133" specimenCount="1" typeStatus="holotype">Lour. (1790: 141)</materialsCitation>
</taxonomicName>
.
<emphasis bold="true" box="[505,563,1350,1371]" pageId="133">Type:</emphasis>
<collectingCountry box="[588,701,1350,1371]" name="Vietnam" pageId="133">VIETNAM</collectingCountry>
.
<emphasis box="[710,841,1350,1372]" italics="true" pageId="133">
<collectorName box="[710,811,1350,1372]" pageId="133">G. Dausus</collectorName>
48
</emphasis>
(
<typeStatus box="[853,939,1350,1372]" pageId="133">holotype</typeStatus>
BM!).
</paragraph>
<paragraph blockId="133.[136,1452,1274,1552]" pageId="133">
<taxonomicName authority="Hook. f. (1880: 189)" authorityName="Hook. f." authorityPageNumber="189" authorityYear="1880" box="[136,628,1386,1408]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="133" phylum="Tracheophyta" rank="species" species="andamanicus">
<emphasis box="[136,389,1386,1407]" italics="true" pageId="133">Lasianthus andamanicus</emphasis>
<materialsCitation box="[408,628,1386,1408]" country="Myanmar" county="Tenasserim" location="Type" municipality="Amherst" pageId="133" specimenCount="1" typeStatus="holotype">Hook.f. (1880: 189)</materialsCitation>
</taxonomicName>
.
<location LSID="urn:lsid:plazi:treatment:43284262FF1AFF585AB6FB5F44ABCD6C:CE5EA5AFFF1AFF5B58B5FACF429AC937" box="[651,700,1386,1407]" country="Myanmar" county="Tenasserim" municipality="Amherst" name="Type" pageId="133">Type</location>
:—
<collectingCountry box="[727,857,1386,1407]" name="Myanmar" pageId="133">MYANMAR</collectingCountry>
.
<collectingMunicipality box="[878,963,1386,1408]" pageId="133">Amherst</collectingMunicipality>
,
<collectingCounty box="[986,1098,1386,1408]" pageId="133">Tenasserim</collectingCounty>
,
<emphasis box="[1120,1296,1386,1407]" italics="true" pageId="133">Wall. Cat. 8309</emphasis>
(
<typeStatus box="[1321,1407,1386,1408]" pageId="133">holotype</typeStatus>
K! [K000031676]).
</paragraph>
<paragraph blockId="133.[136,1452,1274,1552]" pageId="133">
<taxonomicName authority="Pierre ex Pit. (1924: 390)" authorityName="Pit." authorityPageNumber="390" authorityYear="1924" box="[136,607,1458,1480]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="133" phylum="Tracheophyta" rank="species" species="tamirensis">
<emphasis box="[136,348,1458,1479]" italics="true" pageId="133">Lasianthus tamirensis</emphasis>
<materialsCitation box="[355,607,1458,1480]" collectingDate="1870-05" country="Cambodia" county="Mt Tamire" location="Type" municipality="Samrong-tong" pageId="133" specimenCount="1" typeStatus="holotype">Pierre ex Pit. (1924: 390)</materialsCitation>
</taxonomicName>
.
<location LSID="urn:lsid:plazi:treatment:43284262FF1AFF585AB6FB5F44ABCD6C:CE5EA5AFFF1AFF5B5857FA1742BDC98F" box="[617,667,1458,1479]" country="Cambodia" county="Mt Tamire" municipality="Samrong-tong" name="Type" pageId="133">Type</location>
:—
<collectingCountry box="[695,829,1458,1479]" name="Cambodia" pageId="133">CAMBODIA</collectingCountry>
.
<collectingMunicipality box="[840,980,1458,1479]" pageId="133">Samrong-tong</collectingMunicipality>
,
<collectingCounty box="[990,1095,1458,1480]" pageId="133">Mt Tamire</collectingCounty>
,
<date box="[1107,1208,1458,1479]" pageId="133" value="1870-05">
<collectingDate box="[1107,1208,1458,1479]" pageId="133" value="1870-05">May 1870</collectingDate>
</date>
,
<emphasis box="[1219,1323,1458,1479]" italics="true" pageId="133">Pierre 587</emphasis>
(
<typeStatus box="[1337,1423,1458,1480]" pageId="133">holotype</typeStatus>
P! [P02285234]; isotype K! [K000777002]).
</paragraph>
<paragraph blockId="133.[136,1452,1274,1552]" box="[136,1258,1530,1552]" pageId="133">
<taxonomicName authority="King &amp; Gamble (1904: 119)" authorityName="King &amp; Gamble" authorityPageNumber="119" authorityYear="1904" box="[136,583,1530,1552]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="133" phylum="Tracheophyta" rank="species" species="wrayi">
<emphasis box="[136,302,1530,1551]" italics="true" pageId="133">Lasianthus wrayi</emphasis>
<materialsCitation box="[308,583,1530,1552]" country="Malaysia" location="Wray" municipality="Type" pageId="133" specimenCount="1" stateProvince="Perak" typeStatus="holotype">King &amp; Gamble (1904: 119)</materialsCitation>
</taxonomicName>
.
<collectingMunicipality box="[592,641,1530,1551]" pageId="133">Type</collectingMunicipality>
:—
<collectingCountry box="[669,798,1530,1551]" name="Malaysia" pageId="133">MALAYSIA</collectingCountry>
.
<collectingRegion box="[807,864,1530,1552]" country="Malaysia" name="Perak" pageId="133">Perak</collectingRegion>
,
<emphasis box="[873,966,1530,1552]" italics="true" pageId="133">
<location LSID="urn:lsid:plazi:treatment:43284262FF1AFF585AB6FB5F44ABCD6C:CE5EA5AFFF1AFF5B5957FA5E43BACA58" box="[873,924,1531,1552]" country="Malaysia" municipality="Type" name="Wray" pageId="133" stateProvince="Perak">Wray</location>
257
</emphasis>
(
<typeStatus box="[979,1065,1530,1552]" pageId="133">holotype</typeStatus>
K! [K000777043]).
</paragraph>
<paragraph blockId="133.[136,1452,1598,2059]" pageId="133">
<emphasis bold="true" box="[136,396,1598,1624]" pageId="133">
<collectingRegion box="[136,262,1598,1624]" pageId="133">Specimens</collectingRegion>
examined:
</emphasis>
<materialsCitation collectingDate="2018-11-18" collectionCode="VNM" collectorName="Dang V. S. &amp; Naiki A. &amp; Dang &amp; Japan" country="Vietnam" elevation="365" location="Iriomote Station" municipality="Cuc Phuong National Park" pageId="133" specimenCount="1" stateProvince="Ninh Binh">
<collectingCountry box="[425,557,1598,1624]" name="Vietnam" pageId="133">VIETNAM</collectingCountry>
.
<emphasis bold="true" box="[570,691,1598,1624]" pageId="133">Northern:</emphasis>
<collectingRegion box="[701,933,1598,1624]" country="Vietnam" name="Ninh Binh" pageId="133">Ninh Binh Province</collectingRegion>
,
<collectingMunicipality box="[947,1257,1598,1624]" pageId="133">Cuc Phuong National Park</collectingMunicipality>
,
<quantity box="[1272,1345,1598,1625]" metricMagnitude="2" metricUnit="m" metricValue="3.65" pageId="133" unit="m" value="365.0">
<elevation box="[1272,1345,1598,1625]" metricMagnitude="2" metricUnit="m" metricValue="3.65" pageId="133" unit="m" value="365.0">365 m</elevation>
</quantity>
elev.,
<date pageId="133" value="2018-11-18">
<collectingDate pageId="133" value="2018-11-18">18 November 2018</collectingDate>
</date>
,
<emphasis box="[335,861,1634,1661]" italics="true" pageId="133">
<collectorName box="[335,448,1635,1661]" pageId="133">Dang V.S.</collectorName>
&amp;
<collectorName box="[487,582,1634,1661]" pageId="133">Naiki A.</collectorName>
,
<collectorName box="[596,658,1635,1660]" pageId="133">Dang</collectorName>
284 &amp;
<collectorName box="[748,810,1635,1660]" pageId="133">Dang</collectorName>
290
</emphasis>
(
<collectionCode box="[878,948,1634,1660]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="133" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FF1AFF585AB6FB5F44ABCD6C:CE5EA5AFFF1AFF5B5EB9F9C74519CA34" box="[1159,1343,1634,1660]" country="Vietnam" municipality="Cuc Phuong National Park" name="Iriomote Station" pageId="133" stateProvince="Ninh Binh">Iriomote Station</location>
(
<collectorName box="[1361,1430,1634,1660]" pageId="133">Japan</collectorName>
))
</materialsCitation>
.
</paragraph>
<paragraph blockId="133.[136,1452,1598,2059]" pageId="133">
<emphasis bold="true" box="[136,237,1670,1696]" pageId="133">Central:</emphasis>
<materialsCitation collectingDate="2018-07-25" collectionCode="VNM" collectorName="Dang V. S. &amp; Hoang N. T. B. N. &amp; Dang &amp; Southern" elevation="910" location="Hon Ba Nature Reserve" pageId="133" specimenCount="1" stateProvince="Khanh Hoa">
<collectingRegion box="[243,477,1670,1697]" country="Vietnam" name="Khanh Hoa" pageId="133">Khanh Hoa Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FF1AFF585AB6FB5F44ABCD6C:CE5EA5AFFF1AFF5B5BD9F92242D7CAE9" box="[487,753,1671,1697]" name="Hon Ba Nature Reserve" pageId="133" stateProvince="Khanh Hoa">Hon Ba Nature Reserve</location>
,
<quantity box="[762,832,1670,1697]" metricMagnitude="2" metricUnit="m" metricValue="9.1" pageId="133" unit="m" value="910.0">
<elevation box="[762,832,1670,1697]" metricMagnitude="2" metricUnit="m" metricValue="9.1" pageId="133" unit="m" value="910.0">910 m</elevation>
</quantity>
elev.,
<date box="[902,1047,1670,1697]" pageId="133" value="2018-07-25">
<collectingDate box="[902,1047,1670,1697]" pageId="133" value="2018-07-25">25 July 2018</collectingDate>
</date>
,
<emphasis italics="true" pageId="133">
<collectorName box="[1056,1166,1671,1697]" pageId="133">Dang V.S.</collectorName>
&amp;
<collectorName box="[1199,1380,1671,1696]" pageId="133">Hoang N.T.B.N.</collectorName>
,
<collectorName box="[1389,1451,1671,1696]" pageId="133">Dang</collectorName>
197
</emphasis>
(
<collectionCode box="[195,266,1706,1732]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="133" type="Herbarium">VNM</collectionCode>
).
<collectorName box="[286,401,1706,1732]" pageId="133">
<emphasis bold="true" box="[286,401,1706,1732]" pageId="133">Southern</emphasis>
</collectorName>
</materialsCitation>
<emphasis bold="true" box="[401,406,1706,1732]" pageId="133">:</emphasis>
<materialsCitation collectingDate="2020-11-22" collectionCode="VNM" collectorName="Dang V. S. &amp; Hoang N. S. &amp; Dang" location="Logo-Xamat National Park" pageId="133" specimenCount="1" stateProvince="Tay Ninh">
<collectingRegion box="[414,632,1706,1733]" country="Vietnam" name="Tay Ninh" pageId="133">Tay Ninh Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FF1AFF585AB6FB5F44ABCD6C:CE5EA5AFFF1AFF5B58BBF90E439ACA8C" box="[645,956,1706,1733]" name="Logo-Xamat National Park" pageId="133" stateProvince="Tay Ninh">Logo-Xamat National Park</location>
, 10 elev.,
<date box="[1073,1296,1706,1732]" pageId="133" value="2020-11-22">
<collectingDate box="[1073,1296,1706,1732]" pageId="133" value="2020-11-22">22 November 2020</collectingDate>
</date>
,
<emphasis italics="true" pageId="133">
<collectorName box="[1308,1421,1707,1733]" pageId="133">Dang V.S.</collectorName>
&amp;
<collectorName box="[136,269,1743,1769]" pageId="133">Hoang N.S.</collectorName>
,
<collectorName box="[280,342,1743,1768]" pageId="133">Dang</collectorName>
435
</emphasis>
(
<collectionCode box="[406,477,1742,1768]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="133" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="134" pageId="133" type="distribution">
<paragraph blockId="133.[136,1452,1598,2059]" pageId="133">
<emphasis bold="true" box="[189,345,1778,1804]" pageId="133">Distribution:</emphasis>
—Widespread in the tropical and subtropical regions in Asia (
<collectingCountry box="[1048,1168,1778,1804]" name="Cambodia" pageId="133">Cambodia</collectingCountry>
,
<collectingCountry box="[1180,1250,1778,1804]" name="China" pageId="133">China</collectingCountry>
,
<collectingCountry box="[1263,1323,1778,1804]" name="India" pageId="133">India</collectingCountry>
,
<collectingCountry box="[1336,1448,1778,1804]" name="Indonesia" pageId="133">Indonesia</collectingCountry>
,
<collectingCountry box="[136,203,1815,1841]" name="Japan" pageId="133">Japan</collectingCountry>
,
<collectingCountry box="[218,275,1815,1841]" name="Laos" pageId="133">Laos</collectingCountry>
,
<collectingCountry box="[290,395,1814,1840]" name="Malaysia" pageId="133">Malaysia</collectingCountry>
,
<collectingCountry box="[411,522,1815,1841]" name="Myanmar" pageId="133">Myanmar</collectingCountry>
,
<collectingCountry box="[537,764,1814,1841]" name="Papua New Guinea" pageId="133">Papua New Guinea</collectingCountry>
,
<collectingCountry box="[779,908,1814,1840]" name="Philippines" pageId="133">Philippines</collectingCountry>
,
<collectingCountry box="[924,1040,1814,1840]" name="Sri Lanka" pageId="133">Sri Lanka</collectingCountry>
,
<collectingCountry box="[1055,1160,1814,1840]" name="Thailand" pageId="133">Thailand</collectingCountry>
). In
<collectingCountry box="[1216,1314,1814,1840]" name="Vietnam" pageId="133">Vietnam</collectingCountry>
,
<taxonomicName class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="133" phylum="Tracheophyta" rank="species" species="verticillatus">
<emphasis italics="true" pageId="133">Lasianthus verticillatus</emphasis>
</taxonomicName>
occurs in
<collectingRegion box="[387,499,1850,1877]" country="Vietnam" name="Dong Nai" pageId="133">Dong Nai</collectingRegion>
,
<collectingRegion box="[511,605,1850,1877]" country="Vietnam" name="Ha Tinh" pageId="133">Ha Tinh</collectingRegion>
,
<collectingRegion box="[617,747,1850,1877]" country="Vietnam" name="Khanh Hoa" pageId="133">Khanh Hoa</collectingRegion>
,
<collectingRegion box="[759,892,1850,1876]" country="Vietnam" name="Kien Giang" pageId="133">Kien Giang</collectingRegion>
,
<collectingRegion box="[904,1014,1851,1877]" country="Vietnam" name="Kon Tum" pageId="133">Kon Tum</collectingRegion>
,
<collectingRegion box="[1025,1149,1851,1877]" country="Vietnam" name="Lam Dong" pageId="133">Lam Dong</collectingRegion>
,
<collectingRegion box="[1160,1281,1850,1876]" country="Vietnam" name="Ninh Binh" pageId="133">Ninh Binh</collectingRegion>
,
<collectingRegion box="[1292,1400,1850,1877]" country="Vietnam" name="Tay Ninh" pageId="133">Tay Ninh</collectingRegion>
, and Thua Thien Hue.
</paragraph>
<paragraph blockId="133.[136,1452,1598,2059]" pageId="133">
<emphasis bold="true" box="[189,424,1922,1948]" pageId="133">Habitat &amp; Ecology:</emphasis>
—Growing under the shade of primary and secondary forests or in hills and the edge of the evergreen montane forests at
<quantity box="[465,606,1958,1985]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="133" unit="m" value="650.0" valueMax="1000.0" valueMin="300.0">3001000 m</quantity>
elev.
</paragraph>
<paragraph blockId="133.[136,1452,1598,2059]" box="[189,1073,1994,2021]" pageId="133">
<emphasis bold="true" box="[189,323,1994,2020]" pageId="133">Phenology:</emphasis>
—Flowering and fruiting from November to April of the next year.
</paragraph>
<paragraph blockId="133.[136,1452,1598,2059]" box="[189,662,2030,2059]" pageId="133">
<emphasis bold="true" box="[189,405,2030,2056]" pageId="133">Vernacular name:</emphasis>
—Xú hưƠng luân sinh.
</paragraph>
<paragraph blockId="134.[136,1452,158,293]" pageId="134">
<emphasis bold="true" box="[189,265,158,184]" pageId="134">Notes:</emphasis>
—Two Vietnamese species,
<taxonomicName authority="Pierre ex Pit. (1924: 390)" authorityName="Pit." authorityPageNumber="390" authorityYear="1924" box="[585,1028,158,185]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="134" phylum="Tracheophyta" rank="species" species="tamirensis">
<emphasis box="[585,732,159,184]" italics="true" pageId="134">L. tamirensis</emphasis>
Pierre ex Pit. (1924: 390)
</taxonomicName>
and
<taxonomicName authority="King &amp; Gamble (1904: 119)" authorityName="King &amp; Gamble" authorityPageNumber="119" authorityYear="1904" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="134" phylum="Tracheophyta" rank="species" species="wrayi">
<emphasis box="[1085,1179,159,184]" italics="true" pageId="134">L. wrayi</emphasis>
King &amp; Gamble (1904: 119)
</taxonomicName>
were treated as synonyms of
<taxonomicName box="[519,683,194,220]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="134" phylum="Tracheophyta" rank="species" species="verticillatus">
<emphasis box="[519,683,194,220]" italics="true" pageId="134">L. verticillatus</emphasis>
</taxonomicName>
by
<bibRefCitation author="Zhu, H. &amp; Roos, M. C. &amp; Ridsdale, C. E." box="[724,918,194,220]" pageId="134" pagination="1 - 102" refId="ref55608" refString="Zhu, H., Roos, M. C. &amp; Ridsdale, C. E. (2012) A taxonomic revision of the Malesian species of Lasianthus (Rubiaceae). Blumea 57: 1 - 102. https: // doi. org / 10.3767 / 000651912 X 652012" type="journal article" year="2012">
Zhu
<emphasis box="[777,833,194,220]" italics="true" pageId="134">et al.</emphasis>
(2012)
</bibRefCitation>
.
</paragraph>
<paragraph blockId="134.[136,1452,158,293]" pageId="134">This species is easily identified in the field in having oblong to oblong-lanceolate leaves, secondary veins 79 on each side of midrib, calyx cupular truncate, and fruits subglobose, smooth, black when ripe.</paragraph>
</subSubSection>
</treatment>
</document>