322 lines
33 KiB
XML
322 lines
33 KiB
XML
<document id="4DB35FDB58F246ADE602C635BE6A6E1F" ID-DOI="10.11646/zootaxa.4099.1.1" ID-GBIF-Dataset="5d7eab55-66e2-43ed-b764-bac0c6e5b6d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="268274" ID-ZooBank="B73C1BE1-346A-4C89-86B2-51CDACE92877" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460119691825" checkinUser="plazi" docAuthor="François Génier & Patrick Arnaud" docDate="2016" docId="03C68782F057FFC1FF42FF75FB73FCC5" docLanguage="en" docName="zootaxa.4099.1.1.pdf.imf" docOrigin="Zootaxa 4099 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dendropaemon (Nigropaemon) nigritulus Genier & Arnaud, new species" docType="treatment" docVersion="10" lastPageNumber="55" masterDocId="FFFFFFFAF062FFF7FFD5FFEDFFE6FFD0" masterDocTitle="Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)" pageNumber="54" updateTime="1698234910999" updateUser="plazi">
|
||
<mods:mods id="C35AD660FE0182AE6E40FE604D0C1AE6" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="0374A9CD7FD8740B48458829A749E29A">
|
||
<mods:title id="B07FF404171A083001071EE808A7CAA0">Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="6390716D20BCD5527835A2CA0E2D8764" type="personal">
|
||
<mods:role id="D22C276F9EE3F05F428F3F42D7F3A124">
|
||
<mods:roleTerm id="E068AA52574C012D3588AA080142CF88">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C4A9C5D6652DE586FD2AE10C79E104C5">François Génier</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="F39CA40927BBE60502786A0D524A374F" type="personal">
|
||
<mods:role id="0441AE40B1B639CFB5C9FAD03BE0175E">
|
||
<mods:roleTerm id="1D274C3541B1589F85D79D1FD55A83C0">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C52955A5D5F664B76979D604DB0CBFC5">Patrick Arnaud</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="235282FCA232C8A1E74113FEF9BB2241">text</mods:typeOfResource>
|
||
<mods:relatedItem id="D2BA69DF085EFEE0F37FAC0B521333E2" type="host">
|
||
<mods:titleInfo id="70779AB1B9765BF4D25BF9CFB20EF965">
|
||
<mods:title id="F3024453CE0C9D4D6902CB3E28CA7467">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="AB9C5BC6F9B7F2170E291C7FE09F3BD4">
|
||
<mods:date id="8BA3A56877F12CFCF74C4CF81A7E6B32">2016</mods:date>
|
||
<mods:detail id="E8DB25FFF3F0D044D57B62B6BCE60E5A" type="volume">
|
||
<mods:number id="850F71BB786FAFE9119CA5A427F19D49">4099</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="0EA907EB6797F78D5D66D842E749121F" type="issue">
|
||
<mods:number id="A90F6A1996657C72FB8D97D16A10A185">1</mods:number>
|
||
</mods:detail>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="4CE7E5A4EA4265CD36D8FA17200223FD">journal volume</mods:classification>
|
||
<mods:identifier id="F9A0174042B36022FD18534B3A59CCDC" type="DOI">10.11646/zootaxa.4099.1.1</mods:identifier>
|
||
<mods:identifier id="C2CB07C90C46C28478402ABDD647E1FA" type="GBIF-Dataset">5d7eab55-66e2-43ed-b764-bac0c6e5b6d6</mods:identifier>
|
||
<mods:identifier id="BD2C6260953446DBCD1AC93785FDC54B" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="B09DABC81F4A3897DC45785504A0C7ED" type="Zenodo-Dep">268274</mods:identifier>
|
||
<mods:identifier id="3D0D9364B626476BFF837E2F60A8EC6C" type="ZooBank">B73C1BE1-346A-4C89-86B2-51CDACE92877</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03C68782F057FFC1FF42FF75FB73FCC5" ID-DOI="http://doi.org/10.5281/zenodo.5780318" ID-GBIF-Taxon="127631966" ID-Zenodo-Dep="5780318" LSID="urn:lsid:plazi:treatment:03C68782F057FFC1FF42FF75FB73FCC5" httpUri="http://treatment.plazi.org/id/03C68782F057FFC1FF42FF75FB73FCC5" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54">
|
||
<subSubSection id="C375651FF057FFC2FF42FF75FBA4FF62" box="[151,1090,152,178]" pageId="53" pageNumber="54" type="nomenclature">
|
||
<paragraph id="8BD03694F057FFC2FF42FF75FBA4FF62" blockId="53.[151,1090,152,211]" box="[151,1090,152,178]" pageId="53" pageNumber="54">
|
||
<heading id="D09881F8F057FFC2FF42FF75FBA4FF62" bold="true" box="[151,1090,152,178]" fontSize="11" level="1" pageId="53" pageNumber="54" reason="1">
|
||
<emphasis id="B91BEA86F057FFC2FF42FF75FBA4FF62" bold="true" box="[151,1090,152,178]" pageId="53" pageNumber="54">
|
||
26.
|
||
<taxonomicName id="4C6F4D17F057FFC2FF16FF75FC42FF62" ID-CoL="34R2V" authority="Genier & Arnaud" authorityName="Genier & Arnaud" box="[195,932,152,178]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="nigritulus" status="sp. nov." subGenus="Nigropaemon">
|
||
<emphasis id="B91BEA86F057FFC2FF16FF75FD5BFF62" bold="true" box="[195,701,152,178]" italics="true" pageId="53" pageNumber="54">Dendropaemon (Nigropaemon) nigritulus</emphasis>
|
||
Génier & Arnaud
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A22857FDF057FFC2FC67FF75FBA4FF62" box="[946,1090,152,178]" pageId="53" pageNumber="54" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C375651FF057FFC2FF42FF57FE21FF03" box="[151,455,186,211]" pageId="53" pageNumber="54" type="description">
|
||
<paragraph id="8BD03694F057FFC2FF42FF57FE21FF03" blockId="53.[151,1090,152,211]" box="[151,455,186,211]" pageId="53" pageNumber="54">
|
||
(
|
||
<figureCitation id="13542A11F057FFC2FF4AFF57FF1AFF03" box="[159,252,186,211]" captionStart="FIGURES 25 – 30" captionStartId="103.[151,264,1962,1985]" captionTargetBox="[269,1325,201,1926]" captionTargetId="figure@103.[251,1336,193,1941]" captionTargetPageId="103" captionText="FIGURES 25 – 30. Dorsal habitus. 25. D. inemarginatus, female holotype; 26. D. nigritulus, male holotype; 27. D. amyntas, male; 28. D. attalus, male holotype; 29. D. lydiae, male holotype; 30. D. morettoi, male holotype." httpUri="https://zenodo.org/record/268279/files/figure.png" pageId="53" pageNumber="54">Figs. 26</figureCitation>
|
||
, 82–83, 121, 158)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C375651FF057FFC2FF42FF11FC6CFEC5" box="[151,906,252,277]" pageId="53" pageNumber="54" type="materials_examined">
|
||
<paragraph id="8BD03694F057FFC2FF42FF11FC6CFEC5" blockId="53.[151,1437,252,2028]" box="[151,906,252,277]" pageId="53" pageNumber="54">
|
||
<emphasis id="B91BEA86F057FFC2FF42FF11FECBFEC5" bold="true" box="[151,301,252,277]" pageId="53" pageNumber="54">
|
||
<typeStatus id="54D48836F057FFC2FF42FF11FF34FEC5" box="[151,210,252,277]" pageId="53" pageNumber="54">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
. Surumu, Serra Marari, Territorio Rio Branco,
|
||
<collectingCountry id="F3787604F057FFC2FCEAFF11FC60FEC5" box="[831,902,252,277]" name="Brazil" pageId="53" pageNumber="54">Brasil</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C375651FF057FFC2FF12FEF2FA64FE8C" pageId="53" pageNumber="54" type="diagnosis">
|
||
<paragraph id="8BD03694F057FFC2FF12FEF2FA64FE8C" blockId="53.[151,1437,252,2028]" pageId="53" pageNumber="54">
|
||
<emphasis id="B91BEA86F057FFC2FF12FEF2FEA7FEE7" bold="true" box="[199,321,287,312]" pageId="53" pageNumber="54">Diagnosis.</emphasis>
|
||
The small size (less than 12.0 mm) combined with the lack of metallic luster and the elongate second metatarsal segment, which is as long as or slightly longer than the first metatarsus is unique in the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C375651FF057FFC2FF12FE8AFA71F83C" pageId="53" pageNumber="54" type="description">
|
||
<paragraph id="8BD03694F057FFC2FF12FE8AFDA6F88D" blockId="53.[151,1437,252,2028]" pageId="53" pageNumber="54">
|
||
<emphasis id="B91BEA86F057FFC2FF12FE8AFEB7FE50" bold="true" box="[199,337,359,384]" pageId="53" pageNumber="54">Description</emphasis>
|
||
.
|
||
<collectingCountry id="F3787604F057FFC2FEB6FE8AFE7BFE50" box="[355,413,359,384]" name="Malta" pageId="53" pageNumber="54">Male</collectingCountry>
|
||
<typeStatus id="54D48836F057FFC2FE7DFE8AFDECFE50" box="[424,522,359,384]" pageId="53" pageNumber="54" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="13542A11F057FFC2FDCBFE8AFD90FE50" box="[542,630,359,384]" captionStart="FIGURES 25 – 30" captionStartId="103.[151,264,1962,1985]" captionTargetBox="[269,1325,201,1926]" captionTargetId="figure@103.[251,1336,193,1941]" captionTargetPageId="103" captionText="FIGURES 25 – 30. Dorsal habitus. 25. D. inemarginatus, female holotype; 26. D. nigritulus, male holotype; 27. D. amyntas, male; 28. D. attalus, male holotype; 29. D. lydiae, male holotype; 30. D. morettoi, male holotype." httpUri="https://zenodo.org/record/268279/files/figure.png" pageId="53" pageNumber="54">Fig. 26</figureCitation>
|
||
).
