330 lines
46 KiB
XML
330 lines
46 KiB
XML
<document id="8085EEC18538BC976BA356FD798BD6DC" ID-ISSN="0003-0082" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630274637015" checkinUser="carolina" docAuthor="Carvalho, Maria Da Gloria Pires De, Edgecombe, Gregory D. & Smith, Legrand" docDate="2003" docId="0395393C395C3767FCE1FC0ACF710821" docLanguage="en" docName="N3407.pdf" docOrigin="American Museum Novitates 3407" docStyle="DocumentStyle:70DDF1BDF6846024993F05262BAE3001.3:AmMusNovit.2000-2010.journal_article.type1" docStyleId="70DDF1BDF6846024993F05262BAE3001" docStyleName="AmMusNovit.2000-2010.journal_article.type1" docStyleVersion="3" docTitle="Wolfartaspis Cooper 1982" docType="treatment" docVersion="2" lastPageNumber="13" masterDocId="FFAC41443954376AFFC7FFD3CE3C096C" masterDocTitle="New Calmoniid Trilobites (Phacopina: Acastoidea) from the Devonian of Bolivia" masterLastPageNumber="18" masterPageNumber="1" pageNumber="9" updateTime="1699208581363" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="773474391E26B283BB1D670073FF8A12" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B3F3FB71EE21B8DBB2AB8B0C7C18E31A">
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<mods:title id="902975845578E6A73525D23FA56C4ABF">New Calmoniid Trilobites (Phacopina: Acastoidea) from the Devonian of Bolivia</mods:title>
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</mods:titleInfo>
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<mods:name id="9544484D642F3DBC3C419A7E39A72566" type="personal">
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<mods:role id="82D725B4D613105208149036A010CEAA">
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<mods:roleTerm id="8FE03E8138C25D13EA74D49B2B477F4D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DF0166C01AEEB004279FD1BC170DD238">Carvalho, Maria Da Gloria Pires De</mods:namePart>
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<mods:affiliation id="9A979648035FA52476B91C049357CF54">Gregory D. Edgecombe, & Research Associate, Division of Paleontology, American Museum of Natural History. e-mail: carvalho @ amnh. org</mods:affiliation>
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</mods:name>
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<mods:name id="12F64C07758B3FBAA5F760EB750C9D77" type="personal">
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<mods:role id="8634BF0278C7E78DC7A6CAF908133EB3">
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<mods:roleTerm id="EE50744EEE4F0393164A736489AB04B0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4F9EC55781CDD7D66F8671DE0520BCFA">Edgecombe, Gregory D.</mods:namePart>
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<mods:affiliation id="86BD1E442707828CDEA2DB5EA85FEF58">Principal Research Scientist, Australian Museum, 6 College Street, Sydney, NSW 2010, Australia.</mods:affiliation>
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</mods:name>
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<mods:name id="82904059E6D7B2AA30BD89092900EA99" type="personal">
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<mods:roleTerm id="F9E90EAD25155EB1A4FC6ED50380F349">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="57642E2AA54B51DDCC45C92820546B38">Smith, Legrand</mods:namePart>
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<mods:affiliation id="8A86ECB74273B80964564AF29A1D195B">266 Merrimon Avenue, Asheville, NC 28801 - 1218.</mods:affiliation>
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<mods:typeOfResource id="C5C010C72740837957351E5C21CE975C">text</mods:typeOfResource>
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<mods:titleInfo id="25A2C10D86794D468479714AA5B9C771">
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<mods:title id="40521A4DB4446604AD8B1470D9F59A06">American Museum Novitates</mods:title>
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</mods:titleInfo>
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<mods:part id="09F486D9F512BB148AB30D73C29B10DE">
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<mods:date id="8B36D113854ACEB0C0687B98194F507E">2003</mods:date>
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<mods:detail id="0C3189CDD414F327D863BF40B28E842E" type="pubDate">
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<mods:number id="4A9391799D5D5E364D49E827137258F5">2003-05-22</mods:number>
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</mods:detail>
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<mods:detail id="4BB87B6F716A23ED9D11D866984BFCBD" type="volume">
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<mods:number id="58A5DAAEA973E2E995D170C25A4D0588">3407</mods:number>
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<mods:start id="2109D3FBD38B9CFA18B00FF694CA3D7A">1</mods:start>
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<mods:end id="CD535990CF6686B9A5DC84DE70928507">18</mods:end>
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<mods:classification id="21ACCE0AD9821079637D886708D6FAC4">journal article</mods:classification>
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<mods:identifier id="C0EEAECDC6D46242D52132716366DE47" type="ISSN">0003-0082</mods:identifier>
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</mods:mods>
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<treatment id="0395393C395C3767FCE1FC0ACF710821" LSID="urn:lsid:plazi:treatment:0395393C395C3767FCE1FC0ACF710821" httpUri="http://treatment.plazi.org/id/0395393C395C3767FCE1FC0ACF710821" lastPageId="13" lastPageNumber="13" pageId="8" pageNumber="9">
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<subSubSection id="C326DBA1395C3762FCE1FC0ACA5E0A9C" box="[806,1122,985,1008]" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="8B83882A395C3762FCE1FC0ACA5E0A9C" blockId="8.[806,1122,985,1008]" box="[806,1122,985,1008]" pageId="8" pageNumber="9">
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<heading id="D0CB3F46395C3762FCE1FC0ACA5E0A9C" box="[806,1122,985,1008]" centered="true" fontSize="36" level="2" pageId="8" pageNumber="9" reason="8">
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<taxonomicName id="4C3CF3A9395C3762FCE1FC0ACA5E0A9C" authority="Cooper, 1982" authorityName="Cooper" authorityYear="1982" box="[806,1122,985,1008]" class="Trilobita" family="Calmoniidae" genus="Wolfartaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B9485438395C3762FCE1FC0ACD870A9C" box="[806,955,985,1008]" italics="true" pageId="8" pageNumber="9">Wolfartaspis</emphasis>
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<bibRefCitation id="EFADF5DB395C3762FC03FC0ACA5E0A9C" author="Cooper, M. R." box="[964,1122,985,1008]" pageId="8" pageNumber="9" pagination="1 - 174" refId="ref10009" refString="Cooper, M. R. 1982. A revision of the Devonian (Emsian-Eifelian) Trilobita from the Bokkeveld Group of South Africa. Annals of the South African Museum 89: 1 - 174." type="journal article" year="1982">Cooper, 1982</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C326DBA1395C3762FD1EFBD6CDB40C3C" pageId="8" pageNumber="9" type="materials_examined">
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<paragraph id="8B83882A395C3762FD1EFBD6CABE0D18" blockId="8.[700,1229,1029,1140]" pageId="8" pageNumber="9">
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<materialsCitation id="3B548277395C3762FD1EFBD6CA410D18" collectionCode="SPECIES" collectorName="La Paz Department" country="Bolivia" location="Upper Member" pageId="8" pageNumber="9" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54873688395C3762FD1EFBD6CD2A0D77" box="[729,790,1029,1052]" pageId="8" pageNumber="9">TYPE</typeStatus>
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<collectionCode id="ED2D10EF395C3762FCE3FBD6CDBC0D77" box="[804,896,1029,1052]" pageId="8" pageNumber="9">SPECIES</collectionCode>
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:
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<taxonomicName id="4C3CF3A9395C3762FC50FBD6CD5C0D55" authority="Wolfart, 1968" authorityName="Wolfart" authorityYear="1968" class="Trilobita" family="Calmoniidae" genus="Metacryphaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="cornutus">
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<emphasis id="B9485438395C3762FC50FBD6CAF00D70" box="[919,1228,1029,1052]" italics="true" pageId="8" pageNumber="9">Metacryphaeus cornutus</emphasis>
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<bibRefCitation id="EFADF5DB395C3762FD7BFBF1CD5C0D55" author="Wolfart, R." box="[700,864,1058,1081]" pageId="8" pageNumber="9" pagination="5 - 241" refId="ref10594" refString="Wolfart, R. 1968. Die Trilobiten aus dem Devon Boliviens und ihre Bedeutung fur Stratigraphie und Tiergeographie. In R. Wolfart and A. Voges (editors), Beitrage zur Kenntnis des devon von Bolivien. Beihefte zum Geologischen Jahrbuch 74: 5 - 241." type="journal article" year="1968">Wolfart, 1968</bibRefCitation>
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</taxonomicName>
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, from the lower part of the
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<location id="8EE3DEF1395C3762FB58FBF1CD6F0D3B" LSID="urn:lsid:plazi:treatment:0395393C395C3767FCE1FC0ACF710821:8EE3DEF1395C3762FB58FBF1CD6F0D3B" country="Bolivia" name="Upper Member" pageId="8" pageNumber="9">Upper Member</location>
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of the Belén Formation (late Emsian),
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<collectorName id="26C9EDFC395C3762FCE9FB8ECA2B0D18" box="[814,1047,1117,1140]" pageId="8" pageNumber="9">La Paz Department</collectorName>
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,
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<collectingCountry id="F32BC8BA395C3762FBE3FB8ECA410D18" box="[1060,1149,1117,1140]" name="Bolivia" pageId="8" pageNumber="9">Bolivia</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B83882A395C3762FD27FB44CAA30DC3" blockId="8.[736,1183,1175,1199]" box="[736,1183,1175,1199]" pageId="8" pageNumber="9">
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<emphasis id="B9485438395C3762FD27FB44CA3B0DC3" bold="true" box="[736,1031,1175,1199]" pageId="8" pageNumber="9">
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<taxonomicName id="4C3CF3A9395C3762FD27FB44CA3C0DC3" authority="Carvalho & Edgecombe & Smith, 2003" authorityName="Carvalho & Edgecombe & Smith" authorityYear="2003" box="[736,1024,1175,1199]" class="Trilobita" family="Calmoniidae" genus="Wolfartaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="liebermani" status="sp. nov.">
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<emphasis id="B9485438395C3762FD27FB44CA3C0DC3" bold="true" box="[736,1024,1175,1199]" italics="true" pageId="8" pageNumber="9">Wolfartaspis liebermani</emphasis>
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</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A27BE943395C3762FBD6FB4BCAA30DC3" box="[1041,1183,1176,1199]" pageId="8" pageNumber="9" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8B83882A395C3762FCA7FB6ACA140DA0" blockId="8.[864,1064,1205,1228]" box="[864,1064,1205,1228]" pageId="8" pageNumber="9">
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<heading id="D0CB3F46395C3762FCA7FB6ACA140DA0" box="[864,1064,1205,1228]" centered="true" fontSize="36" level="3" pageId="8" pageNumber="9" reason="7">
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<figureCitation id="130794AF395C3762FCA7FB6ACDDE0DA0" box="[864,994,1209,1228]" captionStart="Fig" captionStartId="10.[150,188,1483,1504]" captionTargetBox="[142,1212,186,1461]" captionTargetId="figure-14@10.[142,1212,186,1461]" captionTargetPageId="10" captionText="Fig. 4. Wolfartaspis liebermani, n. sp. Icla Formation, Kochis, Cochabamba Department. All scales 5 mm. A, holotype MHNC 8132, dorsal view of cephalon, latex cast from external mold; B, C, dorsal and lateral views of pygidium, AMNH 47150, latex cast from external mold; D, dorsal view of thorax and pygidium, AMNH 47148, internal mold; E, dorsal view of pygidium and posterior part of thorax, AMNH 47151, latex cast from external mold; F, ventral view of hypostome in situ beneath cephalon, AMNH 47401, latex cast from external mold." pageId="8" pageNumber="9">Figures 4A–F</figureCitation>
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,
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<figureCitation id="130794AF395C3762FC34FB6ACA140DA0" box="[1011,1064,1209,1228]" captionStart="Fig" captionStartId="11.[149,187,1296,1317]" captionTargetBox="[140,1209,186,1275]" captionTargetId="figure-13@11.[140,1209,186,1275]" captionTargetPageId="11" captionText="Fig. 5. Wolfartaspis liebermani, n. sp. Icla Formation, Kochis, Cochabamba Department. All scales 5 mm. A–C, dorsal, lateral, and anterior views of cephalon and part of first thoracic segment, AMNH 47403, internal mold; D–F, dorsal, anterior, and lateral views of cephalon, AMNH 47149, internal mold." pageId="8" pageNumber="9">5A–F</figureCitation>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8B83882A395C3762FD1EFB32CDB40C3C" blockId="8.[700,1229,1249,1477]" pageId="8" pageNumber="9">
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||
DERIVATION OF
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||
<collectionCode id="ED2D10EF395C3762FC50FB32CDE70D9B" box="[919,987,1249,1272]" pageId="8" pageNumber="9">NAME</collectionCode>
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||
: For Bruce S. Lieberman, who has made valuable contributions to systematics and biogeography of calmoniid trilobites.
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</paragraph>
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</subSubSection>
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<subSubSection id="C326DBA1395C3763FD1EFA85CED60AF4" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="diagnosis">
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<paragraph id="8B83882A395C3763FD1EFA85CF330807" blockId="8.[700,1229,1249,1477]" lastBlockId="9.[123,652,193,1741]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">DIAGNOSIS: Cephalic anterior border weakly pointed medially; ventral view of cephalon triangular in outline. Eyes less than onethird glabella length (Large Eye Index 31%); vi sual surface with 25–26 dorsoventral files with a maximum of 10 lenses per file. Occipital ring strongly arched in cross section, wide medially, with pronounced convex anterior margin and robust median spine tapering dorsally.</paragraph>
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<caption id="DF43D8A2395C3762FF51F9E0CA930FA1" pageId="8" pageNumber="9" startId="8.[150,188,1587,1608]">
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||
<paragraph id="8B83882A395C3762FF51F9E0CA930FA1" blockId="8.[124,1229,1586,1741]" pageId="8" pageNumber="9">
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||
Fig. 3.
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<taxonomicName id="4C3CF3A9395C3762FF3DF9E0CFC20F24" authority="Carvalho & Edgecombe & Smith, 2003" authorityName="Carvalho & Edgecombe & Smith" authorityYear="2003" box="[250,510,1587,1608]" class="Trilobita" family="Calmoniidae" genus="Eldredgeia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="eocryphaeus" status="sp. nov.">
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<emphasis id="B9485438395C3762FF3DF9E0CFC20F24" box="[250,510,1587,1608]" italics="true" pageId="8" pageNumber="9">Eldredgeia eocryphaeus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A27BE943395C3762FDD6F9E0CC6C0F24" box="[529,592,1587,1608]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
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||
<emphasis id="B9485438395C3762FD9BF9E1CCE60F2B" bold="true" box="[604,730,1586,1607]" pageId="8" pageNumber="9">A–C, H, I,</emphasis>
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Gamoneda Formation, Cerro Picacho, Tarija Department. All scales 5 mm. A–C, AMNH 47147, dorsal, lateral, and anterodorsal views of cephalon and articulated thorax, internal mold; H, I, AMNH 47146, dorsal and posterior views of pygidium, internal mold.
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<emphasis id="B9485438395C3762FED9F951CF650FFB" bold="true" box="[286,345,1666,1687]" pageId="8" pageNumber="9">D–G,</emphasis>
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holotype MHNC 8129, internal mold of cephalon, Belén Formation, Tikani, La Paz Department. All scales 5 mm. D, ventral view of hypostome; E–G, lateral, anterior, and dorsal views of cephalon;
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<emphasis id="B9485438395C3762FECDF96BCF000FA1" bold="true" box="[266,316,1720,1741]" pageId="8" pageNumber="9">J–L,</emphasis>
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MHNC 12900, dorsal, posterior, and lateral views of pygidium. All scales 5 mm.
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</paragraph>
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</caption>
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<paragraph id="8B83882A395D3763FF5FFEA2CED60AF4" blockId="9.[123,652,193,1741]" pageId="9" pageNumber="10">
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||
<materialsCitation id="3B548277395D3763FF5FFEA2CF050B39" collectionCode="TYPES" country="Bolivia" county="Departments" location="Icla Formation" municipality="Rio Grande" pageId="9" pageNumber="10" specimenCode="MHNC 8132" specimenCount="1" stateProvince="Chuquisaca" typeStatus="holotype">
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||
<collectionCode id="ED2D10EF395D3763FF5FFEA2CEE108EB" box="[152,221,369,392]" pageId="9" pageNumber="10">TYPES</collectionCode>
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||
:
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||
<typeStatus id="54873688395D3763FF33FEA2CF5908E4" box="[244,357,369,392]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
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||
:
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||
<specimenCode id="DB9A2051395D3763FEB0FEA2CC2608E4" box="[375,538,369,392]" collectionCode="MHNC" country="0" httpUri="http://grbio.org/cool/25c9-tsqt" name="Museo de Historia Natural de Concepcion (Chile)" pageId="9" pageNumber="10">MHNC 8132</specimenCode>
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||
, external mold of an incomplete cephalon,
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||
<location id="8EE3DEF1395D3763FDE6FE5DCEF308AF" LSID="urn:lsid:plazi:treatment:0395393C395C3767FCE1FC0ACF710821:8EE3DEF1395D3763FDE6FE5DCEF308AF" country="Bolivia" county="Departments" municipality="Rio Grande" name="Icla Formation" pageId="9" pageNumber="10" stateProvince="Chuquisaca">Icla Formation</location>
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||
, equivalent to basal part of
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<location id="8EE3DEF1395D3763FD85FE7FCEE1088C" LSID="urn:lsid:plazi:treatment:0395393C395C3767FCE1FC0ACF710821:8EE3DEF1395D3763FD85FE7FCEE1088C" country="Bolivia" county="Departments" municipality="Rio Grande" name="Upper Member" pageId="9" pageNumber="10" stateProvince="Chuquisaca">Upper Member</location>
|
||
at
|
||
<location id="8EE3DEF1395D3763FEC2FE1ACF0D088C" LSID="urn:lsid:plazi:treatment:0395393C395C3767FCE1FC0ACF710821:8EE3DEF1395D3763FEC2FE1ACF0D088C" box="[261,305,457,480]" country="Bolivia" county="Departments" municipality="Rio Grande" name="Icla" pageId="9" pageNumber="10" stateProvince="Chuquisaca">Icla</location>
|
||
type locality (see
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||
<location id="8EE3DEF1395D3763FDCBFE1ACCB0088C" LSID="urn:lsid:plazi:treatment:0395393C395C3767FCE1FC0ACF710821:8EE3DEF1395D3763FDCBFE1ACCB0088C" box="[524,652,457,480]" country="Bolivia" county="Departments" municipality="Rio Grande" name="Discussion" pageId="9" pageNumber="10" stateProvince="Chuquisaca">Discussion</location>
|
||
below),
|
||
<location id="8EE3DEF1395D3763FF26FE35CF470891" LSID="urn:lsid:plazi:treatment:0395393C395C3767FCE1FC0ACF710821:8EE3DEF1395D3763FF26FE35CF470891" box="[225,379,486,509]" country="Bolivia" county="Departments" municipality="Rio Grande" name="Kochis Hills" pageId="9" pageNumber="10" stateProvince="Chuquisaca">Kochis Hills</location>
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||
(approximately
|
||
<quantity id="4CC425CF395D3763FD8EFE35CCB70891" box="[585,651,486,509]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="10" unit="km" value="3.0">3 km</quantity>
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||
north of the
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<collectingMunicipality id="6BE71250395D3763FEDFFDD7CF9A0B77" box="[280,422,516,539]" pageId="9" pageNumber="10">Rio Grande</collectingMunicipality>
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||
, on the border between
|
||
<collectingRegion id="49F846C8395D3763FF0CFDF2CF6F0B54" box="[203,339,545,568]" country="Bolivia" name="Chuquisaca" pageId="9" pageNumber="10">Chuquisaca</collectingRegion>
|
||
and
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||
<collectingRegion id="49F846C8395D3763FE56FDF2CC150B54" box="[401,553,545,568]" country="Bolivia" name="Cochabamba" pageId="9" pageNumber="10">Cochabamba</collectingRegion>
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||
<collectingCounty id="62E2F0A6395D3763FDF5FDF2CEF90B39" pageId="9" pageNumber="10">Departments</collectingCounty>
|
||
),
|
||
<collectingCountry id="F32BC8BA395D3763FF27FDEDCF050B39" box="[224,313,574,597]" name="Bolivia" pageId="9" pageNumber="10">Bolivia</collectingCountry>
|
||
</materialsCitation>
|
||
.
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<materialsCitation id="3B548277395D3763FE8BFDEDCCBB0B1F" collectionCode="AMNH" country="Bolivia" pageId="9" pageNumber="10" specimenCode="AMNH 47148" specimenCount="1" stateProvince="Chuquisaca" typeStatus="paratype">
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<typeStatus id="54873688395D3763FE8BFDEDCFFE0B39" box="[332,450,574,597]" pageId="9" pageNumber="10" type="paratype">Paratypes</typeStatus>
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||
:
|
||
<specimenCode id="DB9A2051395D3763FE13FDEDCCBB0B39" box="[468,647,574,597]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="9" pageNumber="10">AMNH 47148</specimenCode>
|
||
, internal mold of thorax and pygidium
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B548277395D3763FFBCFDAACCBB0BFC" box="[123,647,633,656]" collectionCode="AMNH" country="Bolivia" pageId="9" pageNumber="10" specimenCode="AMNH 47149" specimenCount="1" stateProvince="Chuquisaca" typeStatus="paratype">
|
||
<specimenCode id="DB9A2051395D3763FFBCFDAACF160BFC" box="[123,298,633,656]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="9" pageNumber="10">AMNH 47149</specimenCode>
|
||
, internal mold of a cephalon
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B548277395D3763FFBCFD45CCBA0BC1" box="[123,646,662,685]" collectionCode="AMNH" country="Bolivia" pageId="9" pageNumber="10" specimenCode="AMNH 47150" specimenCount="1" stateProvince="Chuquisaca" typeStatus="paratype">
|
||
<specimenCode id="DB9A2051395D3763FFBCFD45CF100BC1" box="[123,300,662,685]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="9" pageNumber="10">AMNH 47150</specimenCode>
|
||
, external mold of pygidium
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B548277395D3763FFBCFD67CF080B84" collectionCode="AMNH" country="Bolivia" pageId="9" pageNumber="10" specimenCode="AMNH 47151" specimenCount="1" stateProvince="Chuquisaca" typeStatus="paratype">
|
||
<specimenCode id="DB9A2051395D3763FFBCFD67CF100BA7" box="[123,300,692,715]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="9" pageNumber="10">AMNH 47151</specimenCode>
|
||
, external mold of an incomplete pygidium
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B548277395D3763FE80FD02CF9D0A69" collectionCode="AMNH" country="Bolivia" pageId="9" pageNumber="10" specimenCode="AMNH 47401" specimenCount="1" stateProvince="Chuquisaca" typeStatus="paratype">
|
||
<specimenCode id="DB9A2051395D3763FE80FD02CFC50B84" box="[327,505,721,744]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="9" pageNumber="10">AMNH 47401</specimenCode>
|
||
, hypostome in situ beneath cephalon
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B548277395D3763FE68FD3DCCBB0A4F" collectionCode="AMNH" country="Bolivia" pageId="9" pageNumber="10" specimenCode="AMNH 47403" specimenCount="1" stateProvince="Chuquisaca" typeStatus="paratype">
|
||
<specimenCode id="DB9A2051395D3763FE68FD3DCC610A69" box="[431,605,750,773]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="9" pageNumber="10">AMNH 47403</specimenCode>
|
||
, internal mold of almost complete cephalon
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B548277395D3763FFBCFCFACEDA0AF4" collectionCode="MHNC" collectorName="All" country="Bolivia" location="Icla Formation" municipality="Kochis" pageId="9" pageNumber="10" specimenCode="MHNC 12749" specimenCount="1" stateProvince="Cochabamba" typeStatus="paratype">
|
||
<specimenCode id="DB9A2051395D3763FFBCFCFACF150A2C" box="[123,297,809,832]" collectionCode="MHNC" country="0" httpUri="http://grbio.org/cool/25c9-tsqt" name="Museo de Historia Natural de Concepcion (Chile)" pageId="9" pageNumber="10">MHNC 12749</specimenCode>
|
||
, internal mold of incomplete pygidium.
|
||
<collectorName id="26C9EDFC395D3763FF39FC95CF1F0A31" box="[254,291,838,861]" pageId="9" pageNumber="10">All</collectorName>
|
||
type specimens are from the
|
||
<location id="8EE3DEF1395D3763FFBCFCB7CF080A17" LSID="urn:lsid:plazi:treatment:0395393C395C3767FCE1FC0ACF710821:8EE3DEF1395D3763FFBCFCB7CF080A17" box="[123,308,868,891]" country="Bolivia" municipality="Kochis" name="Icla Formation" pageId="9" pageNumber="10" stateProvince="Cochabamba">Icla Formation</location>
|
||
,
|
||
<collectingMunicipality id="6BE71250395D3763FE8CFCB7CF9C0A17" box="[331,416,868,891]" pageId="9" pageNumber="10">Kochis</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49F846C8395D3763FE71FCB7CEDA0AF4" country="Bolivia" name="Cochabamba" pageId="9" pageNumber="10">Cochabamba Department</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326DBA1395D3766FF5FFC4DCCDC0A9C" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10" type="description">
|
||
<paragraph id="8B83882A395D3763FF5FFC4DCEEE0C5F" blockId="9.[123,652,193,1741]" pageId="9" pageNumber="10">
|
||
OTHER MATERIAL: Some additional topotype specimens collected within the same stratigraphic interval as the
|
||
<typeStatus id="54873688395D3763FE07FC0ACFC10A9C" box="[448,509,985,1008]" pageId="9" pageNumber="10">types</typeStatus>
|
||
are referred to
|
||
<taxonomicName id="4C3CF3A9395D3763FF5BFC24CF850D62" authorityName="Carvalho & Edgecombe & Smith" authorityYear="2003" box="[156,441,1015,1038]" class="Trilobita" family="Calmoniidae" genus="Wolfartaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="liebermani">
|
||
<emphasis id="B9485438395D3763FF5BFC24CF850D62" box="[156,441,1015,1038]" italics="true" pageId="9" pageNumber="10">Wolfartaspis liebermani</emphasis>
|
||
</taxonomicName>
|
||
, but they are not well preserved. Most are from broken concretions and include three internal molds and two external molds of pygidia, an internal mold of four or five thoracic segments, an external mold of some pleurae, and two internal molds of cephala. In addition, however, there is a single, wellpreserved, small pygidium in part and counterpart. This additional material is housed in the MHNC collection.
|
||
</paragraph>
|
||
<paragraph id="8B83882A395D3761FF5FFAEACCDA0F19" blockId="9.[123,652,193,1741]" lastBlockId="11.[699,1228,1425,1741]" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">
|
||
DESCRIPTION: The cranidial length (sag.) is about
|
||
<quantity id="4CC425CF395D3763FF01FA85CF0C0C01" box="[198,304,1366,1389]" metricMagnitude="-2" metricUnit="m" metricValue="2.9699999999999998" pageId="9" pageNumber="10" unit="mm" value="29.7">29.7 mm</quantity>
|
||
in AMNH 47403. The anterior cephalic border is narrowest medially, where the upturned doublure nearly contacts the cranidial margin; the anterior cephalic border projects only slightly in front to the glabella medially in dorsal view; the anterior border gently widens abaxially, but remains narrow (exsag.) in dorsal view, and is steep along its entire width. The cephalic anterolateral margin is nearly straight in dorsal view and is rounded medially. The axial furrow is deep, wide, straight, slightly divergent forward. Glabellar length (sag., excluding S0) in AMNH 47403 is
|
||
<quantity id="4CC425CF395D3763FC2EFF12CA6409B4" box="[1001,1112,193,216]" metricMagnitude="-2" metricUnit="m" metricValue="2.34" pageId="9" pageNumber="10" unit="mm" value="23.4">23.4 mm</quantity>
|
||
, with the frontal lobe having marked independent convexity from the posterior glabellar region, which has a flat sagittal profile. The frontal glabellar lobe is inflated, about 60% of glabellar length, and gradually expands anteriorly and laterally, with its anteromedian margin rounded. The anterior section of the facial suture circumscribes but does not transect the frontal glabellar lobe. The frontal lobe has a well developed circular to slightly elongated posterior median impression (PMI) and auxiliary impression system (AIS) of muscle scars. The latter is most clearly preserved on the left side and the median part of AMNH 47403 (fig. 5A) as small rounded pits on the internal mold, forming divergent rows. Medially, the AIS does not seem to have a defined pattern. The lateral glabellar furrows are well developed. S3 is wide, deep, oblique, straight, with its proximal part weakly bent posteromedially, lengthening abaxially, and distinctly confluent with the axial furrow. S2 is transverse, narrower and shallower than S1 and S3; it becomes abruptly shallow distally and has only faint impression against the axial furrow on the external cuticular surface (fig. 4A). S1 is deeper than S2 and S3; it sharply narrows distally but is distinctly confluent with the axial furrow on the external mold; the posterior margin of S1 is concave. L3 is wedgeshaped; L2 is approximately trapezoidal; L1 is slightly shorter (exsag.) than L2 and L3, with its posterior margin markedly convex backward and then continuing inward, forming a short (sag.) transglabellar lobe that is weakly curved anteriorly; this feature makes the glabella slightly higher at L1 than at L3. S0 is long and relatively shallow (sag.), becoming shorter and deeper as it approaches the axial furrow. Where the cephalic posterior furrow meets S0, both have approximately the same width; the posterior furrow becomes wider (exsag.) toward the fulcrum, and is then gently flexed forward and shallows distally. L0 is strongly arched in its transverse section, wide medially, with a pronounced convex anterior margin, and bearing a robust median spine that tapers dorsally; the median spine is weakly declined posteriorly. The eye is high, less than onethird glabellar length (Large Eye Index 31%). Its anterior edge is opposite the anterolateral corner of L3, and its posterior edge is opposite the posterolateral margin of L2 and at a considerable distance from the posterior cephalic border furrow (approximately
|
||
<quantity id="4CC425CF395F3761FEB6F9D5CF840F71" box="[369,440,1542,1565]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="12" unit="mm" value="4.0">4 mm</quantity>
|
||
in the
|
||
<typeStatus id="54873688395F3761FDD3F9D5CC430F71" box="[532,639,1542,1565]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
). In anterior view, the visual surface is straight from top to bottom, gradually becoming wid er toward its base, with 25–26 dorsoventral lens files, and a maximum of 10 lenses per file. The interocular fixigena slopes upward as it approaches the narrow, Cshaped pal pebral lobe; the palpebral furrow is moderately deep. The lateral cephalic border furrow is faint. The posterior cephalic border is narrow proximally, about equal in width (exsag.) to the posterior border furrow at the fulcrum, and widened distally; none of the available specimens preserves the genal angle.
|
||
</paragraph>
|
||
<caption id="DF43D8A2395E3760FF51FA18CC550F09" pageId="10" pageNumber="11" startId="10.[150,188,1483,1504]" targetBox="[142,1212,186,1461]" targetPageId="10">
|
||
<paragraph id="8B83882A395E3760FF51FA18CC550F09" blockId="10.[124,1229,1483,1637]" pageId="10" pageNumber="11">
|
||
Fig. 4.
|
||
<taxonomicName id="4C3CF3A9395E3760FF31FA18CFC90C8C" authority="Carvalho & Edgecombe & Smith, 2003" authorityName="Carvalho & Edgecombe & Smith" authorityYear="2003" box="[246,501,1483,1504]" class="Trilobita" family="Calmoniidae" genus="Wolfartaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="liebermani" status="sp. nov.">
|
||
<emphasis id="B9485438395E3760FF31FA18CFC90C8C" box="[246,501,1483,1504]" italics="true" pageId="10" pageNumber="11">Wolfartaspis liebermani</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A27BE943395E3760FDC3FA18CC020C8C" box="[516,574,1483,1504]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
|
||
Icla Formation, Kochis, Cochabamba Department. All scales 5 mm.
|
||
<emphasis id="B9485438395E3760FF0CFA36CED80C96" bold="true" box="[203,228,1509,1530]" pageId="10" pageNumber="11">A,</emphasis>
|
||
holotype MHNC 8132, dorsal view of cephalon, latex cast from external mold;
|
||
<emphasis id="B9485438395E3760FB8DFA36CAB80C96" bold="true" box="[1098,1156,1509,1530]" pageId="10" pageNumber="11">B, C,</emphasis>
|
||
dorsal and lateral views of pygidium, AMNH 47150, latex cast from external mold;
|
||
<emphasis id="B9485438395E3760FC03F9D3CDE10F79" bold="true" box="[964,989,1536,1557]" pageId="10" pageNumber="11">D,</emphasis>
|
||
dorsal view of thorax and pygidium, AMNH 47148, internal mold;
|
||
<emphasis id="B9485438395E3760FDAEF9C9CCBD0F43" bold="true" box="[617,641,1562,1583]" pageId="10" pageNumber="11">E,</emphasis>
|
||
dorsal view of pygidium and posterior part of thorax, AMNH 47151, latex cast from external mold;
|
||
<emphasis id="B9485438395E3760FDB1F9E6CCB10F26" bold="true" box="[630,653,1589,1610]" pageId="10" pageNumber="11">F,</emphasis>
|
||
ventral view of hypostome in situ beneath cephalon, AMNH 47401, latex cast from external mold.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF43D8A2395F3761FF52FAC3CAF00C36" pageId="11" pageNumber="12" startId="11.[149,187,1296,1317]" targetBox="[140,1209,186,1275]" targetPageId="11">
|
||
<paragraph id="8B83882A395F3761FF52FAC3CAF00C36" blockId="11.[123,1228,1296,1370]" pageId="11" pageNumber="12">
|
||
Fig. 5.
|
||
<taxonomicName id="4C3CF3A9395F3761FF33FAC3CFCF0C49" authority="Carvalho & Edgecombe & Smith, 2003" authorityName="Carvalho & Edgecombe & Smith" authorityYear="2003" box="[244,499,1296,1317]" class="Trilobita" family="Calmoniidae" genus="Wolfartaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="liebermani" status="sp. nov.">
|
||
<emphasis id="B9485438395F3761FF33FAC3CFCF0C49" box="[244,499,1296,1317]" italics="true" pageId="11" pageNumber="12">Wolfartaspis liebermani</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A27BE943395F3761FDC4FAC3CC010C49" box="[515,573,1296,1317]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
|
||
Icla Formation, Kochis, Cochabamba Department. All scales 5 mm.
|
||
<emphasis id="B9485438395F3761FF0DFAF9CF3F0C53" bold="true" box="[202,259,1322,1343]" pageId="11" pageNumber="12">A–C,</emphasis>
|
||
dorsal, lateral, and anterior views of cephalon and part of first thoracic segment, AMNH 47403, internal mold;
|
||
<emphasis id="B9485438395F3761FEA1FA96CFA00C36" bold="true" box="[358,412,1349,1370]" pageId="11" pageNumber="12">D–F,</emphasis>
|
||
dorsal, anterior, and lateral views of cephalon, AMNH 47149, internal mold.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B83882A395F3766FD1FF9AFCC490A31" blockId="11.[699,1228,1425,1741]" lastBlockId="12.[124,653,193,1741]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||
The hypostome is preserved in situ beneath the cephalon in
|
||
<specimenCount id="9D3A43A3395F3761FC7AF94ACA610FDC" box="[957,1117,1689,1712]" pageId="11" pageNumber="12" type="generic">one specimen</specimenCount>
|
||
(fig. 4F). Its anterior margin is largely missing, but its course against the moderately wide anterior wings suggests a transverse hypostomal suture. The middle body is roughly circular and gently convex (tr., sag.). The maculae are strong, forming a pair of ovoid protuberances positioned abaxially, near to the junction of the faint middle furrow and the lateral border furrow, anterior to the midlength of the middle body. The border furrow is obscure anterolaterally (against the wings), becoming moderately impressed at the macula, maintaining moderate depth and width posterolaterally and posteromedially. The lateral bor der is narrow, with the lateral margins gently converging between the anterior wing and first of two pairs of short, sharp posterolateral marginal spines; the inner pair of spines (on posterior border) is separated by slightly greater than the distance between the two spine pairs. The posterior border is fairly long and gently convex (sag.); the posteromedian margin is faintly convex backward between the inner pair of marginal spines.
|
||
</paragraph>
|
||
<paragraph id="8B83882A39583766FF5EFCB7CC350CA9" blockId="12.[124,653,193,1741]" pageId="12" pageNumber="13">The thorax consists of 11 segments. The relative width of the axial rings and thorax cannot be determined, as the distal ends of the pleurae are not preserved in any specimens, but in AMNH 47148 (fig. 4D) the axial rings appear to be wider than the pleural fields. The axial furrow is shallow and narrow, weakly diverging back to the sixth ring, then gently converging. In lateral view, the axial rings are faintly convex, raised well above the pleurae. The anteriormost axial rings are spatulate distally, but this is less marked in the more posterior rings. Small but distinct apodemal pits lie between the axial rings abaxially. The pleural furrows are narrow and moderately impressed adjacent to the axial furrow, becoming wider distally, with a straight, oblique course across the rib; the anterior margin of the pleural furrows is gently convex anteriorly, while the posterior margin of the furrows is straight.</paragraph>
|
||
<paragraph id="8B83882A39583766FF5EFA1FCCDC0A9C" blockId="12.[124,653,193,1741]" lastBlockId="12.[700,1229,193,1742]" pageId="12" pageNumber="13">The pygidium is triangular, moderately arched in lateral view, wider than long, with a pointed posterior tip. The axial furrow is narrow and moderately deep. The axis is slightly waisted, the axial furrow moderately convergent against the first five axial rings, weakly converging posterior to this; the axial terminus is blunt, lacking impression of the axial furrow, lying well in advance of the pygidial posterior margin. There are 10–11 axial rings, but the last few are indistinct; the first five rings are gently arched anteriorly, but the subsequent ones are transverse or weakly bowed backward. The axial rings are separated by wide ring furrows that become progressively narrower posteriorly; the first five ring furrows have narrow, groovelike apodemal pits distally. The pleural field includes six pairs of strongly furrowed ribs plus a small seventh pair that parallels the axis. The pleural furrows are deep and wide, though gradually narrowing on more posterior segments; they are moderately curved. The interpleural furrows are narrow, moderately deep proximal to the fulcrum between the anterior few ribs, progressively shallow between more posterior ribs such that fifth interpleural furrow is last distinct one on external mold. The lateral margin of the pygidium is not well preserved in any of the material (fig.4E); the fifth lappet is small, flat, and wide (fig.4B), with a curved anterior margin and straight posterior margin. The posteromedian margin of the pygidium is an elongate, tapering, upturned spine (its basal part is preserved in AMNH 47150; fig. 4B, C).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326DBA139583767FD1EFC25CF710821" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="discussion">
|
||
<paragraph id="8B83882A39583767FD1EFC25CF710821" blockId="12.[700,1229,193,1742]" lastBlockId="13.[123,651,193,334]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
|
||
DISCUSSION: The monotypic
|
||
<taxonomicName id="4C3CF3A939583766FBFFFC24CAF10D62" authorityName="Cooper" authorityYear="1982" box="[1080,1229,1015,1038]" class="Trilobita" family="Calmoniidae" genus="Wolfartaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948543839583766FBFFFC24CAF10D62" box="[1080,1229,1015,1038]" italics="true" pageId="12" pageNumber="13">Wolfartaspis</emphasis>
|
||
</taxonomicName>
|
||
was erected by
|
||
<bibRefCitation id="EFADF5DB39583766FCB2FBC7CA1D0D47" author="Cooper, M. R." box="[885,1057,1044,1067]" pageId="12" pageNumber="13" pagination="1 - 174" refId="ref10009" refString="Cooper, M. R. 1982. A revision of the Devonian (Emsian-Eifelian) Trilobita from the Bokkeveld Group of South Africa. Annals of the South African Museum 89: 1 - 174." type="journal article" year="1982">Cooper (1982)</bibRefCitation>
|
||
as a subgenus of
|
||
<taxonomicName id="4C3CF3A939583766FD18FBE2CDAE0D24" authorityName="Reed" authorityYear="1907" box="[735,914,1073,1096]" class="Trilobita" family="Calmoniidae" genus="Metacryphaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948543839583766FD18FBE2CDAE0D24" box="[735,914,1073,1096]" italics="true" pageId="12" pageNumber="13">Metacryphaeus</emphasis>
|
||
</taxonomicName>
|
||
, but recent usage (
|
||
<bibRefCitation id="EFADF5DB39583766FBB0FBE2CD060D09" author="Lieberman, B. S." pageId="12" pageNumber="13" pagination="549 - 570" refId="ref10359" refString="Lieberman, B. S. 1993. Systematics and biogeography of the '' Metacryphaeus group' ' Calmoniidae (Trilobita, Devonian), with comments on adaptive radiations and the geological history of the Malvinokaffric Realm. Journal of Paleontology 67: 549 - 570." type="journal article" year="1993">Lieberman, 1993</bibRefCitation>
|
||
) recognizes it at the generic rank. According to Lieberman,
|
||
<taxonomicName id="4C3CF3A939583766FBC7FBBFCAA90DEF" authorityName="Cooper" authorityYear="1982" box="[1024,1173,1132,1155]" class="Trilobita" family="Calmoniidae" genus="Wolfartaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948543839583766FBC7FBBFCAA90DEF" box="[1024,1173,1132,1155]" italics="true" pageId="12" pageNumber="13">Wolfartaspis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C3CF3A939583766FCFBFB5ACDD30DCC" authorityName="Reed" authorityYear="1907" box="[828,1007,1161,1184]" class="Trilobita" family="Calmoniidae" genus="Metacryphaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948543839583766FCFBFB5ACDD30DCC" box="[828,1007,1161,1184]" italics="true" pageId="12" pageNumber="13">Metacryphaeus</emphasis>
|
||
</taxonomicName>
|
||
in having an enlarged occipital spine and an upwardly curved pygidial spine. The material described here includes both of these features, and it is therefore assigned to the genus
|
||
<taxonomicName id="4C3CF3A939583766FB53FB2CCD1E0C5F" authorityName="Cooper" authorityYear="1982" class="Trilobita" family="Calmoniidae" genus="Wolfartaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948543839583766FB53FB2CCD1E0C5F" italics="true" pageId="12" pageNumber="13">Wolfartaspis</emphasis>
|
||
</taxonomicName>
|
||
. We place our material in a new species, based on the following differences from
|
||
<taxonomicName id="4C3CF3A939583766FD3AFA84CDB70C02" authorityName="Wolfart" authorityYear="1968" box="[765,907,1367,1390]" class="Trilobita" family="Calmoniidae" genus="Metacryphaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cornutus">
|
||
<emphasis id="B948543839583766FD3AFA84CDB70C02" box="[765,907,1367,1390]" italics="true" pageId="12" pageNumber="13">W. cornutus</emphasis>
|
||
</taxonomicName>
|
||
: (1) the ventral view of the cephalon has a rounded shape in
|
||
<taxonomicName id="4C3CF3A939583766FB93FAA7CCE20CC4" authorityName="Wolfart" authorityYear="1968" class="Trilobita" family="Calmoniidae" genus="Metacryphaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cornutus">
|
||
<emphasis id="B948543839583766FB93FAA7CCE20CC4" italics="true" pageId="12" pageNumber="13">W. cornutus</emphasis>
|
||
</taxonomicName>
|
||
, but is more pointed in our form; (2) the Large Eye Index in
|
||
<taxonomicName id="4C3CF3A939583766FC6AFA7CCA070CAA" authorityName="Wolfart" authorityYear="1968" box="[941,1083,1455,1478]" class="Trilobita" family="Calmoniidae" genus="Metacryphaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cornutus">
|
||
<emphasis id="B948543839583766FC6AFA7CCA070CAA" box="[941,1083,1455,1478]" italics="true" pageId="12" pageNumber="13">W. cornutus</emphasis>
|
||
</taxonomicName>
|
||
is 34%, but is less in our material (31%); (3) the occipital spine in
|
||
<taxonomicName id="4C3CF3A939583766FCE1FA3ACD8A0F6C" authorityName="Wolfart" authorityYear="1968" box="[806,950,1513,1536]" class="Trilobita" family="Calmoniidae" genus="Metacryphaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cornutus">
|
||
<emphasis id="B948543839583766FCE1FA3ACD8A0F6C" box="[806,950,1513,1536]" italics="true" pageId="12" pageNumber="13">W. cornutus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFADF5DB39583766FC0FFA3ACAA40F6C" author="Lieberman, B. S." box="[968,1176,1513,1536]" pageId="12" pageNumber="13" pagination="549 - 570" refId="ref10359" refString="Lieberman, B. S. 1993. Systematics and biogeography of the '' Metacryphaeus group' ' Calmoniidae (Trilobita, Devonian), with comments on adaptive radiations and the geological history of the Malvinokaffric Realm. Journal of Paleontology 67: 549 - 570." type="journal article" year="1993">Lieberman, 1993</bibRefCitation>
|
||
: fig. 2.8) is stouter than in our material (fig. 5E, F) and does not taper from its base. Differences are also noted in the stratigraphic levels of the two forms.
|
||
<taxonomicName id="4C3CF3A939583766FC0FF98CCAF10F1A" baseAuthorityName="Wolfart" baseAuthorityYear="1968" box="[968,1229,1631,1654]" class="Trilobita" family="Calmoniidae" genus="Wolfartaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cornutus">
|
||
<emphasis id="B948543839583766FC0FF98CCAF10F1A" box="[968,1229,1631,1654]" italics="true" pageId="12" pageNumber="13">Wolfartaspis cornutus</emphasis>
|
||
</taxonomicName>
|
||
occurs in the basal part of the Upper Member of the Belén Formation, in strata overlying the layer of
|
||
<taxonomicName id="4C3CF3A939583766FC8CF964CA7E0FA2" authorityName="sensu Wolfart" authorityYear="1968" box="[843,1090,1719,1742]" class="Trilobita" family="Dalmanitidae" genus="Francovichia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="branisi">
|
||
<emphasis id="B948543839583766FC8CF964CA7E0FA2" box="[843,1090,1719,1742]" italics="true" pageId="12" pageNumber="13">Francovichia branisi</emphasis>
|
||
</taxonomicName>
|
||
(= layer of
|
||
<taxonomicName id="4C3CF3A939593767FFBCFF12CC5909B4" authority="sensu Wolfart, 1968" authorityName="Wolfart" authorityYear="1968" box="[123,613,193,216]" class="Trilobita" family="Dalmanitidae" genus="Odontochile" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="branisi">
|
||
<emphasis id="B948543839593767FFBCFF12CF5B09B4" box="[123,359,193,216]" italics="true" pageId="13" pageNumber="14">Odontochile branisi</emphasis>
|
||
sensu
|
||
<bibRefCitation id="EFADF5DB39593767FE79FF12CC5909B4" author="Wolfart, R." box="[446,613,193,216]" pageId="13" pageNumber="14" pagination="5 - 241" refId="ref10594" refString="Wolfart, R. 1968. Die Trilobiten aus dem Devon Boliviens und ihre Bedeutung fur Stratigraphie und Tiergeographie. In R. Wolfart and A. Voges (editors), Beitrage zur Kenntnis des devon von Bolivien. Beihefte zum Geologischen Jahrbuch 74: 5 - 241." type="journal article" year="1968">Wolfart, 1968</bibRefCitation>
|
||
</taxonomicName>
|
||
, in the upper part of Lower Member of Belén Formation). At Kochis,
|
||
<taxonomicName id="4C3CF3A939593767FE5EFF2FCC0A087F" box="[409,566,252,275]" class="Trilobita" family="Dalmanitidae" genus="Francovichia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B948543839593767FE5EFF2FCC0A087F" box="[409,566,252,275]" italics="true" pageId="13" pageNumber="14">Francovichia</emphasis>
|
||
</taxonomicName>
|
||
occurs stratigraphically above the beds with
|
||
<taxonomicName id="4C3CF3A939593767FDF6FECACF7A0822" authorityName="Carvalho & Edgecombe & Smith" authorityYear="2003" class="Trilobita" family="Calmoniidae" genus="Wolfartaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="liebermani">
|
||
<emphasis id="B948543839593767FDF6FECACF7A0822" italics="true" pageId="13" pageNumber="14">Wolfartaspis liebermani</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |