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<document id="E6D54373815312F79CCD298C77DADFF3" ID-DOI="10.11646/zootaxa.4044.2.2" ID-GBIF-Dataset="8b09a6a1-35c3-407c-a4b5-076c4af030db" ID-ISSN="1175-5326" ID-Zenodo-Dep="241241" ID-ZooBank="7000893A-A8BC-4196-B76D-385C97B96FBD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461338858631" checkinUser="plazi" docAuthor="Kanao, Taisuke &amp; Maruyama, Munetoshi" docDate="2015" docId="0395F739FFABBF2AFF021CCBFB95BDB5" docLanguage="en" docName="zt04044p223.pdf" docOrigin="Zootaxa 4044 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Discoxenus minutus Kanao &amp; Maruyama, 2015, n. sp." docType="treatment" docVersion="11" lastPageNumber="216" masterDocId="FFAC8F41FFA5BF25FF951C5CFFC1BC65" masterDocTitle="Eight new species, a new record, and redescription of the genus Discoxenus Wasmann, 1904: The first record of termitophilous rove beetles in Cambodia (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="223" masterPageNumber="201" pageNumber="215" updateTime="1698619795159" updateUser="plazi">
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<mods:title id="EF7BE4397803776CDFEBBE624DB13E6F">Eight new species, a new record, and redescription of the genus Discoxenus Wasmann, 1904: The first record of termitophilous rove beetles in Cambodia (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:name id="0D08326FC6F775A59447631B9CEBE6D3" type="personal">
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<mods:namePart id="A459056E13332D3B39F152B03CE57588">Kanao, Taisuke</mods:namePart>
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<mods:name id="C9F843757923B996898BB591442816C7" type="personal">
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<mods:namePart id="778823D25B4174A1F93FC0B8B68C30EB">Maruyama, Munetoshi</mods:namePart>
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<mods:title id="AA197C9C13A03E8833A6EC690CD80DDC">Zootaxa</mods:title>
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<mods:date id="C9642050768A36FEC7EEB08C3AAED4F1">2015</mods:date>
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<mods:number id="F1CD144519A33BCAC53921D70B61667B">4044</mods:number>
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<mods:identifier id="0C25624320001F1AE3A423C6F262A351" type="DOI">10.11646/zootaxa.4044.2.2</mods:identifier>
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<treatment id="0395F739FFABBF2AFF021CCBFB95BDB5" ID-DOI="http://doi.org/10.5281/zenodo.6111787" ID-GBIF-Taxon="119644725" ID-Zenodo-Dep="6111787" LSID="urn:lsid:plazi:treatment:0395F739FFABBF2AFF021CCBFB95BDB5" httpUri="http://treatment.plazi.org/id/0395F739FFABBF2AFF021CCBFB95BDB5" lastPageId="15" lastPageNumber="216" pageId="14" pageNumber="215">
<subSubSection id="C32615A4FFABBF2BFF021CCBFE5CBCB6" pageId="14" pageNumber="215" type="nomenclature">
<paragraph id="8B83462FFFABBF2BFF021CCBFE16BCD7" blockId="14.[151,471,151,211]" box="[151,471,151,178]" pageId="14" pageNumber="215">
<heading id="D0CBF143FFABBF2BFF021CCBFE16BCD7" bold="true" box="[151,471,151,178]" fontSize="11" level="1" pageId="14" pageNumber="215" reason="1">
<emphasis id="B9489A3DFFABBF2BFF021CCBFE16BCD7" bold="true" box="[151,471,151,178]" pageId="14" pageNumber="215">
<taxonomicName id="4C3C3DACFFABBF2BFF021CCBFE4CBCD4" ID-CoL="6CZBW" box="[151,397,151,177]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="minutus" status="sp. nov.">
<emphasis id="B9489A3DFFABBF2BFF021CCBFE4CBCD4" bold="true" box="[151,397,151,177]" italics="true" pageId="14" pageNumber="215">Discoxenus minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B2746FFABBF2BFE001CC4FE16BCD7" box="[405,471,152,178]" pageId="14" pageNumber="215" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B83462FFFABBF2BFF021CE6FE5CBCB6" blockId="14.[151,471,151,211]" box="[151,413,186,211]" pageId="14" pageNumber="215">
(
<figureCitation id="13075AAAFFABBF2BFF0A1CE6FEE4BCB6" box="[159,293,186,211]" captionStart="FIGURES 5 12" captionStartId="3.[151,264,1680,1703]" captionTargetBox="[151,1436,483,1659]" captionTargetId="figure@3.[151,1436,483,1659]" captionTargetPageId="3" captionText="FIGURES 5 12. Habitus. 5 6. Discoxenus phourini n. sp.: 5. Dorsal view, 6. Ventral view. 7 8. D. kohkongensis n. sp.: 7. Dorsal view, 8. Ventral view. 9 10. D. hirsutus n. sp.: 9. Dorsal view, 10. Ventral view. 11 12. D. minutus n. sp.: 11. Dorsal view, 12. Ventral view." httpUri="https://zenodo.org/record/241243/files/figure.png" pageId="14" pageNumber="215">Figs. 11, 12</figureCitation>
,
<figureCitation id="13075AAAFFABBF2BFEA41CE6FE55BCB6" box="[305,404,186,211]" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">105120</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32615A4FFABBF2BFF021D5EFDF5BE3A" pageId="14" pageNumber="215" type="materials_examined">
<paragraph id="8B83462FFFABBF2BFF021D5EFDEEBD5A" blockId="14.[151,1437,258,2011]" pageId="14" pageNumber="215">
<emphasis id="B9489A3DFFABBF2BFF021D5EFE15BD7E" bold="true" box="[151,468,258,283]" pageId="14" pageNumber="215">
<typeStatus id="5487F88DFFABBF2BFF021D5EFF13BD7E" box="[151,210,258,283]" pageId="14" pageNumber="215">Type</typeStatus>
materials.
<typeStatus id="5487F88DFFABBF2BFEC81D5EFE0FBD7E" box="[349,462,258,283]" pageId="14" pageNumber="215" type="holotype">Holotype</typeStatus>
:
</emphasis>
♀,
<quantity id="4CC4EBCAFFABBF2BFD901D5EFD91BD7F" box="[517,592,258,283]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="215" unit="m" value="500.0">500 m</quantity>
S of Ankor Wat, Siem Reap,
<collectingCountry id="F32B06BFFFABBF2BFC231D5EFBEDBD7E" box="[950,1068,258,283]" name="Cambodia" pageId="14" pageNumber="215">Cambodia</collectingCountry>
,
<date id="FF8260EFFFABBF2BFBAF1D5EFB77BD7E" box="[1082,1206,258,283]" pageId="14" pageNumber="215" value="2012-06-05">5 VI 2012</date>
, Maruyama M. leg. (Colony no. MMCB-T-2012-1-002).
</paragraph>
<paragraph id="8B83462FFFABBF2BFF521D16FDF5BE3A" blockId="14.[151,1437,258,2011]" pageId="14" pageNumber="215">
<emphasis id="B9489A3DFFABBF2BFF521D16FE89BD06" bold="true" box="[199,328,330,355]" pageId="14" pageNumber="215">
<typeStatus id="5487F88DFFABBF2BFF521D16FE82BD06" box="[199,323,330,355]" pageId="14" pageNumber="215" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectingCountry id="F32B06BFFFABBF2BFEC61D16FE0ABD06" box="[339,459,330,355]" name="Cambodia" pageId="14" pageNumber="215">Cambodia</collectingCountry>
: 2♂♂, 3♀♀, same data as
<typeStatus id="5487F88DFFABBF2BFC9B1D16FCB1BD06" box="[782,880,330,355]" pageId="14" pageNumber="215" type="holotype">holotype</typeStatus>
(2♂♂, completely dissected). 2♀♀,
<quantity id="4CC4EBCAFFABBF2BFAB71D17FA9EBD06" box="[1314,1375,330,355]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="215" unit="km" value="1.0">1 km</quantity>
S of Angkor Wat, Siem Reap,
<date id="FF8260EFFFABBF2BFE541D33FD9ABDE2" box="[449,603,367,391]" pageId="14" pageNumber="215" value="2012-08-18">18 VIII 2012</date>
, Maruyama M. leg. (Colony no. MMCB-T-2012-2-003) (1♀, abdominal segments dissected off). 1♀, North Wall of Preah Khan, Siem Reap,
<date id="FF8260EFFFABBF2BFC091DCFFBF3BDCE" box="[924,1074,403,427]" pageId="14" pageNumber="215" value="2012-08-19">19 VIII 2012</date>
, Maruyama M. leg. (Colony no. MMCB-T-2012-2-005).
<specimenCount id="9D3A8DA6FFABBF2BFE3E1DEBFE0EBDAA" box="[427,463,439,463]" pageId="14" pageNumber="215" type="generic">1♂</specimenCount>
, 1♀, Siem Reap, Ankor Thom, Bayon,
<date id="FF8260EFFFABBF2BFC001DEBFBECBDAA" box="[917,1069,439,463]" pageId="14" pageNumber="215" value="2012-08-20">20 VIII 2012</date>
, Maruyama M. leg. (Colony no. MMCB-T-2012-2-009).
<specimenCount id="9D3A8DA6FFABBF2BFE3E1D87FE0EBD96" box="[427,463,475,499]" pageId="14" pageNumber="215" type="generic">1♂</specimenCount>
, 1♀,
<quantity id="4CC4EBCAFFABBF2BFD9A1D87FD8BBD96" box="[527,586,474,499]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="215" unit="km" value="1.0">1 km</quantity>
W of Banteay Prei, Siem Reap,
<date id="FF8260EFFFABBF2BFC551D87FB99BD96" box="[960,1112,475,499]" pageId="14" pageNumber="215" value="2012-08-21">21 VIII 2012</date>
, Maruyama M. leg. (Colony no. MMCB-T-2012-2-0017) (
<specimenCount id="9D3A8DA6FFABBF2BFE711DA3FDC8BE72" box="[484,521,511,535]" pageId="14" pageNumber="215" type="generic">1♂</specimenCount>
, completely dissected).
<specimenCount id="9D3A8DA6FFABBF2BFC821DA3FCFABE72" box="[791,827,511,535]" pageId="14" pageNumber="215" type="generic">1♂</specimenCount>
, 1♀,
<quantity id="4CC4EBCAFFABBF2BFCEC1DA2FC14BE72" box="[889,981,510,535]" metricMagnitude="2" metricUnit="m" metricValue="7.6" pageId="14" pageNumber="215" unit="km" value="0.76">0.76 km</quantity>
NE of Preah Kham, Siem Reap,
<date id="FF8260EFFFABBF2BFADC1DA3FF13BE5E" pageId="14" pageNumber="215" value="2014-08-18">18 VIII 2014</date>
, Kakizoe S. &amp; Maruyama M. leg. (Colony no. SK002). 2♂♂, same locality data to SK002,
<date id="FF8260EFFFABBF2BFB6B1E7FFA59BE5E" box="[1278,1432,547,571]" pageId="14" pageNumber="215" value="2014-08-19">19 VIII 2014</date>
, Kakizoe S. leg. (Colony no. SK007).
</paragraph>
</subSubSection>
<subSubSection id="C32615A4FFABBF2BFF521E36FF0EBF52" pageId="14" pageNumber="215" type="diagnosis">
<paragraph id="8B83462FFFABBF2BFF521E36FF0EBF52" blockId="14.[151,1437,258,2011]" pageId="14" pageNumber="215">
<emphasis id="B9489A3DFFABBF2BFF521E36FE80BEE6" bold="true" box="[199,321,618,643]" pageId="14" pageNumber="215">Diagnosis.</emphasis>
This species is distinguished from other
<taxonomicName id="4C3C3DACFFABBF2BFC9E1E37FC4DBEE7" box="[779,908,619,642]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="B9489A3DFFABBF2BFC9E1E37FC4DBEE7" box="[779,908,619,642]" italics="true" pageId="14" pageNumber="215">Discoxenus</emphasis>
</taxonomicName>
species by a combination of the following four character states: macrochaetotaxy of tergites IIIVIII (6, 6, 6, 6, 6, 6), sternites IIIVII with macrosetae only at each posterior margin, paramere with paramerite more than 3 times wider than condylite (
<figureCitation id="13075AAAFFABBF2BFB7A1EEEFA96BEAE" box="[1263,1367,690,715]" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">Fig. 119</figureCitation>
), and spermatheca with basal part more than 1.5 times longer than apical part (
<figureCitation id="13075AAAFFABBF2BFC5A1E8AFBF3BE8A" box="[975,1074,726,751]" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">Fig. 120</figureCitation>
). This species is most similar to
<taxonomicName id="4C3C3DACFFABBF2BFF021EA7FE93BF76" box="[151,338,763,787]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="kohkongensis">
<emphasis id="B9489A3DFFABBF2BFF021EA7FE93BF76" box="[151,338,763,787]" italics="true" pageId="14" pageNumber="215">D. kohkongensis</emphasis>
</taxonomicName>
, but is distinguishable from it by a male aedeagal median lobe with apically rounded crest (
<figureCitation id="13075AAAFFABBF2BFAFA1EA6FF02BF52" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">Fig. 118</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32615A4FFABBF2BFF521F1EFBB7BB2E" pageId="14" pageNumber="215" type="description">
<paragraph id="8B83462FFFABBF2BFF521F1EFF3FB8A7" blockId="14.[151,1437,258,2011]" pageId="14" pageNumber="215">
<emphasis id="B9489A3DFFABBF2BFF521F1EFE98BF3E" bold="true" box="[199,345,834,859]" pageId="14" pageNumber="215">Description.</emphasis>
Head (
<figureCitation id="13075AAAFFABBF2BFE271F1EFDD7BF3E" box="[434,534,834,859]" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">Fig. 105</figureCitation>
) approximately 1.3 times wider than long. Antenna (
<figureCitation id="13075AAAFFABBF2BFB191F1EFADABF3E" box="[1164,1307,834,859]" captionStart="FIGURES 5 12" captionStartId="3.[151,264,1680,1703]" captionTargetBox="[151,1436,483,1659]" captionTargetId="figure@3.[151,1436,483,1659]" captionTargetPageId="3" captionText="FIGURES 5 12. Habitus. 5 6. Discoxenus phourini n. sp.: 5. Dorsal view, 6. Ventral view. 7 8. D. kohkongensis n. sp.: 7. Dorsal view, 8. Ventral view. 9 10. D. hirsutus n. sp.: 9. Dorsal view, 10. Ventral view. 11 12. D. minutus n. sp.: 11. Dorsal view, 12. Ventral view." httpUri="https://zenodo.org/record/241243/files/figure.png" pageId="14" pageNumber="215">Figs. 11, 12</figureCitation>
,
<figureCitation id="13075AAAFFABBF2BFABE1F1EFA97BF3E" box="[1323,1366,834,859]" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">106</figureCitation>
) with segment I longer than other segments; segment II as long as segment III, with 3 macrosetae; segment III trapezoidal; segment IV transverse; segment V wider than other segments; segments VIXI successively narrowed distally; segments VIVII wider than long; segment VIII subquadrate; segments IXX longer than wide; segment XI approximately 2.5 times longer than wide, widest at middle. Labrum (
<figureCitation id="13075AAAFFABBF2BFC4C1F8EFBFABF8E" box="[985,1083,978,1003]" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">Fig. 107</figureCitation>
, left side) with anterior margin slightly concave at middle; median projection of apodeme short, with apex broadly rounded; 5 lateral setae present in ventral view (
<figureCitation id="13075AAAFFABBF2BFEC81846FE02B856" box="[349,451,1050,1075]" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">Fig. 107</figureCitation>
, right side). Mandibles (
<figureCitation id="13075AAAFFABBF2BFD721846FC52B856" box="[743,915,1050,1075]" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">Figs. 108, 109</figureCitation>
) covered with approximately 30 pores. Left mandible (
<figureCitation id="13075AAAFFABBF2BFE801862FEBBB832" box="[277,378,1086,1111]" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">Fig. 108</figureCitation>
) with adoral margin moderately pointed at apical third. Right mandible (
<figureCitation id="13075AAAFFABBF2BFB7A1862FA94B832" box="[1263,1365,1086,1111]" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">Fig. 109</figureCitation>
) with a distinct tooth. Maxillary palpal segment III approximately two times longer than wide. Mentum (
<figureCitation id="13075AAAFFABBF2BFAA5183EFA55B81E" box="[1328,1428,1122,1147]" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">Fig. 110</figureCitation>
) approximately 2.3 times wider than long, covered with around 40 pores. Labium with prementum covered with 12 14 pores.
</paragraph>
<paragraph id="8B83462FFFABBF2BFF521893FB4DB94A" blockId="14.[151,1437,258,2011]" pageId="14" pageNumber="215">
Pronotum (
<figureCitation id="13075AAAFFABBF2BFEDF1892FE6CB882" box="[330,429,1230,1255]" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">Fig. 111</figureCitation>
) sparsely covered with approximately 44 macrosetae, 6 minute setae present around anterior margin. Elytron (
<figureCitation id="13075AAAFFABBF2BFE2218AEFDD7B96E" box="[439,534,1266,1291]" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">Fig. 112</figureCitation>
) transverse, sparsely covered with several setae at anterolateral outer corner, 12 macrosetae present on disc. Metaventrite approximately 1.3 times as long as mesoventrite.
</paragraph>
<paragraph id="8B83462FFFABBF2BFF521966FDB1B986" blockId="14.[151,1437,258,2011]" pageId="14" pageNumber="215">
Tergites IIIIV (
<figureCitation id="13075AAAFFABBF2BFEE81966FE0AB936" box="[381,459,1338,1363]" captionStart="FIGURES 5 12" captionStartId="3.[151,264,1680,1703]" captionTargetBox="[151,1436,483,1659]" captionTargetId="figure@3.[151,1436,483,1659]" captionTargetPageId="3" captionText="FIGURES 5 12. Habitus. 5 6. Discoxenus phourini n. sp.: 5. Dorsal view, 6. Ventral view. 7 8. D. kohkongensis n. sp.: 7. Dorsal view, 8. Ventral view. 9 10. D. hirsutus n. sp.: 9. Dorsal view, 10. Ventral view. 11 12. D. minutus n. sp.: 11. Dorsal view, 12. Ventral view." httpUri="https://zenodo.org/record/241243/files/figure.png" pageId="14" pageNumber="215">Fig. 11</figureCitation>
) almost without seta. Tergites VVII with a row of setae anteapically. Tergite VIII (
<figureCitation id="13075AAAFFABBF2BFAFA1966FF00B912" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">Fig. 113</figureCitation>
) with posterior margin pointed, 3 pairs of macrosetae present at posterior margin, with 2 pairs of macrosetae around middle. Macrochaetotaxy of abdominal tergites IIIVIII = 6, 6, 6, 6, 6, 6. Sternite III (
<figureCitation id="13075AAAFFABBF2BFB6A19DEFA97B9FE" box="[1279,1366,1410,1435]" captionStart="FIGURES 5 12" captionStartId="3.[151,264,1680,1703]" captionTargetBox="[151,1436,483,1659]" captionTargetId="figure@3.[151,1436,483,1659]" captionTargetPageId="3" captionText="FIGURES 5 12. Habitus. 5 6. Discoxenus phourini n. sp.: 5. Dorsal view, 6. Ventral view. 7 8. D. kohkongensis n. sp.: 7. Dorsal view, 8. Ventral view. 9 10. D. hirsutus n. sp.: 9. Dorsal view, 10. Ventral view. 11 12. D. minutus n. sp.: 11. Dorsal view, 12. Ventral view." httpUri="https://zenodo.org/record/241243/files/figure.png" pageId="14" pageNumber="215">Fig. 12</figureCitation>
) with posterior half densely covered with setae. Sternites IVVIII with a row of yellow setae at middle. Sternites IIIVII with 610 macrosetae at posterior margin.
</paragraph>
<paragraph id="8B83462FFFABBF2BFF5219B2FC9BBAF3" blockId="14.[151,1437,258,2011]" pageId="14" pageNumber="215">
<emphasis id="B9489A3DFFABBF2BFF5219B2FEC9BA63" box="[199,264,1518,1542]" italics="true" pageId="14" pageNumber="215">Male.</emphasis>
Sternite VIII (
<figureCitation id="13075AAAFFABBF2BFE3B19B2FDCFBA62" box="[430,526,1518,1543]" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">Fig. 114</figureCitation>
) with 3 pairs of macrosetae at posterior margin, 2 pairs of macrosetae present at middle. Median lobe of aedeagus moderately narrowed toward apical lobe in ventral view (
<figureCitation id="13075AAAFFABBF2BFB061A4EFB35BA4E" box="[1171,1268,1554,1579]" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">Fig. 117</figureCitation>
); basal capsule with distal crest rounded apically in lateral view (
<figureCitation id="13075AAAFFABBF2BFD581A6AFCF1BA2A" box="[717,816,1590,1615]" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">Fig. 118</figureCitation>
); apical lobe less than half as wide as basal capsule in lateral view (
<figureCitation id="13075AAAFFABBF2BFEA51A06FE54BA16" box="[304,405,1626,1651]" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">Fig. 118</figureCitation>
). Paramere (
<figureCitation id="13075AAAFFABBF2BFDB21A06FD48BA16" box="[551,649,1626,1651]" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">Fig. 119</figureCitation>
) with paramerite more than 3 times wider than condylite; velar sac sclerite with 56 setae; apical lobe with 4 minute setae at apex.
</paragraph>
<paragraph id="8B83462FFFABBF2BFF521AFFFBAABABB" blockId="14.[151,1437,258,2011]" pageId="14" pageNumber="215">
<emphasis id="B9489A3DFFABBF2BFF521AFFFEDABADE" box="[199,283,1699,1723]" italics="true" pageId="14" pageNumber="215">Female</emphasis>
. Sternite VIII (
<figureCitation id="13075AAAFFABBF2BFE411AFEFDF6BADE" box="[468,567,1698,1723]" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">Fig. 115</figureCitation>
) with 2 pairs of macrosetae at posterior margin and middle, respectively. Spermatheca (
<figureCitation id="13075AAAFFABBF2BFEA21A9AFE57BABA" box="[311,406,1734,1759]" captionStart="FIGURES 105 120" captionStartId="15.[151,264,1659,1682]" captionTargetBox="[165,1422,527,1627]" captionTargetId="figure@15.[151,1436,519,1638]" captionTargetPageId="15" captionText="FIGURES 105 120. Discoxenus minutus n. sp.: 105. Head, 106. Antenna, 107. Labrum (left side = dorsal view, right side = ventral view), 108. Mandible, left, 109. Mandible, right, 110. Mentum, 111. Pronotum, 112. Elytron, right, 113. Tergite VIII, 114. Sternite VIII, male, 115. Sternite VIII, female, 116. Tergites IX and X, 117. Median lobe of aedeagus, ventral view, 118. Median lobe of aedeagus, lateral view, 119. Paramere, 120. Spermatheca." httpUri="https://zenodo.org/record/241250/files/figure.png" pageId="14" pageNumber="215">Fig. 120</figureCitation>
) with basal part approximately 1.8 times longer than apical part.
</paragraph>
<paragraph id="8B83462FFFABBF2BFF521AB6FBB7BB2E" blockId="14.[151,1437,258,2011]" pageId="14" pageNumber="215">
<emphasis id="B9489A3DFFABBF2BFF521AB6FEB3BB66" bold="true" box="[199,370,1770,1795]" pageId="14" pageNumber="215">Measurement.</emphasis>
Body length = average
<quantity id="4CC4EBCAFFABBF2BFD141AB6FD27BB67" box="[641,742,1770,1795]" metricMagnitude="-3" metricUnit="m" metricValue="1.62" pageId="14" pageNumber="215" unit="mm" value="1.62">1.62 mm</quantity>
(
<quantity id="4CC4EBCAFFABBF2BFD601AB6FC5DBB67" box="[757,924,1770,1795]" metricMagnitude="-3" metricUnit="m" metricValue="1.63" metricValueMax="1.73" metricValueMin="1.53" pageId="14" pageNumber="215" unit="mm" value="1.63" valueMax="1.73" valueMin="1.53">1.531.73 mm</quantity>
, N = 10), pronotal length = average
<quantity id="4CC4EBCAFFABBF2BFAAD1AB6FA5DBB67" box="[1336,1436,1770,1795]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="14" pageNumber="215" unit="mm" value="0.54">0.54 mm</quantity>
(
<quantity id="4CC4EBCAFFABBF2BFF0B1B52FE84BB43" box="[158,325,1806,1831]" metricMagnitude="-4" metricUnit="m" metricValue="5.35" metricValueMax="5.6" metricValueMin="5.1" pageId="14" pageNumber="215" unit="mm" value="0.535" valueMax="0.56" valueMin="0.51">0.510.56 mm</quantity>
, N = 10), pronotal width = average
<quantity id="4CC4EBCAFFABBF2BFD411B53FCF6BB43" box="[724,823,1807,1831]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="14" pageNumber="215" unit="mm" value="0.73">0.73 mm</quantity>
(
<quantity id="4CC4EBCAFFABBF2BFCD31B52FC2DBB43" box="[838,1004,1806,1831]" metricMagnitude="-4" metricUnit="m" metricValue="7.25" metricValueMax="7.7" metricValueMin="6.8" pageId="14" pageNumber="215" unit="mm" value="0.725" valueMax="0.77" valueMin="0.68">0.680.77 mm</quantity>
, N = 10), elytral length = average
<quantity id="4CC4EBCAFFABBF2BFAFE1B53FF02BB2F" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="14" pageNumber="215" unit="mm" value="0.38">0.38 mm</quantity>
(
<quantity id="4CC4EBCAFFABBF2BFF441B6EFEB9BB2F" box="[209,376,1842,1867]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="3.9" metricValueMin="3.5999999999999996" pageId="14" pageNumber="215" unit="mm" value="0.375" valueMax="0.39" valueMin="0.36">0.360.39 mm</quantity>
, N = 10), elytral width = average
<quantity id="4CC4EBCAFFABBF2BFD661B6FFC96BB2F" box="[755,855,1843,1867]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="14" pageNumber="215" unit="mm" value="0.41">0.41 mm</quantity>
(
<quantity id="4CC4EBCAFFABBF2BFCF01B6FFBCDBB2F" box="[869,1036,1843,1867]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" metricValueMax="4.3" metricValueMin="3.9" pageId="14" pageNumber="215" unit="mm" value="0.41" valueMax="0.43" valueMin="0.39">0.390.43 mm</quantity>
, N = 10).
</paragraph>
</subSubSection>
<subSubSection id="C32615A4FFABBF2BFF521B0AFAC7BBD2" pageId="14" pageNumber="215" type="etymology">
<paragraph id="8B83462FFFABBF2BFF521B0AFEB1BBF6" blockId="14.[151,1437,258,2011]" pageId="14" pageNumber="215">
<emphasis id="B9489A3DFFABBF2BFF521B0AFE8FBB0A" bold="true" box="[199,334,1878,1903]" pageId="14" pageNumber="215">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C3C3DACFFABBF2BFDA91B0BFD54BB0B" box="[572,661,1879,1902]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B9489A3DFFABBF2BFDA91B0BFD54BB0B" box="[572,661,1879,1902]" italics="true" pageId="14" pageNumber="215">minutus</emphasis>
</taxonomicName>
is a Latin adjective meaning “small,” in reference to the small body size of this species.
</paragraph>
<paragraph id="8B83462FFFABBF2BFF521BC2FAC7BBD2" blockId="14.[151,1437,258,2011]" box="[199,1286,1950,1975]" pageId="14" pageNumber="215">
<emphasis id="B9489A3DFFABBF2BFF521BC2FE9EBBD2" bold="true" box="[199,351,1950,1975]" pageId="14" pageNumber="215">Host species.</emphasis>
<taxonomicName id="4C3C3DACFFABBF2BFEF31BC3FCEABBD2" authority="Ahmad, 1965" authorityName="Ahmad" authorityYear="1965" box="[358,811,1950,1975]" class="Insecta" family="Termitidae" genus="Hypotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="makhamensis">
<emphasis id="B9489A3DFFABBF2BFEF31BC3FD4DBBD3" box="[358,652,1950,1974]" italics="true" pageId="14" pageNumber="215">Hypotermes makhamensis</emphasis>
<bibRefCitation id="EFAD3BDEFFABBF2BFD071BC2FCEABBD2" author="Ahmad" box="[658,811,1950,1975]" pageId="14" pageNumber="215" refString="Ahmad, M. (1965) Termites (Isoptera) of Thailand. Bulletin of the American Museum of Natural History, 131 (1), 1 - 114." type="journal article" year="1965">Ahmad, 1965</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C3C3DACFFABBF2BFCF41BC3FAC3BBD2" authority="Wasmann, 1896" authorityName="Wasmann" authorityYear="1896" box="[865,1282,1950,1975]" class="Insecta" family="Termitidae" genus="Hypotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="xenotermitis">
<emphasis id="B9489A3DFFABBF2BFCF41BC3FCBCBBD3" box="[865,893,1951,1974]" italics="true" pageId="14" pageNumber="215">H.</emphasis>
cf.
<emphasis id="B9489A3DFFABBF2BFC321BC3FBF3BBD3" box="[935,1074,1951,1974]" italics="true" pageId="14" pageNumber="215">xenotermitis</emphasis>
(
<bibRefCitation id="EFAD3BDEFFABBF2BFBD51BC3FB38BBD2" author="Wasmann" box="[1088,1273,1950,1975]" pageId="14" pageNumber="215" refString="Wasmann, E. (1896) Neue Termitophilen und Termiten aus Indien. Annali del Museo civico di storia naturale di Genova, 2, 613 - 630." type="journal article" year="1896">Wasmann, 1896</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C32615A4FFABBF2AFF521B9EFB95BDB5" lastPageId="15" lastPageNumber="216" pageId="14" pageNumber="215" type="discussion">
<paragraph id="8B83462FFFABBF2AFF521B9EFBA0BD78" blockId="14.[151,1437,258,2011]" lastBlockId="15.[151,1436,151,464]" lastPageId="15" lastPageNumber="216" pageId="14" pageNumber="215">
<emphasis id="B9489A3DFFABBF2BFF521B9EFE8EBBBE" bold="true" box="[199,335,1986,2011]" pageId="14" pageNumber="215">Comments.</emphasis>
Total of
<specimenCount id="9D3A8DA6FFABBF2BFE221B9FFD91BBBE" box="[439,592,1986,2011]" pageId="14" pageNumber="215" type="generic">13 specimens</specimenCount>
of
<taxonomicName id="4C3C3DACFFABBF2BFDE31B9FFD32BBBF" box="[630,755,1987,2010]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B9489A3DFFABBF2BFDE31B9FFD32BBBF" box="[630,755,1987,2010]" italics="true" pageId="14" pageNumber="215">D. minutus</emphasis>
</taxonomicName>
were collected and morphologically observed in the present study. Six beetle specimens were collected from only one colony (Colony no. MMCB-T-2012-1-002) of
<taxonomicName id="4C3C3DACFFAABF2AFACF1CC5FEE3BCB1" class="Insecta" family="Termitidae" genus="Hypotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="xenotermitis">
<emphasis id="B9489A3DFFAABF2AFACF1CC5FAAFBCD5" box="[1370,1390,153,176]" italics="true" pageId="15" pageNumber="216">H</emphasis>
. cf.
<emphasis id="B9489A3DFFAABF2AFF021CE1FEE3BCB1" box="[151,290,189,212]" italics="true" pageId="15" pageNumber="216">xenotermitis</emphasis>
</taxonomicName>
. The other seven beetle specimens were found in four different nests of
<taxonomicName id="4C3C3DACFFAABF2AFB311CE1FAA2BCB1" box="[1188,1379,188,212]" class="Insecta" family="Termitidae" genus="Hypotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="makhamensis">
<emphasis id="B9489A3DFFAABF2AFB311CE1FAA2BCB1" box="[1188,1379,188,212]" italics="true" pageId="15" pageNumber="216">H. makhamensis</emphasis>
</taxonomicName>
. All specimens of
<taxonomicName id="4C3C3DACFFAABF2AFEA11CBDFE71BC92" box="[308,432,224,248]" class="Insecta" family="Staphylinidae" genus="Discoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B9489A3DFFAABF2AFEA11CBDFE71BC92" box="[308,432,224,248]" italics="true" pageId="15" pageNumber="216">D. minutus</emphasis>
</taxonomicName>
were directly collected from the fungus gardens of their host termites, and it is unlikely that the beetles may have accidentally intruded in the nest of one of these host species.
</paragraph>
<paragraph id="8B83462FFFAABF2AFF521D7BFB95BDB5" blockId="15.[151,1436,151,464]" pageId="15" pageNumber="216">
There are small morphological differences between the specimens collected from the nests of
<taxonomicName id="4C3C3DACFFAABF2AFB761D75FA5DBD25" box="[1251,1436,296,320]" class="Insecta" family="Termitidae" genus="Hypotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="makhamensis">
<emphasis id="B9489A3DFFAABF2AFB761D75FA5DBD25" box="[1251,1436,296,320]" italics="true" pageId="15" pageNumber="216">H. makhamensis</emphasis>
</taxonomicName>
and
<emphasis id="B9489A3DFFAABF2AFF5D1D11FF22BD01" box="[200,227,333,356]" italics="true" pageId="15" pageNumber="216">H.</emphasis>
cf.
<emphasis id="B9489A3DFFAABF2AFE841D11FE70BD01" box="[273,433,333,356]" italics="true" pageId="15" pageNumber="216">xenotermitinis</emphasis>
. The specimens collected from
<taxonomicName id="4C3C3DACFFAABF2AFC8A1D11FC1BBD01" box="[799,986,332,356]" class="Insecta" family="Termitidae" genus="Hypotermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="makhamensis">
<emphasis id="B9489A3DFFAABF2AFC8A1D11FC1BBD01" box="[799,986,332,356]" italics="true" pageId="15" pageNumber="216">H. makhamensis</emphasis>
</taxonomicName>
nests have slightly narrower elytra and broader paramerites of paramere than those of the specimens collected from
<emphasis id="B9489A3DFFAABF2AFC6C1D2DFBD4BDED" box="[1017,1045,369,392]" italics="true" pageId="15" pageNumber="216">H.</emphasis>
cf.
<emphasis id="B9489A3DFFAABF2AFBD51D2CFB21BDE2" box="[1088,1248,368,391]" italics="true" pageId="15" pageNumber="216">xenotermitinis</emphasis>
nests. However, given the present data, we consider these differences to be insufficient for a species-level distinction. Examination of further material and DNA analysis will provide more data for species delimitation.
</paragraph>
</subSubSection>
</treatment>
</document>