|
||
<emphasis id="B91BEA86F057FFC2FD58FE8AFD2FFE50" bold="true" box="[653,713,359,384]" pageId="53" pageNumber="54">Body</emphasis>
|
||
. Body small, length
|
||
<quantity id="4C979B71F057FFC2FC11FE8AFBC6FEAF" box="[964,1056,359,384]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="53" pageNumber="54" unit="mm" value="9.5">9.5 mm</quantity>
|
||
, maximum width 4.0 mm; body subrectangular in dorsal view; dorsum narrowly flat.
|
||
<emphasis id="B91BEA86F057FFC2FD2EFE67FCDAFE73" bold="true" box="[763,828,394,419]" pageId="53" pageNumber="54">Color</emphasis>
|
||
. Dorsal surface dark brown to black, glossy, lacking metallic sheen; ventrum dark brown to black; pygidium dark brown; legs reddish brown to dark brown.
|
||
<emphasis id="B91BEA86F057FFC2FA80FE43FA73FE17" bold="true" box="[1365,1429,430,455]" pageId="53" pageNumber="54">Head</emphasis>
|
||
. Clypeus semicircular, anterior portion slightly upturned; clypeal teeth triangular; clypeal median emargination broadly v-shaped, clypeal edge emarginate on external side of each clypeal tooth, clypeal teeth ventral surface with a fine v-shaped carina, clypeal margin ill-defined, lacking sharp carina posteriorly, clypeal surface with transverse blunt rugulae; clypeogenal suture ill-defined, bluntly carinate internally; genal surface with ill-defined rugulae anteriorly and minute scattered tubercles posteriorly on disc, with a long blunt transverse carina; clypeofrontal carina rather low, approximately 4 times wider than high, slightly arcuate in dorsal view, simply carinate, clypeofrontal carina apical edge straight in frontal view; eyes moderately large in dorsal view, interocular ratio 4.4.
|
||
<emphasis id="B91BEA86F057FFC2FF42FD21FEF6FD35" bold="true" box="[151,272,716,741]" pageId="53" pageNumber="54">Pronotum</emphasis>
|
||
. Pronotum transverse in dorsal view, pronotal width/length ratio 1.3; disc of pronotum minutely punctate basally, puncture becoming fine anteriorly, with a fine longitudinal sulcus on posterior two-third; pronotal anterior margin wider and flat lateral to eyes; anterior portion with a broad and fine tectiform carina tuberculate medially; anterior angles surface with few ill-defined rugulae, slightly but distinctly sulcate along posterior edge of anterior margin; lateral fossae simply rounded, concave; lateral portions unmodified; pronotal basal fossae very small and more or less rounded; posterior margin well-defined, lacking crenulation and setae.
|
||
<emphasis id="B91BEA86F057FFC2FB2DFC92FAA2FC48" bold="true" box="[1272,1348,895,920]" pageId="53" pageNumber="54">Elytra</emphasis>
|
||
. Elytra approximately as long as wide in dorsal view, elytral combined width/length ratio 0.9; elytral base lacking distinct margin, simply convex; elytral striae 1–4 fine, not bordered by a minute carina laterally, evenly impressed throughout, elytral striae 5 similar to 4 on disc, strial punctures ill-defined, adjacent strial edge encroaching on interval, stria 1 weakly impressed apically, going straight to elytral apical margin; interstriae slightly convex, minutely punctate throughout, surface glossy.
|
||
<emphasis id="B91BEA86F057FFC2FD7CFBDCFC8FFB9A" bold="true" box="[681,873,1073,1098]" pageId="53" pageNumber="54">Thoracic sterna</emphasis>
|
||
. Proepisternal carina absent; metasternal median lobe angularly produced anteromedially, ventral ridge ill-defined, keel shaped.
|
||
<emphasis id="B91BEA86F057FFC2FBF7FBB8FBBCFBBE" bold="true" box="[1058,1114,1109,1134]" pageId="53" pageNumber="54">Legs</emphasis>
|
||
. Profemur posterior surface convex, glabrous and punctate internally, posterointernal margin rather thin, evenly developed, internal edge rather wide, with a contiguous row of setae along anterointernal edge and few scattered long setae on anterior half, remaining surface smooth. Protibia with four teeth on lateral edge; internal basal angle lobate; anterior surface with long aligned row of setae internally, surface glossy or feebly microsculptured between punctures; posterior surface with some well-defined punctures externally to median carina, surface glossy between punctures, with a single interrupted setal row along lateral teeth. Mesofemur unmodified on anterointernal edge apically. Mesotibia rather short, gradually widening toward apex in anterior view; anteroapical edge slightly sinuate in anterior view, anteroapical row of setae complete; apicoanterior edge circularly indented internally; external edge more or less flat, with irregular setiferous punctures and microsculpture. Mesotarsus similar in shape to metatarsus, 3- segmented, first segment moderately elongate, approximately two times as long as wide at apex. Metafemur internal edge nearly straight and lateral edge arcuate, lacking distinct depressed area anterointernally before apex, apicoposterior edge unmodified, anterior surface with a well-defined sulcus on median half. Metatibia moderately slender, slightly widening toward apex in anterior view, anterior surface with distinct row of setae, with moderately dense minute punctures on a glossy surface, metatibial posterior surface flat between longitudinal row of setae and lateral edge, with ill-defined microsculpture. Metatarsus 3-segmented (
|
||
<figureCitation id="13542A11F057FFC2FC3BF980FBB1F956" box="[1006,1111,1645,1670]" captionStart="FIGURES 118 – 125" captionStartId="113.[151,264,1924,1947]" captionTargetBox="[217,1363,200,1895]" captionTargetId="figure@113.[211,1376,193,1903]" captionTargetPageId="113" captionText="FIGURES 118 – 125. Metatarsus, ventral view. 118. D. fredericki, holotype; 119. D. angustipennis; 120. D. bahianus; 121. D. nigritulus, paratype; 122. D. amyntas; 123. D. attalus; 124. D. lydiae, paratype; 125. D. morettoi, paratype." httpUri="https://zenodo.org/record/268283/files/figure.png" pageId="53" pageNumber="54">Fig. 121</figureCitation>
|
||
), first segment moderately elongate, approximately two times as long as wide at apex, with anterointernal carina well defined and almost reaching apical edge.
|
||
<emphasis id="B91BEA86F057FFC2FE4BF958FD70F91E" bold="true" box="[414,662,1717,1742]" pageId="53" pageNumber="54">Abdominal sternites</emphasis>
|
||
. Sternites 3–6 longitudinally slightly convex, segment 6 slightly concave; sternites 4–6 with 1–3 unaligned rows of setae laterally, glabrous medially; sternite 7 slightly longitudinally concave medially, shorter than segment 6 along midline; pygidium with moderate to large punctures, punctures separated by one diameter or less.
|
||
<emphasis id="B91BEA86F057FFC2FD70F8CDFCB5F8E9" bold="true" box="[677,851,1824,1849]" pageId="53" pageNumber="54">
|
||
<collectingCountry id="F3787604F057FFC2FD70F8CDFD07F8E9" box="[677,737,1824,1849]" name="Malta" pageId="53" pageNumber="54">Male</collectingCountry>
|
||
genitalia
|
||
</emphasis>
|
||
(Figs. 82–83). Parameres laterally concave before apex; surface smooth, glossy apically.
|
||
</paragraph>
|
||
<paragraph id="8BD03694F057FFC2FF12F88AFAB2F8AF" blockId="53.[151,1437,252,2028]" box="[199,1364,1895,1920]" pageId="53" pageNumber="54">
|
||
<emphasis id="B91BEA86F057FFC2FF12F88AFE90F850" bold="true" box="[199,374,1895,1920]" pageId="53" pageNumber="54">Measurements</emphasis>
|
||
(
|
||
<specimenCount id="9D69FD1DF057FFC2FE50F885FE39F850" box="[389,479,1895,1920]" pageId="53" pageNumber="54" type="male">4 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D69FD1DF057FFC2FE3EF885FDBFF850" box="[491,601,1895,1920]" pageId="53" pageNumber="54" type="female">9 females</specimenCount>
|
||
). Length: male 9.0–11.0 (9.8±0.9), female 9.5–12.0 (10.5±0.8) mm.
|
||
</paragraph>
|
||
<paragraph id="8BD03694F057FFC2FF12F866FA71F83C" blockId="53.[151,1437,252,2028]" pageId="53" pageNumber="54">
|
||
<emphasis id="B91BEA86F057FFC2FF12F866FE42F874" bold="true" box="[199,420,1931,1956]" pageId="53" pageNumber="54">
|
||
Primary
|
||
<typeStatus id="54D48836F057FFC2FEE1F866FE81F874" box="[308,359,1931,1956]" pageId="53" pageNumber="54">type</typeStatus>
|
||
data
|
||
</emphasis>
|
||
.
|
||
<typeStatus id="54D48836F057FFC2FE66F866FDFDF874" box="[435,539,1931,1956]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
|
||
male (CMNC): [Sept.956/
|
||
<collectingCountry id="F3787604F057FFC2FC86F861FC50F874" box="[851,950,1932,1956]" name="Brazil" pageId="53" pageNumber="54">BRASIL</collectingCountry>
|
||
/ T. Rio Branco/ Surumu/ Sa. Marari/ Coll. Martínez] handwritten; [H. & A. HOWDEN/ COLLECTION/ ex. A. Martínez coll.]; [WORLD/ SCARAB./ DATABASE/ WSD00017011]; [
|
||
<typeStatus id="54D48836F057FFC2FDD4F83EFD71F83B" box="[513,663,2003,2027]" pageId="53" pageNumber="54" type="holotype">HOLOTYPE</typeStatus>
|
||
/
|
||
<taxonomicName id="4C6F4D17F057FFC2FD72F83EFCB2F83C" box="[679,852,2003,2028]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91BEA86F057FFC2FD72F83EFCB2F83C" box="[679,852,2003,2028]" italics="true" pageId="53" pageNumber="54">Dendropaemon</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C6F4D17F057FFC2FCB6F83EFC36F83C" box="[867,976,2003,2028]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="nigritulus" status="sp. nov.">
|
||
<emphasis id="B91BEA86F057FFC2FCB6F83EFC36F83C" box="[867,976,2003,2028]" italics="true" pageId="53" pageNumber="54">nigritulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22857FDF057FFC2FC03F83EFBEFF83B" box="[982,1033,2003,2027]" pageId="53" pageNumber="54" rank="species">n.sp.</taxonomicNameLabel>
|
||
/ Génier & Arnaud, 2014] red card.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C375651FF054FFC1FF12FF7AFE2AFDB0" pageId="54" pageNumber="55" type="materials_examined">
|
||
<paragraph id="8BD03694F054FFC1FF12FF7AFE2AFDB0" blockId="54.[151,1436,151,789]" pageId="54" pageNumber="55">
|
||
<emphasis id="B91BEA86F054FFC1FF12FF7AFE4EFF60" bold="true" box="[199,424,151,176]" pageId="54" pageNumber="55">Material examined</emphasis>
|
||
.
|
||
<collectingCountry id="F3787604F054FFC1FE63FF7AFDC7FF60" box="[438,545,151,176]" name="Brazil" pageId="54" pageNumber="55">
|
||
<emphasis id="B91BEA86F054FFC1FE63FF7AFDC7FF60" bold="true" box="[438,545,151,176]" pageId="54" pageNumber="55">BRAZIL</emphasis>
|
||
</collectingCountry>
|
||
: AMAZONAS, Reserva Florestal Adolpho Ducke,
|
||
<quantity id="4C979B71F054FFC1FBB6FF7AFB4CFF60" box="[1123,1194,151,176]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="54" pageNumber="55" unit="km" value="26.0">26 km</quantity>
|
||
NE Manaus, (
|
||
<geoCoordinate id="EE5B5053F054FFC1FA9EFF7AFA71FF60" box="[1355,1431,151,176]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="-2.95">2°57'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5B5053F054FFC1FF42FF51FF1BFF05" box="[151,253,188,213]" direction="west" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="-59.95">59°57'W</geoCoordinate>
|
||
),
|
||
<date id="FFD11054F054FFC1FEC4FF51FE77FF05" box="[273,401,188,213]" pageId="54" pageNumber="55" value="1995" valueMax="1996">1995–1996</date>
|
||
, coll. [anonymous]—
|
||
<specimenCount id="9D69FD1DF054FFC1FD50FF51FD17FF05" box="[645,753,188,213]" pageId="54" pageNumber="55" type="female">2 females</specimenCount>
|
||
(incl. 2
|
||
<typeStatus id="54D48836F054FFC1FC9BFF51FC5DFF04" box="[846,955,188,212]" pageId="54" pageNumber="55" type="paratype">paratypes</typeStatus>
|
||
) (
|
||
<collectionCode id="ED7EAE51F054FFC1FC07FF51FBCAFF04" box="[978,1068,188,212]" pageId="54" pageNumber="55">BMNH</collectionCode>
|
||
); Rio Uatumã, (
|
||
<geoCoordinate id="EE5B5053F054FFC1FB0AFF51FACDFF04" box="[1247,1323,188,212]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="-2.5666666">2°34'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5B5053F054FFC1FAE2FF51FA69FF05" box="[1335,1423,188,213]" direction="west" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="-58.15">58°9'W</geoCoordinate>
|
||
),
|
||
<date id="FFD11054F054FFC1FF42FF32FF06FF28" box="[151,224,223,248]" pageId="54" pageNumber="55" value="1944-01">i.1944</date>
|
||
, coll. [anonymous]—
|
||
<specimenCount id="9D69FD1DF054FFC1FE0DFF0DFDD5FF28" box="[472,563,223,248]" pageId="54" pageNumber="55" type="female">1 female</specimenCount>
|
||
allotype (
|
||
<collectionCode id="ED7EAE51F054FFC1FD76FF0DFD1AFF28" box="[675,764,224,248]" pageId="54" pageNumber="55">CMNC</collectionCode>
|
||
); RORAIMA, [Rio] Surumu, Serra Marari [= Serra Saporá], (
|
||
<geoCoordinate id="EE5B5053F054FFC1FF4AFEE9FF16FECC" box="[159,240,260,284]" direction="north" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="4.383333">4°23'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5B5053F054FFC1FED5FEE9FE80FECD" box="[256,358,260,285]" direction="west" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="-60.75">60°45'W</geoCoordinate>
|
||
),
|
||
<date id="FFD11054F054FFC1FEA8FEE9FE33FECD" box="[381,469,260,285]" pageId="54" pageNumber="55" value="1956-09">ix.1956</date>
|
||
, coll. [anonymous]—
|
||
<specimenCount id="9D69FD1DF054FFC1FD01FEE9FCFBFECD" box="[724,797,260,285]" pageId="54" pageNumber="55" type="male">1 male</specimenCount>
|
||
(
|
||
<typeStatus id="54D48836F054FFC1FCE5FEE9FC70FECD" box="[816,918,260,285]" pageId="54" pageNumber="55" type="holotype">holotype</typeStatus>
|
||
) (
|
||
<collectionCode id="ED7EAE51F054FFC1FC7AFEE9FBEEFECC" box="[943,1032,260,284]" pageId="54" pageNumber="55">CMNC</collectionCode>
|
||
);
|
||
<collectingCountry id="F3787604F054FFC1FBCBFEE9FB5BFECD" box="[1054,1213,260,285]" name="Colombia" pageId="54" pageNumber="55">
|
||
<emphasis id="B91BEA86F054FFC1FBCBFEE9FB5BFECD" bold="true" box="[1054,1213,260,285]" pageId="54" pageNumber="55">COLOMBIA</emphasis>
|
||
</collectingCountry>
|
||
: ARAUCA, Tame, (
|
||
<geoCoordinate id="EE5B5053F054FFC1FF4AFECAFEF1FE90" box="[159,279,295,320]" direction="north" orientation="latitude" pageId="54" pageNumber="55" precision="15" value="6.4591665">6°27'33''N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5B5053F054FFC1FEFFFEC5FE50FE90" box="[298,438,296,320]" direction="west" orientation="longitude" pageId="54" pageNumber="55" precision="15" value="-71.736664">71°44'12''W</geoCoordinate>
|
||
),
|
||
<date id="FFD11054F054FFC1FE04FECAFDB5FE90" box="[465,595,295,320]" pageId="54" pageNumber="55" value="1976-07-27">27.vii.1976</date>
|
||
, coll. M. Cooper—
|
||
<specimenCount id="9D69FD1DF054FFC1FC92FEC5FC4CFE90" box="[839,938,295,320]" pageId="54" pageNumber="55" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus id="54D48836F054FFC1FC15FEC5FBC2FE90" box="[960,1060,296,320]" pageId="54" pageNumber="55" type="paratype">paratype</typeStatus>
|
||
) (
|
||
<collectionCode id="ED7EAE51F054FFC1FB97FEC5FB7AFE90" box="[1090,1180,296,320]" pageId="54" pageNumber="55">BMNH</collectionCode>
|
||
);
|
||
<collectingCountry id="F3787604F054FFC1FB63FECAFAC8FE90" box="[1206,1326,295,320]" name="Guyana" pageId="54" pageNumber="55">
|
||
<emphasis id="B91BEA86F054FFC1FB63FECAFAC8FE90" bold="true" box="[1206,1326,295,320]" pageId="54" pageNumber="55">GUYANA</emphasis>
|
||
</collectingCountry>
|
||
:
|
||
<collectionCode id="ED7EAE51F054FFC1FA90FEC5FA7AFEEF" box="[1349,1436,296,319]" pageId="54" pageNumber="55">UPPER</collectionCode>
|
||
DEMERARA-BERBICE, kurupukari, (
|
||
<geoCoordinate id="EE5B5053F054FFC1FD88FEA1FD49FEB4" box="[605,687,332,356]" direction="north" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="4.6666665">4°40'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5B5053F054FFC1FD6AFEA1FCC0FEB5" box="[703,806,332,357]" direction="west" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="-58.666668">58°40'W</geoCoordinate>
|
||
),
|
||
<date id="FFD11054F054FFC1FCEBFEA1FC5CFEB5" box="[830,954,332,357]" pageId="54" pageNumber="55" value="1992-04" valueMax="1992-11-31" valueMin="1992-04-00">iv–xi.1992</date>
|
||
, coll. [anonymous]—
|
||
<specimenCount id="9D69FD1DF054FFC1FB69FEA1FAFBFEB5" box="[1212,1309,332,357]" pageId="54" pageNumber="55" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus id="54D48836F054FFC1FAE4FEA1FA73FEB4" box="[1329,1429,332,356]" pageId="54" pageNumber="55" type="paratype">paratype</typeStatus>
|
||
) (
|
||
<collectionCode id="ED7EAE51F054FFC1FF4AFE9DFF1FFE58" box="[159,249,368,392]" pageId="54" pageNumber="55">BMNH</collectionCode>
|
||
);
|
||
<emphasis id="B91BEA86F054FFC1FEDAFE82FDD2FE58" bold="true" box="[271,564,367,392]" pageId="54" pageNumber="55">
|
||
<collectingCountry id="F3787604F054FFC1FEDAFE82FE60FE58" box="[271,390,367,392]" name="French Guiana" pageId="54" pageNumber="55">GUYANE</collectingCountry>
|
||
FRANÇAISE
|
||
</emphasis>
|
||
: PK 5, Piste KM 25, Route Régina-Saint-Georges, (
|
||
<geoCoordinate id="EE5B5053F054FFC1FB4DFE82FAE4FE58" box="[1176,1282,367,392]" direction="north" orientation="latitude" pageId="54" pageNumber="55" precision="15" value="4.115">4°6'54''N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5B5053F054FFC1FAC4FE82FA69FE58" box="[1297,1423,367,392]" direction="west" orientation="longitude" pageId="54" pageNumber="55" precision="15" value="-52.12111">52°7'16''W</geoCoordinate>
|
||
),
|
||
<date id="FFD11054F054FFC1FF42FE79FF18FE7D" box="[151,254,404,429]" pageId="54" pageNumber="55" value="2008-08">viii.2008</date>
|
||
, coll. J.L. Giuglaris—
|
||
<specimenCount id="9D69FD1DF054FFC1FE29FE79FDBEFE7D" box="[508,600,404,429]" pageId="54" pageNumber="55" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus id="54D48836F054FFC1FDB2FE79FD2DFE7C" box="[615,715,404,428]" pageId="54" pageNumber="55" type="paratype">paratype</typeStatus>
|
||
) (
|
||
<collectionCode id="ED7EAE51F054FFC1FD34FE79FCD7FE7C" box="[737,817,404,428]" pageId="54" pageNumber="55">ATHC</collectionCode>
|
||
); Réserve naturelle des Nouragues, (
|
||
<geoCoordinate id="EE5B5053F054FFC1FB19FE79FAFBFE7C" box="[1228,1309,404,428]" direction="north" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="4.3166666">4°19'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5B5053F054FFC1FAFCFE79FA76FE7D" box="[1321,1424,404,429]" direction="west" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="-52.366665">52°22'W</geoCoordinate>
|
||
),
|
||
<date id="FFD11054F054FFC1FF42FE5AFEC6FE00" box="[151,288,439,464]" pageId="54" pageNumber="55" value="2010-08-26">26.viii.2010</date>
|
||
, coll. [anonymous]—
|
||
<specimenCount id="9D69FD1DF054FFC1FDF2FE55FD95FE00" box="[551,627,439,464]" pageId="54" pageNumber="55" type="male">1 male</specimenCount>
|
||
(
|
||
<typeStatus id="54D48836F054FFC1FD5CFE55FD0BFE00" box="[649,749,440,464]" pageId="54" pageNumber="55" type="paratype">paratype</typeStatus>
|
||
) (
|
||
<collectionCode id="ED7EAE51F054FFC1FCDFFE55FCBFFE00" box="[778,857,440,464]" pageId="54" pageNumber="55">COBF</collectionCode>
|
||
); same locality,
|
||
<date id="FFD11054F054FFC1FBF4FE5AFB68FE00" box="[1057,1166,439,464]" pageId="54" pageNumber="55" value="2010-09-02">2.ix.2010</date>
|
||
, coll. [anonymous]—
|
||
<specimenCount id="9D69FD1DF054FFC1FA5AFE55FF08FE25" pageId="54" pageNumber="55" type="female">2 females</specimenCount>
|
||
(incl. 2
|
||
<typeStatus id="54D48836F054FFC1FE98FE31FE5CFE24" box="[333,442,476,500]" pageId="54" pageNumber="55" type="paratype">paratypes</typeStatus>
|
||
) (
|
||
<collectionCode id="ED7EAE51F054FFC1FE07FE31FDC7FE24" box="[466,545,476,500]" pageId="54" pageNumber="55">COBF</collectionCode>
|
||
);
|
||
<emphasis id="B91BEA86F054FFC1FDE0FE31FD2BFE25" bold="true" box="[565,717,476,501]" pageId="54" pageNumber="55">
|
||
<collectingCountry id="F3787604F054FFC1FDE0FE31FD2BFE25" box="[565,717,476,501]" name="Suriname" pageId="54" pageNumber="55">SURINAME</collectingCountry>
|
||
</emphasis>
|
||
: SIPALIWINI, Kutari River, Kwamalasamutu region, elev.
|
||
<quantity id="4C979B71F054FFC1FAA7FE31FF4BFDC7" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="54" pageNumber="55" unit="m" value="250.0">250 m</quantity>
|
||
(
|
||
<geoCoordinate id="EE5B5053F054FFC1FF17FDEDFEF2FDC8" box="[194,276,512,536]" direction="north" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="2.1666667">2°10'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5B5053F054FFC1FEF0FE12FE6DFDC8" box="[293,395,511,536]" direction="west" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="-56.783333">56°47'W</geoCoordinate>
|
||
),
|
||
<date id="FFD11054F054FFC1FE70FE12FDBFFDC8" box="[421,601,511,536]" pageId="54" pageNumber="55" value="2010-08-18" valueMax="2010-08-24" valueMin="2010-08-18">18–24.viii.2010</date>
|
||
, coll. T. Larsen—
|
||
<specimenCount id="9D69FD1DF054FFC1FCE1FDEDFC69FDC8" box="[820,911,511,536]" pageId="54" pageNumber="55" type="male">2 males</specimenCount>
|
||
(incl. 2
|
||
<typeStatus id="54D48836F054FFC1FC2BFDEDFB8DFDC8" box="[1022,1131,512,536]" pageId="54" pageNumber="55" type="paratype">paratypes</typeStatus>
|
||
) (
|
||
<collectionCode id="ED7EAE51F054FFC1FB52FDEDFB2BFDC8" box="[1159,1229,512,536]" pageId="54" pageNumber="55">AFIC</collectionCode>
|
||
);
|
||
<collectingCountry id="F3787604F054FFC1FB33FE12FA72FDC8" box="[1254,1428,511,536]" name="Venezuela" pageId="54" pageNumber="55">
|
||
<emphasis id="B91BEA86F054FFC1FB33FE12FA72FDC8" bold="true" box="[1254,1428,511,536]" pageId="54" pageNumber="55">VENEZUELA</emphasis>
|
||
</collectingCountry>
|
||
: BOLIVAR, Río Sipao,
|
||
<quantity id="4C979B71F054FFC1FE7DFDC9FDE7FDED" box="[424,513,548,573]" metricMagnitude="5" metricUnit="m" metricValue="1.1" pageId="54" pageNumber="55" unit="km" value="110.0">110 km</quantity>
|
||
E Caicara, (
|
||
<geoCoordinate id="EE5B5053F054FFC1FD4CFDC9FCF7FDEC" box="[665,785,548,572]" direction="north" orientation="latitude" pageId="54" pageNumber="55" precision="15" value="7.4130554">7°24'47''N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5B5053F054FFC1FCF5FDC9FC4BFDED" box="[800,941,548,573]" direction="west" orientation="longitude" pageId="54" pageNumber="55" precision="15" value="-65.206665">65°12'24''W</geoCoordinate>
|
||
),
|
||
<date id="FFD11054F054FFC1FC10FDC9FB69FDED" box="[965,1167,548,573]" pageId="54" pageNumber="55" value="1987-06-17" valueMax="1987-08-04" valueMin="1987-06-17">17.vi.–4.viii.1987</date>
|
||
, coll. S. & J. Peck—
|
||
<specimenCount id="9D69FD1DF054FFC1FA41FDC9FF05FDB0" pageId="54" pageNumber="55" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus id="54D48836F054FFC1FF27FDA5FEB0FDB0" box="[242,342,584,608]" pageId="54" pageNumber="55" type="paratype">paratype</typeStatus>
|
||
) (
|
||
<collectionCode id="ED7EAE51F054FFC1FEB9FDA5FE26FDB0" box="[364,448,584,608]" pageId="54" pageNumber="55">BDGC</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C375651FF054FFC1FF12FD81FB0DFD78" pageId="54" pageNumber="55" type="etymology">
|
||
<paragraph id="8BD03694F054FFC1FF12FD81FAEEFD55" blockId="54.[151,1436,151,789]" box="[199,1288,620,645]" pageId="54" pageNumber="55">
|
||
<emphasis id="B91BEA86F054FFC1FF12FD81FEA1FD55" bold="true" box="[199,327,620,645]" pageId="54" pageNumber="55">Etymology</emphasis>
|
||
.
|
||
<taxonomicName id="4C6F4D17F054FFC1FE81FD81FE23FD55" box="[340,453,620,645]" class="Insecta" family="Scarabaeidae" genus="Dendropaemon" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="119" phylum="Arthropoda" rank="species" species="nigritulus">
|
||
<emphasis id="B91BEA86F054FFC1FE81FD81FE23FD55" box="[340,453,620,645]" italics="true" pageId="54" pageNumber="55">Nigritulus</emphasis>
|
||
</taxonomicName>
|
||
, a Latin adjective referring to the uniform black coloration of this species.
|
||
</paragraph>
|
||
<paragraph id="8BD03694F054FFC1FF12FD62FB0DFD78" blockId="54.[151,1436,151,789]" box="[199,1259,655,680]" pageId="54" pageNumber="55">
|
||
<emphasis id="B91BEA86F054FFC1FF12FD62FE9BFD78" bold="true" box="[199,381,655,680]" pageId="54" pageNumber="55">Natural history</emphasis>
|
||
. Some specimens collected using flight interception traps set in gallery forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C375651FF054FFC1FF12FD59FB73FCC5" pageId="54" pageNumber="55" type="discussion">
|
||
<paragraph id="8BD03694F054FFC1FF12FD59FC57FD20" blockId="54.[151,1436,151,789]" pageId="54" pageNumber="55">
|
||
<emphasis id="B91BEA86F054FFC1FF12FD59FED2FD1D" bold="true" box="[199,308,692,717]" pageId="54" pageNumber="55">Remarks</emphasis>
|
||
. Females differs by the shape of the anterior pronotal carina is not tuberculate medially and by the lower and wider clypeofrontal carina and the finer pygidial punctation.
|
||
</paragraph>
|
||
<paragraph id="8BD03694F054FFC1FF12FD11FB73FCC5" blockId="54.[151,1436,151,789]" box="[199,1173,764,789]" pageId="54" pageNumber="55">Variation is limited to size and coloration which is reddish brown in teneral specimens.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |