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<document id="77F5284EDE13D779A932DB9FA35D049E" ID-CLB-Dataset="271660" ID-DOI="10.5852/ejt.2023.900.2299" ID-GBIF-Dataset="8b15583e-a9c6-4c32-8438-f02b6e4490d6" ID-ISSN="2118-9773" ID-Zenodo-Dep="8420708" ID-ZooBank="0B4BF469-77A8-4CF9-A006-3B98FF4187D5" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1696845197903" checkinUser="plazi" docAuthor="Cole, Mary L. &amp; Herbert, David G." docDate="2023" docId="E60087E6FFB6FFD7FD245056FD0DF8B9" docLanguage="en" docName="EJT.2023.900.1-31.pdf" docOrigin="European Journal of Taxonomy 900 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2299/9967" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Gulella mystica Cole &amp; Herbert 2023, sp. nov." docType="treatment" docUuid="6837690E-1542-46A0-BA98-B09F59D56501" docUuidSource="ZooBank" docVersion="6" lastPageNumber="23" masterDocId="1A39FF9EFFA2FFC1FFAF547EFF99FFD9" masterDocTitle="Seven new narrowly endemic species of Gulella Pfeiffer, 1856 from eastern South Africa (Gastropoda, Streptaxidae) and status revision for another" masterLastPageNumber="31" masterPageNumber="1" pageNumber="21" updateTime="1699694286568" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1255C92F3E6B8735455743ED737AEDAC">Seven new narrowly endemic species of Gulella Pfeiffer, 1856 from eastern South Africa (Gastropoda, Streptaxidae) and status revision for another</mods:title>
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<mods:name id="14A7EAB72312B3EDD883F3FF06B9369B" type="personal">
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<mods:namePart id="35EE90B965F9909EE3279174AE5D4BA5">Cole, Mary L.</mods:namePart>
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<mods:affiliation id="A7765929B7689A16955EF2303AD96B33">East London Museum, 319 Oxford St, East London, 5201, South Africa. &amp; Department of Zoology and Entomology, Rhodes University, Makhanda, 6140, South Africa.</mods:affiliation>
<mods:nameIdentifier id="7EFA9865EA4A5A1C2826D733EA679463" type="email">marybursey@elmuseum.za.org</mods:nameIdentifier>
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<mods:namePart id="99A9520351F5CAF67B7796DDF9AAFE29">Herbert, David G.</mods:namePart>
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<mods:affiliation id="B85D672EECF631C9AB29C2150A587AC2">Department of Natural Sciences, National Museum Wales, Cathays Park, Cardiff, CF 10 3 NP, United Kingdom.</mods:affiliation>
<mods:nameIdentifier id="004B6A4F08D2EA1594AD58281413ED75" type="email">phasianella@gmail.com</mods:nameIdentifier>
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<mods:title id="BE6DBCF7DAA8FBB7244A65ADB9FD4864">European Journal of Taxonomy</mods:title>
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<mods:date id="F24E778D164E7B222DBD68789FC068A2">2023</mods:date>
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<treatment id="E60087E6FFB6FFD7FD245056FD0DF8B9" ID-DOI="http://doi.org/10.5281/zenodo.8423120" ID-GBIF-Taxon="213740312" ID-Zenodo-Dep="8423120" LSID="urn:lsid:zoobank.org:act:6837690E-1542-46A0-BA98-B09F59D56501" httpUri="http://treatment.plazi.org/id/E60087E6FFB6FFD7FD245056FD0DF8B9" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21">
<subSubSection id="26B3657BFFB6FFD5FD245056FB25FBBE" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="6E1636F0FFB6FFD5FD245056FC30FB9A" blockId="20.[376,1212,1064,1161]" box="[651,937,1064,1091]" pageId="20" pageNumber="21">
<heading id="355E819CFFB6FFD5FD245056FC30FB9A" box="[651,937,1064,1091]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="21" reason="2">
<taxonomicName id="A9A94D73FFB6FFD5FD245056FCDEFB9A" authority="Cole &amp; Herbert, 2023" authorityName="Cole &amp; Herbert" authorityYear="2023" box="[651,839,1064,1091]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="mystica" status="sp. nov.">
<emphasis id="5CDDEAE2FFB6FFD5FD245056FCDEFB9A" bold="true" box="[651,839,1064,1091]" italics="true" pageId="20" pageNumber="21">Gulella mystica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47EE5799FFB6FFD5FCE15057FC30FB9A" box="[846,937,1065,1091]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="26B3657BFFB6FFD5FD4F5011FCCAFB50" box="[736,851,1135,1161]" pageId="20" pageNumber="21" type="description">
<paragraph id="6E1636F0FFB6FFD5FD4F5011FCCAFB50" blockId="20.[376,1212,1064,1161]" box="[736,851,1135,1161]" pageId="20" pageNumber="21">
<figureCitation id="F6922A75FFB6FFD5FD4F5011FCB2FB50" box="[736,811,1135,1161]" captionStart="Fig" captionStartId="17.[189,232,1928,1954]" captionTargetBox="[189,1398,968,1884]" captionTargetId="figure-178@17.[189,1398,968,1884]" captionTargetPageId="17" captionText="Fig. 9. Distribution map of Gulella hlathikhulu sp. nov. (green circles), G. nkandla sp. nov. (yellow triangles), G. mystica sp. nov. (red squares) and G. libertas sp. nov. (blue circle) in north-central KwaZulu-Natal. Contours at 1000 m and 2000 m." figureDoi="http://doi.org/10.5281/zenodo.8420730" httpUri="https://zenodo.org/record/8420730/files/figure.png" pageId="20" pageNumber="21">Figs 9</figureCitation>
,
<figureCitation id="F6922A75FFB6FFD5FC985011FCCAFB50" box="[823,851,1135,1161]" captionStart="Fig" captionStartId="21.[189,232,1928,1954]" captionTargetBox="[192,1395,466,1880]" captionTargetId="figure-39@21.[189,1398,453,1886]" captionTargetPageId="21" captionText="Fig. 11. Gulella mystica sp. nov., holotype (NMSA W1181/T4600), length 3.3 mm, width 1.7 mm. A. Aperture view. B. Side view. C. Oblique view into aperture. D. Oblique view of base showing umbilicus." figureDoi="http://doi.org/10.5281/zenodo.8420734" httpUri="https://zenodo.org/record/8420734/files/figure.png" pageId="20" pageNumber="21">11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="26B3657BFFB6FFD5FF1250CEFCA2FA15" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="6E1636F0FFB6FFD5FF1250CEFEAFFB13" blockId="20.[189,310,1200,1226]" box="[189,310,1200,1226]" pageId="20" pageNumber="21">
<heading id="355E819CFFB6FFD5FF1250CEFEAFFB13" bold="true" box="[189,310,1200,1226]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="5CDDEAE2FFB6FFD5FF1250CEFEAFFB13" bold="true" box="[189,310,1200,1226]" pageId="20" pageNumber="21">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFB6FFD5FF1250A1FCA2FA15" blockId="20.[188,1401,1247,1485]" pageId="20" pageNumber="21">Shell very small, sub-cylindrical; sculptured with axial ribs extending from suture to suture with spiral microsculpture between them; apertural dentition eight-fold, including a parietal lamella running into the aperture with a sinuous lower margin bearing a notch, a low sinular denticle, a large triangular labral tooth with a peg-like tooth at its upper limit, a low, deeply inset transverse basal tooth to right of centre, a deeply inset trigonal basal tooth to left of centre and a large inset columella lamella consisting of a vertical slab with two ridge-like teeth, pointing obliquely downwards, in its lower half; apertural tube collar-like behind columella lip, umbilicus very small.</paragraph>
</subSubSection>
<subSubSection id="26B3657BFFB6FFD5FF12518AFC37F9B9" pageId="20" pageNumber="21" type="etymology">
<paragraph id="6E1636F0FFB6FFD5FF12518AFEDDF9D7" blockId="20.[189,324,1524,1550]" box="[189,324,1524,1550]" pageId="20" pageNumber="21">
<heading id="355E819CFFB6FFD5FF12518AFEDDF9D7" bold="true" box="[189,324,1524,1550]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="5CDDEAE2FFB6FFD5FF12518AFEDDF9D7" bold="true" box="[189,324,1524,1550]" pageId="20" pageNumber="21">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFB6FFD5FF12525DFC37F9B9" blockId="20.[189,1398,1570,1632]" pageId="20" pageNumber="21">
From the Greek
<emphasis id="5CDDEAE2FFB6FFD5FE3A525CFE63F9E5" box="[405,506,1570,1596]" italics="true" pageId="20" pageNumber="21">mystikos</emphasis>
(
<emphasis id="5CDDEAE2FFB6FFD5FD8B525CFD13F9E4" box="[548,650,1570,1597]" italics="true" pageId="20" pageNumber="21">μυσΤΙΚός</emphasis>
): a mystery, mysterious; with reference to the environs of Nkandla long considered a region of mystery in Zulu folklore.
</paragraph>
</subSubSection>
<subSubSection id="26B3657BFFB6FFD7FF1252F9FD93FE42" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21" type="materials_examined">
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<heading id="355E819CFFB6FFD5FF1252F9FEF5F978" bold="true" box="[189,364,1671,1697]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="5CDDEAE2FFB6FFD5FF1252F9FEF5F978" bold="true" box="[189,364,1671,1697]" pageId="20" pageNumber="21">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFB6FFD5FF4552C8FDFDF8E0" blockId="20.[189,1398,1718,1849]" pageId="20" pageNumber="21">
<materialsCitation id="DEC13CADFFB6FFD5FF4552C8FDFDF8E0" ID-GBIF-Occurrence="4423011326" collectedFrom="under logs and in leaf-litter" collectingDate="2003-10-20" collectionCode="NMSA" collectorName="D. Herbert" country="South Africa" elevation="1175" latitude="-28.7227" location="Chibini area" longLatPrecision="6" longitude="31.1283" municipality="Nkandla Forest Reserve" pageId="20" pageNumber="21" specimenCode="W1181" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="holotype">
<emphasis id="5CDDEAE2FFB6FFD5FF4552C8FEC5F909" bold="true" box="[234,348,1718,1744]" pageId="20" pageNumber="21">
<typeStatus id="B1128852FFB6FFD5FF4552C8FEC5F909" box="[234,348,1718,1744]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="16BE7660FFB6FFD5FF1252A7FE09F92A" box="[189,400,1753,1779]" name="South Africa" pageId="20" pageNumber="21">SOUTH AFRICA</collectingCountry>
<emphasis id="5CDDEAE2FFB6FFD5FE1E52A7FDEFF92A" bold="true" box="[433,630,1753,1779]" pageId="20" pageNumber="21">
<collectingRegion id="AC6DF812FFB6FFD5FE1E52A7FDEFF92A" box="[433,630,1753,1779]" country="South Africa" name="KwaZulu-Natal" pageId="20" pageNumber="21">KwaZulu-Natal</collectingRegion>
</emphasis>
<collectingMunicipality id="8E72AC8AFFB6FFD5FD3D52A7FC2BF92A" box="[658,946,1753,1779]" pageId="20" pageNumber="21">Nkandla Forest Reserve</collectingMunicipality>
,
<locationDeviation id="7FA42FD2FFB6FFD5FC6F52A7FB49F92A" box="[960,1232,1753,1779]" location="Eshowe" pageId="20" pageNumber="21">
<quantity id="A9519B15FFB6FFD5FC6F52A7FB95F92A" box="[960,1036,1753,1779]" metricMagnitude="4" metricUnit="m" metricValue="3.8" pageId="20" pageNumber="21" unit="km" value="38.0">38 km</quantity>
NW of Eshowe
</locationDeviation>
,
<location id="6B76602BFFB6FFD5FB7252A7FAEBF92A" LSID="urn:lsid:plazi:treatment:E60087E6FFB6FFD7FD245056FD0DF8B9:6B76602BFFB6FFD5FB7252A7FAEBF92A" box="[1245,1394,1753,1779]" country="South Africa" latitude="-28.7227" longLatPrecision="6" longitude="31.1283" municipality="Nkandla Forest Reserve" name="Chibini area" pageId="20" pageNumber="21" stateProvince="KwaZulu-Natal">Chibini area</location>
,
<location id="6B76602BFFB6FFD5FF125282FEF3F8CF" LSID="urn:lsid:plazi:treatment:E60087E6FFB6FFD7FD245056FD0DF8B9:6B76602BFFB6FFD5FF125282FEF3F8CF" box="[189,362,1788,1814]" country="South Africa" latitude="-28.7227" longLatPrecision="6" longitude="31.1283" municipality="Nkandla Forest Reserve" name="mistbelt forest" pageId="20" pageNumber="21" stateProvince="KwaZulu-Natal">mistbelt forest</location>
;
<geoCoordinate id="0B9D5037FFB6FFD5FED95282FE62F8CF" box="[374,507,1788,1814]" degrees="28.7227" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="5" value="-28.7227">28.7227° S</geoCoordinate>
,
<geoCoordinate id="0B9D5037FFB6FFD5FDA65282FD08F8CE" box="[521,657,1788,1815]" degrees="31.1283" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="5" value="31.1283">31.1283° E</geoCoordinate>
; ±
<elevation id="E584D1C3FFB6FFD5FD195282FCD6F8CF" box="[694,847,1788,1815]" metricMagnitude="3" metricUnit="m" metricValue="1.175" pageId="20" pageNumber="21" unit="m" value="1175.0">
<quantity id="A9519B15FFB6FFD5FD195282FC96F8CE" box="[694,783,1788,1815]" metricMagnitude="3" metricUnit="m" metricValue="1.175" pageId="20" pageNumber="21" unit="m" value="1175.0">1175 m</quantity>
a.s.l.
</elevation>
;
<date id="1A171030FFB6FFD5FCF25282FC63F8CF" box="[861,1018,1788,1814]" pageId="20" pageNumber="21" value="2003-10-20">
<collectingDate id="0A53E9D8FFB6FFD5FCF25282FC63F8CF" box="[861,1018,1788,1814]" pageId="20" pageNumber="21" value="2003-10-20">20 Oct. 2003</collectingDate>
</date>
;
<collectorName id="C35C5326FFB6FFD5FBA85283FB1FF8CF" box="[1031,1158,1788,1815]" pageId="20" pageNumber="21">D. Herbert</collectorName>
leg.;
<collectedFrom id="DDB5750DFFB6FFD5FB675282FED4F8E0" pageId="20" pageNumber="21">under logs and in leaf-litter</collectedFrom>
;
<collectionCode id="08B8AE35FFB6FFD5FEF75361FE32F8E0" box="[344,427,1823,1849]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="20" pageNumber="21" type="Museum">NMSA</collectionCode>
<specimenCode id="3E0F9E8BFFB6FFD5FE1D5361FD9EF8E0" box="[434,519,1823,1849]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="20" pageNumber="21" type="Museum">W1181</specimenCode>
/T4600.
</materialsCitation>
</paragraph>
<paragraph id="6E1636F0FFB6FFD4FF45531FFD46FE9F" blockId="20.[189,1398,1889,2021]" lastBlockId="21.[189,1398,265,327]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<materialsCitation id="DEC13CADFFB6FFD5FF45531FFE30F83D" ID-GBIF-Occurrence="4423011322" collectingDate="2015-04-09" collectorName="M. and K. Cole" country="South Africa" elevation="1175" latitude="-28.7227" location="Chibini area" longLatPrecision="6" longitude="31.1283" municipality="Nkandla Forest Reserve" pageId="20" pageNumber="21" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<emphasis id="5CDDEAE2FFB6FFD5FF45531FFEFEF8A2" bold="true" box="[234,359,1889,1915]" pageId="20" pageNumber="21">
<typeStatus id="B1128852FFB6FFD5FF45531FFEFEF8A2" box="[234,359,1889,1915]" pageId="20" pageNumber="21" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="16BE7660FFB6FFD5FF1253FAFE08F847" box="[189,401,1924,1950]" name="South Africa" pageId="20" pageNumber="21">SOUTH AFRICA</collectingCountry>
<emphasis id="5CDDEAE2FFB6FFD5FE1C53FAFDE1F847" bold="true" box="[435,632,1924,1950]" pageId="20" pageNumber="21">
<collectingRegion id="AC6DF812FFB6FFD5FE1C53FAFDE1F847" box="[435,632,1924,1950]" country="South Africa" name="KwaZulu-Natal" pageId="20" pageNumber="21">KwaZulu-Natal</collectingRegion>
</emphasis>
<specimenCount id="78AFFD79FFB6FFD5FD3A53FAFD74F847" box="[661,749,1924,1950]" pageId="20" pageNumber="21" type="generic">1 spec.</specimenCount>
;
<collectingMunicipality id="8E72AC8AFFB6FFD5FD5453FAFB85F847" box="[763,1052,1924,1950]" pageId="20" pageNumber="21">Nkandla Forest Reserve</collectingMunicipality>
,
<location id="6B76602BFFB6FFD5FB8553FAFB59F847" LSID="urn:lsid:plazi:treatment:E60087E6FFB6FFD7FD245056FD0DF8B9:6B76602BFFB6FFD5FB8553FAFB59F847" box="[1066,1216,1924,1950]" country="South Africa" latitude="-28.7227" longLatPrecision="6" longitude="31.1283" municipality="Nkandla Forest Reserve" name="Chibini area" pageId="20" pageNumber="21" stateProvince="KwaZulu-Natal">Chibini area</location>
,
<locationDeviation id="7FA42FD2FFB6FFD5FB6153FAFE84F818" location="Eshowe" pageId="20" pageNumber="21">
<quantity id="A9519B15FFB6FFD5FB6153FAFA83F847" box="[1230,1306,1924,1950]" metricMagnitude="4" metricUnit="m" metricValue="3.8" pageId="20" pageNumber="21" unit="km" value="38.0">38 km</quantity>
NW of Eshowe
</locationDeviation>
,
<location id="6B76602BFFB6FFD5FE8453D9FE43F818" LSID="urn:lsid:plazi:treatment:E60087E6FFB6FFD7FD245056FD0DF8B9:6B76602BFFB6FFD5FE8453D9FE43F818" box="[299,474,1959,1985]" country="South Africa" latitude="-28.7227" longLatPrecision="6" longitude="31.1283" municipality="Nkandla Forest Reserve" name="mistbelt forest" pageId="20" pageNumber="21" stateProvince="KwaZulu-Natal">mistbelt forest</location>
;
<geoCoordinate id="0B9D5037FFB6FFD5FE4753D9FDF7F818" box="[488,622,1959,1985]" degrees="28.7227" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="5" value="-28.7227">28.7227° S</geoCoordinate>
,
<geoCoordinate id="0B9D5037FFB6FFD5FDD153D9FC91F81B" box="[638,776,1959,1986]" degrees="31.1283" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="5" value="31.1283">31.1283° E</geoCoordinate>
; ±
<elevation id="E584D1C3FFB6FFD5FC9F53D9FC54F818" box="[816,973,1959,1986]" metricMagnitude="3" metricUnit="m" metricValue="1.175" pageId="20" pageNumber="21" unit="m" value="1175.0">
<quantity id="A9519B15FFB6FFD5FC9F53D9FC13F81B" box="[816,906,1959,1986]" metricMagnitude="3" metricUnit="m" metricValue="1.175" pageId="20" pageNumber="21" unit="m" value="1175.0">1175 m</quantity>
a.s.l.
</elevation>
;
<date id="1A171030FFB6FFD5FC7353D9FBF7F81B" box="[988,1134,1959,1986]" pageId="20" pageNumber="21" value="2015-04-09">
<collectingDate id="0A53E9D8FFB6FFD5FC7353D9FBF7F81B" box="[988,1134,1959,1986]" pageId="20" pageNumber="21" value="2015-04-09">9 Apr. 2015</collectingDate>
</date>
;
<collectorName id="C35C5326FFB6FFD5FBD253D6FAA0F818" box="[1149,1337,1959,1986]" pageId="20" pageNumber="21">M. and K. Cole</collectorName>
leg.; ELMD 18701/T244
</materialsCitation>
<materialsCitation id="DEC13CADFFB6FFD4FE6C53B4FD46FE9F" ID-GBIF-Occurrence="4423011321" collectingDate="2015-04-09" collectionCode="ELMD" collectorName="M. &amp; K. Cole" elevation="1040" lastPageId="21" lastPageNumber="22" latitude="-28.7347" location="mistbelt forest" longLatPrecision="6" longitude="31.1439" municipality="Nkandla Forest Reserve" pageId="20" pageNumber="21" specimenCode="ELMD 18622" specimenCount="2">
<specimenCount id="78AFFD79FFB6FFD5FE6C53B4FD8CF83C" box="[451,533,1994,2021]" pageId="20" pageNumber="21" type="generic">1 spec.</specimenCount>
;
<collectingMunicipality id="8E72AC8AFFB6FFD5FD8E53B4FCA7F83C" box="[545,830,1994,2021]" pageId="20" pageNumber="21">Nkandla Forest Reserve</collectingMunicipality>
,
<locationDeviation id="7FA42FD2FFB6FFD4FCE553B4FEAEFEFA" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<quantity id="A9519B15FFB6FFD5FCE553B4FC0DF83D" box="[842,916,1994,2020]" metricMagnitude="4" metricUnit="m" metricValue="3.7" pageId="20" pageNumber="21" unit="km" value="37.0">37 km</quantity>
NW of Eshowe, near picnic site north of main road
</locationDeviation>
,
<location id="6B76602BFFB7FFD4FEEC5577FE68FEFA" LSID="urn:lsid:plazi:treatment:E60087E6FFB6FFD7FD245056FD0DF8B9:6B76602BFFB7FFD4FEEC5577FE68FEFA" box="[323,497,265,291]" latitude="-28.7347" longLatPrecision="6" longitude="31.1439" municipality="Nkandla Forest Reserve" name="mistbelt forest" pageId="21" pageNumber="22">mistbelt forest</location>
;
<geoCoordinate id="0B9D5037FFB7FFD4FE525577FD1AFEFA" box="[509,643,265,291]" degrees="28.7347" direction="south" orientation="latitude" pageId="21" pageNumber="22" precision="5" value="-28.7347">28.7347° S</geoCoordinate>
,
<geoCoordinate id="0B9D5037FFB7FFD4FD3F5577FC80FEFA" box="[656,793,265,291]" degrees="31.1439" direction="east" orientation="longitude" pageId="21" pageNumber="22" precision="5" value="31.1439">31.1439° E</geoCoordinate>
; ±
<quantity id="A9519B15FFB7FFD4FC915577FC01FEFA" box="[830,920,265,291]" metricMagnitude="3" metricUnit="m" metricValue="1.04" pageId="21" pageNumber="22" unit="m" value="1040.0">
<elevation id="E584D1C3FFB7FFD4FC915577FC01FEFA" box="[830,920,265,291]" metricMagnitude="3" metricUnit="m" metricValue="1.04" pageId="21" pageNumber="22" unit="m" value="1040.0">1040 m</elevation>
</quantity>
a.s.l.;
<date id="1A171030FFB7FFD4FC485577FBEFFEFD" box="[999,1142,265,292]" pageId="21" pageNumber="22" value="2015-04-09">
<collectingDate id="0A53E9D8FFB7FFD4FC485577FBEFFEFD" box="[999,1142,265,292]" pageId="21" pageNumber="22" value="2015-04-09">9 Apr. 2015</collectingDate>
</date>
;
<collectorName id="C35C5326FFB7FFD4FB2C5577FB3DFEFA" box="[1155,1188,265,291]" pageId="21" pageNumber="22">M.</collectorName>
and
<collectorName id="C35C5326FFB7FFD4FB705577FAA2FEFA" box="[1247,1339,265,291]" pageId="21" pageNumber="22">K. Cole</collectorName>
leg.; NMSA-Mol 0P2362/T4611,
<specimenCount id="78AFFD79FFB7FFD4FDBE5553FDB4FE9E" box="[529,557,301,327]" pageId="21" pageNumber="22" type="generic">ex</specimenCount>
<specimenCode id="3E0F9E8BFFB7FFD4FD9B5553FD42FE9F" box="[564,731,300,327]" pageId="21" pageNumber="22">ELMD 18622</specimenCode>
.
</materialsCitation>
</paragraph>
<caption id="3AD66678FFB7FFD4FF1253F6FEA0F831" ID-DOI="http://doi.org/10.5281/zenodo.8420734" ID-Zenodo-Dep="8420734" httpUri="https://zenodo.org/record/8420734/files/figure.png" pageId="21" pageNumber="22" startId="21.[189,232,1928,1954]" targetBox="[192,1395,466,1880]" targetPageId="21">
<paragraph id="6E1636F0FFB7FFD4FF1253F6FEA0F831" blockId="21.[189,1399,1928,2024]" pageId="21" pageNumber="22">
<emphasis id="5CDDEAE2FFB7FFD4FF1253F6FE84F87B" bold="true" box="[189,285,1928,1954]" pageId="21" pageNumber="22">Fig. 11.</emphasis>
<taxonomicName id="A9A94D73FFB7FFD4FE8753F6FE7CF87B" authority="Cole &amp; Herbert, 2023" authorityName="Cole &amp; Herbert" authorityYear="2023" box="[296,485,1928,1954]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="mystica" status="sp. nov.">
<emphasis id="5CDDEAE2FFB7FFD4FE8753F6FE7CF87B" box="[296,485,1928,1954]" italics="true" pageId="21" pageNumber="22">Gulella mystica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47EE5799FFB7FFD4FE4053F6FDC9F87B" box="[495,592,1928,1954]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (NMSA W1181/T4600), length 3.3 mm, width 1.7 mm.
<emphasis id="5CDDEAE2FFB7FFD4FF1253D5FF4BF81C" bold="true" box="[189,210,1963,1989]" pageId="21" pageNumber="22">A</emphasis>
. Aperture view.
<emphasis id="5CDDEAE2FFB7FFD4FE0D53D5FE2FF81C" bold="true" box="[418,438,1963,1989]" pageId="21" pageNumber="22">B</emphasis>
. Side view.
<emphasis id="5CDDEAE2FFB7FFD4FDFC53D5FDF1F81C" bold="true" box="[595,616,1963,1989]" pageId="21" pageNumber="22">C</emphasis>
. Oblique view into aperture.
<emphasis id="5CDDEAE2FFB7FFD4FC7853D5FC75F81C" bold="true" box="[983,1004,1963,1989]" pageId="21" pageNumber="22">D</emphasis>
. Oblique view of base showing umbilicus.
</paragraph>
</caption>
<paragraph id="6E1636F0FFB4FFD7FF125576FE60FEFB" blockId="22.[189,505,264,290]" box="[189,505,264,290]" pageId="22" pageNumber="23">
<heading id="355E819CFFB4FFD7FF125576FE60FEFB" bold="true" box="[189,505,264,290]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="5CDDEAE2FFB4FFD7FF125576FE60FEFB" bold="true" box="[189,505,264,290]" pageId="22" pageNumber="23">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFB4FFD7FF125546FD93FE42" blockId="22.[188,1398,312,411]" pageId="22" pageNumber="23">
<materialsCitation id="DEC13CADFFB4FFD7FF125546FD9FFE42" ID-GBIF-Occurrence="4423011358" collectingDate="2010-01-14" collectorName="M. Cole" country="South Africa" elevation="1050" latitude="-28.7453" location="Nkandla Forest Reserve" longLatPrecision="6" longitude="31.1361" municipality="Nkandla Forest Reserve" pageId="22" pageNumber="23" specimenCount="1" stateProvince="KwaZulu-Natal">
<collectingCountry id="16BE7660FFB4FFD7FF125546FE0BFE8B" box="[189,402,312,338]" name="South Africa" pageId="22" pageNumber="23">SOUTH AFRICA</collectingCountry>
<emphasis id="5CDDEAE2FFB4FFD7FE165546FDE7FE8B" bold="true" box="[441,638,312,338]" pageId="22" pageNumber="23">
<collectingRegion id="AC6DF812FFB4FFD7FE165546FDE7FE8B" box="[441,638,312,338]" country="South Africa" name="KwaZulu-Natal" pageId="22" pageNumber="23">KwaZulu-Natal</collectingRegion>
</emphasis>
<specimenCount id="78AFFD79FFB4FFD7FD325546FD6FFE8A" box="[669,758,312,339]" pageId="22" pageNumber="23" type="generic">1 spec.</specimenCount>
;
<collectingMunicipality id="8E72AC8AFFB4FFD7FCA95546FBB3FE8A" box="[774,1066,312,339]" pageId="22" pageNumber="23">Nkandla Forest Reserve</collectingMunicipality>
,
<locationDeviation id="7FA42FD2FFB4FFD7FB955546FDA4FEAF" pageId="22" pageNumber="23">
Mdonini area near eastern boundary,
<quantity id="A9519B15FFB4FFD7FE975522FE18FEAF" box="[312,385,348,374]" metricMagnitude="4" metricUnit="m" metricValue="3.6" pageId="22" pageNumber="23" unit="km" value="36.0">36 km</quantity>
NW of Eshowe
</locationDeviation>
, mistbelt forest;
<geoCoordinate id="0B9D5037FFB4FFD7FD505522FC1BFEAF" box="[767,898,348,375]" degrees="28.7453" direction="south" orientation="latitude" pageId="22" pageNumber="23" precision="5" value="-28.7453">28.7453° S</geoCoordinate>
,
<geoCoordinate id="0B9D5037FFB4FFD7FC215522FB8CFEAE" box="[910,1045,348,375]" degrees="31.1361" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="5" value="31.1361">31.1361° E</geoCoordinate>
; ±
<elevation id="E584D1C3FFB4FFD7FB985522FB57FEAF" box="[1079,1230,348,375]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="22" pageNumber="23" unit="m" value="1050.0">
<quantity id="A9519B15FFB4FFD7FB985522FB16FEAE" box="[1079,1167,348,375]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="22" pageNumber="23" unit="m" value="1050.0">1050 m</quantity>
a.s.l.
</elevation>
;
<date id="1A171030FFB4FFD7FB745522FAEBFEAF" box="[1243,1394,348,375]" pageId="22" pageNumber="23" value="2010-01-14">
<collectingDate id="0A53E9D8FFB4FFD7FB745522FAEBFEAF" box="[1243,1394,348,375]" pageId="22" pageNumber="23" value="2010-01-14">14 Jan. 2010</collectingDate>
</date>
;
<collectorName id="C35C5326FFB4FFD7FF1355FFFE85FE42" box="[188,284,385,411]" pageId="22" pageNumber="23">M. Cole</collectorName>
leg.; ELMD 18888
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="26B3657BFFB4FFD7FF1255BDFDC1FA80" pageId="22" pageNumber="23" type="description">
<paragraph id="6E1636F0FFB4FFD7FF1255BDFED7FE04" blockId="22.[189,334,451,477]" box="[189,334,451,477]" pageId="22" pageNumber="23">
<heading id="355E819CFFB4FFD7FF1255BDFED7FE04" bold="true" box="[189,334,451,477]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="5CDDEAE2FFB4FFD7FF1255BDFED7FE04" bold="true" box="[189,334,451,477]" pageId="22" pageNumber="23">Description</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFB4FFD7FF12558DFE15FBF4" blockId="22.[187,1401,499,1069]" pageId="22" pageNumber="23">
<smallCapsWord id="68F0A02CFFB4FFD7FF12558DFE9BFDD5" baselines="518,519" box="[189,258,499,525]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Shell" pageId="22" pageNumber="23">SHELL</smallCapsWord>
(
<figureCitation id="F6922A75FFB4FFD7FEB8558DFEE9FDD4" box="[279,368,499,525]" captionStart="Fig" captionStartId="21.[189,232,1928,1954]" captionTargetBox="[192,1395,466,1880]" captionTargetId="figure-39@21.[189,1398,453,1886]" captionTargetPageId="21" captionText="Fig. 11. Gulella mystica sp. nov., holotype (NMSA W1181/T4600), length 3.3 mm, width 1.7 mm. A. Aperture view. B. Side view. C. Oblique view into aperture. D. Oblique view of base showing umbilicus." figureDoi="http://doi.org/10.5281/zenodo.8420734" httpUri="https://zenodo.org/record/8420734/files/figure.png" pageId="22" pageNumber="23">Fig. 11</figureCitation>
). Shell very small, sub-cylindrical to squat, length
<quantity id="A9519B15FFB4FFD7FC45558DFBE6FDD4" box="[1002,1151,499,525]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="22" pageNumber="23" unit="mm" value="3.25" valueMax="3.3" valueMin="3.2">3.23.3 mm</quantity>
, width
<quantity id="A9519B15FFB4FFD7FB72558DFAEBFDD4" box="[1245,1394,499,525]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="22" pageNumber="23" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.71.8 mm</quantity>
, L:W 1.882.00 (n = 4). Protoconch approx.
<quantity id="A9519B15FFB4FFD7FD175669FC88FDEB" box="[696,785,535,562]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="22" pageNumber="23" unit="mm" value="0.8">0.8 mm</quantity>
in diameter, comprising approx. 2.5 whorls, smooth; junction between protoconch and teleoconch distinct. Teleoconch comprising approx. 5.5 convex whorls; sculptured with well-developed axial ribs, extending from suture to suture, prosocline on spire whorls and almost orthocline on last whorl, rib intervals with spiral microsculpture (
<figureCitation id="F6922A75FFB4FFD7FB7A56FAFAF0FD47" box="[1237,1385,644,670]" captionStart="Fig" captionStartId="21.[189,232,1928,1954]" captionTargetBox="[192,1395,466,1880]" captionTargetId="figure-39@21.[189,1398,453,1886]" captionTargetPageId="21" captionText="Fig. 11. Gulella mystica sp. nov., holotype (NMSA W1181/T4600), length 3.3 mm, width 1.7 mm. A. Aperture view. B. Side view. C. Oblique view into aperture. D. Oblique view of base showing umbilicus." figureDoi="http://doi.org/10.5281/zenodo.8420734" httpUri="https://zenodo.org/record/8420734/files/figure.png" pageId="22" pageNumber="23">Fig. 11AB</figureCitation>
). Aperture sub-quadrate with rounded base; outer lip shallowly indented, columella straight (in aperture view); peristome thickened and reflected; dentition eight-fold (
<figureCitation id="F6922A75FFB4FFD7FC0C56B2FB96FD3F" box="[931,1039,716,742]" captionStart="Fig" captionStartId="21.[189,232,1928,1954]" captionTargetBox="[192,1395,466,1880]" captionTargetId="figure-39@21.[189,1398,453,1886]" captionTargetPageId="21" captionText="Fig. 11. Gulella mystica sp. nov., holotype (NMSA W1181/T4600), length 3.3 mm, width 1.7 mm. A. Aperture view. B. Side view. C. Oblique view into aperture. D. Oblique view of base showing umbilicus." figureDoi="http://doi.org/10.5281/zenodo.8420734" httpUri="https://zenodo.org/record/8420734/files/figure.png" pageId="22" pageNumber="23">Fig. 11C</figureCitation>
): 1) an almost vertical parietal lamella, its lower margin sinuous and notched, 2) a minute sinular denticle, 34) a large triangular labral tooth extending into aperture, with a small peg-like tooth at its upper limit near lip edge, 5) a deeply inset low, transverse basal tooth to right of centre, 6) a deeply inset pointed trigonal tooth to left of centre, 78) a large deeply inset columella lamella consisting of a truncate vertical slab set with two ridge-like downward-pointing teeth in lower half, upper one larger. Aperture indented in region of labral teeth and with a deep pit behind outer lip corresponding to labral tooth base (
<figureCitation id="F6922A75FFB4FFD7FB2757D8FB6BFC19" box="[1160,1266,934,960]" captionStart="Fig" captionStartId="21.[189,232,1928,1954]" captionTargetBox="[192,1395,466,1880]" captionTargetId="figure-39@21.[189,1398,453,1886]" captionTargetPageId="21" captionText="Fig. 11. Gulella mystica sp. nov., holotype (NMSA W1181/T4600), length 3.3 mm, width 1.7 mm. A. Aperture view. B. Side view. C. Oblique view into aperture. D. Oblique view of base showing umbilicus." figureDoi="http://doi.org/10.5281/zenodo.8420734" httpUri="https://zenodo.org/record/8420734/files/figure.png" pageId="22" pageNumber="23">Fig. 11B</figureCitation>
). Apertural tube behind columella lip collar-like; umbilicus very small, elongate-ovate, approx.
<quantity id="A9519B15FFB4FFD7FB3257B4FA9FFC3C" box="[1181,1286,970,997]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="22" pageNumber="23" unit="mm" value="0.08">0.08 mm</quantity>
in length (
<figureCitation id="F6922A75FFB4FFD7FF6A5790FEAFFBD0" box="[197,310,1006,1033]" captionStart="Fig" captionStartId="21.[189,232,1928,1954]" captionTargetBox="[192,1395,466,1880]" captionTargetId="figure-39@21.[189,1398,453,1886]" captionTargetPageId="21" captionText="Fig. 11. Gulella mystica sp. nov., holotype (NMSA W1181/T4600), length 3.3 mm, width 1.7 mm. A. Aperture view. B. Side view. C. Oblique view into aperture. D. Oblique view of base showing umbilicus." figureDoi="http://doi.org/10.5281/zenodo.8420734" httpUri="https://zenodo.org/record/8420734/files/figure.png" pageId="22" pageNumber="23">Fig. 11D</figureCitation>
). Shell almost transparent when fresh, yellow-orange coloration of dried tissue of animal visible internally.
</paragraph>
<paragraph id="6E1636F0FFB4FFD7FF12502BFE2CFBB6" blockId="22.[189,437,1109,1135]" box="[189,437,1109,1135]" pageId="22" pageNumber="23">
<emphasis id="5CDDEAE2FFB4FFD7FF12502BFECFFBB6" bold="true" box="[189,342,1109,1135]" pageId="22" pageNumber="23">Distribution</emphasis>
(
<figureCitation id="F6922A75FFB4FFD7FEC9502BFE35FBB6" box="[358,428,1109,1135]" captionStart="Fig" captionStartId="17.[189,232,1928,1954]" captionTargetBox="[189,1398,968,1884]" captionTargetId="figure-178@17.[189,1398,968,1884]" captionTargetPageId="17" captionText="Fig. 9. Distribution map of Gulella hlathikhulu sp. nov. (green circles), G. nkandla sp. nov. (yellow triangles), G. mystica sp. nov. (red squares) and G. libertas sp. nov. (blue circle) in north-central KwaZulu-Natal. Contours at 1000 m and 2000 m." figureDoi="http://doi.org/10.5281/zenodo.8420730" httpUri="https://zenodo.org/record/8420730/files/figure.png" pageId="22" pageNumber="23">Fig. 9</figureCitation>
)
</paragraph>
<paragraph id="6E1636F0FFB4FFD7FF1250FBFEE0FB1A" blockId="22.[189,1399,1157,1220]" pageId="22" pageNumber="23">
Known only from Nkandla Forest Reserve, in northern
<collectingRegion id="AC6DF812FFB4FFD7FCFC50FBFB96FB46" box="[851,1039,1157,1183]" country="South Africa" name="KwaZulu-Natal" pageId="22" pageNumber="23">KwaZulu-Natal</collectingRegion>
; between
<quantity id="A9519B15FFB4FFD7FB2950FBFB78FB46" box="[1158,1249,1157,1183]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="23" unit="m" value="1000.0">1000 m</quantity>
and
<quantity id="A9519B15FFB4FFD7FAB350FBFAEEFB46" box="[1308,1399,1157,1183]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="22" pageNumber="23" unit="m" value="1200.0">1200 m</quantity>
above sea level.
</paragraph>
<paragraph id="6E1636F0FFB4FFD7FF125095FE84FADC" blockId="22.[189,285,1259,1285]" box="[189,285,1259,1285]" pageId="22" pageNumber="23">
<heading id="355E819CFFB4FFD7FF125095FE84FADC" bold="true" box="[189,285,1259,1285]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="5CDDEAE2FFB4FFD7FF125095FE84FADC" bold="true" box="[189,285,1259,1285]" pageId="22" pageNumber="23">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFB4FFD7FF125165FDC1FA80" blockId="22.[189,1399,1307,1370]" pageId="22" pageNumber="23">
Patches of
<collectingRegion id="AC6DF812FFB4FFD7FEE95165FE67FAEC" box="[326,510,1307,1333]" country="South Africa" name="KwaZulu-Natal" pageId="22" pageNumber="23">KwaZulu-Natal</collectingRegion>
Scarp forest of the
<collectingRegion id="AC6DF812FFB4FFD7FD555165FC2BFAEC" box="[762,946,1307,1333]" country="South Africa" name="KwaZulu-Natal" pageId="22" pageNumber="23">KwaZulu-Natal</collectingRegion>
Inland Scarp subtype (
<bibRefCitation id="0A384B01FFB4FFD7FB7E5165FF65FA83" author="Mucina L. &amp; Lotter M. C. &amp; Tichy L. &amp; Siebert S. J. &amp; Scott-Shaw C. R." pageId="22" pageNumber="23" pagination="125 - 225" refId="ref14157" refString="Mucina L., Lotter M. C., Tichy L., Siebert S. J. &amp; Scott-Shaw C. R. 2018. Classification of the Eastern scarp forests. In: Mucina L. (ed.) Vegetation Survey and Classification of Subtropical Forests of Southern Africa: 125 - 225. Geobotany Studies series, Springer, Cham, Switzerland. https: // doi. org / 10.1007 / 978 - 3 - 319 - 67831 - 3 _ 6" type="book chapter" year="2018">
Mucina
<emphasis id="5CDDEAE2FFB4FFD7FA975162FAEEFAEC" box="[1336,1399,1307,1333]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
2018
</bibRefCitation>
); in leaf-litter and under logs.
</paragraph>
</subSubSection>
<subSubSection id="26B3657BFFB4FFD7FF1251FCFD0DF8B9" pageId="22" pageNumber="23" type="discussion">
<paragraph id="6E1636F0FFB4FFD7FF1251FCFEB6FA45" blockId="22.[189,303,1410,1436]" box="[189,303,1410,1436]" pageId="22" pageNumber="23">
<heading id="355E819CFFB4FFD7FF1251FCFEB6FA45" bold="true" box="[189,303,1410,1436]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="5CDDEAE2FFB4FFD7FF1251FCFEB6FA45" bold="true" box="[189,303,1410,1436]" pageId="22" pageNumber="23">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFB4FFD7FF1251CFFAEDF910" blockId="22.[189,1400,1457,1738]" pageId="22" pageNumber="23">
<taxonomicName id="A9A94D73FFB4FFD7FF1251CFFEEEFA12" authority="Cole &amp; Herbert, 2023" authorityName="Cole &amp; Herbert" authorityYear="2023" box="[189,375,1457,1483]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="mystica" status="sp. nov.">
<emphasis id="5CDDEAE2FFB4FFD7FF1251CFFEEEFA12" box="[189,375,1457,1483]" italics="true" pageId="22" pageNumber="23">Gulella mystica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47EE5799FFB4FFD7FE2F51CCFE42FA15" box="[384,475,1458,1484]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
does not bear close resemblance to any other species. The truncate columella lamella with downward-pointing ridge-like teeth at its lower end is unique. Its apertural dentition bears superficial resemblance to certain features of
<taxonomicName id="A9A94D73FFB4FFD7FD5F5185FBEAF9CC" authority="Burnup, 1925" authorityName="Burnup" authorityYear="1925" box="[752,1139,1530,1557]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="mfongosiensis">
<emphasis id="5CDDEAE2FFB4FFD7FD5F5185FC26F9CD" box="[752,959,1530,1557]" italics="true" pageId="22" pageNumber="23">G. mfongosiensis</emphasis>
Burnup, 1925
</taxonomicName>
and
<taxonomicName id="A9A94D73FFB4FFD7FB195185FEEEF9E0" authority="(Sturany, 1898)" baseAuthorityName="Sturany" baseAuthorityYear="1898" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="perissodonta">
<emphasis id="5CDDEAE2FFB4FFD7FB195185FAEEF9CD" box="[1206,1399,1530,1557]" italics="true" pageId="22" pageNumber="23">G. perissodonta</emphasis>
(Sturany, 1898)
</taxonomicName>
, namely the large labral tooth with a small tooth on its upper base, basal teeth to right and left of centre and the columella lamella with two ridges. Both
<taxonomicName id="A9A94D73FFB4FFD7FC14523DFB1DF984" authorityName="Burnup" authorityYear="1925" box="[955,1156,1603,1629]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="mfongosiensis">
<emphasis id="5CDDEAE2FFB4FFD7FC14523DFB1DF984" box="[955,1156,1603,1629]" italics="true" pageId="22" pageNumber="23">G. mfongosiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="A9A94D73FFB4FFD7FB14523DFAEFF984" baseAuthorityName="Sturany" baseAuthorityYear="1898" box="[1211,1398,1603,1629]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="perissodonta">
<emphasis id="5CDDEAE2FFB4FFD7FB14523DFAEFF984" box="[1211,1398,1603,1629]" italics="true" pageId="22" pageNumber="23">G. perissodonta</emphasis>
</taxonomicName>
have an additional parietal denticle and the former also has a denticle in the middle of the columella lip.
<taxonomicName id="A9A94D73FFB4FFD7FF4252F5FE33F97C" authorityName="Cole &amp; Herbert" authorityYear="2023" box="[237,426,1675,1701]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="mystica">
<emphasis id="5CDDEAE2FFB4FFD7FF4252F5FE33F97C" box="[237,426,1675,1701]" italics="true" pageId="22" pageNumber="23">Gulella mystica</emphasis>
</taxonomicName>
occurs in the same broad vicinity as these other two species (inland in northern
<collectingRegion id="AC6DF812FFB4FFD7FF1252D1FEE3F910" box="[189,378,1711,1737]" country="South Africa" name="KwaZulu-Natal" pageId="22" pageNumber="23">KwaZulu-Natal</collectingRegion>
), but they occur in drier bushveld habitats while
<taxonomicName id="A9A94D73FFB4FFD7FC1952CEFBADF910" authorityName="Cole &amp; Herbert" authorityYear="2023" box="[950,1076,1712,1738]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="mystica">
<emphasis id="5CDDEAE2FFB4FFD7FC1952CEFBADF910" box="[950,1076,1712,1738]" italics="true" pageId="22" pageNumber="23">G. mystica</emphasis>
</taxonomicName>
occurs in tall, moist forest.
</paragraph>
<paragraph id="6E1636F0FFB4FFD7FF12528CFEFCF8D5" blockId="22.[189,357,1778,1804]" box="[189,357,1778,1804]" pageId="22" pageNumber="23">
<heading id="355E819CFFB4FFD7FF12528CFEFCF8D5" bold="true" box="[189,357,1778,1804]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="5CDDEAE2FFB4FFD7FF12528CFEFCF8D5" bold="true" box="[189,357,1778,1804]" pageId="22" pageNumber="23">Conservation</emphasis>
</heading>
</paragraph>
<paragraph id="6E1636F0FFB4FFD7FF12535FFD0DF8B9" blockId="22.[189,1399,1825,1888]" pageId="22" pageNumber="23">
<taxonomicName id="A9A94D73FFB4FFD7FF12535FFEE0F8E2" authority="Cole &amp; Herbert, 2023" authorityName="Cole &amp; Herbert" authorityYear="2023" box="[189,377,1825,1851]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="mystica" status="sp. nov.">
<emphasis id="5CDDEAE2FFB4FFD7FF12535FFEE0F8E2" box="[189,377,1825,1851]" italics="true" pageId="22" pageNumber="23">Gulella mystica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47EE5799FFB4FFD7FE2C535CFE78F8E5" box="[387,481,1826,1852]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
has been found only at Nkandla Forest Reserve, a protected area under the jurisdiction of Ezemvelo-KZN Wildlife.
</paragraph>
</subSubSection>
</treatment>
</document>