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<document id="A0C985C0871C21AA9C95C2EC70964B82" ID-DOI="10.17161/jom.i110.17001" ID-ISSN="2325-4467" ID-Zenodo-Dep="13146150" ID-ZooBank="urn:lsid:zoobank.org:pub:53C3B699-96AD-4692-9AB3-05058738EADB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722456082791" checkinUser="felipe" docAuthor="Engel, Michael S." docDate="2022" docId="926F87F592100D1715831E78B132F9C0" docLanguage="en" docName="JMelittology.2022.110.1-51.pdf" docOrigin="Journal of Melittology 2022 (110)" docSource="http://dx.doi.org/10.17161/jom.i110.17001" docStyle="DocumentStyle:51747547F162FCF7F4DBBB3D518F67EC.3:JMelittology.2013-.journal_article.2cover" docStyleId="51747547F162FCF7F4DBBB3D518F67EC" docStyleName="JMelittology.2013-.journal_article.2cover" docStyleVersion="3" docTitle="Scaptotrigona rosellae Engel 2022, new species" docType="treatment" docVersion="2" lastPageNumber="22" masterDocId="6E56FF8D92050D00141C1A19B220FFD9" masterDocTitle="Notes on South American stingless bees of the genus Scaptotrigona (Hymenoptera: Apidae), Part II: Subgroup A of the postica species group" masterLastPageNumber="51" masterPageNumber="1" pageNumber="20" updateTime="1722456440860" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
<mods:mods id="852FA974522D03AEB2FB829826A4CDFF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AC916B2046DA6F26A593AC6761C2D5FD">
<mods:title id="C4AFEEFBA082F530DB38193448C1A7F1">Notes on South American stingless bees of the genus Scaptotrigona (Hymenoptera: Apidae), Part II: Subgroup A of the postica species group</mods:title>
</mods:titleInfo>
<mods:name id="0CF390A0CC589CDB03768CFB5325A526" type="personal">
<mods:role id="F64C79FE8BFBC1469C3F41D7FAA90011">
<mods:roleTerm id="12B92FE1356F3FA9C28EC94A21B6298B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1E0A1357713B4193DE96CBB17BE28CCB">Engel, Michael S.</mods:namePart>
<mods:affiliation id="C9A7D24A51EC7CCE60859CD423A65146">Division of Entomology, Natural History Museum and Department of Ecology &amp; Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 - 4415, USA (msengel @ ku. edu). &amp; Division of Invertebrate Zoölogy, American Museum of Natural History, Central Park West at 79 Street, New York, New York 10024 - 5192, USA.</mods:affiliation>
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<mods:title id="C1524D560ECEF7B1840043E2ED7FF77A">Journal of Melittology</mods:title>
</mods:titleInfo>
<mods:part id="572E366DF2CC1896CEB9E991E0F33A7F">
<mods:date id="0BAD6C31CEF9D6F57F88831EA4988627">2022</mods:date>
<mods:detail id="7202BFD3BFE406100AB811E3D05BDCE1" type="pubDate">
<mods:number id="4B6E7ADDA6D055E671E6D64BF3664FFC">2022-03-14</mods:number>
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<mods:detail id="498485D02D8112B8F9605BD692F72C18" type="volume">
<mods:number id="436FA0DBD67091B1294D4771A6D1D743">2022</mods:number>
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<mods:detail id="BABDC4DD455105FEEE40F6DE5B3A5585" type="issue">
<mods:number id="CC909E1FC60CE6C1B1244F421EE16A06">110</mods:number>
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<mods:start id="D4DA074381AAA2EC4B849F33F2FEF920">1</mods:start>
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<mods:url id="52586DDD986C6EBD21334864703BF3C3">http://dx.doi.org/10.17161/jom.i110.17001</mods:url>
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<mods:classification id="1682AD37EA2B2B29CB32899B3EA52C4F">journal article</mods:classification>
<mods:identifier id="DB2D78C6B671D0446430D692C45751ED" type="DOI">10.17161/jom.i110.17001</mods:identifier>
<mods:identifier id="DF4006A387297FAAB04BF9814AD26BBB" type="ISSN">2325-4467</mods:identifier>
<mods:identifier id="CC18B76A7450D954A28C40E74E1C5215" type="Zenodo-Dep">13146150</mods:identifier>
<mods:identifier id="E2A592B8538F178A42AFAC833041D826" type="ZooBank">urn:lsid:zoobank.org:pub:53C3B699-96AD-4692-9AB3-05058738EADB</mods:identifier>
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<treatment id="926F87F592100D1715831E78B132F9C0" LSID="urn:lsid:plazi:treatment:926F87F592100D1715831E78B132F9C0" httpUri="http://treatment.plazi.org/id/926F87F592100D1715831E78B132F9C0" lastPageId="23" lastPageNumber="22" pageId="21" pageNumber="20">
<subSubSection id="52DC656892100D1515831E78B618FB40" pageId="21" pageNumber="20" type="nomenclature">
<paragraph id="1A7936E392100D1515831E78B1B1FBA0" blockId="21.[415,913,1121,1145]" box="[415,913,1121,1145]" pageId="21" pageNumber="20">
<heading id="4131818F92100D1515831E78B1B1FBA0" box="[415,913,1121,1145]" centered="true" fontSize="10" level="2" pageId="21" pageNumber="20" reason="2">
<taxonomicName id="DDC64D6092100D1515831E78B0D9FBA0" authority="Engel" authorityName="Engel" authorityYear="2022" box="[415,761,1121,1145]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="rosellae" status="sp. nov.">
<emphasis id="28B2EAF192100D1515831E78B08CFBA0" bold="true" box="[415,684,1121,1145]" italics="true" pageId="21" pageNumber="20">Scaptotrigona rosellae</emphasis>
Engel
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92100D1517181E7BB1B1FBA0" box="[772,913,1121,1145]" pageId="21" pageNumber="20" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="1A7936E392100D1514E41E9AB618FB40" blockId="21.[248,1080,1155,1209]" box="[248,1080,1155,1177]" pageId="21" pageNumber="20">
ZooBank:
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urn:lsid:zoobank.org:act:
<uuid id="6E600C3692100D1516451E9AB618FB40" box="[601,1080,1155,1177]" pageId="21" pageNumber="20">115BA1F9-566A-43A4-B717-6130B3324BCD</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="52DC656892100D1516531EB8B0C1FB60" box="[591,737,1185,1209]" pageId="21" pageNumber="20" type="description">
<paragraph id="1A7936E392100D1516531EB8B0C1FB60" blockId="21.[248,1080,1155,1209]" box="[591,737,1185,1209]" pageId="21" pageNumber="20">
(
<figureCitation id="82FD2A6692100D15164B1EB8B0F9FB60" box="[599,729,1185,1209]" captionStart="Figures 26–28" captionStartId="22.[160,241,1387,1409]" captionTargetBox="[165,1159,241,1371]" captionTargetId="figure-139@22.[160,1168,224,1381]" captionTargetPageId="22" captionText="Figures 26–28. Worker of Scaptotrigona rosellae, new species. 26. Lateral habitus. 27. Facial view. 28. Dorsal view of vertex and mesosoma." figureDoi="http://doi.org/10.5281/zenodo.13146170" httpUri="https://zenodo.org/record/13146170/files/figure.png" pageId="21" pageNumber="20">Figs. 26–28</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="52DC656892100D1514CC1EF8B0DEFAA0" pageId="21" pageNumber="20" type="diagnosis">
<paragraph id="1A7936E392100D1514CC1EF8B0DEFAA0" blockId="21.[160,1175,1249,1785]" pageId="21" pageNumber="20">
DIAGNOSIS: This species is currently known only from the lower Anchicayá River Valley of
<collectingCountry id="62D1767392100D15150F1F18B3A9FAC0" box="[275,393,1281,1305]" name="Colombia" pageId="21" pageNumber="20">Colombia</collectingCountry>
, an area known for its considerable biodiversity and endemism. The species superficially resembles a lighter form of
<taxonomicName id="DDC64D6092100D1517161F3BB17AFAE0" authorityName="Engel" authorityYear="2022" box="[778,858,1313,1337]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192100D1517161F3BB17AFAE0" box="[778,858,1313,1337]" italics="true" pageId="21" pageNumber="20">S. ederi</emphasis>
</taxonomicName>
but is immediately distin- guished by the presence of a genal tooth, the presence of weak striping on the mesos- cutum, and the generally yellow to fulvous bristles.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892100D1714CC1F98B30FFA80" lastPageId="23" lastPageNumber="22" pageId="21" pageNumber="20" type="description">
<paragraph id="1A7936E392100D1514CC1F98B1ABF920" blockId="21.[160,1175,1249,1785]" pageId="21" pageNumber="20">
DESCRIPTION: As described for
<taxonomicName id="DDC64D6092100D1516331F9BB05DFA40" authorityName="Engel" authorityYear="2022" box="[559,637,1409,1433]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192100D1516331F9BB05DFA40" box="[559,637,1409,1433]" italics="true" pageId="21" pageNumber="20">S. ederi</emphasis>
</taxonomicName>
(
<emphasis id="28B2EAF192100D1516971F98B0D9FA40" box="[651,761,1409,1433]" italics="true" pageId="21" pageNumber="20">vide supra</emphasis>
), with the following modifications: ⚲: Total body length approximately
<quantity id="DD3E9B0692100D1516521FB8B0F7FA60" box="[590,727,1441,1465]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="6.2" metricValueMin="6.1" pageId="21" pageNumber="20" unit="mm" value="6.15" valueMax="6.2" valueMin="6.1">6.1–6.2 mm</quantity>
, forewing length (to base of humeral sclerite) 6.0–
<quantity id="DD3E9B0692100D15152B1FD8B3B3FA00" box="[311,403,1473,1497]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="21" pageNumber="20" unit="mm" value="6.1">6.1 mm</quantity>
. Head wider than long, width
<quantity id="DD3E9B0692100D1517071FD8B1E1FA00" box="[795,961,1473,1497]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.48" metricValueMin="2.42" pageId="21" pageNumber="20" unit="mm" value="2.45" valueMax="2.48" valueMin="2.42">2.42–2.48 mm</quantity>
, length
<quantity id="DD3E9B0692100D1510391FD8B2F1FA20" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.03" metricValueMin="1.97" pageId="21" pageNumber="20" unit="mm" value="2.0" valueMax="2.03" valueMin="1.97">1.97–2.03 mm</quantity>
; compound eye length
<quantity id="DD3E9B0692100D1515F11FF8B0B1FA20" box="[493,657,1505,1529]" metricMagnitude="-3" metricUnit="m" metricValue="1.39" metricValueMax="1.42" metricValueMin="1.36" pageId="21" pageNumber="20" unit="mm" value="1.39" valueMax="1.42" valueMin="1.36">1.36–1.42 mm</quantity>
; upper interorbital distance
<quantity id="DD3E9B0692100D1517FB1FF8B6ABFA20" box="[999,1163,1505,1529]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.58" metricValueMin="1.52" pageId="21" pageNumber="20" unit="mm" value="1.55" valueMax="1.58" valueMin="1.52">1.52–1.58 mm</quantity>
, lower interorbital distance
<quantity id="DD3E9B0692100D1515C61C18B05DF9C0" box="[474,637,1537,1561]" metricMagnitude="-3" metricUnit="m" metricValue="1.4849999999999999" metricValueMax="1.52" metricValueMin="1.45" pageId="21" pageNumber="20" unit="mm" value="1.4849999999999999" valueMax="1.52" valueMin="1.45">1.45–1.52 mm</quantity>
. Scape length
<quantity id="DD3E9B0692100D15173A1C18B1ABF9C0" box="[806,907,1537,1561]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" pageId="21" pageNumber="20" unit="mm" value="0.91">0.91 mm</quantity>
, about as long as toru- locellar distance, torulocellar distance
<quantity id="DD3E9B0692100D15167F1C38B126F9E0" box="[611,774,1569,1593]" metricMagnitude="-4" metricUnit="m" metricValue="8.95" metricValueMax="9.1" metricValueMin="8.8" pageId="21" pageNumber="20" unit="mm" value="0.895" valueMax="0.91" valueMin="0.88">0.88–0.91 mm</quantity>
. Clypeus approximately 1.7–1.8× as wide as long, length
<quantity id="DD3E9B0692100D1515A81C58B03BF980" box="[436,539,1601,1625]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="21" pageNumber="20" unit="mm" value="0.64">0.64 mm</quantity>
, width
<quantity id="DD3E9B0692100D15166F1C58B137F980" box="[627,791,1601,1625]" metricMagnitude="-3" metricUnit="m" metricValue="1.13" metricValueMax="1.15" metricValueMin="1.11" pageId="21" pageNumber="20" unit="mm" value="1.13" valueMax="1.15" valueMin="1.11">1.11–1.15 mm</quantity>
. Malar area approximately 1.8× flagellar diameter or 0.82× basal mandibular width. Preoccipital ridge carinate, lamel- late dorsally, with bordering medial and lateral indentations deep, carina interrupted laterally by deep concavity, lower margin of concavity with carina forming acutely rounded angle projecting upward as a sharp, lamellate tooth, upper margin of concav- ity with lamella extending ventrally a third to half of concavity.
</paragraph>
<caption id="4EB9666B92130D1614BC1F72B073FA47" ID-DOI="http://doi.org/10.5281/zenodo.13146170" ID-Zenodo-Dep="13146170" httpUri="https://zenodo.org/record/13146170/files/figure.png" pageId="22" pageNumber="21" startId="22.[160,241,1387,1409]" targetBox="[165,1159,241,1371]" targetPageId="22" targetType="figure">
<paragraph id="1A7936E392130D1614BC1F72B073FA47" blockId="22.[160,1174,1387,1438]" pageId="22" pageNumber="21">
<emphasis id="28B2EAF192130D1614BC1F72B318FA58" bold="true" box="[160,312,1387,1409]" pageId="22" pageNumber="21">Figures 26–28.</emphasis>
Worker of
<taxonomicName id="DDC64D6092130D1615AE1F75B0A6FA58" authorityName="Engel" authorityYear="2022" box="[434,646,1388,1409]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="rosellae" status="sp. nov.">
<emphasis id="28B2EAF192130D1615AE1F75B0A6FA58" box="[434,646,1388,1409]" italics="true" pageId="22" pageNumber="21">Scaptotrigona rosellae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3381578A92130D16168D1F75B132FA5B" box="[657,786,1388,1410]" pageId="22" pageNumber="21" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="28B2EAF192130D1617031F75B11DFA58" bold="true" box="[799,829,1388,1409]" pageId="22" pageNumber="21">26.</emphasis>
Lateral habitus.
<emphasis id="28B2EAF192130D1617EC1F75B62EFA58" bold="true" box="[1008,1038,1388,1409]" pageId="22" pageNumber="21">27.</emphasis>
Facial view.
<emphasis id="28B2EAF192130D1614BC1F91B29EFA44" bold="true" box="[160,190,1416,1437]" pageId="22" pageNumber="21">28.</emphasis>
Dorsal view of vertex and mesosoma.
</paragraph>
</caption>
<paragraph id="1A7936E392130D1714CC1FD0B0DCFD40" blockId="22.[160,1168,1480,1793]" lastBlockId="23.[160,1168,225,1562]" lastPageId="23" lastPageNumber="22" pageId="22" pageNumber="21">
Integument of labiomaxillary complex yellow orange; labrum yellow orange; mandible yellow to reddish orange; clypeus yellow orange, epistomal sulcus well de- lineated by orange brown to brown; supraclypeal area yellow orange; malar space yellow orange; face below upper tangent of antennal toruli yellow orange; lower frons yellow orange blending to reddish orange to brown on upper frons, separated from in- ner and upper orbit by strip of yellow orange to reddish orange; vertex reddish brown to dark brown; ocellocular area reddish brown to dark brown except laterally where separated from upper orbit; posterior of head orange with strips of reddish brown to dark brown extending from ocellocular area to preoccipital carina, preoccipital carina dark brown to black; gena and postgena yellow orange to orange; scape yellow or- ange except sometimes with brown dorsoapically; pedicel and flagellum brown, with flagellum more orange ventrally. Mesosoma largely yellow orange to orange except mesoscutum black with reddish brown laterally and posteriorly, faint longitudinal strips of reddish brown from notauli to posterior border; mesepisternum with brown to dark brown patch ventrally; basal area and posterior surface of propodeum reddish brown to dark brown; metepisternum sometimes stained brown; tegula wholly yellow orange and semi-translucent; legs yellow orange. Metasoma generally brown to dark brown except anterior-facing surface of tergum I yellow orange, dorsal-facing disc of tergum I largely yellow orange with yellow brown laterally, sometimes lateral areas dark brown and more encroaching on yellow orange disc; tergum II largely yellow orange with small areas of brown laterally, although sometimes largely dark brown and blending to orange posteriorly before broad yellow marginal zone; terga
<collectionCode id="7CD7AE2692120D171057185BB646FD80" box="[1099,1126,578,601]" pageId="23" pageNumber="22">III</collectionCode>
–
<collectionCode id="7CD7AE2692120D171068185BB6B0FD80" box="[1140,1168,578,601]" pageId="23" pageNumber="22">IV</collectionCode>
brown to dark brown with broad yellow marginal zones, largely obscured by tomen- tum; sterna largely yellow orange to yellow brown.
</paragraph>
<paragraph id="1A7936E392120D1714CC18BBB31BFAC0" blockId="23.[160,1168,225,1562]" pageId="23" pageNumber="22">
As for
<taxonomicName id="DDC64D6092120D17150218BBB34EFD60" authorityName="Engel" authorityYear="2022" box="[286,366,673,697]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192120D17150218BBB34EFD60" box="[286,366,673,697]" italics="true" pageId="23" pageNumber="22">S. ederi</emphasis>
</taxonomicName>
except areas of short, plumose setae yellow to pale yellow instead of white; metepisternum and lateral surface of propodeum obscured by dense, plumose, yellow setae; dorsal-facing surface of tergum I with largely simple setae. Bristles of vertex erect, simple, yellow to fulvous; pronotal dorsolateral margin near and anterior to pronotal lobe with long, arched, yellow bristles; mesoscutum anterior and lateral borders with erect bristles, those on lateral border short, anterior margin with largely fulvous bristles, sometimes intermixed with yellow to dark fulvous bristles; tegula with short, suberect, yellow bristles anteriorly; mesoscutellum with abundant, elon- gate, erect bristles posteriorly, such bristles minutely pectinate apically, discal bristles shorter, mesoscutellar bristles yellow to fulvous; mesepisternal bristles as in
<taxonomicName id="DDC64D6092120D17102119DBB6B0FC00" authorityName="Engel" authorityYear="2022" box="[1085,1168,961,985]" class="Insecta" family="Apidae" genus="Scaptotrigona" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="ederi">
<emphasis id="28B2EAF192120D17102119DBB6B0FC00" box="[1085,1168,961,985]" italics="true" pageId="23" pageNumber="22">S. ederi</emphasis>
</taxonomicName>
except yellow to fulvous. Legs with abundant bristles, bristles typically yellow to ful- vous except sometimes noticeably intermixed with fuscous to black bristles on prolat- eral surface of mesotibia apically, prolateral surface of mesobasitarsus, metafemur at extreme apex; corbicular bristles fuscous to black, marginal bristles intermixed with some yellow to fulvous bristles; prolateral surface of metabasitarsus with scatered fine, simple, yellow setae (noticeably thinner than marginal bristles), but without black setae, margins with black bristles intermixed with yellow to fulvous bristles; penicil- lum dark fulvous to fuscous; rastellum fulvous. Metasoma with numerous, subde- cumbent, yellow bristles on terga
<collectionCode id="7CD7AE2692120D1716351EFBB064FB20" box="[553,580,1250,1273]" pageId="23" pageNumber="22">III</collectionCode>
–
<collectionCode id="7CD7AE2692120D17164D1EFBB044FB20" box="[593,612,1250,1273]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="23" pageNumber="22" type="Museum">V</collectionCode>
arising from tomentum, similar bristles sparser on tergum II.
</paragraph>
<paragraph id="1A7936E392120D1714CC1F3BB30FFAE0" blockId="23.[160,1168,225,1562]" box="[208,303,1314,1340]" pageId="23" pageNumber="22">
<specimenCount id="0CC0FD6A92120D1714CC1F3BB2C4FAE5" box="[208,228,1314,1340]" count="1" pageId="23" pageNumber="22" type="female">♀</specimenCount>
:
<emphasis id="28B2EAF192120D1714ED1F3BB308FAE0" box="[241,296,1314,1337]" italics="true" pageId="23" pageNumber="22">Latet</emphasis>
.
</paragraph>
<paragraph id="1A7936E392120D1714CC1F5BB30FFA80" blockId="23.[160,1168,225,1562]" box="[208,303,1346,1372]" pageId="23" pageNumber="22">
<specimenCount id="0CC0FD6A92120D1714CC1F5BB2C4FA85" box="[208,228,1346,1372]" count="1" pageId="23" pageNumber="22" type="male">♂</specimenCount>
:
<emphasis id="28B2EAF192120D1714ED1F5BB308FA80" box="[241,296,1346,1369]" italics="true" pageId="23" pageNumber="22">Latet</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892120D1714CC1F78B3F1FA00" pageId="23" pageNumber="22" type="materials_examined">
<paragraph id="1A7936E392120D1714CC1F78B03CFA40" blockId="23.[160,1168,225,1562]" pageId="23" pageNumber="22">
<materialsCitation id="AAAE3CBE92120D1714CC1F78B03CFA40" collectingDate="1977-02-11" collectionCode="SEMC" collectorName="M. D. Breed &amp; C. D. Michener" country="Colombia" elevation="400" location="Rio Anchicaya" pageId="23" pageNumber="22" specimenCount="1" typeStatus="holotype">
<typeStatus id="C57D884192120D1714CC1F78B368FAA0" box="[208,328,1377,1401]" pageId="23" pageNumber="22" type="holotype">HOLOTYPE</typeStatus>
: ⚲,
<collectingCountry id="62D1767392120D17156A1F78B3CCFAA0" box="[374,492,1377,1401]" name="Colombia" pageId="23" pageNumber="22">Colombia</collectingCountry>
: Prov. Valle,
<location id="1F19603892120D17168D1F78B161FAA0" LSID="urn:lsid:plazi:treatment:926F87F592100D1715831E78B132F9C0:1F19603892120D17168D1F78B161FAA0" box="[657,833,1377,1401]" country="Colombia" name="Rio Anchicaya" pageId="23" pageNumber="22">Rio Anchicaya</location>
,
<quantity id="DD3E9B0692120D17174C1F78B1BDFAA0" box="[848,925,1377,1401]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="23" pageNumber="22" unit="m" value="400.0">
<elevation id="91EBD1D092120D17174C1F78B1BDFAA0" box="[848,925,1377,1401]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="23" pageNumber="22" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<date id="6E78102392120D1717B01F78B666FAA0" box="[940,1094,1377,1401]" pageId="23" pageNumber="22" value="1977-02-11">
<collectingDate id="7E3CE9CB92120D1717B01F78B666FAA0" box="[940,1094,1377,1401]" pageId="23" pageNumber="22" value="1977-02-11">11 Feb. 1977</collectingDate>
</date>
,
<collectorName id="B733533592120D1710491F7BB2C5FA40" pageId="23" pageNumber="22">M.D. Breed</collectorName>
&amp;
<collectorName id="B733533592120D17151B1F98B392FA40" box="[263,434,1409,1433]" pageId="23" pageNumber="22">C.D. Michener</collectorName>
(
<collectionCode id="7CD7AE2692120D1715DD1F98B031FA40" box="[449,529,1409,1433]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="23" pageNumber="22" type="Museum">SEMC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="1A7936E392120D1714CC1FBBB3F1FA00" blockId="23.[160,1168,225,1562]" pageId="23" pageNumber="22">
<materialsCitation id="AAAE3CBE92120D1714CC1FBBB3EDFA00" collectingDate="1977-02-10" collectionCode="SEMC" collectorName="M. D. Breed &amp; C. D. Michener" country="Colombia" elevation="400" location="Rio Anchicaya" pageId="23" pageNumber="22" specimenCount="1" typeStatus="paratype">
<typeStatus id="C57D884192120D1714CC1FBBB363FA63" box="[208,323,1442,1466]" pageId="23" pageNumber="22" type="paratype">PARATYPE</typeStatus>
: ⚲,
<collectingCountry id="62D1767392120D1715791FB8B3FBFA60" box="[357,475,1441,1465]" name="Colombia" pageId="23" pageNumber="22">Colombia</collectingCountry>
: Prov. Valle,
<location id="1F19603892120D1716731FB8B139FA60" LSID="urn:lsid:plazi:treatment:926F87F592100D1715831E78B132F9C0:1F19603892120D1716731FB8B139FA60" box="[623,793,1441,1465]" country="Colombia" name="Rio Anchicaya" pageId="23" pageNumber="22">Rio Anchicaya</location>
,
<quantity id="DD3E9B0692120D17173F1FB8B14AFA60" box="[803,874,1441,1465]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="23" pageNumber="22" unit="m" value="400.0">
<elevation id="91EBD1D092120D17173F1FB8B14AFA60" box="[803,874,1441,1465]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="23" pageNumber="22" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<date id="6E78102392120D1717681FB8B622FA60" box="[884,1026,1441,1465]" pageId="23" pageNumber="22" value="1977-02-10">
<collectingDate id="7E3CE9CB92120D1717681FB8B622FA60" box="[884,1026,1441,1465]" pageId="23" pageNumber="22" value="1977-02-10">10 Feb. 1977</collectingDate>
</date>
,
<collectorName id="B733533592120D1710171FBBB6B0FA60" box="[1035,1168,1441,1465]" pageId="23" pageNumber="22">M.D. Breed</collectorName>
&amp;
<collectorName id="B733533592120D1714A71FD8B347FA00" box="[187,359,1473,1497]" pageId="23" pageNumber="22">C.D. Michener</collectorName>
(
<collectionCode id="7CD7AE2692120D1715691FD8B3E5FA00" box="[373,453,1473,1497]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="23" pageNumber="22" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="52DC656892120D1714CC1FF8B132F9C0" pageId="23" pageNumber="22" type="etymology">
<paragraph id="1A7936E392120D1714CC1FF8B132F9C0" blockId="23.[160,1168,225,1562]" pageId="23" pageNumber="22">ETYMOLOGY: The specific epithet honors my goddaughter, Ms. Rosella Amparo Be- tancourt, a brilliant young poet and budding scholar.</paragraph>
</subSubSection>
</treatment>
</document>
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