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<document id="317B26A6F2EDB5ECC0534B7988850EBF" ID-DOI="10.11646/zootaxa.4819.1.11" ID-ISSN="1175-5326" ID-Zenodo-Dep="3955808" ID-ZooBank="B2D8A697-9473-47C6-BEF8-A510444210E9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1595405865904" checkinUser="plazi" docAuthor="Kamal, N. S. S., Tan, H. H. &amp; Ng, Casey K. C." docDate="2020" docId="0386E26AFFF0FFF9DAC448DDFD091F5A" docLanguage="en" docName="zootaxa.4819.1.11.pdf" docOrigin="Zootaxa 4819 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Betta nuluhon Kamal &amp; Tan &amp; Ng 2020, new species" docType="treatment" docUuid="76DF3D63-C008-47A5-949C-A8FFF3554D9C" docUuidSource="ZooBank" docVersion="3" lastPageNumber="192" masterDocId="FFBF9A12FFF1FFFCDA534D5DFFAC1D41" masterDocTitle="Betta nuluhon, a new species of fighting fish from western Sabah, Malaysia (Teleostei: Osphronemidae)" masterLastPageNumber="194" masterPageNumber="187" pageNumber="188" updateTime="1698784451639" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="16D8BB83C0FCD9FD73BCEDFB1BF819FD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5C5BEA14C9BF77BB257BCB9B152F989E">
<mods:title id="57E38E101A97CB4BD89C13A77399931B">Betta nuluhon, a new species of fighting fish from western Sabah, Malaysia (Teleostei: Osphronemidae)</mods:title>
</mods:titleInfo>
<mods:name id="6A4BF79AB0852C0F82F6487A8A7456EC" type="personal">
<mods:role id="4102C0E9E1F5E88E08A4B6DB038CEEA6">
<mods:roleTerm id="995083C0A8E219BE267F1F3592D33DE4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B16591C8E80D840EDF7192CD36139B7B">Kamal, N. S. S.</mods:namePart>
<mods:affiliation id="3C836325B12946D2FD81DB6E3B41D2B8">Forest Research Centre, Sabah Forestry Department, P. O. Box 1407, 90715 Sandakan, Sabah, Malaysia. NurSyafiqah. Shkamal @ sabah. gov. my; https: // orcid. org / 0000 - 0003 - 1236 - 3848</mods:affiliation>
</mods:name>
<mods:name id="40E1C6C9A52DCCF39CE560424F420AB2" type="personal">
<mods:role id="BF270067C32D2C1184895EBE862730E3">
<mods:roleTerm id="648DE80E5D6FE15D44C57AE3A52F2AA0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D20ECFC27797A5F1B2491E35F6103F1A">Tan, H. H.</mods:namePart>
<mods:affiliation id="34316BA78422E6685862D926925EE1FA">National University of Singapore, Lee Kong Chian Natural History Museum, 2 Conservatory Drive, Singapore 117377. heokhui @ nus. edu. sg; https: // orcid. org / 0000 - 0003 - 2227 - 8866</mods:affiliation>
</mods:name>
<mods:name id="00563984529943ECEB2EFEE2650D44DD" type="personal">
<mods:role id="AD90ECC4031A6716975DFFAF1761DC1C">
<mods:roleTerm id="28763433FF2C573B4A8F9A55BDD051BA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="927AC71C4B067B19B47DD93C60A83774">Ng, Casey K. C.</mods:namePart>
<mods:affiliation id="96A10784FDB161F0CE6BF4316B772725">Forever Sabah, H 30, Gaya Park, Lorong Muntahan 1 C, Jalan Penampang, 88300 Kota Kinabalu, Sabah, Malaysia. casey @ foreversabah. org; https: // orcid. org / 0000 - 0003 - 1758 - 3268</mods:affiliation>
</mods:name>
<mods:typeOfResource id="3EBEAFAB51AE767F2B9A9C10A58AE717">text</mods:typeOfResource>
<mods:relatedItem id="BD7EF1118B0EC77732BBE9B5950A95F9" type="host">
<mods:titleInfo id="12E923A4E326234BA59625E9157CD871">
<mods:title id="CA30A3D3C53EDF80895FE45279B380A3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="50DBC6B1EAFBC5E597777771C8822E1C">
<mods:date id="90BADB615254CC43DB93984682A6F2E9">2020</mods:date>
<mods:detail id="872F09739A516D7B09833838E7D020E1" type="pubDate">
<mods:number id="8DAB7BF20EF1B52D6206D833DFB90059">2020-07-22</mods:number>
</mods:detail>
<mods:detail id="E878B1448FEE513EB3826C3683A157D3" type="volume">
<mods:number id="FA2BF824893246F52077C6BFC3BA5A59">4819</mods:number>
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<mods:number id="2F51F5864D0B9A0FF1F2EF3A956ECEE0">1</mods:number>
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<mods:start id="D5D768E84720E08452990E46AA4F20CF">187</mods:start>
<mods:end id="EA973E34AB1E58B4AA529EB99E207984">194</mods:end>
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<mods:classification id="9F16A36DC3087C2E7405937D5FF3EAEE">journal article</mods:classification>
<mods:identifier id="C896D9378A665ABD5413181C5E7E9624" type="DOI">10.11646/zootaxa.4819.1.11</mods:identifier>
<mods:identifier id="8464095FCA0A5E60693A2527A92F7E90" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1944A182EFB2C904DA778AB1246FAF04" type="Zenodo-Dep">3955808</mods:identifier>
<mods:identifier id="C9BB6BCDE74BD1E1772DAEEA63EDEED6" type="ZooBank">B2D8A697-9473-47C6-BEF8-A510444210E9</mods:identifier>
</mods:mods>
<treatment id="0386E26AFFF0FFF9DAC448DDFD091F5A" LSID="urn:lsid:zoobank.org:act:76DF3D63-C008-47A5-949C-A8FFF3554D9C" httpUri="http://treatment.plazi.org/id/0386E26AFFF0FFF9DAC448DDFD091F5A" lastPageId="5" lastPageNumber="192" pageId="1" pageNumber="188">
<subSubSection id="C33500F7FFF0FFFDDAC448DDFE4918D8" box="[151,485,1407,1434]" pageId="1" pageNumber="188" type="nomenclature">
<paragraph id="8B90537CFFF0FFFDDAC448DDFE4918D8" blockId="1.[151,485,1407,1470]" box="[151,485,1407,1434]" pageId="1" pageNumber="188">
<heading id="D0D8E410FFF0FFFDDAC448DDFE4918D8" bold="true" box="[151,485,1407,1434]" fontSize="11" level="1" pageId="1" pageNumber="188" reason="1">
<emphasis id="B95B8F6EFFF0FFFDDAC448DDFE4918D8" bold="true" box="[151,485,1407,1434]" pageId="1" pageNumber="188">
<emphasis id="B95B8F6EFFF0FFFDDAC448DDFEE118DB" bold="true" box="[151,333,1407,1434]" italics="true" pageId="1" pageNumber="188">
<taxonomicName id="4C2F28FFFFF0FFFDDAC448DDFEEB18DB" authority="Kamal &amp; Tan &amp; Ng, 2020" authorityName="Kamal &amp; Tan &amp; Ng" authorityYear="2020" box="[151,327,1407,1434]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="188" phylum="Chordata" rank="species" species="nuluhon" status="sp. nov.">Betta nuluhon</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A2683215FFF0FFFDDB0648DDFE4918D8" box="[341,485,1407,1434]" pageId="1" pageNumber="188" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33500F7FFF0FFFDDAC448F9FEAC18FF" box="[151,256,1444,1470]" pageId="1" pageNumber="188" type="description">
<paragraph id="8B90537CFFF0FFFDDAC448F9FEAC18FF" blockId="1.[151,485,1407,1470]" box="[151,256,1444,1470]" pageId="1" pageNumber="188">
<figureCitation id="13144FF9FFF0FFFDDAC448F9FEAC18FF" box="[151,256,1444,1470]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,787,812]" captionStartId-1="3.[151,250,1708,1733]" captionStartId-2="4.[151,250,1952,1977]" captionStartId-3="5.[151,250,1353,1378]" captionTargetBox-0="[149,1438,184,761]" captionTargetBox-1="[150,1437,834,1684]" captionTargetBox-2="[151,1436,1160,1927]" captionTargetBox-3="[151,1436,561,1325]" captionTargetId-0="figure@3.[149,1438,181,761]" captionTargetId-1="figure@3.[150,1437,834,1684]" captionTargetId-2="figure@4.[151,1436,1101,1927]" captionTargetId-3="figure@5.[151,1436,561,1328]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Betta nuluhon, new species, ca. 50 mm SL, freshly caught male specimen, not preserved." captionText-1="FIGURE 2. Betta nuluhon, new species, male (top), female (bottom); freshly caught, not preserved." captionText-2="FIGURE 3. Betta nuluhon, new species, ZRC 61247, holotype, 62.6 mm SL (above with white background, bottom with black background)." captionText-3="FIGURE 4. Top—Betta chini, ZRC 61155, 58.6 mm SL male; bottom—Betta nuluhon, holotype, ZRC 61247, 62.6 mm SL male; both freshly preserved." figureDoi-0="http://doi.org/10.5281/zenodo.3955810" figureDoi-1="http://doi.org/10.5281/zenodo.3955812" figureDoi-2="http://doi.org/10.5281/zenodo.3955814" figureDoi-3="http://doi.org/10.5281/zenodo.3955816" httpUri-0="https://zenodo.org/record/3955810/files/figure.png" httpUri-1="https://zenodo.org/record/3955812/files/figure.png" httpUri-2="https://zenodo.org/record/3955814/files/figure.png" httpUri-3="https://zenodo.org/record/3955816/files/figure.png" pageId="1" pageNumber="188">Figs. 1–4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33500F7FFF0FFFDDAC448B8FF7B1BD1" pageId="1" pageNumber="188" type="materials_examined">
<paragraph id="8B90537CFFF0FFFDDAC448B8FE231B65" blockId="1.[151,1437,1509,2041]" pageId="1" pageNumber="188">
<emphasis id="B95B8F6EFFF0FFFDDAC448B8FE2C1B41" bold="true" box="[151,384,1509,1536]" pageId="1" pageNumber="188">Material examined.</emphasis>
Holotype—ZRC 61247,
<quantity id="4CD7FE99FFF0FFFDD8CF48BBFD531B40" box="[668,767,1510,1537]" metricMagnitude="-2" metricUnit="m" metricValue="6.26" pageId="1" pageNumber="188" unit="mm" value="62.6">62.6 mm</quantity>
SL, male;
<collectingRegion id="49EB9D9EFFF0FFFDD92A48BBFC6E1B41" box="[889,962,1510,1536]" country="Malaysia" name="Sabah" pageId="1" pageNumber="188">Sabah</collectingRegion>
: Crocker Range Forest Reserve; Kamal N. S. S.
<emphasis id="B95B8F6EFFF0FFFDDA814B56FEA81B65" box="[210,260,1546,1572]" italics="true" pageId="1" pageNumber="188">et al</emphasis>
.,
<date id="FF9175BCFFF0FFFDDB4B4B56FE201B65" box="[280,396,1546,1573]" pageId="1" pageNumber="188" value="2019-11">Nov 2019</date>
.
</paragraph>
<paragraph id="8B90537CFFF0FFFDDA944B72FF7B1BD1" blockId="1.[151,1437,1509,2041]" pageId="1" pageNumber="188">
<materialsCitation id="3B475921FFF0FFFDDA944B72FE531B08" box="[199,511,1581,1609]" collectionCode="FRC" pageId="1" pageNumber="188" specimenCode="FRC 0162" specimenCount="5" typeStatus="paratype">
<typeStatus id="5494EDDEFFF0FFFDDA944B72FE941B08" box="[199,312,1583,1609]" pageId="1" pageNumber="188" type="paratype">Paratypes</typeStatus>
.
<specimenCode id="DB89FB07FFF0FFFDDB104B73FE161B09" box="[323,442,1582,1608]" collectionCode="FRC" country="USA" httpUri="http://grbio.org/cool/z9c1-g2b0" name="Fusarium Research Center" pageId="1" pageNumber="188">FRC 0162</specimenCode>
,
<specimenCount id="9D2998F5FFF0FFFDDB954B70FE531B08" box="[454,511,1581,1609]" pageId="1" pageNumber="188" type="generic">5 ex.</specimenCount>
</materialsCitation>
,
<materialsCitation id="3B475921FFF0FFFDD85F4B73FD7B1B08" box="[524,727,1581,1609]" collectionCode="ZRC" pageId="1" pageNumber="188" specimenCount="5" typeStatus="paratype">
<collectionCode id="ED3ECBB9FFF0FFFDD85F4B73FDEE1B09" box="[524,578,1582,1608]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="188">ZRC</collectionCode>
61154,
<specimenCount id="9D2998F5FFF0FFFDD8CE4B70FD7B1B08" box="[669,727,1581,1609]" pageId="1" pageNumber="188" type="generic">5 ex.</specimenCount>
</materialsCitation>
,
<materialsCitation id="3B475921FFF0FFFDD8B04B70FABA1B09" box="[739,1302,1581,1609]" collectionCode="ZRC" pageId="1" pageNumber="188" specimenCount="1" typeStatus="holotype">
39.8–54.0 mm SL; same locality data as
<typeStatus id="5494EDDEFFF0FFFDDEE34B73FABA1B09" box="[1200,1302,1582,1608]" pageId="1" pageNumber="188" type="holotype">holotype</typeStatus>
</materialsCitation>
.
<materialsCitation id="3B475921FFF0FFFDDF724B73FF7E1B2C" collectionCode="FRC" pageId="1" pageNumber="188" specimenCode="FRC 0161" specimenCount="3" typeStatus="paratype">
<specimenCode id="DB89FB07FFF0FFFDDF724B73FA341B09" box="[1313,1432,1582,1608]" collectionCode="FRC" country="USA" httpUri="http://grbio.org/cool/z9c1-g2b0" name="Fusarium Research Center" pageId="1" pageNumber="188">FRC 0161</specimenCode>
,
<specimenCount id="9D2998F5FFF0FFFDDAC44B0FFF7E1B2C" box="[151,210,1618,1645]" pageId="1" pageNumber="188" type="generic">3 ex.</specimenCount>
</materialsCitation>
,
<materialsCitation id="3B475921FFF0FFFDDAB34B0FFE1C1B2C" box="[224,432,1617,1645]" collectionCode="ZRC" pageId="1" pageNumber="188" specimenCount="3" typeStatus="paratype">
<collectionCode id="ED3ECBB9FFF0FFFDDAB34B0FFEBA1B2D" box="[224,278,1618,1644]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="188">ZRC</collectionCode>
57026,
<specimenCount id="9D2998F5FFF0FFFDDB254B0FFE1C1B2C" box="[374,432,1618,1645]" pageId="1" pageNumber="188" type="generic">3 ex.</specimenCount>
</materialsCitation>
,
<materialsCitation id="3B475921FFF0FFFDDBED4B0CFF7F1BD1" collectingDate="2018-04" collectorName="Kamal N. S. S." location="Crocker Range Forest Reserve" pageId="1" pageNumber="188" specimenCount="1" stateProvince="Sabah" typeStatus="paratype">
46.0–
<quantity id="4CD7FE99FFF0FFFDDBAD4B0CFDCF1B2C" box="[510,611,1617,1645]" metricMagnitude="-2" metricUnit="m" metricValue="5.74" pageId="1" pageNumber="188" unit="mm" value="57.4">57.4 mm</quantity>
SL;
<collectingRegion id="49EB9D9EFFF0FFFDD8CE4B0FFD4A1B2D" box="[669,742,1618,1644]" country="Malaysia" name="Sabah" pageId="1" pageNumber="188">Sabah</collectingRegion>
:
<location id="8EF005A7FFF0FFFDD8A04B0FFBF81B2C" LSID="urn:lsid:plazi:treatment:0386E26AFFF0FFF9DAC448DDFD091F5A:8EF005A7FFF0FFFDD8A04B0FFBF81B2C" box="[755,1108,1618,1645]" name="Crocker Range Forest Reserve" pageId="1" pageNumber="188" stateProvince="Sabah">Crocker Range Forest Reserve</location>
;
<collectorName id="26DA36AAFFF0FFFDDE324B0FFAA31B2D" box="[1121,1295,1618,1645]" pageId="1" pageNumber="188">Kamal N. S. S.</collectorName>
<emphasis id="B95B8F6EFFF0FFFDDF4B4B0EFAE01B2D" box="[1304,1356,1618,1644]" italics="true" pageId="1" pageNumber="188">et al</emphasis>
.,
<date id="FF9175BCFFF0FFFDDF324B0FFF7F1BD1" pageId="1" pageNumber="188" value="2018-04">
<collectingDate id="EFD58C54FFF0FFFDDF324B0FFF7F1BD1" pageId="1" pageNumber="188" value="2018-04">April 2018</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33500F7FFF0FFFDDA944BC4FE831A05" pageId="1" pageNumber="188" type="diagnosis">
<paragraph id="8B90537CFFF0FFFDDA944BC4FE831A05" blockId="1.[151,1437,1509,2041]" pageId="1" pageNumber="188">
<emphasis id="B95B8F6EFFF0FFFDDA944BC4FE961BF5" bold="true" box="[199,314,1689,1716]" pageId="1" pageNumber="188">Diagnosis</emphasis>
.
<taxonomicName id="4C2F28FFFFF0FFFDDB144BC6FE481BF5" authorityName="Kamal &amp; Tan &amp; Ng" authorityYear="2020" box="[327,484,1690,1716]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="188" phylum="Chordata" rank="species" species="nuluhon">
<emphasis id="B95B8F6EFFF0FFFDDB144BC6FE481BF5" box="[327,484,1690,1716]" italics="true" pageId="1" pageNumber="188">Betta nuluhon</emphasis>
</taxonomicName>
is distinct from its congeners in the
<taxonomicName id="4C2F28FFFFF0FFFDD9214BC6FC561BF5" authorityName="Regan" authorityYear="1910" box="[882,1018,1690,1716]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="188" phylum="Chordata" rank="species" species="akarensis">
<emphasis id="B95B8F6EFFF0FFFDD9214BC6FC561BF5" box="[882,1018,1690,1716]" italics="true" pageId="1" pageNumber="188">B. akarensis</emphasis>
</taxonomicName>
group in having the following unique combination of characters: yellow iris when live; mature males with greenish-blue iridescence on opercle when live; mature fish with distinct transverse bars on caudal fin; slender body (body depth 22.1–25.2 % SL); belly area with faint reticulate pattern (posterior margin of scales rimmed with black); anal-fin plain; lateral scales 29–31; predorsal scales 20–21.
</paragraph>
</subSubSection>
<subSubSection id="C33500F7FFF0FFF8DA944A10FE871CCD" lastPageId="4" lastPageNumber="191" pageId="1" pageNumber="188" type="description">
<paragraph id="8B90537CFFF0FFFDDA944A10FF721ACC" blockId="1.[151,1437,1509,2041]" pageId="1" pageNumber="188">
<emphasis id="B95B8F6EFFF0FFFDDA944A10FEFD1A29" bold="true" box="[199,337,1869,1896]" pageId="1" pageNumber="188">Description</emphasis>
. See
<figureCitation id="13144FF9FFF0FFFDDBC24A13FE571A29" box="[401,507,1870,1896]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,787,812]" captionStartId-1="3.[151,250,1708,1733]" captionStartId-2="4.[151,250,1952,1977]" captionStartId-3="5.[151,250,1353,1378]" captionTargetBox-0="[149,1438,184,761]" captionTargetBox-1="[150,1437,834,1684]" captionTargetBox-2="[151,1436,1160,1927]" captionTargetBox-3="[151,1436,561,1325]" captionTargetId-0="figure@3.[149,1438,181,761]" captionTargetId-1="figure@3.[150,1437,834,1684]" captionTargetId-2="figure@4.[151,1436,1101,1927]" captionTargetId-3="figure@5.[151,1436,561,1328]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Betta nuluhon, new species, ca. 50 mm SL, freshly caught male specimen, not preserved." captionText-1="FIGURE 2. Betta nuluhon, new species, male (top), female (bottom); freshly caught, not preserved." captionText-2="FIGURE 3. Betta nuluhon, new species, ZRC 61247, holotype, 62.6 mm SL (above with white background, bottom with black background)." captionText-3="FIGURE 4. Top—Betta chini, ZRC 61155, 58.6 mm SL male; bottom—Betta nuluhon, holotype, ZRC 61247, 62.6 mm SL male; both freshly preserved." figureDoi-0="http://doi.org/10.5281/zenodo.3955810" figureDoi-1="http://doi.org/10.5281/zenodo.3955812" figureDoi-2="http://doi.org/10.5281/zenodo.3955814" figureDoi-3="http://doi.org/10.5281/zenodo.3955816" httpUri-0="https://zenodo.org/record/3955810/files/figure.png" httpUri-1="https://zenodo.org/record/3955812/files/figure.png" httpUri-2="https://zenodo.org/record/3955814/files/figure.png" httpUri-3="https://zenodo.org/record/3955816/files/figure.png" pageId="1" pageNumber="188">Figs. 1–4</figureCitation>
for general appearance. See
<tableCitation id="C6AD66C7FFF0FFFDD9104A13FC3A1A29" box="[835,918,1870,1896]" captionStart="TABLE 1" captionStartId="2.[151,239,442,466]" captionText="TABLE 1. Meristic and morphometric data of Betta nuluhon, new species" pageId="1" pageNumber="188">Table 1</tableCitation>
for meristic and morphometric data of
<taxonomicName id="4C2F28FFFFF0FFFDDF044A12FF7B1ACD" authorityName="Kamal &amp; Tan &amp; Ng" authorityYear="2020" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="188" phylum="Chordata" rank="species" species="nuluhon">
<emphasis id="B95B8F6EFFF0FFFDDF044A12FF7B1ACD" italics="true" pageId="1" pageNumber="188">B. nuluhon</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B90537CFFF0FFFEDA944ACBFE211CCD" blockId="1.[151,1437,1509,2041]" lastBlockId="2.[151,1436,152,396]" lastPageId="2" lastPageNumber="189" pageId="1" pageNumber="188">
Head broad, blunt, short (head length 31.7–35.2 % SL); body cylindrical in cross-section anteriorly, becoming compressed at dorsal-fin origin to caudal region; body deepest at dorsal-fin origin (22.1–25.2 % SL), shallowest in caudal peduncle (16.4–20.0 % SL); dorsal-fin rays I, 7 (total 8), pointed, situated at about ⅔ head-and-body length (predorsal length 65.8–71.3 % SL), dorsal-fin base short (10.6–13.5 % SL); caudal-fin lanceolate, with 5+6 principal rays, middle rays elongated slightly beyond interradial membrane margin; anal-fin with I–II, 25–27 (total 26–28) rays, pointed, situated around ½ body (preanal length 45.4–49.7 % SL), anal-fin base long (49.5–54.6 % SL); pelvic-fin with i, 5 rays, rounded, with filamentous second ray, adpressed filamentous ray reaching 13
<superScript id="7C5AFE34FFF3FFFEDF034C59FAF11C53" attach="left" box="[1360,1373,260,274]" fontSize="6" pageId="2" pageNumber="189">th</superScript>
analfin ray (pelvic-fin length 31.5–45.1 % SL); pectoral-fin rays 13–14, rounded; lateral scales 29–31, predorsal scales 20–21, subdorsal scales 5½–6½, postdorsal scales 10–11, transverse scales 9½. Vertebral count 11–12 + 19–20, total vertebrae = 31 (n =6).
</paragraph>
<caption id="DF5003F4FFF3FFFEDAC44CE7FC031C92" box="[151,943,442,467]" pageId="2" pageNumber="189">
<paragraph id="8B90537CFFF3FFFEDAC44CE7FC031C92" blockId="2.[151,943,442,467]" box="[151,943,442,467]" pageId="2" pageNumber="189">
<emphasis id="B95B8F6EFFF3FFFEDAC44CE7FEAF1C93" bold="true" box="[151,259,442,466]" pageId="2" pageNumber="189">TABLE 1</emphasis>
. Meristic and morphometric data of
<taxonomicName id="4C2F28FFFFF3FFFED8D84CE6FC8D1C92" authority="Kamal &amp; Tan &amp; Ng, 2020" authorityName="Kamal &amp; Tan &amp; Ng" authorityYear="2020" box="[651,801,443,467]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="189" phylum="Chordata" rank="species" species="nuluhon" status="sp. nov.">
<emphasis id="B95B8F6EFFF3FFFED8D84CE6FC8D1C92" box="[651,801,443,467]" italics="true" pageId="2" pageNumber="189">Betta nuluhon</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2683215FFF3FFFED97C4CE6FC031C92" box="[815,943,443,467]" pageId="2" pageNumber="189" rank="species">new species</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B90537CFFF3FFFEDAF54CBFFA8F1B97" pageId="2" pageNumber="189">
<table id="F92FA1DCFFF30003DAF54CBFFAF31B97" box="[166,1375,482,1750]" gridcols="5" gridrows="31" pageId="2" pageNumber="189">
<tr id="351F513EFFF30003DAF54CBFFAF31F5E" box="[166,1375,482,543]" gridrow="0" pageId="2" pageNumber="189">
<th id="76CE3842FFF30003DAF54CBFFD861F5E" box="[166,554,482,543]" gridcol="0" gridrow="0" pageId="2" pageNumber="189">Meristics</th>
<th id="76CE3842FFF30003D83F4CBFFD4F1F5E" box="[620,739,482,543]" gridcol="1" gridrow="0" pageId="2" pageNumber="189">Holotype ZRC 61247</th>
<th id="76CE3842FFF30003D9714CBFFAF31F5E" box="[802,1375,482,543]" colspan="3" colspanRight="2" gridcol="2" gridrow="0" pageId="2" pageNumber="189">Paratypes (+ holotype data, n = 17, SL = 39.8–54.0 mm) ZRC 57026+FRC, 61154+FRC</th>
</tr>
<tr id="351F513EFFF30003DAF54F73FAF31F06" box="[166,1375,558,583]" gridrow="1" pageId="2" pageNumber="189" rowspan-4="1">
<th id="76CE3842FFF30003DAF54F73FD861F06" box="[166,554,558,583]" gridcol="0" gridrow="1" pageId="2" pageNumber="189">anal fin rays</th>
<td id="76CE3842FFF30003D83F4F73FD4F1F06" box="[620,739,558,583]" gridcol="1" gridrow="1" pageId="2" pageNumber="189">I,25</td>
<td id="76CE3842FFF30003D9714F73FB351F06" box="[802,1177,558,583]" colspan="2" colspanRight="1" gridcol="2" gridrow="1" pageId="2" pageNumber="189">I–II,25–27 (total 26–28, mode 27)</td>
</tr>
<tr id="351F513EFFF30003DAF54F0AFAF31F31" box="[166,1375,599,624]" gridrow="2" pageId="2" pageNumber="189" rowspan-3="1" rowspan-4="1">
<th id="76CE3842FFF30003DAF54F0AFD861F31" box="[166,554,599,624]" gridcol="0" gridrow="2" pageId="2" pageNumber="189">dorsal fin rays</th>
<td id="76CE3842FFF30003D83F4F0AFD4F1F31" box="[620,739,599,624]" gridcol="1" gridrow="2" pageId="2" pageNumber="189">I,7</td>
<td id="76CE3842FFF30003D9714F0AFC521F31" box="[802,1022,599,624]" gridcol="2" gridrow="2" pageId="2" pageNumber="189">I,7</td>
</tr>
<tr id="351F513EFFF30003DAF54F22FAF31FD9" box="[166,1375,639,664]" gridrow="3" pageId="2" pageNumber="189" rowspan-3="1" rowspan-4="1">
<th id="76CE3842FFF30003DAF54F22FD861FD9" box="[166,554,639,664]" gridcol="0" gridrow="3" pageId="2" pageNumber="189">caudal fin rays</th>
<td id="76CE3842FFF30003D83F4F22FD4F1FD9" box="[620,739,639,664]" gridcol="1" gridrow="3" pageId="2" pageNumber="189">iii,5+6,ii</td>
<td id="76CE3842FFF30003D9714F22FC521FD9" box="[802,1022,639,664]" gridcol="2" gridrow="3" pageId="2" pageNumber="189">iii,5+6,ii</td>
</tr>
<tr id="351F513EFFF30003DAF54FFAFAF31F81" box="[166,1375,679,704]" gridrow="4" pageId="2" pageNumber="189" rowspan-3="1" rowspan-4="1">
<th id="76CE3842FFF30003DAF54FFAFD861F81" box="[166,554,679,704]" gridcol="0" gridrow="4" pageId="2" pageNumber="189">pelvic fin rays</th>
<td id="76CE3842FFF30003D83F4FFAFD4F1F81" box="[620,739,679,704]" gridcol="1" gridrow="4" pageId="2" pageNumber="189">I,5</td>
<td id="76CE3842FFF30003D9714FFAFC521F81" box="[802,1022,679,704]" gridcol="2" gridrow="4" pageId="2" pageNumber="189">I,5</td>
</tr>
<tr id="351F513EFFF30003DAF54F92FAF31FA9" box="[166,1375,719,744]" gridrow="5" pageId="2" pageNumber="189" rowspan-3="1" rowspan-4="1">
<th id="76CE3842FFF30003DAF54F92FD861FA9" box="[166,554,719,744]" gridcol="0" gridrow="5" pageId="2" pageNumber="189">pectoral fin rays</th>
<td id="76CE3842FFF30003D83F4F92FD4F1FA9" box="[620,739,719,744]" gridcol="1" gridrow="5" pageId="2" pageNumber="189">14</td>
<td id="76CE3842FFF30003D9714F92FC521FA9" box="[802,1022,719,744]" gridcol="2" gridrow="5" pageId="2" pageNumber="189">13–14 (mode 13)</td>
</tr>
<tr id="351F513EFFF30003DAF54FA5FAF31E50" box="[166,1375,760,785]" gridrow="6" pageId="2" pageNumber="189" rowspan-3="1" rowspan-4="1">
<th id="76CE3842FFF30003DAF54FA5FD861E50" box="[166,554,760,785]" gridcol="0" gridrow="6" pageId="2" pageNumber="189">subdorsal scales</th>
<td id="76CE3842FFF30003D83F4FA5FD4F1E50" box="[620,739,760,785]" gridcol="1" gridrow="6" pageId="2" pageNumber="189">6</td>
<td id="76CE3842FFF30003D9714FA5FC521E50" box="[802,1022,760,785]" gridcol="2" gridrow="6" pageId="2" pageNumber="189">5½–6½ (mode 6)</td>
</tr>
<tr id="351F513EFFF30003DAF54E7DFAF31E78" box="[166,1375,800,825]" gridrow="7" pageId="2" pageNumber="189" rowspan-3="1" rowspan-4="1">
<th id="76CE3842FFF30003DAF54E7DFD861E78" box="[166,554,800,825]" gridcol="0" gridrow="7" pageId="2" pageNumber="189">dorsal depth scales</th>
<td id="76CE3842FFF30003D83F4E7DFD4F1E78" box="[620,739,800,825]" gridcol="1" gridrow="7" pageId="2" pageNumber="189">9½</td>
<td id="76CE3842FFF30003D9714E7DFC521E78" box="[802,1022,800,825]" gridcol="2" gridrow="7" pageId="2" pageNumber="189">9½</td>
</tr>
<tr id="351F513EFFF30003DAF54E15FAF31E20" box="[166,1375,840,865]" gridrow="8" pageId="2" pageNumber="189" rowspan-3="1" rowspan-4="1">
<th id="76CE3842FFF30003DAF54E15FD861E20" box="[166,554,840,865]" gridcol="0" gridrow="8" pageId="2" pageNumber="189">lateral scales</th>
<td id="76CE3842FFF30003D83F4E15FD4F1E20" box="[620,739,840,865]" gridcol="1" gridrow="8" pageId="2" pageNumber="189">31</td>
<td id="76CE3842FFF30003D9714E15FC521E20" box="[802,1022,840,865]" gridcol="2" gridrow="8" pageId="2" pageNumber="189">29–31 (mode 30)</td>
</tr>
<tr id="351F513EFFF30003DAF54E2DFAF31EC8" box="[166,1375,880,905]" gridrow="9" pageId="2" pageNumber="189" rowspan-3="1" rowspan-4="1">
<th id="76CE3842FFF30003DAF54E2DFD861EC8" box="[166,554,880,905]" gridcol="0" gridrow="9" pageId="2" pageNumber="189">Lateral scale count at dorsal-fin origin</th>
<td id="76CE3842FFF30003D83F4E2DFD4F1EC8" box="[620,739,880,905]" gridcol="1" gridrow="9" pageId="2" pageNumber="189">16</td>
<td id="76CE3842FFF30003D9714E2DFC521EC8" box="[802,1022,880,905]" gridcol="2" gridrow="9" pageId="2" pageNumber="189">14–16 (mode 15)</td>
</tr>
<tr id="351F513EFFF30003DAF54EC5FAF31EF0" box="[166,1375,920,945]" gridrow="10" pageId="2" pageNumber="189" rowspan-3="1" rowspan-4="1">
<th id="76CE3842FFF30003DAF54EC5FD861EF0" box="[166,554,920,945]" gridcol="0" gridrow="10" pageId="2" pageNumber="189">Lateral scale count at anal-fin origin</th>
<td id="76CE3842FFF30003D83F4EC5FD4F1EF0" box="[620,739,920,945]" gridcol="1" gridrow="10" pageId="2" pageNumber="189">6</td>
<td id="76CE3842FFF30003D9714EC5FC521EF0" box="[802,1022,920,945]" gridcol="2" gridrow="10" pageId="2" pageNumber="189">6–7 (mode 6)</td>
</tr>
<tr id="351F513EFFF30003DAF54E9CFAF31E9B" box="[166,1375,961,986]" gridrow="11" pageId="2" pageNumber="189" rowspan-3="1" rowspan-4="1">
<th id="76CE3842FFF30003DAF54E9CFD861E9B" box="[166,554,961,986]" gridcol="0" gridrow="11" pageId="2" pageNumber="189">predorsal scales</th>
<td id="76CE3842FFF30003D83F4E9CFD4F1E9B" box="[620,739,961,986]" gridcol="1" gridrow="11" pageId="2" pageNumber="189">21</td>
<td id="76CE3842FFF30003D9714E9CFC521E9B" box="[802,1022,961,986]" gridcol="2" gridrow="11" pageId="2" pageNumber="189">20–21 (mode 20)</td>
</tr>
<tr id="351F513EFFF30003DAF54EB4FAF31943" box="[166,1375,1001,1026]" gridrow="12" pageId="2" pageNumber="189" rowspan-3="1" rowspan-4="1">
<th id="76CE3842FFF30003DAF54EB4FD861943" box="[166,554,1001,1026]" gridcol="0" gridrow="12" pageId="2" pageNumber="189">postdorsal scales</th>
<td id="76CE3842FFF30003D83F4EB4FD4F1943" box="[620,739,1001,1026]" gridcol="1" gridrow="12" pageId="2" pageNumber="189">11</td>
<td id="76CE3842FFF30003D9714EB4FC521943" box="[802,1022,1001,1026]" gridcol="2" gridrow="12" pageId="2" pageNumber="189">10–11 (mode 11)</td>
</tr>
<tr id="351F513EFFF30003DAF5494CFAF3196B" box="[166,1375,1041,1066]" gridrow="13" pageId="2" pageNumber="189">
<th id="76CE3842FFF30003DAF5494CFAF3196B" box="[166,1375,1041,1066]" colspan="5" colspanRight="4" gridcol="0" gridrow="13" pageId="2" pageNumber="189">Morphometrics</th>
</tr>
<tr id="351F513EFFF30003DAF54964FAF31913" box="[166,1375,1081,1106]" gridrow="14" pageId="2" pageNumber="189" rowspan-1="1">
<th id="76CE3842FFF30003DAF54964FD861913" box="[166,554,1081,1106]" gridcol="0" gridrow="14" pageId="2" pageNumber="189">% standard length</th>
<td id="76CE3842FFF30003D9714964FC521913" box="[802,1022,1081,1106]" gridcol="2" gridrow="14" pageId="2" pageNumber="189">min max</td>
<td id="76CE3842FFF30003DE334964FB351913" box="[1120,1177,1081,1106]" gridcol="3" gridrow="14" pageId="2" pageNumber="189">mean</td>
<td id="76CE3842FFF30003DEA44964FAF31913" box="[1271,1375,1081,1106]" gridcol="4" gridrow="14" pageId="2" pageNumber="189">SD</td>
</tr>
<tr id="351F513EFFF30003DAF5493FFAF3193A" box="[166,1375,1122,1147]" gridrow="15" pageId="2" pageNumber="189">
<th id="76CE3842FFF30003DAF5493FFD86193A" box="[166,554,1122,1147]" gridcol="0" gridrow="15" pageId="2" pageNumber="189">total length</th>
<td id="76CE3842FFF30003D83F493FFD4F193A" box="[620,739,1122,1147]" gridcol="1" gridrow="15" pageId="2" pageNumber="189">145.4</td>
<td id="76CE3842FFF30003D971493FFC52193A" box="[802,1022,1122,1147]" gridcol="2" gridrow="15" pageId="2" pageNumber="189">139.6 148.4</td>
<td id="76CE3842FFF30003DE33493FFB35193A" box="[1120,1177,1122,1147]" gridcol="3" gridrow="15" pageId="2" pageNumber="189">143.1</td>
<td id="76CE3842FFF30003DEA4493FFAF3193A" box="[1271,1375,1122,1147]" gridcol="4" gridrow="15" pageId="2" pageNumber="189">2.75</td>
</tr>
<tr id="351F513EFFF30003DAF549D7FAF319E2" box="[166,1375,1162,1187]" gridrow="16" pageId="2" pageNumber="189">
<th id="76CE3842FFF30003DAF549D7FD8619E2" box="[166,554,1162,1187]" gridcol="0" gridrow="16" pageId="2" pageNumber="189">body length</th>
<td id="76CE3842FFF30003D83F49D7FD4F19E2" box="[620,739,1162,1187]" gridcol="1" gridrow="16" pageId="2" pageNumber="189">70.6</td>
<td id="76CE3842FFF30003D97149D7FC5219E2" box="[802,1022,1162,1187]" gridcol="2" gridrow="16" pageId="2" pageNumber="189">65.8 70.6</td>
<td id="76CE3842FFF30003DE3349D7FB3519E2" box="[1120,1177,1162,1187]" gridcol="3" gridrow="16" pageId="2" pageNumber="189">68.4</td>
<td id="76CE3842FFF30003DEA449D7FAF319E2" box="[1271,1375,1162,1187]" gridcol="4" gridrow="16" pageId="2" pageNumber="189">1.51</td>
</tr>
<tr id="351F513EFFF30003DAF549EFFAF3198A" box="[166,1375,1202,1227]" gridrow="17" pageId="2" pageNumber="189">
<th id="76CE3842FFF30003DAF549EFFD86198A" box="[166,554,1202,1227]" gridcol="0" gridrow="17" pageId="2" pageNumber="189">predorsal length</th>
<td id="76CE3842FFF30003D83F49EFFD4F198A" box="[620,739,1202,1227]" gridcol="1" gridrow="17" pageId="2" pageNumber="189">68.4</td>
<td id="76CE3842FFF30003D97149EFFC52198A" box="[802,1022,1202,1227]" gridcol="2" gridrow="17" pageId="2" pageNumber="189">65.8 71.3</td>
<td id="76CE3842FFF30003DE3349EFFB35198A" box="[1120,1177,1202,1227]" gridcol="3" gridrow="17" pageId="2" pageNumber="189">68.3</td>
<td id="76CE3842FFF30003DEA449EFFAF3198A" box="[1271,1375,1202,1227]" gridcol="4" gridrow="17" pageId="2" pageNumber="189">1.45</td>
</tr>
<tr id="351F513EFFF30003DAF54987FAF319B2" box="[166,1375,1242,1267]" gridrow="18" pageId="2" pageNumber="189">
<th id="76CE3842FFF30003DAF54987FD8619B2" box="[166,554,1242,1267]" gridcol="0" gridrow="18" pageId="2" pageNumber="189">postdorsal length</th>
<td id="76CE3842FFF30003D83F4987FD4F19B2" box="[620,739,1242,1267]" gridcol="1" gridrow="18" pageId="2" pageNumber="189">22.0</td>
<td id="76CE3842FFF30003D9714987FC5219B2" box="[802,1022,1242,1267]" gridcol="2" gridrow="18" pageId="2" pageNumber="189">18.1 24.2</td>
<td id="76CE3842FFF30003DE334987FB3519B2" box="[1120,1177,1242,1267]" gridcol="3" gridrow="18" pageId="2" pageNumber="189">20.9</td>
<td id="76CE3842FFF30003DEA44987FAF319B2" box="[1271,1375,1242,1267]" gridcol="4" gridrow="18" pageId="2" pageNumber="189">1.66</td>
</tr>
<tr id="351F513EFFF30003DAF5485EFAF3185D" box="[166,1375,1283,1308]" gridrow="19" pageId="2" pageNumber="189">
<th id="76CE3842FFF30003DAF5485EFD86185D" box="[166,554,1283,1308]" gridcol="0" gridrow="19" pageId="2" pageNumber="189">caudal peduncle depth</th>
<td id="76CE3842FFF30003D83F485EFD4F185D" box="[620,739,1283,1308]" gridcol="1" gridrow="19" pageId="2" pageNumber="189">20.0</td>
<td id="76CE3842FFF30003D971485EFC52185D" box="[802,1022,1283,1308]" gridcol="2" gridrow="19" pageId="2" pageNumber="189">16.4 20.0</td>
<td id="76CE3842FFF30003DE33485EFB35185D" box="[1120,1177,1283,1308]" gridcol="3" gridrow="19" pageId="2" pageNumber="189">17.7</td>
<td id="76CE3842FFF30003DEA4485EFAF3185D" box="[1271,1375,1283,1308]" gridcol="4" gridrow="19" pageId="2" pageNumber="189">0.79</td>
</tr>
<tr id="351F513EFFF30003DAF54876FAF31805" box="[166,1375,1323,1348]" gridrow="20" pageId="2" pageNumber="189">
<th id="76CE3842FFF30003DAF54876FD861805" box="[166,554,1323,1348]" gridcol="0" gridrow="20" pageId="2" pageNumber="189">preanal length</th>
<td id="76CE3842FFF30003D83F4876FD4F1805" box="[620,739,1323,1348]" gridcol="1" gridrow="20" pageId="2" pageNumber="189">46.6</td>
<td id="76CE3842FFF30003D9714876FC521805" box="[802,1022,1323,1348]" gridcol="2" gridrow="20" pageId="2" pageNumber="189">45.4 49.7</td>
<td id="76CE3842FFF30003DE334876FB351805" box="[1120,1177,1323,1348]" gridcol="3" gridrow="20" pageId="2" pageNumber="189">47.7</td>
<td id="76CE3842FFF30003DEA44876FAF31805" box="[1271,1375,1323,1348]" gridcol="4" gridrow="20" pageId="2" pageNumber="189">1.31</td>
</tr>
<tr id="351F513EFFF30003DAF5480EFAF3182D" box="[166,1375,1363,1388]" gridrow="21" pageId="2" pageNumber="189">
<th id="76CE3842FFF30003DAF5480EFD86182D" box="[166,554,1363,1388]" gridcol="0" gridrow="21" pageId="2" pageNumber="189">head length</th>
<td id="76CE3842FFF30003D83F480EFD4F182D" box="[620,739,1363,1388]" gridcol="1" gridrow="21" pageId="2" pageNumber="189">33.5</td>
<td id="76CE3842FFF30003D971480EFC52182D" box="[802,1022,1363,1388]" gridcol="2" gridrow="21" pageId="2" pageNumber="189">31.7 35.2</td>
<td id="76CE3842FFF30003DE33480EFB35182D" box="[1120,1177,1363,1388]" gridcol="3" gridrow="21" pageId="2" pageNumber="189">33.6</td>
<td id="76CE3842FFF30003DEA4480EFAF3182D" box="[1271,1375,1363,1388]" gridcol="4" gridrow="21" pageId="2" pageNumber="189">0.95</td>
</tr>
<tr id="351F513EFFF30003DAF54826FAF318D5" box="[166,1375,1403,1428]" gridrow="22" pageId="2" pageNumber="189">
<th id="76CE3842FFF30003DAF54826FD8618D5" box="[166,554,1403,1428]" gridcol="0" gridrow="22" pageId="2" pageNumber="189">dorsal depth</th>
<td id="76CE3842FFF30003D83F4826FD4F18D5" box="[620,739,1403,1428]" gridcol="1" gridrow="22" pageId="2" pageNumber="189">25.2</td>
<td id="76CE3842FFF30003D9714826FC5218D5" box="[802,1022,1403,1428]" gridcol="2" gridrow="22" pageId="2" pageNumber="189">22.1 25.2</td>
<td id="76CE3842FFF30003DE334826FB3518D5" box="[1120,1177,1403,1428]" gridcol="3" gridrow="22" pageId="2" pageNumber="189">23.6</td>
<td id="76CE3842FFF30003DEA44826FAF318D5" box="[1271,1375,1403,1428]" gridcol="4" gridrow="22" pageId="2" pageNumber="189">0.98</td>
</tr>
<tr id="351F513EFFF30003DAF548F9FAF318FC" box="[166,1375,1444,1469]" gridrow="23" pageId="2" pageNumber="189">
<th id="76CE3842FFF30003DAF548F9FD8618FC" box="[166,554,1444,1469]" gridcol="0" gridrow="23" pageId="2" pageNumber="189">pelvic fin length</th>
<td id="76CE3842FFF30003D83F48F9FD4F18FC" box="[620,739,1444,1469]" gridcol="1" gridrow="23" pageId="2" pageNumber="189">36.4</td>
<td id="76CE3842FFF30003D97148F9FC5218FC" box="[802,1022,1444,1469]" gridcol="2" gridrow="23" pageId="2" pageNumber="189">31.5 45.1</td>
<td id="76CE3842FFF30003DE3348F9FB3518FC" box="[1120,1177,1444,1469]" gridcol="3" gridrow="23" pageId="2" pageNumber="189">35.6</td>
<td id="76CE3842FFF30003DEA448F9FAF318FC" box="[1271,1375,1444,1469]" gridcol="4" gridrow="23" pageId="2" pageNumber="189">3.36</td>
</tr>
<tr id="351F513EFFF30003DAF54891FAF318A4" box="[166,1375,1484,1509]" gridrow="24" pageId="2" pageNumber="189">
<th id="76CE3842FFF30003DAF54891FD8618A4" box="[166,554,1484,1509]" gridcol="0" gridrow="24" pageId="2" pageNumber="189">anal fin base length</th>
<td id="76CE3842FFF30003D83F4891FD4F18A4" box="[620,739,1484,1509]" gridcol="1" gridrow="24" pageId="2" pageNumber="189">51.1</td>
<td id="76CE3842FFF30003D9714891FC5218A4" box="[802,1022,1484,1509]" gridcol="2" gridrow="24" pageId="2" pageNumber="189">49.5 54.6</td>
<td id="76CE3842FFF30003DE334891FB3518A4" box="[1120,1177,1484,1509]" gridcol="3" gridrow="24" pageId="2" pageNumber="189">51.9</td>
<td id="76CE3842FFF30003DEA44891FAF318A4" box="[1271,1375,1484,1509]" gridcol="4" gridrow="24" pageId="2" pageNumber="189">1.40</td>
</tr>
<tr id="351F513EFFF30003DAF548A9FAF31B4C" box="[166,1375,1524,1549]" gridrow="25" pageId="2" pageNumber="189">
<th id="76CE3842FFF30003DAF548A9FD861B4C" box="[166,554,1524,1549]" gridcol="0" gridrow="25" pageId="2" pageNumber="189">dorsal fin base length</th>
<td id="76CE3842FFF30003D83F48A9FD4F1B4C" box="[620,739,1524,1549]" gridcol="1" gridrow="25" pageId="2" pageNumber="189">12.8</td>
<td id="76CE3842FFF30003D97148A9FC521B4C" box="[802,1022,1524,1549]" gridcol="2" gridrow="25" pageId="2" pageNumber="189">10.6 13.5</td>
<td id="76CE3842FFF30003DE3348A9FB351B4C" box="[1120,1177,1524,1549]" gridcol="3" gridrow="25" pageId="2" pageNumber="189">12.3</td>
<td id="76CE3842FFF30003DEA448A9FAF31B4C" box="[1271,1375,1524,1549]" gridcol="4" gridrow="25" pageId="2" pageNumber="189">0.82</td>
</tr>
<tr id="351F513EFFF30003DAF54B41FAF31B74" box="[166,1375,1564,1589]" gridrow="26" pageId="2" pageNumber="189">
<th id="76CE3842FFF30003DAF54B41FAF31B74" box="[166,1375,1564,1589]" colspan="5" colspanRight="4" gridcol="0" gridrow="26" pageId="2" pageNumber="189">% head length</th>
</tr>
<tr id="351F513EFFF30003DAF54B18FAF31B1F" box="[166,1375,1605,1630]" gridrow="27" pageId="2" pageNumber="189">
<th id="76CE3842FFF30003DAF54B18FD861B1F" box="[166,554,1605,1630]" gridcol="0" gridrow="27" pageId="2" pageNumber="189">orbit diameter</th>
<td id="76CE3842FFF30003D83F4B18FD4F1B1F" box="[620,739,1605,1630]" gridcol="1" gridrow="27" pageId="2" pageNumber="189">23.3</td>
<td id="76CE3842FFF30003D9714B18FC521B1F" box="[802,1022,1605,1630]" gridcol="2" gridrow="27" pageId="2" pageNumber="189">23.3 29.7</td>
<td id="76CE3842FFF30003DE334B18FB351B1F" box="[1120,1177,1605,1630]" gridcol="3" gridrow="27" pageId="2" pageNumber="189">26.2</td>
<td id="76CE3842FFF30003DEA44B18FAF31B1F" box="[1271,1375,1605,1630]" gridcol="4" gridrow="27" pageId="2" pageNumber="189">1.75</td>
</tr>
<tr id="351F513EFFF30003DAF54B30FAF31BC7" box="[166,1375,1645,1670]" gridrow="28" pageId="2" pageNumber="189">
<th id="76CE3842FFF30003DAF54B30FD861BC7" box="[166,554,1645,1670]" gridcol="0" gridrow="28" pageId="2" pageNumber="189">postorbital length</th>
<td id="76CE3842FFF30003D83F4B30FD4F1BC7" box="[620,739,1645,1670]" gridcol="1" gridrow="28" pageId="2" pageNumber="189">50.5</td>
<td id="76CE3842FFF30003D9714B30FC521BC7" box="[802,1022,1645,1670]" gridcol="2" gridrow="28" pageId="2" pageNumber="189">46.4 51.6</td>
<td id="76CE3842FFF30003DE334B30FB351BC7" box="[1120,1177,1645,1670]" gridcol="3" gridrow="28" pageId="2" pageNumber="189">48.9</td>
<td id="76CE3842FFF30003DEA44B30FAF31BC7" box="[1271,1375,1645,1670]" gridcol="4" gridrow="28" pageId="2" pageNumber="189">1.37</td>
</tr>
<tr id="351F513EFFF30003DAF54BC8FAF31BEF" box="[166,1375,1685,1710]" gridrow="29" pageId="2" pageNumber="189">
<th id="76CE3842FFF30003DAF54BC8FD861BEF" box="[166,554,1685,1710]" gridcol="0" gridrow="29" pageId="2" pageNumber="189">interorbital width</th>
<td id="76CE3842FFF30003D83F4BC8FD4F1BEF" box="[620,739,1685,1710]" gridcol="1" gridrow="29" pageId="2" pageNumber="189">35.7</td>
<td id="76CE3842FFF30003D9714BC8FC521BEF" box="[802,1022,1685,1710]" gridcol="2" gridrow="29" pageId="2" pageNumber="189">30.4 35.7</td>
<td id="76CE3842FFF30003DE334BC8FB351BEF" box="[1120,1177,1685,1710]" gridcol="3" gridrow="29" pageId="2" pageNumber="189">33.0</td>
<td id="76CE3842FFF30003DEA44BC8FAF31BEF" box="[1271,1375,1685,1710]" gridcol="4" gridrow="29" pageId="2" pageNumber="189">1.35</td>
</tr>
<tr id="351F513EFFF30003DAF54BE0FAF31B97" box="[166,1375,1725,1750]" gridrow="30" pageId="2" pageNumber="189">
<th id="76CE3842FFF30003DAF54BE0FD861B97" box="[166,554,1725,1750]" gridcol="0" gridrow="30" pageId="2" pageNumber="189">snout length</th>
<td id="76CE3842FFF30003D83F4BE0FD4F1B97" box="[620,739,1725,1750]" gridcol="1" gridrow="30" pageId="2" pageNumber="189">21.9</td>
<td id="76CE3842FFF30003D9714BE0FC521B97" box="[802,1022,1725,1750]" gridcol="2" gridrow="30" pageId="2" pageNumber="189">20.0 25.8</td>
<td id="76CE3842FFF30003DE334BE0FB351B97" box="[1120,1177,1725,1750]" gridcol="3" gridrow="30" pageId="2" pageNumber="189">23.1</td>
<td id="76CE3842FFF30003DEA44BE0FAF31B97" box="[1271,1375,1725,1750]" gridcol="4" gridrow="30" pageId="2" pageNumber="189">1.64</td>
</tr>
</table>
</paragraph>
<paragraph id="8B90537CFFF3FFFEDA944A5BFCFB1AB9" blockId="2.[151,1436,1797,2040]" pageId="2" pageNumber="189">
<emphasis id="B95B8F6EFFF3FFFEDA944A5BFE301A61" bold="true" box="[199,412,1797,1824]" pageId="2" pageNumber="189">Fresh colouration</emphasis>
. See
<figureCitation id="13144FF9FFF3FFFEDB8F4A5BFD861A61" box="[476,554,1798,1824]" captionStart="FIGURE 1" captionStartId="3.[151,250,787,812]" captionTargetBox="[149,1438,184,761]" captionTargetId="figure@3.[149,1438,181,761]" captionTargetPageId="3" captionText="FIGURE 1. Betta nuluhon, new species, ca. 50 mm SL, freshly caught male specimen, not preserved." figureDoi="http://doi.org/10.5281/zenodo.3955810" httpUri="https://zenodo.org/record/3955810/files/figure.png" pageId="2" pageNumber="189">Figs. 1</figureCitation>
,
<figureCitation id="13144FF9FFF3FFFED86A4A5BFDE51A61" box="[569,585,1798,1824]" captionStart="FIGURE 2" captionStartId="3.[151,250,1708,1733]" captionTargetBox="[150,1437,834,1684]" captionTargetId="figure@3.[150,1437,834,1684]" captionTargetPageId="3" captionText="FIGURE 2. Betta nuluhon, new species, male (top), female (bottom); freshly caught, not preserved." figureDoi="http://doi.org/10.5281/zenodo.3955812" httpUri="https://zenodo.org/record/3955812/files/figure.png" pageId="2" pageNumber="189">2</figureCitation>
,
<figureCitation id="13144FF9FFF3FFFED8054A5BFDCB1A61" box="[598,615,1798,1824]" captionStart="FIGURE 4" captionStartId="5.[151,250,1353,1378]" captionTargetBox="[151,1436,561,1325]" captionTargetId="figure@5.[151,1436,561,1328]" captionTargetPageId="5" captionText="FIGURE 4. Top—Betta chini, ZRC 61155, 58.6 mm SL male; bottom—Betta nuluhon, holotype, ZRC 61247, 62.6 mm SL male; both freshly preserved." figureDoi="http://doi.org/10.5281/zenodo.3955816" httpUri="https://zenodo.org/record/3955816/files/figure.png" pageId="2" pageNumber="189">4</figureCitation>
. Head brown, dorsum dark brown; dark brown to black preorbital stripe running from upper jaw through eye to postorbital to opercle edge; black chin bar present, dark brown suborbital stripe present; eye with dorsal region of iris yellow, lateral regions black, bottom region reddish; mature male with whole throat area dark brown, sometimes with greenish-blue iridescence; female and juvenile with brown throat; body brown, dorsum dark brown, posterior edge of body scales along flank rimmed with bright blue; all fins brownish, dorsal fin with 4 to 6 black transverse bars, caudal fin with 12 to 16 black transverse bars, anal fin with broad reddish-brown margin, pelvic fin with whitish filamentous ray.
</paragraph>
<caption id="DF5003F4FFF2FFFFDAC44E4EFB0D1E6D" ID-DOI="http://doi.org/10.5281/zenodo.3955810" ID-Zenodo-Dep="3955810" box="[151,1185,787,812]" httpUri="https://zenodo.org/record/3955810/files/figure.png" pageId="3" pageNumber="190" startId="3.[151,250,787,812]" targetBox="[149,1438,184,761]" targetPageId="3">
<paragraph id="8B90537CFFF2FFFFDAC44E4EFB0D1E6D" blockId="3.[151,1185,787,812]" box="[151,1185,787,812]" pageId="3" pageNumber="190">
<emphasis id="B95B8F6EFFF2FFFFDAC44E4EFEBF1E6D" bold="true" box="[151,275,787,812]" pageId="3" pageNumber="190">FIGURE 1.</emphasis>
<taxonomicName id="4C2F28FFFFF2FFFFDB4A4E49FE041E6A" authority="Kamal &amp; Tan &amp; Ng, 2020" authorityName="Kamal &amp; Tan &amp; Ng" authorityYear="2020" box="[281,424,787,811]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="190" phylum="Chordata" rank="species" species="nuluhon" status="sp. nov.">
<emphasis id="B95B8F6EFFF2FFFFDB4A4E49FE041E6A" box="[281,424,787,811]" italics="true" pageId="3" pageNumber="190">Betta nuluhon</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2683215FFF2FFFFDBE64E49FD9D1E6D" box="[437,561,787,812]" pageId="3" pageNumber="190" rank="species">new species</taxonomicNameLabel>
, ca. 50 mm SL, freshly caught male specimen, not preserved.
</paragraph>
</caption>
<caption id="DF5003F4FFF2FFFFDAC44BF1FB3C1B84" ID-DOI="http://doi.org/10.5281/zenodo.3955812" ID-Zenodo-Dep="3955812" box="[151,1168,1708,1733]" httpUri="https://zenodo.org/record/3955812/files/figure.png" pageId="3" pageNumber="190" startId="3.[151,250,1708,1733]" targetBox="[150,1437,834,1684]" targetPageId="3">
<paragraph id="8B90537CFFF2FFFFDAC44BF1FB3C1B84" blockId="3.[151,1168,1708,1733]" box="[151,1168,1708,1733]" pageId="3" pageNumber="190">
<emphasis id="B95B8F6EFFF2FFFFDAC44BF1FEBF1B84" bold="true" box="[151,275,1708,1733]" pageId="3" pageNumber="190">FIGURE 2.</emphasis>
<taxonomicName id="4C2F28FFFFF2FFFFDB4A4BF3FE041B84" authority="Kamal &amp; Tan &amp; Ng, 2020" authorityName="Kamal &amp; Tan &amp; Ng" authorityYear="2020" box="[281,424,1709,1733]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="190" phylum="Chordata" rank="species" species="nuluhon" status="sp. nov.">
<emphasis id="B95B8F6EFFF2FFFFDB4A4BF3FE041B84" box="[281,424,1709,1733]" italics="true" pageId="3" pageNumber="190">Betta nuluhon</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2683215FFF2FFFFDBE64BF3FD9D1B84" box="[437,561,1708,1733]" pageId="3" pageNumber="190" rank="species">new species</taxonomicNameLabel>
, male (top), female (bottom); freshly caught, not preserved.
</paragraph>
</caption>
<paragraph id="8B90537CFFF2FFFFDA944BA9FBED1AA7" blockId="3.[151,1436,1779,2022]" pageId="3" pageNumber="190">
<emphasis id="B95B8F6EFFF2FFFFDA944BA9FE631A4F" bold="true" box="[199,463,1779,1806]" pageId="3" pageNumber="190">Preserved colouration</emphasis>
. See
<figureCitation id="13144FF9FFF2FFFFD85F4BAEFDFC1A4F" box="[524,592,1779,1806]" captionStart="FIGURE 3" captionStartId="4.[151,250,1952,1977]" captionTargetBox="[151,1436,1160,1927]" captionTargetId="figure@4.[151,1436,1101,1927]" captionTargetPageId="4" captionText="FIGURE 3. Betta nuluhon, new species, ZRC 61247, holotype, 62.6 mm SL (above with white background, bottom with black background)." figureDoi="http://doi.org/10.5281/zenodo.3955814" httpUri="https://zenodo.org/record/3955814/files/figure.png" pageId="3" pageNumber="190">Fig. 3</figureCitation>
. Head brown, dorsum dark brown, black preorbital stripe running from upper jaw through eye to postorbital to opercular edge, black chin bar present, dark brown suborbital stripe present with pigments along opercle edge (pattern similar to
<taxonomicName id="4C2F28FFFFF2FFFFD88C4A61FCF41A14" authorityName="Ng" authorityYear="1993" box="[735,856,1851,1877]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="190" phylum="Chordata" rank="species" species="chini">
<emphasis id="B95B8F6EFFF2FFFFD88C4A61FCF41A14" box="[735,856,1851,1877]" italics="true" pageId="3" pageNumber="190">Betta chini</emphasis>
</taxonomicName>
, see
<bibRefCitation id="EFBE2E8DFFF2FFFFD9DD4A61FBC11A17" author="Tan, H. H. &amp; Ng, P. K. L." box="[910,1133,1851,1878]" pageId="3" pageNumber="190" pagination="43 - 99" refId="ref4540" refString="Tan, H. H. &amp; Ng, P. K. L. (2005) The fighting fishes (Teleostei: Osphronemidae: genus Betta) of Singapore, Malaysia and Brunei. Raffles Bulletin of Zoology, Supplement 13, 43 - 99." type="journal article" year="2005">Tan &amp; Ng, 2005: 49</bibRefCitation>
, Fig. 7c); mature male with whole throat area diffused with brown; female and juvenile with throat area light brown; body brown, dorsum dark brown; belly area cream with light brown reticulate pattern from posterior edge of scales bearing darker pigments; all fins brownish except hyaline pectoral fin, dorsal fin with 4 to 6 black transverse bars, caudal fin with 12 to 16 black transverse bars, anal fin reddish-brown, pelvic fin with white filamentous ray.
</paragraph>
<paragraph id="8B90537CFFF5FFF8DA944DC5FE871CCD" blockId="4.[151,1437,152,1080]" pageId="4" pageNumber="191">
<emphasis id="B95B8F6EFFF5FFF8DA944DC5FEE61DF2" bold="true" box="[199,330,152,179]" pageId="4" pageNumber="191">Field notes</emphasis>
. Specimens were collected from the velocity refugia of a shallow (knee level) clear-water stream with overhanging bank vegetation in Crocker Range Forest Reserve’s closed-canopy forest fringe area. The substrate was composed mainly of pebbles, sand and silt. Some water parameters were recorded: dissolved oxygen
<quantity id="4CD7FE99FFF5FFF8DAC44C59FF5F1C5E" box="[151,243,260,287]" metricMagnitude="-6" metricUnit="kg" metricValue="6.25" pageId="4" pageNumber="191" unit="mg" value="6.25">6.25 mg</quantity>
/L, pH 6.57, suspended solids
<quantity id="4CD7FE99FFF5FFF8D8134C58FD221C5E" box="[576,654,261,287]" metricMagnitude="-7" metricUnit="kg" metricValue="4.0" pageId="4" pageNumber="191" unit="mg" value="0.4">0.4 mg</quantity>
/L, turbidity 28.0 FAU, temperature 24.0° C. Syntopic species include:
<taxonomicName id="4C2F28FFFFF5FFF8DAC44C74FD581C05" authority="Quoy &amp; Gaimard" authorityName="Quoy &amp; Gaimard" box="[151,756,297,324]" class="Actinopterygii" family="Anguillidae" genus="Anguilla" kingdom="Animalia" order="Anguilliformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="marmorata">
<emphasis id="B95B8F6EFFF5FFF8DAC44C74FED31C02" box="[151,383,297,323]" italics="true" pageId="4" pageNumber="191">Anguilla marmorata</emphasis>
Quoy &amp; Gaimard (Anguillidae)
</taxonomicName>
,
<taxonomicName id="4C2F28FFFFF5FFF8D9514C74FBA81C02" authority="Herre" authorityName="Herre" authorityYear="1933" box="[770,1028,297,323]" class="Actinopterygii" family="Cyprinidae" genus="Barbodes" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="sealei">
<emphasis id="B95B8F6EFFF5FFF8D9514C74FC1B1C02" box="[770,951,297,323]" italics="true" pageId="4" pageNumber="191">Barbodes sealei</emphasis>
Herre
</taxonomicName>
,
<taxonomicName id="4C2F28FFFFF5FFF8DE424C74FF7C1C26" authority="Inger &amp; Chin" authorityName="Inger &amp; Chin" authorityYear="1962" class="Actinopterygii" family="Cyprinidae" genus="Nematabramis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="borneensis">
<emphasis id="B95B8F6EFFF5FFF8DE424C74FA941C02" box="[1041,1336,297,323]" italics="true" pageId="4" pageNumber="191">Nematabramis borneensis</emphasis>
Inger &amp; Chin
</taxonomicName>
,
<taxonomicName id="4C2F28FFFFF5FFF8DA8E4C13FC6B1C29" authority="Valenciennes, in Cuvier &amp; Valenciennes" authorityName="Valenciennes, in Cuvier &amp; Valenciennes" box="[221,967,333,360]" class="Actinopterygii" family="Cyprinidae" genus="Tor" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="tambra">
<emphasis id="B95B8F6EFFF5FFF8DA8E4C13FEF01C26" box="[221,348,333,359]" italics="true" pageId="4" pageNumber="191">Tor tambra</emphasis>
Valenciennes, in Cuvier &amp; Valenciennes (Cyprinidae)
</taxonomicName>
, and
<taxonomicName id="4C2F28FFFFF5FFF8DE574C10FE8B1CCD" authority="Tan" authorityName="Tan" authorityYear="2006" class="Actinopterygii" family="Balitoridae" genus="Gastromyzon" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="introrsus">
<emphasis id="B95B8F6EFFF5FFF8DE574C10FAA91C26" box="[1028,1285,333,359]" italics="true" pageId="4" pageNumber="191">Gastromyzon introrsus</emphasis>
Tan (Gastromyzontidae)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33500F7FFF5FFF8DA944CC8FD541C92" pageId="4" pageNumber="191" type="etymology">
<paragraph id="8B90537CFFF5FFF8DA944CC8FD541C92" blockId="4.[151,1437,152,1080]" pageId="4" pageNumber="191">
<emphasis id="B95B8F6EFFF5FFF8DA944CC8FEE41CEE" bold="true" box="[199,328,405,431]" pageId="4" pageNumber="191">Etymology</emphasis>
. This new species is named for the Kadazandusun vernacular
<taxonomicName id="4C2F28FFFFF5FFF8DE564CC8FBCD1CEE" authorityName="Kamal &amp; Tan &amp; Ng" authorityYear="2020" box="[1029,1121,405,431]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="nuluhon">
<emphasis id="B95B8F6EFFF5FFF8DE564CC8FBCD1CEE" box="[1029,1121,405,431]" italics="true" pageId="4" pageNumber="191">nuluhon</emphasis>
</taxonomicName>
, meaning hill. This pertains to its hill stream habitat. Used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C33500F7FFF5FFF9DA944C80FD091F5A" lastPageId="5" lastPageNumber="192" pageId="4" pageNumber="191" type="discussion">
<paragraph id="8B90537CFFF5FFF8DA944C80FEB41F8E" blockId="4.[151,1437,152,1080]" pageId="4" pageNumber="191">
<emphasis id="B95B8F6EFFF5FFF8DA944C80FE971CB6" bold="true" box="[199,315,477,503]" pageId="4" pageNumber="191">Remarks.</emphasis>
<taxonomicName id="4C2F28FFFFF5FFF8DB114C83FE4C1CB6" authorityName="Kamal &amp; Tan &amp; Ng" authorityYear="2020" box="[322,480,477,503]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="nuluhon">
<emphasis id="B95B8F6EFFF5FFF8DB114C83FE4C1CB6" box="[322,480,477,503]" italics="true" pageId="4" pageNumber="191">Betta nuluhon</emphasis>
</taxonomicName>
can be distinguished from similar species from the geographic vicinity (
<taxonomicName id="4C2F28FFFFF5FFF8DF5F4C83FA391CB6" authorityName="Regan" authorityYear="1910" box="[1292,1429,477,503]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="akarensis">
<emphasis id="B95B8F6EFFF5FFF8DF5F4C83FA391CB6" box="[1292,1429,477,503]" italics="true" pageId="4" pageNumber="191">B. akarensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2F28FFFFF5FFF8DAC44F5FFEBF1F5A" authorityName="Herre" authorityYear="1940" box="[151,275,513,539]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="balunga">
<emphasis id="B95B8F6EFFF5FFF8DAC44F5FFEBF1F5A" box="[151,275,513,539]" italics="true" pageId="4" pageNumber="191">B. balunga</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2F28FFFFF5FFF8DB194F5FFE0E1F5A" authorityName="Ng" authorityYear="1993" box="[330,418,513,539]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="chini">
<emphasis id="B95B8F6EFFF5FFF8DB194F5FFE0E1F5A" box="[330,418,513,539]" italics="true" pageId="4" pageNumber="191">B. chini</emphasis>
</taxonomicName>
) in the following characters: gently sloping anterior body dorsum profile (vs. convex profile of
<taxonomicName id="4C2F28FFFFF5FFF8DAB74F7BFECD1F7E" authorityName="Herre" authorityYear="1940" box="[228,353,549,575]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="balunga">
<emphasis id="B95B8F6EFFF5FFF8DAB74F7BFECD1F7E" box="[228,353,549,575]" italics="true" pageId="4" pageNumber="191">B. balunga</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2F28FFFFF5FFF8DBCA4F78FE5F1F7E" authorityName="Ng" authorityYear="1993" box="[409,499,549,575]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="192" phylum="Chordata" rank="species" species="chini">
B.
<emphasis id="B95B8F6EFFF5FFF8DBE84F78FE5F1F7E" box="[443,499,549,575]" italics="true" pageId="4" pageNumber="191">chini</emphasis>
</taxonomicName>
); absence of tiny black spots on anal fin (vs. presence of many tiny black spots in
<taxonomicName id="4C2F28FFFFF5FFF8DAC44F17FF431F22" authorityName="Ng" authorityYear="1993" box="[151,239,585,611]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="chini">
<emphasis id="B95B8F6EFFF5FFF8DAC44F17FF431F22" box="[151,239,585,611]" italics="true" pageId="4" pageNumber="191">B. chini</emphasis>
</taxonomicName>
); shallower body depth than
<taxonomicName id="4C2F28FFFFF5FFF8D86F4F17FD6B1F22" authorityName="Regan" authorityYear="1910" box="[572,711,585,611]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="akarensis">
<emphasis id="B95B8F6EFFF5FFF8D86F4F17FD6B1F22" box="[572,711,585,611]" italics="true" pageId="4" pageNumber="191">B. akarensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2F28FFFFF5FFF8D9534F17FCD01F22" authorityName="Herre" authorityYear="1940" box="[768,892,585,611]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="balunga">
<emphasis id="B95B8F6EFFF5FFF8D9534F17FCD01F22" box="[768,892,585,611]" italics="true" pageId="4" pageNumber="191">B. balunga</emphasis>
</taxonomicName>
(22.1–25.2 % SL, vs. 23.5–27.9 and 28.0–31.5, respectively); shorter preanal length than
<taxonomicName id="4C2F28FFFFF5FFF8D8304F33FD711FC6" authorityName="Herre" authorityYear="1940" box="[611,733,621,647]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="balunga">
<emphasis id="B95B8F6EFFF5FFF8D8304F33FD711FC6" box="[611,733,621,647]" italics="true" pageId="4" pageNumber="191">B. balunga</emphasis>
</taxonomicName>
(45.4–49.7 % SL, vs. 49.2–52.2); shorter dorsal-fin base length than
<taxonomicName id="4C2F28FFFFF5FFF8DA824FCFFEE31FEA" authorityName="Herre" authorityYear="1940" box="[209,335,657,683]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="balunga">
<emphasis id="B95B8F6EFFF5FFF8DA824FCFFEE31FEA" box="[209,335,657,683]" italics="true" pageId="4" pageNumber="191">B. balunga</emphasis>
</taxonomicName>
(10.6–13.5 % SL, vs. 13.7–14.1); smaller orbit diameter than
<taxonomicName id="4C2F28FFFFF5FFF8DE734FCFFB321FEA" authorityName="Herre" authorityYear="1940" box="[1056,1182,657,683]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="balunga">
<emphasis id="B95B8F6EFFF5FFF8DE734FCFFB321FEA" box="[1056,1182,657,683]" italics="true" pageId="4" pageNumber="191">B. balunga</emphasis>
</taxonomicName>
(23.3–29.7 % SL, vs. 29.5–33.3).
</paragraph>
<paragraph id="8B90537CFFF5FFF8DA944F87FF641E56" blockId="4.[151,1437,152,1080]" pageId="4" pageNumber="191">
<taxonomicName id="4C2F28FFFFF5FFF8DA944F87FEC91FB2" authorityName="Kamal &amp; Tan &amp; Ng" authorityYear="2020" box="[199,357,729,755]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="nuluhon">
<emphasis id="B95B8F6EFFF5FFF8DA944F87FEC91FB2" box="[199,357,729,755]" italics="true" pageId="4" pageNumber="191">Betta nuluhon</emphasis>
</taxonomicName>
can be differentiated from the other members of the
<taxonomicName id="4C2F28FFFFF5FFF8D9E54F87FBEC1FB2" authorityName="Regan" authorityYear="1910" box="[950,1088,729,755]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="akarensis">
<emphasis id="B95B8F6EFFF5FFF8D9E54F87FBEC1FB2" box="[950,1088,729,755]" italics="true" pageId="4" pageNumber="191">B. akarensis</emphasis>
</taxonomicName>
group in the following characters:
</paragraph>
<paragraph id="8B90537CFFF5FFF8DA944E7CFC581979" blockId="4.[151,1437,152,1080]" pageId="4" pageNumber="191">
From
<taxonomicName id="4C2F28FFFFF5FFF8DB584E7FFEDE1E7A" authorityName="Tan &amp; Ng" authorityYear="2006" box="[267,370,802,827]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="antoni">
<emphasis id="B95B8F6EFFF5FFF8DB584E7FFEDE1E7A" box="[267,370,802,827]" italics="true" pageId="4" pageNumber="191">B. antoni</emphasis>
</taxonomicName>
—deeper body (22.1–25.2 % SL, vs. 20.6–22.9); fewer lateral scales (29–31, vs. 31–33); fewer predorsal scales (20–21, vs. 23–24). From
<taxonomicName id="4C2F28FFFFF5FFF8D8D34E1BFCA91E1E" authorityName="Tan &amp; Lim" authorityYear="2004" box="[640,773,838,863]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="aurigans">
<emphasis id="B95B8F6EFFF5FFF8D8D34E1BFCA91E1E" box="[640,773,838,863]" italics="true" pageId="4" pageNumber="191">B. aurigans</emphasis>
</taxonomicName>
—shallower body (22.1–25.2 % SL, vs. 25.2–27.4); fewer lateral scales (29–31, vs. 33–33½); fewer predorsal scales (20–21, vs. 24–26); fewer anal-fin rays (26–28, vs. 29–30). From
<taxonomicName id="4C2F28FFFFF5FFF8DB134ED3FE7D1EE6" authorityName="Tan &amp; Ng" authorityYear="2004" box="[320,465,909,935]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="ibanorum">
<emphasis id="B95B8F6EFFF5FFF8DB134ED3FE7D1EE6" box="[320,465,909,935]" italics="true" pageId="4" pageNumber="191">B. ibanorum</emphasis>
</taxonomicName>
—more patterned opercle (vs. less patterned); lower range of lateral scales (29–31, vs. 30–32); fewer predorsal scales (20–21, vs. 21–23).From
<taxonomicName id="4C2F28FFFFF5FFF8D9074EEFFC7C1E8A" authorityName="Tan &amp; Ng" authorityYear="2005" box="[852,976,945,971]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="obscura">
<emphasis id="B95B8F6EFFF5FFF8D9074EEFFC7C1E8A" box="[852,976,945,971]" italics="true" pageId="4" pageNumber="191">B. obscura</emphasis>
</taxonomicName>
—shallower body (22.1–25.2 % SL, vs. 28.7–32.7). From
<taxonomicName id="4C2F28FFFFF5FFF8DB364E8BFE741EAE" authorityName="Tan &amp; Kottelat" authorityYear="1998" box="[357,472,982,1007]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="pinguis">
<emphasis id="B95B8F6EFFF5FFF8DB364E8BFE741EAE" box="[357,472,982,1007]" italics="true" pageId="4" pageNumber="191">B. pinguis</emphasis>
</taxonomicName>
—less pigmented opercle (vs. heavily pigmented); shallower body (22.1–25.2 % SL, vs. 30.6–31.6); fewer lateral scales (29–31, vs. 31½–32½); fewer predorsal scales (20–21, vs. 23–24) (data obtained from
<bibRefCitation id="EFBE2E8DFFF5FFF8DA874940FE691976" author="Tan, H. H. &amp; Kottelat, M." box="[212,453,1053,1079]" pageId="4" pageNumber="191" pagination="41 - 51" refId="ref4633" refString="Tan, H. H. &amp; Kottelat, M. (1998) Two new species of Betta (Teleostei: Osphronemidae) from the Kapuas basin, Kalimantan Barat, Borneo. Raffles Bulletin of Zoology, 46 (1), 41 - 51." type="journal article" year="1998">Tan &amp; Kottelat, 1998</bibRefCitation>
;
<bibRefCitation id="EFBE2E8DFFF5FFF8DB834940FD3A1976" author="Tan, H. H. &amp; Lim, K. K. P." box="[464,662,1053,1079]" pageId="4" pageNumber="191" pagination="107 - 115" refId="ref4387" refString="Tan, H. H. &amp; Lim, K. K. P. (2004) Inland fishes from the Anambas and Natuna Islands, South China Sea, with description of a new species of Betta (Teleostei: Osphronemidae). Raffles Bulletin of Zoology, Supplement 11, 107 - 115." type="journal article" year="2004">Tan &amp; Lim, 2004</bibRefCitation>
;
<bibRefCitation id="EFBE2E8DFFF5FFF8D8F34940FCF61976" author="Tan, H. H. &amp; Ng, P. K. L." box="[672,858,1053,1079]" pageId="4" pageNumber="191" pagination="267 - 284" refId="ref4444" refString="Tan, H. H. &amp; Ng, P. K. L. (2004) Two new species of freshwater fish (Teleostei: Balitoridae, Osphronemidae) from southern Sarawak. In: Yong, H. S., Ng, F. S. P. &amp; Yen, E. E. L. (Eds.), Sarawak Bau Limestone Biodiversity Sarawak Museum Journal, New Series, Special Issue No. 6, LIX (No. 80), pp. 267 - 284." type="book chapter" year="2004">Tan &amp; Ng, 2004</bibRefCitation>
;
<bibRefCitation id="EFBE2E8DFFF5FFF8D9364941FC0C1976" author="Tan, H. H. &amp; Ng, P. K. L." box="[869,928,1052,1079]" pageId="4" pageNumber="191" pagination="43 - 99" refId="ref4540" refString="Tan, H. H. &amp; Ng, P. K. L. (2005) The fighting fishes (Teleostei: Osphronemidae: genus Betta) of Singapore, Malaysia and Brunei. Raffles Bulletin of Zoology, Supplement 13, 43 - 99." type="journal article" year="2005">2005</bibRefCitation>
;
<bibRefCitation id="EFBE2E8DFFF5FFF8D9F84940FC441979" author="Tan, H. H. &amp; Ng, P. K. L." box="[939,1000,1053,1080]" pageId="4" pageNumber="191" pagination="97 - 114" refId="ref4588" refString="Tan, H. H. &amp; Ng, P. K. L. (2006) Six new species of Betta (Teleostei: Osphronemidae) from Borneo. Ichthyological Exploration of Freshwaters, 17 (2), 97 - 114." type="journal article" year="2006">2006</bibRefCitation>
).
</paragraph>
<caption id="DF5003F4FFF5FFF8DAC44AFDFEB31A9C" ID-DOI="http://doi.org/10.5281/zenodo.3955814" ID-Zenodo-Dep="3955814" httpUri="https://zenodo.org/record/3955814/files/figure.png" pageId="4" pageNumber="191" startId="4.[151,250,1952,1977]" targetBox="[151,1436,1160,1927]" targetPageId="4">
<paragraph id="8B90537CFFF5FFF8DAC44AFDFEB31A9C" blockId="4.[151,1436,1952,2013]" pageId="4" pageNumber="191">
<emphasis id="B95B8F6EFFF5FFF8DAC44AFDFEBF1AF8" bold="true" box="[151,275,1952,1977]" pageId="4" pageNumber="191">FIGURE 3.</emphasis>
<taxonomicName id="4C2F28FFFFF5FFF8DB4A4AFFFE0B1AF8" authority="Kamal &amp; Tan &amp; Ng, 2020" authorityName="Kamal &amp; Tan &amp; Ng" authorityYear="2020" box="[281,423,1953,1977]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="191" phylum="Chordata" rank="species" species="nuluhon" status="sp. nov.">
<emphasis id="B95B8F6EFFF5FFF8DB4A4AFFFE0B1AF8" box="[281,423,1953,1977]" italics="true" pageId="4" pageNumber="191">Betta nuluhon</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2683215FFF5FFF8DBE74AFCFD9C1AF8" box="[436,560,1952,1977]" pageId="4" pageNumber="191" rank="species">new species</taxonomicNameLabel>
, ZRC 61247, holotype, 62.6 mm SL (above with white background, bottom with black background).
</paragraph>
</caption>
<paragraph id="8B90537CFFF4FFF9DA944DC4FB7A1C5E" blockId="5.[151,1437,152,539]" pageId="5" pageNumber="192">
The recent collection of
<taxonomicName id="4C2F28FFFFF4FFF9DB894DC7FDF81DF2" authorityName="Ng" authorityYear="1993" box="[474,596,153,179]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="192" phylum="Chordata" rank="species" species="chini">
<emphasis id="B95B8F6EFFF4FFF9DB894DC7FDF81DF2" box="[474,596,153,179]" italics="true" pageId="5" pageNumber="192">Betta chini</emphasis>
</taxonomicName>
(ZRC 61155; see
<figureCitation id="13144FF9FFF4FFF9D9724DC4FC271DF2" box="[801,907,152,179]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="5.[151,250,1353,1378]" captionStartId-1="6.[151,250,1124,1149]" captionTargetBox-0="[151,1436,561,1325]" captionTargetBox-1="[151,1436,220,1099]" captionTargetId-0="figure@5.[151,1436,561,1328]" captionTargetId-1="figure@6.[151,1436,181,1099]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 4. Top—Betta chini, ZRC 61155, 58.6 mm SL male; bottom—Betta nuluhon, holotype, ZRC 61247, 62.6 mm SL male; both freshly preserved." captionText-1="FIGURE 5. Betta chini, ZRC 61155, 58.6 mm SL; Sabah: Klias Forest Reserve (right side, laterally inverted) (above with white background, bottom with black background)." figureDoi-0="http://doi.org/10.5281/zenodo.3955816" figureDoi-1="http://doi.org/10.5281/zenodo.3955818" httpUri-0="https://zenodo.org/record/3955816/files/figure.png" httpUri-1="https://zenodo.org/record/3955818/files/figure.png" pageId="5" pageNumber="192">Figs. 4–5</figureCitation>
) from Klias Forest Reserve indicates that there is still a viable population. The heath and peat swamp forest habitat had been badly burnt previously, during the El- Niño fires of 1998 (
<bibRefCitation id="EFBE2E8DFFF4FFF9DB274DBCFD9B1DBA" author="Phua, M. - H. &amp; Tsyuti, S. &amp; Lee, J. S. &amp; Sasakaw, H." box="[372,567,225,251]" pageId="5" pageNumber="192" pagination="103 - 116" refId="ref4086" refString="Phua, M. - H., Tsyuti, S., Lee, J. S. &amp; Sasakaw, H. (2007) Detection of burned peat swamp forest in a hetrogenous tropical landscape: A case study of the Klias Peninsula, Sabah, Malaysia. Landscape &amp; Urban Planning, 82 (3), 103 - 116. https: // doi. org / 10.1016 / j. landurbplan. 2007.01.021" type="journal article" year="2007">
Phua
<emphasis id="B95B8F6EFFF4FFF9DBE64DBFFE4A1DBA" box="[437,486,225,251]" italics="true" pageId="5" pageNumber="192">et al</emphasis>
., 2007
</bibRefCitation>
), and there was suspicion the species may have become extinct.
<taxonomicName id="4C2F28FFFFF4FFF9DF5B4DBFFA2F1DBA" authorityName="Ng" authorityYear="1993" box="[1288,1411,225,251]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="192" phylum="Chordata" rank="species" species="chini">
<emphasis id="B95B8F6EFFF4FFF9DF5B4DBFFA2F1DBA" box="[1288,1411,225,251]" italics="true" pageId="5" pageNumber="192">Betta chini</emphasis>
</taxonomicName>
is now assessed as Endangered by the International Union for Conservation of Nature (
<bibRefCitation id="EFBE2E8DFFF4FFF9DE1F4C58FB661C5E" author="Low, B. W." box="[1100,1226,261,287]" pageId="5" pageNumber="192" pagination="2781" refId="ref3904" refString="Low, B. W. (2019) Betta chini. The IUCN Red List of Threatened Species, 2019, e. T 2781 A 91307844. https: // doi. org / 10.2305 / IUCN. UK. 2019 - 2. RLTS. T 2781 A 91307844. en" type="journal article" year="2019">Low, 2019</bibRefCitation>
).
</paragraph>
<paragraph id="8B90537CFFF4FFF9DA944C74FD091F5A" blockId="5.[151,1437,152,539]" pageId="5" pageNumber="192">
When
<taxonomicName id="4C2F28FFFFF4FFF9DB474C77FEC21C02" authorityName="Ng" authorityYear="1993" box="[276,366,297,323]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="192" phylum="Chordata" rank="species" species="chini">
<emphasis id="B95B8F6EFFF4FFF9DB474C77FEC21C02" box="[276,366,297,323]" italics="true" pageId="5" pageNumber="192">B. chini</emphasis>
</taxonomicName>
was described, the largest specimen recorded was 56.0 mm SL (ZRC 35088; see
<bibRefCitation id="EFBE2E8DFFF4FFF9DF7A4C74FF6A1C26" author="Ng, P. K. L." pageId="5" pageNumber="192" pagination="289 - 294" refId="ref4039" refString="Ng, P. K. L. (1993) On a new species of Betta (Teleostei: Belontiidae) from peat swamps in Sabah, East Malaysia, Borneo. Ichthyological Exploration of Freshwaters, 4 (4), 289 - 294." type="journal article" year="1993">Ng, 1993: 290</bibRefCitation>
). The recent series contains five specimens (out of ten) ranging from
<quantity id="4CD7FE99FFF4FFF9D9864C11FB3C1C26" box="[981,1168,332,359]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="6.25" metricValueMin="5.75" pageId="5" pageNumber="192" unit="mm" value="60.0" valueMax="62.5" valueMin="57.5">57.5 to 62.5 mm</quantity>
SL, which indicate that the present series maybe occupying a preferred micro-habitat; as the
<typeStatus id="5494EDDEFFF4FFF9D9CF4C2CFC601CCA" box="[924,972,369,395]" pageId="5" pageNumber="192">type</typeStatus>
series had been obtained from peripheral streams of the reserve. All specimens of the recently-collected series have stocky bodies, with a convex body dorsum profile (see
<figureCitation id="13144FF9FFF4FFF9DB004CE4FE3A1C92" box="[339,406,440,467]" captionStart="FIGURE 5" captionStartId="6.[151,250,1124,1149]" captionTargetBox="[151,1436,220,1099]" captionTargetId="figure@6.[151,1436,181,1099]" captionTargetPageId="6" captionText="FIGURE 5. Betta chini, ZRC 61155, 58.6 mm SL; Sabah: Klias Forest Reserve (right side, laterally inverted) (above with white background, bottom with black background)." figureDoi="http://doi.org/10.5281/zenodo.3955818" httpUri="https://zenodo.org/record/3955818/files/figure.png" pageId="5" pageNumber="192">Fig. 5</figureCitation>
), similar to the range of values of body depth and caudal peduncle depth of
<taxonomicName id="4C2F28FFFFF4FFF9DEBE4CE7FAC41C92" authorityName="Kamal &amp; Tan &amp; Ng" authorityYear="2020" box="[1261,1384,441,467]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="192" phylum="Chordata" rank="species" species="nuluhon">
<emphasis id="B95B8F6EFFF4FFF9DEBE4CE7FAC41C92" box="[1261,1384,441,467]" italics="true" pageId="5" pageNumber="192">B. nuluhon</emphasis>
</taxonomicName>
(see
<tableCitation id="C6AD66C7FFF4FFF9DAC44C80FF421CB6" box="[151,238,477,503]" captionStart="TABLE 2" captionStartId="5.[151,239,1462,1486]" captionText="TABLE 2. Comparison of percentages (% SL) of body depth and caudal peduncle depth of type series of B. nuluhon, and comparative material of B. akarensis, B. balunga and B. chini (BD [blue]: body depth at dorsal-fin origin; CPD [grey]: caudal peduncle depth; red represents mean value) (values have been rounded up or down to integer values for depiction). Range (%SL) 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32" pageId="5" pageNumber="192">Table 2</tableCitation>
). From these values, the body shapes of both
<taxonomicName id="4C2F28FFFFF4FFF9D8A24C83FCE51CB6" authorityName="Ng" authorityYear="1993" box="[753,841,477,503]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="192" phylum="Chordata" rank="species" species="chini">
<emphasis id="B95B8F6EFFF4FFF9D8A24C83FCE51CB6" box="[753,841,477,503]" italics="true" pageId="5" pageNumber="192">B. chini</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2F28FFFFF4FFF9D9D24C83FC501CB6" authorityName="Kamal &amp; Tan &amp; Ng" authorityYear="2020" box="[897,1020,477,503]" class="Actinopterygii" family="Osphronemidae" genus="Betta" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="192" phylum="Chordata" rank="species" species="nuluhon">
<emphasis id="B95B8F6EFFF4FFF9D9D24C83FC501CB6" box="[897,1020,477,503]" italics="true" pageId="5" pageNumber="192">B. nuluhon</emphasis>
</taxonomicName>
may be expected to be more similar, but they are distinctly different (see
<figureCitation id="13144FF9FFF4FFF9D87E4F5CFD341F5A" box="[557,664,512,539]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="4.[151,250,1952,1977]" captionStartId-1="5.[151,250,1353,1378]" captionStartId-2="6.[151,250,1124,1149]" captionTargetBox-0="[151,1436,1160,1927]" captionTargetBox-1="[151,1436,561,1325]" captionTargetBox-2="[151,1436,220,1099]" captionTargetId-0="figure@4.[151,1436,1101,1927]" captionTargetId-1="figure@5.[151,1436,561,1328]" captionTargetId-2="figure@6.[151,1436,181,1099]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 3. Betta nuluhon, new species, ZRC 61247, holotype, 62.6 mm SL (above with white background, bottom with black background)." captionText-1="FIGURE 4. Top—Betta chini, ZRC 61155, 58.6 mm SL male; bottom—Betta nuluhon, holotype, ZRC 61247, 62.6 mm SL male; both freshly preserved." captionText-2="FIGURE 5. Betta chini, ZRC 61155, 58.6 mm SL; Sabah: Klias Forest Reserve (right side, laterally inverted) (above with white background, bottom with black background)." figureDoi-0="http://doi.org/10.5281/zenodo.3955814" figureDoi-1="http://doi.org/10.5281/zenodo.3955816" figureDoi-2="http://doi.org/10.5281/zenodo.3955818" httpUri-0="https://zenodo.org/record/3955814/files/figure.png" httpUri-1="https://zenodo.org/record/3955816/files/figure.png" httpUri-2="https://zenodo.org/record/3955818/files/figure.png" pageId="5" pageNumber="192">Figs. 3–5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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