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<mods:title id="5459771E7A498299404A39BC90E10753">Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data</mods:title>
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<mods:namePart id="D4BE8F8978484FE51A5CFC9599E6B861">Gao, Chuan-bu</mods:namePart>
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<mods:affiliation id="435DC87CDB43167B529502DC8462D6DF">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang Road W., Guangzhou 510260, China. Unidad de Protección Vegetal. Centro de Investigación y Tecnología Agroalimentaria (CITA) e Instituto Agroalimentario de Aragón (IA 2) (CITA-Universidad de Zaragoza), Avenida Montañana, 930. 50059 Zaragoza, Spain. &amp; Buzh.</mods:affiliation>
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<mods:namePart id="FB2EEBCB90C4639EF704AA5207B6D11A">Coca-Abia, María Milagro</mods:namePart>
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<mods:title id="8A359C4D8C247FB76898B0664A957080">European Journal of Taxonomy</mods:title>
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Genus
<taxonomicName id="4C1CADD6A631062EFD45FCA8FC2AE72A" authority="Reitter, 1902" authorityName="Reitter" authorityYear="1902" box="[702,968,769,796]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A631062EFD45FCA8FCCAE72D" bold="true" box="[702,808,769,796]" italics="true" pageId="4" pageNumber="5">Miridiba</emphasis>
<bibRefCitation id="EF8DABA4A631062EFCD4FCA8FC2AE72A" author="Reitter E." box="[815,968,769,795]" pageId="4" pageNumber="5" pagination="93 - 303" refId="ref45541" refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303." type="journal article" year="1902">Reitter, 1902</bibRefCitation>
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<treatmentCitation id="0ABDF044A631062EFF46FCEAFDE1E76C" author="Reitter E." box="[189,515,835,861]" page="170" pageId="4" pageNumber="5" year="1902">
<taxonomicName id="4C1CADD6A631062EFF46FCEAFDE1E76C" authority="Reitter, 1902: 170" authorityName="Reitter" authorityPageNumber="170" authorityYear="1902" box="[189,515,835,861]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A631062EFF46FCEAFEC7E76C" box="[189,293,835,861]" italics="true" pageId="4" pageNumber="5">Miridiba</emphasis>
<bibRefCitation id="EF8DABA4A631062EFED6FCEAFDE1E76C" author="Reitter E." box="[301,515,835,861]" pageId="4" pageNumber="5" pagination="93 - 303" refId="ref45541" refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303." type="journal article" year="1902">Reitter, 1902: 170</bibRefCitation>
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.
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</paragraph>
<paragraph id="8BA3D655A631062EFF46FCCFFCA3E7B1" blockId="4.[189,833,835,1038]" box="[189,833,870,897]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="AB0CF17BA631062EFF46FCCFFCA3E7B1" box="[189,833,870,897]" pageId="4" pageNumber="5">
<treatmentCitation id="0ABDF044A631062EFF46FCCFFDDBE7B0" author="Brenske E." box="[189,569,870,897]" page="75" pageId="4" pageNumber="5" year="1894">
<taxonomicName id="4C1CADD6A631062EFF46FCCFFDDBE7B0" ID-CoL="8VRQX" authority="Brenske, 1894 a: 75" authorityName="Brenske" authorityPageNumber="75" authorityYear="1894" box="[189,569,870,897]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A631062EFF46FCCFFEADE7B1" box="[189,335,870,896]" italics="true" pageId="4" pageNumber="5">Holotrochus</emphasis>
<bibRefCitation id="EF8DABA4A631062EFEADFCCFFDDBE7B0" author="Brenske E." box="[342,569,870,897]" pageId="4" pageNumber="5" pagination="3 - 87" refId="ref43282" refString="Brenske E. 1894 a. Die Melolonthiden der palaearktischen und orientalischen Region im koniglichen naturhistorischen Museum zu Brussel. Beschreibung neuer arten und Bemerkungen zu bekannten. Memoires de la Societe entomologique de Belgique 2: 3 - 87." type="journal article" year="1894">Brenske, 1894a: 75</bibRefCitation>
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(non Erichson, 1840).
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<treatmentCitationGroup id="AB0CF17BA631062EFF46FC20FD7CE792" box="[189,670,905,932]" pageId="4" pageNumber="5">
<treatmentCitation id="0ABDF044A631062EFF46FC20FD78E792" author="Reitter E." box="[189,666,905,932]" page="173" pageId="4" pageNumber="5" year="1902">
<taxonomicName id="4C1CADD6A631062EFF46FC20FD78E792" authority="Reitter, 1902: 173" authorityName="Reitter" authorityPageNumber="173" authorityYear="1902" box="[189,666,905,932]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subGenus" subGenus="Pledina">
<emphasis id="B9680A47A631062EFF46FC20FEA4E792" box="[189,326,905,931]" italics="true" pageId="4" pageNumber="5">Holotrichia</emphasis>
(
<emphasis id="B9680A47A631062EFEACFC20FE50E792" box="[343,434,905,931]" italics="true" pageId="4" pageNumber="5">Pledina</emphasis>
)
<bibRefCitation id="EF8DABA4A631062EFE38FC20FD78E792" author="Reitter E." box="[451,666,905,931]" pageId="4" pageNumber="5" pagination="93 - 303" refId="ref45541" refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303." type="journal article" year="1902">Reitter, 1902: 173</bibRefCitation>
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.
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<paragraph id="8BA3D655A631062EFF46FC05FDE4E7F6" blockId="4.[189,833,835,1038]" box="[189,518,940,967]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="AB0CF17BA631062EFF46FC05FDE4E7F6" box="[189,518,940,967]" pageId="4" pageNumber="5">
<treatmentCitation id="0ABDF044A631062EFF46FC05FDE0E7F6" author="Lucas R." box="[189,514,940,967]" page="592" pageId="4" pageNumber="5" year="1920">
<taxonomicName id="4C1CADD6A631062EFF46FC05FDE0E7F6" authority="Lucas, 1920: 592" authorityName="Lucas" authorityPageNumber="592" authorityYear="1920" box="[189,514,940,967]" class="Insecta" family="Melolonthidae" genus="Shangaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A631062EFF46FC05FECEE7F7" box="[189,300,940,966]" italics="true" pageId="4" pageNumber="5">Shangaia</emphasis>
<bibRefCitation id="EF8DABA4A631062EFEC8FC04FDE0E7F6" author="Lucas R." box="[307,514,940,967]" pageId="4" pageNumber="5" refId="ref45049" refString="Lucas R. 1920. Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (famil., trib., subtr., sect. incl.). Pars 1. Nicolaische Verlagsbuchhandlung R. Stricker, Berlin." type="book" year="1920">Lucas, 1920: 592</bibRefCitation>
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.
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</paragraph>
<paragraph id="8BA3D655A631062EFF46FC66FDA8E7DB" blockId="4.[189,833,835,1038]" box="[189,586,975,1002]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="AB0CF17BA631062EFF46FC66FDA8E7DB" box="[189,586,975,1002]" pageId="4" pageNumber="5">
<treatmentCitation id="0ABDF044A631062EFF46FC66FDA4E7DB" author="Arrow G. J." box="[189,582,975,1002]" page="50" pageId="4" pageNumber="5" year="1948">
<taxonomicName id="4C1CADD6A631062EFF46FC66FDA4E7DB" authority="Arrow, 1948: 50" authorityName="Arrow" authorityPageNumber="50" authorityYear="1948" box="[189,582,975,1002]" class="Insecta" family="Melolonthidae" genus="Neodontocnema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A631062EFF46FC66FE9EE7D8" box="[189,380,975,1001]" italics="true" pageId="4" pageNumber="5">Neodontocnema</emphasis>
<bibRefCitation id="EF8DABA4A631062EFE7AFC79FDA4E7DB" author="Arrow G. J." box="[385,582,975,1002]" pageId="4" pageNumber="5" pagination="49 - 54" refId="ref43061" refString="Arrow G. J. 1948. Further notes on the beetle genus Lachnosterna (Col.: Melolonthinae), with descriptions of three new genera. The Proceedings of the Royal Entomological Society of London. Series B, Taxonomy 17: 49 - 54. https: // doi. org / 10.1111 / j. 1365 - 3113.1948. tb 00885. x" type="journal article" year="1948">Arrow, 1948: 50</bibRefCitation>
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.
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</paragraph>
<paragraph id="8BA3D655A631062EFF46FC5AFDC0E03F" blockId="4.[189,833,835,1038]" box="[189,546,1011,1038]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="AB0CF17BA631062EFF46FC5AFDC0E03F" box="[189,546,1011,1038]" pageId="4" pageNumber="5">
<treatmentCitation id="0ABDF044A631062EFF46FC5AFDFCE03F" author="Arrow G. J." box="[189,542,1011,1038]" page="51" pageId="4" pageNumber="5" year="1948">
<taxonomicName id="4C1CADD6A631062EFF46FC5AFDFCE03F" ID-CoL="7PKFL" authority="Arrow, 1948: 51" authorityName="Arrow" authorityPageNumber="51" authorityYear="1948" box="[189,542,1011,1038]" class="Insecta" family="Scarabaeidae" genus="Hippotrichia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A631062EFF46FC5AFEB7E03C" box="[189,341,1011,1037]" italics="true" pageId="4" pageNumber="5">Hippotrichia</emphasis>
<bibRefCitation id="EF8DABA4A631062EFEA1FC5AFDFCE03F" author="Arrow G. J." box="[346,542,1011,1038]" pageId="4" pageNumber="5" pagination="49 - 54" refId="ref43061" refString="Arrow G. J. 1948. Further notes on the beetle genus Lachnosterna (Col.: Melolonthinae), with descriptions of three new genera. The Proceedings of the Royal Entomological Society of London. Series B, Taxonomy 17: 49 - 54. https: // doi. org / 10.1111 / j. 1365 - 3113.1948. tb 00885. x" type="journal article" year="1948">Arrow, 1948: 51</bibRefCitation>
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.
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<taxonomicName id="4C1CADD6A631062EFF46FB9DFEC7E07F" authorityName="Reitter" authorityYear="1902" box="[189,293,1076,1102]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A631062EFF46FB9DFEC7E07F" box="[189,293,1076,1102]" italics="true" pageId="4" pageNumber="5">Miridiba</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABDF044A631062EFEB0FB9DFDBBE07F" author="Dalla Torre K. W. von" box="[331,601,1076,1103]" page="248" pageId="4" pageNumber="5" year="1912">
<bibRefCitation id="EF8DABA4A631062EFEB0FB9DFDBBE07F" author="Dalla Torre K. W. von" box="[331,601,1076,1103]" pageId="4" pageNumber="5" pagination="135 - 290" refId="ref43620" refString="Dalla Torre K. W. von. 1912. Pars 49. Scarabaeidae: Melolonthidae III. In: Junk W. &amp; Schenkling S. (eds) Coleopterorum Catalogus. Vol. XX: 135 - 290. W. Junk, Berlin, Germany." type="book chapter" year="1912">Dalla Torre 1912: 248</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="0ABDF044A631062EFCDFFB9CFC11E07F" author="Lucas R." box="[804,1011,1076,1103]" page="420" pageId="4" pageNumber="5" year="1920">
<bibRefCitation id="EF8DABA4A631062EFCDFFB9CFC11E07F" author="Lucas R." box="[804,1011,1076,1103]" pageId="4" pageNumber="5" refId="ref45049" refString="Lucas R. 1920. Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (famil., trib., subtr., sect. incl.). Pars 1. Nicolaische Verlagsbuchhandlung R. Stricker, Berlin." type="book" year="1920">Lucas 1920: 420</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0ABDF044A631062EFBD1FB9DFA94E07F" author="Smetana A. &amp; Kral D." box="[1066,1398,1076,1102]" page="222" pageId="4" pageNumber="5" year="2006">
<bibRefCitation id="EF8DABA4A631062EFBD1FB9DFA94E07F" author="Smetana A. &amp; Kral D." box="[1066,1398,1076,1102]" pageId="4" pageNumber="5" pagination="207 - 228" refId="ref45912" refString="Smetana A. &amp; Kral D. 2006. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea: 207 - 228. Apollo Books, Stenstrup, Denmark." type="book chapter" year="2006">Smetana &amp; Král 2006: 222</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="0ABDF044A631062EFE5CFBFEFD4BE040" author="Coca-Abia M. M." box="[423,681,1111,1138]" page="673" pageId="4" pageNumber="5" year="2008">
<bibRefCitation id="EF8DABA4A631062EFE5CFBFEFD4BE040" author="Coca-Abia M. M." box="[423,681,1111,1138]" pageId="4" pageNumber="5" pagination="673 - 685" refId="ref43499" refString="Coca-Abia M. M. 2008. Revision of the genus Miridiba Reitter, 1901 (Coleoptera, Scarabaeidae, Melolonthinae). Zoological Science 25 (6): 673 - 685. https: // doi. org / 10.2108 / zsj. 25.673" type="journal article" year="2008">Coca-Abia 2008: 673</bibRefCitation>
</treatmentCitation>
(review). —
<treatmentCitation id="0ABDF044A631062EFCB4FBFEFBD6E043" ID-DOI="http://doi.org/10.5281/zenodo.6115513" ID-Zenodo-Dep="6115513" author="Li C. - L. &amp; Yang P. - S. &amp; Wang C. - C." box="[847,1076,1111,1138]" httpUri="http://treatment.plazi.org/id/03CA87D3FFB8FF8B98808CB2FE41C842" page="522" pageId="4" pageNumber="5" year="2015">
<bibRefCitation id="EF8DABA4A631062EFCB4FBFEFBD6E043" author="Li C. - L. &amp; Yang P. - S. &amp; Wang C. - C." box="[847,1076,1111,1138]" pageId="4" pageNumber="5" pagination="521 - 536" refId="ref44907" refString="Li C. - L., Yang P. - S. &amp; Wang C. - C. 2015. A review of the genus Miridiba Reitter (Coleoptera: Scarabaeidae: Melolonthinae) of Taiwan. Zootaxa 3955 (4): 521 - 536. https: // doi. org / 10.11646 / zootaxa. 3955.4.4" type="journal article" year="2015">
Li
<emphasis id="B9680A47A631062EFC88FBF1FC48E040" box="[883,938,1111,1137]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
. 2015: 522
</bibRefCitation>
</treatmentCitation>
(redescription; in key). —
<treatmentCitation id="0ABDF044A631062EFF11FBD2FE22E0A4" author="Bezdek A." box="[234,448,1146,1173]" page="271" pageId="4" pageNumber="5" year="2016">
<bibRefCitation id="EF8DABA4A631062EFF11FBD2FE22E0A4" author="Bezdek A." box="[234,448,1146,1173]" pageId="4" pageNumber="5" pagination="249 - 280" refId="ref43171" refString="Bezdek A. 2016. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 249 - 280. Koninklijke Brill NV, Leiden, The Netherlands." type="book chapter" year="2016">Bezděk 2016: 271</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="0ABDF044A631062EFD85FBD2FC82E0A4" ID-DOI="http://doi.org/10.5281/zenodo.5971102" ID-Zenodo-Dep="5971102" author="Gao C. - B. &amp; Bai M. &amp; Fang H. &amp; Yu Z. - G." box="[638,864,1147,1173]" httpUri="http://treatment.plazi.org/id/03B587C5ED3EF34928EDFAE85A4DD931" page="12" pageId="4" pageNumber="5" year="2018">
<bibRefCitation id="EF8DABA4A631062EFD85FBD2FC82E0A4" author="Gao C. - B. &amp; Bai M. &amp; Fang H. &amp; Yu Z. - G." box="[638,864,1147,1173]" pageId="4" pageNumber="5" pagination="1 - 20" refId="ref44032" refString="Gao C. - B., Bai M., Fang H. &amp; Yu Z. - G. 2018. Four new species of the genus Miridiba Reitter (Coleoptera: Scarabaeidae: Melolonthinae) from China. Zootaxa 4527 (1): 1 - 20. https: // doi. org / 10.11646 / zootaxa. 4527.1.1" type="journal article" year="2018">
Gao
<emphasis id="B9680A47A631062EFD4CFBD2FD08E0A4" box="[695,746,1147,1173]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
. 2018: 12
</bibRefCitation>
</treatmentCitation>
(catalogue); 2019: 461 (species list; in key).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A631062EFF46FB37FD12E0CE" blockId="4.[189,1401,1076,1595]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="AB0CF17BA631062EFF46FB37FD12E0CE" pageId="4" pageNumber="5">
<taxonomicName id="4C1CADD6A631062EFF46FB37FEADE089" ID-CoL="8VRQX" authorityName="Brenske" authorityYear="1894" box="[189,335,1182,1208]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A631062EFF46FB37FEADE089" box="[189,335,1182,1208]" italics="true" pageId="4" pageNumber="5">Holotrochus</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABDF044A631062EFE8BFB37FD9BE089" author="Dalla Torre K. W. von" box="[368,633,1182,1208]" page="219" pageId="4" pageNumber="5" year="1912">
<bibRefCitation id="EF8DABA4A631062EFE8BFB37FD9BE089" author="Dalla Torre K. W. von" box="[368,633,1182,1208]" pageId="4" pageNumber="5" pagination="135 - 290" refId="ref43620" refString="Dalla Torre K. W. von. 1912. Pars 49. Scarabaeidae: Melolonthidae III. In: Junk W. &amp; Schenkling S. (eds) Coleopterorum Catalogus. Vol. XX: 135 - 290. W. Junk, Berlin, Germany." type="book chapter" year="1912">Dalla Torre 1912: 219</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="0ABDF044A631062EFCC6FB37FBE7E089" author="Lucas R." box="[829,1029,1182,1208]" page="332" pageId="4" pageNumber="5" year="1920">
<bibRefCitation id="EF8DABA4A631062EFCC6FB37FBE7E089" author="Lucas R." box="[829,1029,1182,1208]" pageId="4" pageNumber="5" refId="ref45049" refString="Lucas R. 1920. Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (famil., trib., subtr., sect. incl.). Pars 1. Nicolaische Verlagsbuchhandlung R. Stricker, Berlin." type="book" year="1920">Lucas 1920: 332</bibRefCitation>
</treatmentCitation>
(as nom. nov. for
<taxonomicName id="4C1CADD6A631062EFB1CFB37FEACE0EA" ID-CoL="8VRQX" authority="Brenske" authorityName="Brenske" authorityYear="1894" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A631062EFB1CFB37FA9BE089" box="[1255,1401,1182,1208]" italics="true" pageId="4" pageNumber="5">Holotrochus</emphasis>
Brenske
</taxonomicName>
; catalogue). —
<treatmentCitation id="0ABDF044A631062EFDE8FB68FC82E0EA" author="Smetana A. &amp; Kral D." box="[531,864,1217,1243]" page="222" pageId="4" pageNumber="5" year="2006">
<bibRefCitation id="EF8DABA4A631062EFDE8FB68FC82E0EA" author="Smetana A. &amp; Kral D." box="[531,864,1217,1243]" pageId="4" pageNumber="5" pagination="207 - 228" refId="ref45912" refString="Smetana A. &amp; Kral D. 2006. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea: 207 - 228. Apollo Books, Stenstrup, Denmark." type="book chapter" year="2006">Smetana &amp; Král 2006: 222</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="0ABDF044A631062EFBD1FB68FACDE0EA" author="Coca-Abia M. M." box="[1066,1327,1217,1243]" page="674" pageId="4" pageNumber="5" year="2008">
<bibRefCitation id="EF8DABA4A631062EFBD1FB68FACDE0EA" author="Coca-Abia M. M." box="[1066,1327,1217,1243]" pageId="4" pageNumber="5" pagination="673 - 685" refId="ref43499" refString="Coca-Abia M. M. 2008. Revision of the genus Miridiba Reitter, 1901 (Coleoptera, Scarabaeidae, Melolonthinae). Zoological Science 25 (6): 673 - 685. https: // doi. org / 10.2108 / zsj. 25.673" type="journal article" year="2008">Coca-Abia 2008: 674</bibRefCitation>
</treatmentCitation>
(
<typeStatus id="54A768F7A631062EFAB9FB68FA94E0EA" box="[1346,1398,1217,1243]" pageId="4" pageNumber="5">type</typeStatus>
material). —
<treatmentCitation id="0ABDF044A631062EFE73FB4DFDBFE0CF" author="Bezdek A." box="[392,605,1252,1278]" page="271" pageId="4" pageNumber="5" year="2016">
<bibRefCitation id="EF8DABA4A631062EFE73FB4DFDBFE0CF" author="Bezdek A." box="[392,605,1252,1278]" pageId="4" pageNumber="5" pagination="249 - 280" refId="ref43171" refString="Bezdek A. 2016. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 249 - 280. Koninklijke Brill NV, Leiden, The Netherlands." type="book chapter" year="2016">Bezděk 2016: 271</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A631062EFF46FAAEFACCE177" blockId="4.[189,1401,1076,1595]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="AB0CF17BA631062EFF46FAAEFACCE177" pageId="4" pageNumber="5">
<taxonomicName id="4C1CADD6A631062EFF46FAAEFE5CE113" authority="" authorityName="Reitter" authorityYear="1902" box="[189,446,1287,1314]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subGenus" subGenus="Pledina">
<emphasis id="B9680A47A631062EFF46FAAEFEA4E110" box="[189,326,1287,1313]" italics="true" pageId="4" pageNumber="5">Holotrichia</emphasis>
(
<emphasis id="B9680A47A631062EFEA2FAAEFE56E110" box="[345,436,1287,1313]" italics="true" pageId="4" pageNumber="5">Pledina</emphasis>
)
</taxonomicName>
–
<treatmentCitation id="0ABDF044A631062EFE1BFAAEFD0BE113" author="Dalla Torre K. W. von" box="[480,745,1287,1314]" page="200" pageId="4" pageNumber="5" year="1912">
<bibRefCitation id="EF8DABA4A631062EFE1BFAAEFD0BE113" author="Dalla Torre K. W. von" box="[480,745,1287,1314]" pageId="4" pageNumber="5" pagination="135 - 290" refId="ref43620" refString="Dalla Torre K. W. von. 1912. Pars 49. Scarabaeidae: Melolonthidae III. In: Junk W. &amp; Schenkling S. (eds) Coleopterorum Catalogus. Vol. XX: 135 - 290. W. Junk, Berlin, Germany." type="book chapter" year="1912">Dalla Torre 1912: 200</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="0ABDF044A631062EFC55FAAEFB9CE113" author="Chang Y. - W." box="[942,1150,1287,1314]" page="145" pageId="4" pageNumber="5" year="1964">
<bibRefCitation id="EF8DABA4A631062EFC55FAAEFB9CE113" author="Chang Y. - W." box="[942,1150,1287,1314]" pageId="4" pageNumber="5" pagination="139 - 152" refId="ref43461" refString="Chang Y. - W. 1964. Revision of Chinese may beetles of the genus Holotrichia Hope I. (Coleoptera, Scarabaeidae). Acta Zootaxonomica Sinica 1 (1): 139 - 152." type="journal article" year="1964">Chang 1964: 145</bibRefCitation>
</treatmentCitation>
, 148 (species list; in key). —
<bibRefCitation id="EF8DABA4A631062EFEAFFA82FE14E174" author="Nomura S." box="[340,502,1323,1349]" pageId="4" pageNumber="5" pagination="81 - 95" refId="ref45479" refString="Nomura S. 1977. On the Melolonthini of Taiwan (Coleoptera, Scarabaeidae). Toho-Gakuho 27: 85 - 109. Nonfried A. F. 1892. Verzeichniss der um Nienghali in Sudchina gesammelten Lucanoiden, Scarabaeiden, Buprestiden, und Cerambyciden, nebst Beschreibung neuer arten. Entomologische Nachrichten 18 (6): 81 - 95." type="journal article" year="1977">Nomura 1977</bibRefCitation>
(as subgenus of
<taxonomicName id="4C1CADD6A631062EFD42FA82FCC3E174" authorityName="Reitter" authorityYear="1902" box="[697,801,1323,1349]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A631062EFD42FA82FCC3E174" box="[697,801,1323,1349]" italics="true" pageId="4" pageNumber="5">Miridiba</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0ABDF044A631062EFCA5FA82FB79E174" author="Smetana A. &amp; Kral D." box="[862,1179,1322,1349]" page="219" pageId="4" pageNumber="5" year="2006">
<bibRefCitation id="EF8DABA4A631062EFCA5FA82FB79E174" author="Smetana A. &amp; Kral D." box="[862,1179,1322,1349]" pageId="4" pageNumber="5" pagination="207 - 228" refId="ref45912" refString="Smetana A. &amp; Kral D. 2006. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea: 207 - 228. Apollo Books, Stenstrup, Denmark." type="book chapter" year="2006">Smetana &amp; Král 2006: 219</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A631062EFF46FAE7FE8AE19F" blockId="4.[189,1401,1076,1595]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="AB0CF17BA631062EFF46FAE7FE8AE19F" pageId="4" pageNumber="5">
<taxonomicName id="4C1CADD6A631062EFF46FAE7FE9EE159" authorityName="Arrow" authorityYear="1948" box="[189,380,1358,1384]" class="Insecta" family="Melolonthidae" genus="Neodontocnema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A631062EFF46FAE7FE9EE159" box="[189,380,1358,1384]" italics="true" pageId="4" pageNumber="5">Neodontocnema</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABDF044A631062EFE66FAE7FD8EE158" author="Chang Y. - W." box="[413,620,1358,1385]" page="145" pageId="4" pageNumber="5" year="1964">
<bibRefCitation id="EF8DABA4A631062EFE66FAE7FD8EE158" author="Chang Y. - W." box="[413,620,1358,1385]" pageId="4" pageNumber="5" pagination="139 - 152" refId="ref43461" refString="Chang Y. - W. 1964. Revision of Chinese may beetles of the genus Holotrichia Hope I. (Coleoptera, Scarabaeidae). Acta Zootaxonomica Sinica 1 (1): 139 - 152." type="journal article" year="1964">Chang 1964: 145</bibRefCitation>
</treatmentCitation>
(as synonym of
<taxonomicName id="4C1CADD6A631062EFCC1FAE7FC23E159" authorityName="Hope" authorityYear="1837" box="[826,961,1358,1384]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A631062EFCC1FAE7FC23E159" box="[826,961,1358,1384]" italics="true" pageId="4" pageNumber="5">Holotrichia</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0ABDF044A631062EFBF9FAE7FB37E159" author="Nomura S." box="[1026,1237,1358,1384]" page="88" pageId="4" pageNumber="5" year="1977">
<bibRefCitation id="EF8DABA4A631062EFBF9FAE7FB37E159" author="Nomura S." box="[1026,1237,1358,1384]" pageId="4" pageNumber="5" pagination="81 - 95" refId="ref45479" refString="Nomura S. 1977. On the Melolonthini of Taiwan (Coleoptera, Scarabaeidae). Toho-Gakuho 27: 85 - 109. Nonfried A. F. 1892. Verzeichniss der um Nienghali in Sudchina gesammelten Lucanoiden, Scarabaeiden, Buprestiden, und Cerambyciden, nebst Beschreibung neuer arten. Entomologische Nachrichten 18 (6): 81 - 95." type="journal article" year="1977">Nomura 1977: 88</bibRefCitation>
</treatmentCitation>
(as synonym of
<taxonomicName id="4C1CADD6A631062EFEF1FAD8FE90E1BA" authorityName="Reitter" authorityYear="1902" box="[266,370,1393,1419]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A631062EFEF1FAD8FE90E1BA" box="[266,370,1393,1419]" italics="true" pageId="4" pageNumber="5">Miridiba</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0ABDF044A631062EFE4BFADBFD86E1BA" author="Frey G." box="[432,612,1393,1420]" page="223" pageId="4" pageNumber="5" year="1971">
<bibRefCitation id="EF8DABA4A631062EFE4BFADBFD86E1BA" author="Frey G." box="[432,612,1393,1420]" pageId="4" pageNumber="5" pagination="206 - 225" refId="ref43999" refString="Frey G. 1971. Bestimmungstabelle der Indischen und Ceylonesischen arten der Gattung Holotrichia Hope. Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 22: 206 - 225." type="journal article" year="1971">Frey 1971: 223</bibRefCitation>
</treatmentCitation>
(in key). —
<treatmentCitation id="0ABDF044A631062EFD07FAD8FBDBE1BA" author="Smetana A. &amp; Kral D." box="[764,1081,1393,1419]" page="222" pageId="4" pageNumber="5" year="2006">
<bibRefCitation id="EF8DABA4A631062EFD07FAD8FBDBE1BA" author="Smetana A. &amp; Kral D." box="[764,1081,1393,1419]" pageId="4" pageNumber="5" pagination="207 - 228" refId="ref45912" refString="Smetana A. &amp; Kral D. 2006. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea: 207 - 228. Apollo Books, Stenstrup, Denmark." type="book chapter" year="2006">Smetana &amp; Král 2006: 222</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="0ABDF044A631062EFB02FAD8FE86E19F" author="Sabatinelli G." page="123" pageId="4" pageNumber="5" year="1983">
<bibRefCitation id="EF8DABA4A631062EFB02FAD8FE86E19F" author="Sabatinelli G." pageId="4" pageNumber="5" pagination="211 - 214" refId="ref45580" refString="Sabatinelli G. 1983. Una nuova Neodontocnema Arrow dell'Himalaya (Coleoptera, Scarabaeidae, Melolonthinae). Entomologica Basiliensia 8: 211 - 214." type="journal article" year="1983">Sabatinelli 1983: 123</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A631062EFF46FA1EFC3EE1C4" blockId="4.[189,1401,1076,1595]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="AB0CF17BA631062EFF46FA1EFC3EE1C4" pageId="4" pageNumber="5">
<taxonomicName id="4C1CADD6A631062EFF46FA1EFEB7E1E0" ID-CoL="7PKFL" authorityName="Arrow" authorityYear="1948" box="[189,341,1463,1489]" class="Insecta" family="Scarabaeidae" genus="Hippotrichia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A631062EFF46FA1EFEB7E1E0" box="[189,341,1463,1489]" italics="true" pageId="4" pageNumber="5">Hippotrichia</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABDF044A631062EFE8CFA1EFDA4E1E3" author="Chang Y. - W." box="[375,582,1463,1490]" page="145" pageId="4" pageNumber="5" year="1964">
<bibRefCitation id="EF8DABA4A631062EFE8CFA1EFDA4E1E3" author="Chang Y. - W." box="[375,582,1463,1490]" pageId="4" pageNumber="5" pagination="139 - 152" refId="ref43461" refString="Chang Y. - W. 1964. Revision of Chinese may beetles of the genus Holotrichia Hope I. (Coleoptera, Scarabaeidae). Acta Zootaxonomica Sinica 1 (1): 139 - 152." type="journal article" year="1964">Chang 1964: 145</bibRefCitation>
</treatmentCitation>
(as synonym of
<taxonomicName id="4C1CADD6A631062EFCEFFA1EFC7FE1E0" authorityName="Hope" authorityYear="1837" box="[788,925,1463,1489]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A631062EFCEFFA1EFC7FE1E0" box="[788,925,1463,1489]" italics="true" pageId="4" pageNumber="5">Holotrichia</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0ABDF044A631062EFC24FA11FB50E1E3" author="Nomura S." box="[991,1202,1464,1490]" page="88" pageId="4" pageNumber="5" year="1977">
<bibRefCitation id="EF8DABA4A631062EFC24FA11FB50E1E3" author="Nomura S." box="[991,1202,1464,1490]" pageId="4" pageNumber="5" pagination="81 - 95" refId="ref45479" refString="Nomura S. 1977. On the Melolonthini of Taiwan (Coleoptera, Scarabaeidae). Toho-Gakuho 27: 85 - 109. Nonfried A. F. 1892. Verzeichniss der um Nienghali in Sudchina gesammelten Lucanoiden, Scarabaeiden, Buprestiden, und Cerambyciden, nebst Beschreibung neuer arten. Entomologische Nachrichten 18 (6): 81 - 95." type="journal article" year="1977">Nomura 1977: 88</bibRefCitation>
</treatmentCitation>
(as synonym of
<taxonomicName id="4C1CADD6A631062EFF11FA72FEB0E1C4" authorityName="Reitter" authorityYear="1902" box="[234,338,1499,1525]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A631062EFF11FA72FEB0E1C4" box="[234,338,1499,1525]" italics="true" pageId="4" pageNumber="5">Miridiba</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0ABDF044A631062EFE6BFA72FD32E1C4" author="Smetana A. &amp; Kral D." box="[400,720,1498,1525]" page="222" pageId="4" pageNumber="5" year="2006">
<bibRefCitation id="EF8DABA4A631062EFE6BFA72FD32E1C4" author="Smetana A. &amp; Kral D." box="[400,720,1498,1525]" pageId="4" pageNumber="5" pagination="207 - 228" refId="ref45912" refString="Smetana A. &amp; Kral D. 2006. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea: 207 - 228. Apollo Books, Stenstrup, Denmark." type="book chapter" year="2006">Smetana &amp; Král 2006: 222</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0ABDF044A631062EFD04FA72FC3AE1C4" author="Bezdek A." box="[767,984,1498,1525]" page="271" pageId="4" pageNumber="5" year="2016">
<bibRefCitation id="EF8DABA4A631062EFD04FA72FC3AE1C4" author="Bezdek A." box="[767,984,1498,1525]" pageId="4" pageNumber="5" pagination="249 - 280" refId="ref43171" refString="Bezdek A. 2016. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 249 - 280. Koninklijke Brill NV, Leiden, The Netherlands." type="book chapter" year="2016">Bezděk 2016: 271</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A631062EFF46FA57FE25E20A" blockId="4.[189,1401,1076,1595]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="AB0CF17BA631062EFF46FA57FE25E20A" pageId="4" pageNumber="5">
<taxonomicName id="4C1CADD6A631062EFF46FA57FECEE229" authorityName="Lucas" authorityYear="1920" box="[189,300,1534,1560]" class="Insecta" family="Melolonthidae" genus="Shangaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A631062EFF46FA57FECEE229" box="[189,300,1534,1560]" italics="true" pageId="4" pageNumber="5">Shangaia</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABDF044A631062EFEAAFA57FE0BE228" author="Itoh T." box="[337,489,1534,1561]" page="5" pageId="4" pageNumber="5" year="1990">
<bibRefCitation id="EF8DABA4A631062EFEAAFA57FE0BE228" author="Itoh T." box="[337,489,1534,1561]" pageId="4" pageNumber="5" pagination="5 - 6" refId="ref44413" refString="Itoh T. 1990. On Miridiba trichophora (Fairmaire, 1891). Lamellicornia, Tokyo 6: 5 - 6." type="journal article" year="1990">Itoh 1990: 5</bibRefCitation>
</treatmentCitation>
(as synonym of
<taxonomicName id="4C1CADD6A631062EFD46FA57FCC7E229" authorityName="Reitter" authorityYear="1902" box="[701,805,1534,1560]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A631062EFD46FA57FCC7E229" box="[701,805,1534,1560]" italics="true" pageId="4" pageNumber="5">Miridiba</emphasis>
</taxonomicName>
). —
<treatmentCitation id="0ABDF044A631062EFC91FA57FB55E229" author="Smetana A. &amp; Kral D." box="[874,1207,1534,1560]" page="222" pageId="4" pageNumber="5" year="2006">
<bibRefCitation id="EF8DABA4A631062EFC91FA57FB55E229" author="Smetana A. &amp; Kral D." box="[874,1207,1534,1560]" pageId="4" pageNumber="5" pagination="207 - 228" refId="ref45912" refString="Smetana A. &amp; Kral D. 2006. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea: 207 - 228. Apollo Books, Stenstrup, Denmark." type="book chapter" year="2006">Smetana &amp; Král 2006: 222</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="0ABDF044A631062EFF11F988FE21E20A" author="Bezdek A." box="[234,451,1569,1595]" page="271" pageId="4" pageNumber="5" year="2016">
<bibRefCitation id="EF8DABA4A631062EFF11F988FE21E20A" author="Bezdek A." box="[234,451,1569,1595]" pageId="4" pageNumber="5" pagination="249 - 280" refId="ref43171" refString="Bezdek A. 2016. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 249 - 280. Koninklijke Brill NV, Leiden, The Netherlands." type="book chapter" year="2016">Bezděk 2016: 271</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA631062EFF46F9CAFB9FE29A" pageId="4" pageNumber="5" type="type_taxon">
<paragraph id="8BA3D655A631062EFF46F9CAFEBAE24D" blockId="4.[189,344,1634,1661]" box="[189,344,1634,1661]" pageId="4" pageNumber="5">
<heading id="D0EB6139A631062EFF46F9CAFEBAE24D" bold="true" box="[189,344,1634,1661]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B9680A47A631062EFF46F9CAFEBAE24D" bold="true" box="[189,344,1634,1661]" pageId="4" pageNumber="5">
<typeStatus id="54A768F7A631062EFF46F9CAFF18E24C" box="[189,250,1635,1661]" pageId="4" pageNumber="5">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A631062EFF46F938FB9FE29A" blockId="4.[189,1149,1681,1707]" box="[189,1149,1681,1707]" pageId="4" pageNumber="5">
<taxonomicName id="4C1CADD6A631062EFF46F938FD4CE29A" authority="Fairmaire, 1891" authorityName="Fairmaire" authorityYear="1891" box="[189,686,1681,1707]" class="Insecta" family="Melolonthidae" genus="Rhizotrogus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="trichophorus">
<emphasis id="B9680A47A631062EFF46F938FE0BE29A" box="[189,489,1681,1707]" italics="true" pageId="4" pageNumber="5">Rhizotrogus trichophorus</emphasis>
<bibRefCitation id="EF8DABA4A631062EFE0BF938FD4CE29A" author="Fairmaire L." box="[496,686,1681,1707]" pageId="4" pageNumber="5" pagination="187 - 219" refId="ref43912" refString="Fairmaire L. 1891. Coleopteres de l'interieur de la Chine (Suite: 7 e partie). Bulletin ou Compte Rendus des Seances de la Societe entomologique de Belgique 35: 187 - 219." type="journal article" year="1891">Fairmaire, 1891</bibRefCitation>
</taxonomicName>
, by original designation (
<bibRefCitation id="EF8DABA4A631062EFC23F938FB92E29A" author="Reitter E." box="[984,1136,1681,1707]" pageId="4" pageNumber="5" pagination="93 - 303" refId="ref45541" refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303." type="journal article" year="1902">Reitter 1902</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA631062FFF46F97BFDE9E628" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BA3D655A631062EFF46F97BFED4E2DD" blockId="4.[189,310,1746,1772]" box="[189,310,1746,1772]" pageId="4" pageNumber="5">
<heading id="D0EB6139A631062EFF46F97BFED4E2DD" bold="true" box="[189,310,1746,1772]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B9680A47A631062EFF46F97BFED4E2DD" bold="true" box="[189,310,1746,1772]" pageId="4" pageNumber="5">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A631062FFF46F8A8FDE9E628" blockId="4.[189,1401,1793,1996]" lastBlockId="5.[188,1400,265,537]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
Species of
<taxonomicName id="4C1CADD6A631062EFEB1F8A8FE51E32A" authorityName="Reitter" authorityYear="1902" box="[330,435,1793,1819]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A631062EFEB1F8A8FE51E32A" box="[330,435,1793,1819]" italics="true" pageId="4" pageNumber="5">Miridiba</emphasis>
</taxonomicName>
are distinguished from other melolonthines by the following combination of features. Head densely punctate. Antennal club in both sexes shorter than stem.
<collectingRegion id="49D818B7A631062EFB7CF88CFB09E30E" box="[1159,1259,1829,1855]" country="Myanmar" name="Chin" pageId="4" pageNumber="5">Mentum</collectingRegion>
with dense pubescence between its palpi. Labrum strongly depressed at middle of anterior part. Posterior frontal carina transverse and well developed. Vertex finely punctate with short decumbent pubescence. Pronotal anterior margin widely flanged, lateral margins smooth (not serrated) or serrated. Foretibia with a pubescent longitudinal carina. Apices of first four tarsomeres of fore- and mesotarsi with a tuft of setae ventrally. First ventrite under metacoxa, just its thickened posterior edge visible and not at the same level as next ventrite, with strong and short pubescence. Second ventrite densely punctate, with conspicuous and decumbent pubescence. Third and fourth ventrites with punctures and pubescence irregularly distributed, sparsely at middle and denser at sides. Fifth ventrite more or less depressed at posterior half, densely punctate and with conspicuous pubescence of different lengths. Sixth ventrite more or less concave, densely punctate and with conspicuous pubescence. Parameres with branches of variable shape. Genital chamber with reduced sternites or without them, median oviduct with wrinkled and/or hardened epithelium.
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA630062DFF46FDE9FDC2E721" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="description">
<paragraph id="8BA3D655A630062FFF46FDE9FE89E66B" blockId="5.[189,363,576,602]" box="[189,363,576,602]" pageId="5" pageNumber="6">
<heading id="D0EB6139A630062FFF46FDE9FE89E66B" bold="true" box="[189,363,576,602]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B9680A47A630062FFF46FDE9FE89E66B" bold="true" box="[189,363,576,602]" pageId="5" pageNumber="6">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A630062FFF46FDC6FB16E6B8" blockId="5.[189,1268,623,649]" box="[189,1268,623,649]" pageId="5" pageNumber="6">
External morphology of
<taxonomicName id="4C1CADD6A630062FFE25FDC6FDA4E6B8" authorityName="Reitter" authorityYear="1902" box="[478,582,623,649]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A630062FFE25FDC6FDA4E6B8" box="[478,582,623,649]" italics="true" pageId="5" pageNumber="6">Miridiba</emphasis>
</taxonomicName>
is distinguished by the following combination of features.
</paragraph>
<paragraph id="8BA3D655A630062FFF46FD18FB8CE6FA" blockId="5.[189,1134,689,715]" box="[189,1134,689,715]" pageId="5" pageNumber="6">COLORATION AND BODY SHAPE. Brown, body strong, oblong; length 11.0–26.0 mm.</paragraph>
<paragraph id="8BA3D655A630062FFF46FD5AFE7AE032" blockId="5.[189,1401,754,1027]" pageId="5" pageNumber="6">
HEAD. Antenna 9, or 10 segmented, antennal club with 3 lamellae shorter than stem (funiculus and scape). Mouthparts: mentum with dense pubescence between palpi, with a notch at middle of anterior edge; galea of maxilla with well-developed teeth; mandible with strong wrinkled molar lobe; edge of incisor lobe depressed in dorsal view; labrum strongly depressed at middle (
<bibRefCitation id="EF8DABA4A630062FFBC3FCF5FB1DE747" author="Coca-Abia M. M." box="[1080,1279,860,886]" pageId="5" pageNumber="6" pagination="673 - 685" refId="ref43499" refString="Coca-Abia M. M. 2008. Revision of the genus Miridiba Reitter, 1901 (Coleoptera, Scarabaeidae, Melolonthinae). Zoological Science 25 (6): 673 - 685. https: // doi. org / 10.2108 / zsj. 25.673" type="journal article" year="2008">Coca-Abia 2008</bibRefCitation>
: figs 1–5). Clypeus densely punctate, shorter and wider than frons, with anterior edge emarginate to a greater or lesser extent or depressed at middle, arcuate or oblique laterally. Ocular canthus pubescent, flat, short (
<quantity id="4CE47BB0A630062FFF3EFC6CFECAE7D1" box="[197,296,965,992]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="mm" value="0.5">0.5 mm</quantity>
), start out below the clypeus. Frons densely punctate with frontal carina. Vertex with short conspicuous setae.
</paragraph>
<paragraph id="8BA3D655A630062FFF46FB82FB43E09F" blockId="5.[189,1399,1066,1198]" pageId="5" pageNumber="6">PRONOTUM. Widest at posterior half; pronotal surface more or less densely punctate; anterior margin widely flanged, narrowing towards lateral sides; posterior margin not flanged at least at middle; lateral margins smooth (not serrated) or serrated to a greater or lesser extent; anterior angles obtuse or acute, more or less projected; posterior angles obtuse. Prosternal process varying in shapes.</paragraph>
<paragraph id="8BA3D655A630062FFF46FB7CFC14E0DE" blockId="5.[189,1014,1237,1264]" box="[189,1014,1237,1264]" pageId="5" pageNumber="6">SCUTELLUM. With punctures more or less densely distributed, glabrous.</paragraph>
<paragraph id="8BA3D655A630062FFF46FABEFE59E146" blockId="5.[189,1399,1302,1399]" pageId="5" pageNumber="6">ELYTRA. Elytral surface without strong striations, at most with weak ones; sutural costa a little more developed than the other ones; punctures less defined than those of pronotum; epipleuron with or without marginal pubescence.</paragraph>
<paragraph id="8BA3D655A630062FFF46FA36FD59E2E3" blockId="5.[188,1401,1438,1746]" pageId="5" pageNumber="6">LEGS. Dorsal surface of foretibia with longitudinal carina developed to a greater or lesser extent and with a longitudinal row of pubescent punctures; three outer strong teeth spaced equally; insertion of inner spur between second and third outer tooth. Meso- and metatibia with transverse carina on outer surface complete or interrupted at middle; dorsal surface with or without spines. Metatibial plates with two articulated free spurs, proximal one behind tarsomere 1 and longer than distal one; the distal spur below tarsomere 1. Fore- and mesotarsi with a tuft of setae on apex underside of tarsomeres 1–4. Metafemora thick, surface with a row of coarse punctures near posterior margin with long and thick pubescence. Hind coxal plate rectangular shaped, with external free vertex forming a right angle. Tarsal claws sickleshaped with one tooth developed at medial.</paragraph>
<paragraph id="8BA3D655A630062CFF46F953FC5FE577" blockId="5.[187,1399,1785,1988]" lastBlockId="6.[189,1399,265,327]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">ABDOMEN. Pygidium with surface punctate and pubescent or glabrous. Ventrite 1 almost completely under metacoxa, just its thickened posterior edge visible, not at the same level as next ventrite, with short pubescence more or less dense. Ventrite 2 densely punctate and with conspicuous, short and decumbent pubescence. Ventrites 3 and 4 with punctures and pubescence irregularly distributed, punctures denser and deeper at sides together with short and sparse pubescence. Ventrite 5 moderately depressed at posterior half, densely punctate and with conspicuous pubescence in different lengths. Ventrite 6 (anal plate) moderately concave, densely punctate with conspicuous pubescence, articulating by a complete suture. Ventrites 2–4 with the suture almost obliterated at middle.</paragraph>
<paragraph id="8BA3D655A633062CFF46FEC7FE53E7A9" blockId="6.[188,1401,366,920]" pageId="6" pageNumber="7">
MALE GENITALIA. Consisting of five parts: 1) Genital segment, derived from abdominal segment 9, resting on ventral wall of abdominal segments 7 and 8 (
<bibRefCitation id="EF8DABA4A633062CFCA3FE38FC3CE59D" author="Krell F. - T." box="[856,990,401,428]" pageId="6" pageNumber="7" pagination="1 - 101" refId="ref44858" refString="Krell F. - T. 1996. Die Kopulationsorgane des maikafers Melolontha melolontha (Insecta: Coleoptera: Scarabaeidae). - Ein Beitrag zur vergleichenden und funktionellen Anatomie der ektodermalen Genitalien der Coleoptera. Stuttgarter Beitrage zur Naturkunde, Serie A 537: 1 - 101." type="journal article" year="1996">Krell 1996</bibRefCitation>
), consisting of a Y-shaped sclerite (spiculum gastrale), a vestigial sternite and a vestigial tergite, which acting as an anchor for aedeagus to abdominal wall through a connective membrane. 2) Tegmen derived from abdominal segment 10 (
<bibRefCitation id="EF8DABA4A633062CFF3CFE53FEABE624" author="Krell F. - T." box="[199,329,506,533]" pageId="6" pageNumber="7" pagination="1 - 101" refId="ref44858" refString="Krell F. - T. 1996. Die Kopulationsorgane des maikafers Melolontha melolontha (Insecta: Coleoptera: Scarabaeidae). - Ein Beitrag zur vergleichenden und funktionellen Anatomie der ektodermalen Genitalien der Coleoptera. Stuttgarter Beitrage zur Naturkunde, Serie A 537: 1 - 101." type="journal article" year="1996">Krell 1996</bibRefCitation>
) consisting of parameres and phallobase (basal piece), both pieces articulated to each other. Parameres distinctively varying in shapes of dorsal and ventral branches among
<taxonomicName id="4C1CADD6A633062CFB5FFDB7FA93E609" box="[1188,1393,542,568]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9680A47A633062CFB5FFDB7FAEFE609" box="[1188,1293,542,568]" italics="true" pageId="6" pageNumber="7">Miridiba</emphasis>
species
</taxonomicName>
, apices of branches delimitating apical ostium (caudal); phallobase, a ventrally open structure partly obliterated by connective membrane, without sclerotized plates and with a gently dorsal strangulation as attachment point for connective membrane (
<bibRefCitation id="EF8DABA4A633062CFD16FD2EFC96E693" author="Krell F. - T." box="[749,884,647,674]" pageId="6" pageNumber="7" pagination="1 - 101" refId="ref44858" refString="Krell F. - T. 1996. Die Kopulationsorgane des maikafers Melolontha melolontha (Insecta: Coleoptera: Scarabaeidae). - Ein Beitrag zur vergleichenden und funktionellen Anatomie der ektodermalen Genitalien der Coleoptera. Stuttgarter Beitrage zur Naturkunde, Serie A 537: 1 - 101." type="journal article" year="1996">Krell 1996</bibRefCitation>
). 3) Endophallus (internal sac of aedeagus) lying within tegmen anchored through connective membrane and temones, evaginated during copulation through apical ostium. Epithelium covered with soft sensillae, occasionally, with areas termed raspulae, with dense sensillae and with spines of different sizes and shapes probably acting as a mechanism to anchor endophallus inside female genital chamber during copulation (
<bibRefCitation id="EF8DABA4A633062CFB2DFCBDFE92E763" author="D'Hotman D. &amp; Scholtz C. H." pageId="6" pageNumber="7" pagination="1 - 51" refId="ref43703" refString="D'Hotman D. &amp; Scholtz C. H. 1990 b. Phylogenetic significance of the structure of the external male genitalia in the Scarabaeoidea (Coleoptera). Entomology Memoir - Department of Agricultural Development 77: 1 - 51." type="journal article" year="1990">D’Hotman &amp; Scholtz 1990b</bibRefCitation>
). 4) Temones, two dorsal elongate apophysis extending into basal piece; distal ends surrounding endophallus partially or completely. 5) Vesicula seminalis or caudal diverticulum, serving as a sperm reservoir.
</paragraph>
<paragraph id="8BA3D655A633062DFF46FC69FDC2E721" blockId="6.[188,1400,959,1232]" lastBlockId="7.[188,1400,265,784]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
FEMALE GENITALIA. Tubular
<typeStatus id="54A768F7A633062CFDE8FC69FDA4E7EB" box="[531,582,960,986]" pageId="6" pageNumber="7">type</typeStatus>
(
<bibRefCitation id="EF8DABA4A633062CFDA6FC16FC40E7E8" author="Lindroth C. H. &amp; Palmen E." box="[605,930,959,985]" pageId="6" pageNumber="7" pagination="80 - 88" refId="ref44968" refString="Lindroth C. H. &amp; Palmen E. 1970. Coleoptera. In: Tuxen S. L. (ed.) Taxonomist's Glossary of Genitalia in Insects. Second revised and enlarged Edition: 80 - 88. Munksgarrd, Copenhagen." type="book chapter" year="1970">Lindroth &amp; Palmén 1970</bibRefCitation>
), usually membranous, with parts of structures sclerotized, consisting of five parts: 1) Genital chamber at posterior extension of oviduct behind sternum 8, symmetrical bilaterally, opens through vaginal ostium or vulva (
<bibRefCitation id="EF8DABA4A633062CFB5FFBACFA8AE011" author="Snodgrass R. E." box="[1188,1384,1029,1056]" pageId="6" pageNumber="7" refId="ref45968" refString="Snodgrass R. E. 1935. Principles of Insect Morphology. McGraw-Hill Book Company, New York." type="book" year="1935">Snodgrass 1935</bibRefCitation>
); dorsal wall with two folds (anal and gonopore folds), allowing its extension and retraction (
<figureCitation id="1327CAD0A633062CFB11FB80FACCE072" box="[1258,1326,1065,1091]" captionStart="Fig" captionStartId="6.[188,231,1958,1984]" captionTargetBox="[319,1226,1314,1918]" captionTargetId="figure-381@6.[319,1226,1314,1918]" captionTargetPageId="6" captionText="Fig. 1. Outline of the female genitalia of Miridiba Reitter, 1902 in lateral section. Abbreviations: see Material and methods throughout the figure captions." figureDoi="http://doi.org/10.5281/zenodo.4770295" httpUri="https://zenodo.org/record/4770295/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
); anal fold at posterior position with two pairs of vestigial tergites, one pair at each side of rectum; gonopore fold at anterior position above opening of median oviduct (gonopore) and below of anal fold (
<figureCitation id="1327CAD0A633062CFAE6FBC6FA81E0B8" box="[1309,1379,1135,1161]" captionStart="Fig" captionStartId="6.[188,231,1958,1984]" captionTargetBox="[319,1226,1314,1918]" captionTargetId="figure-381@6.[319,1226,1314,1918]" captionTargetPageId="6" captionText="Fig. 1. Outline of the female genitalia of Miridiba Reitter, 1902 in lateral section. Abbreviations: see Material and methods throughout the figure captions." figureDoi="http://doi.org/10.5281/zenodo.4770295" httpUri="https://zenodo.org/record/4770295/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
); ventral wall of gonopore fold with sensory structures (genital palps or styli) or plates located behind gonopore (
<figureCitation id="1327CAD0A633062CFEC5FB1CFE60E0E1" box="[318,386,1205,1232]" captionStart="Fig" captionStartId="6.[188,231,1958,1984]" captionTargetBox="[319,1226,1314,1918]" captionTargetId="figure-381@6.[319,1226,1314,1918]" captionTargetPageId="6" captionText="Fig. 1. Outline of the female genitalia of Miridiba Reitter, 1902 in lateral section. Abbreviations: see Material and methods throughout the figure captions." figureDoi="http://doi.org/10.5281/zenodo.4770295" httpUri="https://zenodo.org/record/4770295/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
); ventral anterior end of genital chamber without or with one pair of vestigial sternites weakly developed, sometimes with sensory setae. 2) Accessory glands, on each side of anterior end of genital chamber with a dark area; probably produce pheromone for sexual attraction (
<bibRefCitation id="EF8DABA4A632062DFB15FE85FF1DE558" author="Zamotailov A. S." pageId="7" pageNumber="8" pagination="61 - 64" refId="ref46108" refString="Zamotailov A. S. 1988. On sex attractants and sexual behavior in scarabaeid beetles (Coleoptera: Scarabaeidae). Vestnik Zoologii 6: 61 - 64." type="journal article" year="1988">Zamotailov 1988</bibRefCitation>
). 3) Oviducts lead gonads from ovaries and opens ventrally to median oviduct, which is a wide chamber with wrinkled and/or hardened epithelium sclerotized to a greater or lesser extent, median oviduct is opened to genital chamber through gonopore (
<figureCitation id="1327CAD0A632062DFCAEFE3FFC7AE581" box="[853,920,406,432]" captionStart="Fig" captionStartId="6.[188,231,1958,1984]" captionTargetBox="[319,1226,1314,1918]" captionTargetId="figure-381@6.[319,1226,1314,1918]" captionTargetPageId="6" captionText="Fig. 1. Outline of the female genitalia of Miridiba Reitter, 1902 in lateral section. Abbreviations: see Material and methods throughout the figure captions." figureDoi="http://doi.org/10.5281/zenodo.4770295" httpUri="https://zenodo.org/record/4770295/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
) locating beneath genital chamber when the female genitalia rest. 4) Bursa copulatrix, diverticulum with function of storage and enzymatic digestion of spermatophore to deliver sperm (
<bibRefCitation id="EF8DABA4A632062DFD0BFE75FC9BE5C6" author="Krell F. - T." box="[752,889,476,503]" pageId="7" pageNumber="8" pagination="1 - 101" refId="ref44858" refString="Krell F. - T. 1996. Die Kopulationsorgane des maikafers Melolontha melolontha (Insecta: Coleoptera: Scarabaeidae). - Ein Beitrag zur vergleichenden und funktionellen Anatomie der ektodermalen Genitalien der Coleoptera. Stuttgarter Beitrage zur Naturkunde, Serie A 537: 1 - 101." type="journal article" year="1996">Krell 1996</bibRefCitation>
), pedunculated, opens dorsally to median oviduct, with a pouch at proximal position with plicate epithelium and small setae of pressosensorial function (
<bibRefCitation id="EF8DABA4A632062DFECDFD8AFD25E60C" author="Sanmartin I. &amp; Martin-Piera F." box="[310,711,546,573]" pageId="7" pageNumber="8" pagination="2 - 46" refId="ref45647" refString="Sanmartin I. &amp; Martin-Piera F. 2003. First phylogenetic analysis of the subfamily Pachydeminae (Coleoptera, Scarabaeoidea, Melolonthidae): the Palearctic Pachydeminae. Journal of Zoological Systematics and Evolutionary Research 41: 2 - 46. https: // doi. org / 10.1046 / j. 1439 - 0469.2003.00179. x" type="journal article" year="2003">Sanmartín &amp; Martín-Piera 2003</bibRefCitation>
), some species with sclerotized structures in peduncle. 5) Spermatheca (receptaculum seminis or seminal receptacle) with function of sperm storage after releasing from bursa copulatrix, pedunculated with a pouch at proximal position, opens to median oviduct ventrally. Spermathecal gland, longer than spermatheca, opens at spermathecal tract just below pouch, its secretion leads sperm chemotactically through a concentration gradient from spermatophore to spermatheca (
<bibRefCitation id="EF8DABA4A632062DFE86FD7BFDE1E6DC" author="Krell F. - T." box="[381,515,722,749]" pageId="7" pageNumber="8" pagination="1 - 101" refId="ref44858" refString="Krell F. - T. 1996. Die Kopulationsorgane des maikafers Melolontha melolontha (Insecta: Coleoptera: Scarabaeidae). - Ein Beitrag zur vergleichenden und funktionellen Anatomie der ektodermalen Genitalien der Coleoptera. Stuttgarter Beitrage zur Naturkunde, Serie A 537: 1 - 101." type="journal article" year="1996">Krell 1996</bibRefCitation>
). Both spermathecal and its gland have a distal tubular duct and a proximal caecum more or less globular.
</paragraph>
</subSubSection>
<caption id="DF6386DDA633062CFF47F80FFCCDE3D2" ID-DOI="http://doi.org/10.5281/zenodo.4770295" ID-Zenodo-Dep="4770295" httpUri="https://zenodo.org/record/4770295/files/figure.png" pageId="6" pageNumber="7" startId="6.[188,231,1958,1984]" targetBox="[319,1226,1314,1918]" targetPageId="6">
<paragraph id="8BA3D655A633062CFF47F80FFCCDE3D2" blockId="6.[188,1398,1958,2019]" pageId="6" pageNumber="7">
<emphasis id="B9680A47A633062CFF47F80FFEEFE3F1" bold="true" box="[188,269,1958,1984]" pageId="6" pageNumber="7">Fig. 1.</emphasis>
Outline of the female genitalia of
<taxonomicName id="4C1CADD6A633062CFD4AF80FFC59E3F1" authority="Reitter, 1902" authorityName="Reitter" authorityYear="1902" box="[689,955,1958,1984]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A633062CFD4AF80FFCFBE3F1" box="[689,793,1958,1984]" italics="true" pageId="6" pageNumber="7">Miridiba</emphasis>
<bibRefCitation id="EF8DABA4A633062CFCDAF80FFC59E3F1" author="Reitter E." box="[801,955,1958,1984]" pageId="6" pageNumber="7" pagination="93 - 303" refId="ref45541" refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303." type="journal article" year="1902">Reitter, 1902</bibRefCitation>
</taxonomicName>
in lateral section. Abbreviations: see Material and methods throughout the figure captions.
</paragraph>
</caption>
<subSubSection id="C30685DEA632062DFF46FC9EFED0E7F6" pageId="7" pageNumber="8" type="distribution">
<paragraph id="8BA3D655A632062DFF46FC9EFEB4E760" blockId="7.[189,342,823,849]" box="[189,342,823,849]" pageId="7" pageNumber="8">
<heading id="D0EB6139A632062DFF46FC9EFEB4E760" bold="true" box="[189,342,823,849]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B9680A47A632062DFF46FC9EFEB4E760" bold="true" box="[189,342,823,849]" pageId="7" pageNumber="8">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A632062DFF46FCCFFED0E7F6" blockId="7.[188,1398,870,967]" pageId="7" pageNumber="8">
Species of
<taxonomicName id="4C1CADD6A632062DFEC7FCCFFE46E7B1" authorityName="Reitter" authorityYear="1902" box="[316,420,870,896]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A632062DFEC7FCCFFE46E7B1" box="[316,420,870,896]" italics="true" pageId="7" pageNumber="8">Miridiba</emphasis>
</taxonomicName>
are documented in the eastern Palaearctic Region (
<collectingCountry id="F30B96C5A632062DFC02FCCFFBA6E7B1" box="[1017,1092,870,896]" name="China" pageId="7" pageNumber="8">China</collectingCountry>
,
<collectingCountry id="F30B96C5A632062DFBB4FCCFFB77E7B1" box="[1103,1173,870,896]" name="Japan" pageId="7" pageNumber="8">Japan</collectingCountry>
, Korean Peninsula, Far East of
<collectingCountry id="F30B96C5A632062DFEBEFC20FE77E795" box="[325,405,905,932]" name="Russia" pageId="7" pageNumber="8">Russia</collectingCountry>
) and Oriental Region (
<collectingCountry id="F30B96C5A632062DFD59FC20FD01E792" box="[674,739,905,931]" name="India" pageId="7" pageNumber="8">India</collectingCountry>
,
<collectingCountry id="F30B96C5A632062DFD14FC20FC86E792" box="[751,868,905,931]" name="Indonesia" pageId="7" pageNumber="8">Indonesia</collectingCountry>
,
<collectingCountry id="F30B96C5A632062DFC8BFC20FC4EE792" box="[880,940,905,931]" name="Laos" pageId="7" pageNumber="8">Laos</collectingCountry>
,
<collectingCountry id="F30B96C5A632062DFC43FC20FBC5E792" box="[952,1063,905,931]" name="Malaysia" pageId="7" pageNumber="8">Malaysia</collectingCountry>
,
<collectingCountry id="F30B96C5A632062DFBC9FC20FB44E792" box="[1074,1190,905,931]" name="Myanmar" pageId="7" pageNumber="8">Myanmar</collectingCountry>
,
<collectingCountry id="F30B96C5A632062DFB49FC20FB19E792" box="[1202,1275,905,931]" name="Nepal" pageId="7" pageNumber="8">Nepal</collectingCountry>
,
<collectingCountry id="F30B96C5A632062DFAFDFC20FA93E792" box="[1286,1393,905,931]" name="Thailand" pageId="7" pageNumber="8">Thailand</collectingCountry>
,
<collectingCountry id="F30B96C5A632062DFF46FC05FEC4E7F6" box="[189,294,940,967]" name="Vietnam" pageId="7" pageNumber="8">Vietnam</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6320625FF46FC47FCE1E086" lastPageId="15" lastPageNumber="16" pageId="7" pageNumber="8" type="description">
<paragraph id="8BA3D655A632062DFF46FC47FE32E03A" blockId="7.[189,464,1006,1035]" box="[189,464,1006,1035]" pageId="7" pageNumber="8">
<emphasis id="B9680A47A632062DFF46FC47FE32E03A" bold="true" box="[189,464,1006,1035]" italics="true" pageId="7" pageNumber="8">Genital morphotypes</emphasis>
</paragraph>
<paragraph id="8BA3D655A632062DFF46FB8AFC00E051" blockId="7.[189,1397,1059,1121]" pageId="7" pageNumber="8">
Based on the examination of
<typeStatus id="54A768F7A632062DFDF4FB8DFDA0E00F" box="[527,578,1060,1086]" pageId="7" pageNumber="8">type</typeStatus>
series of twenty-eight species and eight synonymous taxa of
<emphasis id="B9680A47A632062DFAFDFB8AFA97E00C" box="[1286,1397,1059,1085]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C1CADD6A632062DFAFDFB8AFA93E00C" authorityName="Reitter" authorityYear="1902" box="[1286,1393,1059,1085]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Miridiba</taxonomicName>
,
</emphasis>
nine genital morphotypes were established and described as follows.
</paragraph>
<paragraph id="8BA3D655A632062DFD9FFB21FC32E090" blockId="7.[612,976,1159,1221]" box="[612,976,1159,1186]" pageId="7" pageNumber="8">
<heading id="D0EB6139A632062DFD9FFB21FC32E090" box="[612,976,1159,1186]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="8" reason="2">
<emphasis id="B9680A47A632062DFD9FFB21FC32E090" bold="true" box="[612,976,1159,1186]" pageId="7" pageNumber="8">
Morphotype I “
<taxonomicName id="4C1CADD6A632062DFCD2FB2EFC23E093" box="[809,961,1159,1186]" class="Insecta" family="Tachinidae" genus="Trichophora" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A632062DFCD2FB2EFC23E093" bold="true" box="[809,961,1159,1186]" italics="true" pageId="7" pageNumber="8">Trichophora</emphasis>
</taxonomicName>
”
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A632062DFD1CFB02FCAEE0F4" blockId="7.[612,976,1159,1221]" box="[743,844,1195,1221]" pageId="7" pageNumber="8">
<figureCitation id="1327CAD0A632062DFD1CFB02FCAEE0F4" box="[743,844,1195,1221]" captionStart="Figs 2–4" captionStartId="8.[189,241,1020,1046]" captionTargetBox="[277,1310,274,986]" captionTargetId="figure-18@8.[264,1323,262,990]" captionTargetPageId="8" captionText="Figs 2–4. Morphotype I “Trichophora”. 2–3. Male genitalia of M. trichophora (Fairmaire, 1891), paralectotype (MNHN). 2. Parameres in dorsal (A), ventral (B) and lateral (C) views. 3. Endophallus in lateral view. 4. Female genitalia of M. abdominalis (Hope, 1831), lectotype (BMNH), in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770299" httpUri="https://zenodo.org/record/4770299/files/figure.png" pageId="7" pageNumber="8">Figs 2–4</figureCitation>
</paragraph>
<paragraph id="8BA3D655A632062DFF46FB45FE92E137" blockId="7.[189,368,1260,1286]" box="[189,368,1260,1286]" pageId="7" pageNumber="8">
<heading id="D0EB6139A632062DFF46FB45FE92E137" bold="true" box="[189,368,1260,1286]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B9680A47A632062DFF46FB45FE92E137" bold="true" box="[189,368,1260,1286]" pageId="7" pageNumber="8">Male genitalia</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A632062DFF46FAB2FAC2E239" blockId="7.[187,1400,1307,1545]" pageId="7" pageNumber="8">
Parameres (
<figureCitation id="1327CAD0A632062DFEBDFAB2FE69E104" box="[326,395,1307,1333]" captionStart="Figs 2–4" captionStartId="8.[189,241,1020,1046]" captionTargetBox="[277,1310,274,986]" captionTargetId="figure-18@8.[264,1323,262,990]" captionTargetPageId="8" captionText="Figs 2–4. Morphotype I “Trichophora”. 2–3. Male genitalia of M. trichophora (Fairmaire, 1891), paralectotype (MNHN). 2. Parameres in dorsal (A), ventral (B) and lateral (C) views. 3. Endophallus in lateral view. 4. Female genitalia of M. abdominalis (Hope, 1831), lectotype (BMNH), in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770299" httpUri="https://zenodo.org/record/4770299/files/figure.png" pageId="7" pageNumber="8">Fig. 2</figureCitation>
) consisting of two dorsal and two ventral thin branches separated in dorsal, ventral and lateral views; dorsal branches longer than ventral ones; branches start from a collar-shaped structure, termed
<emphasis id="B9680A47A632062DFEECFAC8FE85E14A" box="[279,359,1377,1403]" italics="true" pageId="7" pageNumber="8">collum</emphasis>
at proximal part of parameres close to phallobase; collum short, with a suture dorsally and ventrally sclerotized to a greater or lesser extent. Endophallus (
<figureCitation id="1327CAD0A632062DFC35FA2DFBF1E1AE" box="[974,1043,1412,1439]" captionStart="Figs 2–4" captionStartId="8.[189,241,1020,1046]" captionTargetBox="[277,1310,274,986]" captionTargetId="figure-18@8.[264,1323,262,990]" captionTargetPageId="8" captionText="Figs 2–4. Morphotype I “Trichophora”. 2–3. Male genitalia of M. trichophora (Fairmaire, 1891), paralectotype (MNHN). 2. Parameres in dorsal (A), ventral (B) and lateral (C) views. 3. Endophallus in lateral view. 4. Female genitalia of M. abdominalis (Hope, 1831), lectotype (BMNH), in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770299" httpUri="https://zenodo.org/record/4770299/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
) slim and graceful with spines becoming larger towards distal end and in some species arranged helicoidally around internal walls of the sac, some species with large spines or raspulae in distal end. Temones (
<figureCitation id="1327CAD0A632062DFBCAFA62FB95E1D4" box="[1073,1143,1483,1509]" captionStart="Figs 2–4" captionStartId="8.[189,241,1020,1046]" captionTargetBox="[277,1310,274,986]" captionTargetId="figure-18@8.[264,1323,262,990]" captionTargetPageId="8" captionText="Figs 2–4. Morphotype I “Trichophora”. 2–3. Male genitalia of M. trichophora (Fairmaire, 1891), paralectotype (MNHN). 2. Parameres in dorsal (A), ventral (B) and lateral (C) views. 3. Endophallus in lateral view. 4. Female genitalia of M. abdominalis (Hope, 1831), lectotype (BMNH), in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770299" httpUri="https://zenodo.org/record/4770299/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
) laid at proximal half of endophallus; gently developed, seldom strongly developed, apophysis fused at proximal end.
</paragraph>
<paragraph id="8BA3D655A632062DFF46F999FE09E27B" blockId="7.[189,491,1583,1610]" box="[189,491,1583,1610]" pageId="7" pageNumber="8">
<emphasis id="B9680A47A632062DFF46F999FE6EE278" bold="true" box="[189,396,1583,1610]" pageId="7" pageNumber="8">Female genitalia</emphasis>
(
<figureCitation id="1327CAD0A632062DFE60F986FE00E27B" box="[411,482,1583,1610]" captionStart="Figs 2–4" captionStartId="8.[189,241,1020,1046]" captionTargetBox="[277,1310,274,986]" captionTargetId="figure-18@8.[264,1323,262,990]" captionTargetPageId="8" captionText="Figs 2–4. Morphotype I “Trichophora”. 2–3. Male genitalia of M. trichophora (Fairmaire, 1891), paralectotype (MNHN). 2. Parameres in dorsal (A), ventral (B) and lateral (C) views. 3. Endophallus in lateral view. 4. Female genitalia of M. abdominalis (Hope, 1831), lectotype (BMNH), in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770299" httpUri="https://zenodo.org/record/4770299/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
)
</paragraph>
<paragraph id="8BA3D655A632062DFF46F9F7FAD3E2D3" blockId="7.[188,1400,1630,1762]" pageId="7" pageNumber="8">Anal fold with one tergite of each pair distinctly reduced; gonopore fold with a pair of sensory plates in oval-transverse shaped, plates with sensilla at posterior margin; vestigial sternites absent or weakly developed with a few sensillae. Median oviduct with wrinkled and/or hardened epithelium sclerotized to a greater or lesser extent. Bursa copulatrix with striated pouch. Spermatheca with short peduncle.</paragraph>
<paragraph id="8BA3D655A632062DFD83F8A0FC5EE312" blockId="7.[632,956,1801,1863]" box="[632,956,1801,1827]" pageId="7" pageNumber="8">
<heading id="D0EB6139A632062DFD83F8A0FC5EE312" box="[632,956,1801,1827]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="8" reason="2">
<emphasis id="B9680A47A632062DFD83F8A0FC5EE312" bold="true" box="[632,956,1801,1827]" pageId="7" pageNumber="8">
Morphotype II “
<emphasis id="B9680A47A632062DFCB3F8A0FC4FE312" bold="true" box="[840,941,1801,1827]" italics="true" pageId="7" pageNumber="8">Gressitti</emphasis>
”
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A632062DFD1CF885FCAEE376" blockId="7.[632,956,1801,1863]" box="[743,844,1836,1863]" pageId="7" pageNumber="8">
<figureCitation id="1327CAD0A632062DFD1CF885FCAEE376" box="[743,844,1836,1863]" captionStart="Figs 5–7" captionStartId="8.[189,241,1914,1940]" captionTargetBox="[277,1310,1238,1870]" captionTargetId="figure-79@8.[264,1323,1229,1878]" captionTargetPageId="8" captionText="Figs 5–7. Morphotype II “Gressitti”. 5–6. Male genitalia of Miridiba herteli (Frey, 1971), paratype (NHMB). 5. Parameres in lateral view. 6. Endophallus and temones in dorsal view. 7. Female genitalia of M. gressitti (Frey, 1970), paratype (NHMB) in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770301" httpUri="https://zenodo.org/record/4770301/files/figure.png" pageId="7" pageNumber="8">Figs 5–7</figureCitation>
</paragraph>
<paragraph id="8BA3D655A632062DFF46F8C6FE92E3B8" blockId="7.[189,368,1903,1929]" box="[189,368,1903,1929]" pageId="7" pageNumber="8">
<heading id="D0EB6139A632062DFF46F8C6FE92E3B8" bold="true" box="[189,368,1903,1929]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B9680A47A632062DFF46F8C6FE92E3B8" bold="true" box="[189,368,1903,1929]" pageId="7" pageNumber="8">Male genitalia</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6320623FF46F836FDF1E55D" blockId="7.[189,1399,1950,2013]" lastBlockId="9.[189,1398,265,364]" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
Parameres (
<figureCitation id="1327CAD0A632062DFEB3F837FE6DE388" box="[328,399,1950,1977]" captionStart="Figs 5–7" captionStartId="8.[189,241,1914,1940]" captionTargetBox="[277,1310,1238,1870]" captionTargetId="figure-79@8.[264,1323,1229,1878]" captionTargetPageId="8" captionText="Figs 5–7. Morphotype II “Gressitti”. 5–6. Male genitalia of Miridiba herteli (Frey, 1971), paratype (NHMB). 5. Parameres in lateral view. 6. Endophallus and temones in dorsal view. 7. Female genitalia of M. gressitti (Frey, 1970), paratype (NHMB) in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770301" httpUri="https://zenodo.org/record/4770301/files/figure.png" pageId="7" pageNumber="8">Fig. 5</figureCitation>
) with a dorsal complex divided into three parts, upper part continuous and joined to collum by a membrane, intermediate and lower parts each with two branches. Two ventral branches simple and similar to those of morphotype I. Endophallus slim and raspulae conspicuous or gently at distal end, spines absent (
<figureCitation id="1327CAD0A63C0623FE03FE84FDA3E579" box="[504,577,301,328]" captionStart="Figs 5–7" captionStartId="8.[189,241,1914,1940]" captionTargetBox="[277,1310,1238,1870]" captionTargetId="figure-79@8.[264,1323,1229,1878]" captionTargetPageId="8" captionText="Figs 5–7. Morphotype II “Gressitti”. 5–6. Male genitalia of Miridiba herteli (Frey, 1971), paratype (NHMB). 5. Parameres in lateral view. 6. Endophallus and temones in dorsal view. 7. Female genitalia of M. gressitti (Frey, 1970), paratype (NHMB) in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770301" httpUri="https://zenodo.org/record/4770301/files/figure.png" pageId="9" pageNumber="10">Fig. 6</figureCitation>
). Temones laid at proximal half of endophallus, thin and apophysis separated each other (
<figureCitation id="1327CAD0A63C0623FE47FEF8FDE6E55D" box="[444,516,337,364]" captionStart="Figs 5–7" captionStartId="8.[189,241,1914,1940]" captionTargetBox="[277,1310,1238,1870]" captionTargetId="figure-79@8.[264,1323,1229,1878]" captionTargetPageId="8" captionText="Figs 5–7. Morphotype II “Gressitti”. 5–6. Male genitalia of Miridiba herteli (Frey, 1971), paratype (NHMB). 5. Parameres in lateral view. 6. Endophallus and temones in dorsal view. 7. Female genitalia of M. gressitti (Frey, 1970), paratype (NHMB) in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770301" httpUri="https://zenodo.org/record/4770301/files/figure.png" pageId="9" pageNumber="10">Fig. 6</figureCitation>
).
</paragraph>
<caption id="DF6386DDA63D0622FF46FC55FD6DE0B0" ID-DOI="http://doi.org/10.5281/zenodo.4770299" ID-Zenodo-Dep="4770299" httpUri="https://zenodo.org/record/4770299/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,241,1020,1046]" targetBox="[277,1310,274,986]" targetPageId="8">
<paragraph id="8BA3D655A63D0622FF46FC55FD6DE0B0" blockId="8.[189,1399,1020,1153]" pageId="8" pageNumber="9">
<emphasis id="B9680A47A63D0622FF46FC55FED3E027" bold="true" box="[189,305,1020,1046]" pageId="8" pageNumber="9">Figs 2–4.</emphasis>
Morphotype I “
<taxonomicName id="4C1CADD6A63D0622FDFBFC55FD71E027" box="[512,659,1020,1046]" class="Insecta" family="Tachinidae" genus="Trichophora" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A63D0622FDFBFC55FD71E027" box="[512,659,1020,1046]" italics="true" pageId="8" pageNumber="9">Trichophora</emphasis>
</taxonomicName>
”.
<emphasis id="B9680A47A63D0622FD4FFC55FD02E027" bold="true" box="[692,736,1020,1046]" pageId="8" pageNumber="9">2–3</emphasis>
. Male genitalia of
<taxonomicName id="4C1CADD6A63D0622FC2EFC54FA90E026" authority="(Fairmaire, 1891)" authorityName="CH. Bu Gao &amp; M. Coca-Abia" baseAuthorityName="Fairmaire" baseAuthorityYear="1891" box="[981,1394,1020,1047]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="trichophora">
<emphasis id="B9680A47A63D0622FC2EFC54FB6FE027" box="[981,1165,1020,1046]" italics="true" pageId="8" pageNumber="9">M. trichophora</emphasis>
(
<bibRefCitation id="EF8DABA4A63D0622FB59FC55FA8BE026" author="Fairmaire L." box="[1186,1385,1020,1047]" pageId="8" pageNumber="9" pagination="187 - 219" refId="ref43912" refString="Fairmaire L. 1891. Coleopteres de l'interieur de la Chine (Suite: 7 e partie). Bulletin ou Compte Rendus des Seances de la Societe entomologique de Belgique 35: 187 - 219." type="journal article" year="1891">Fairmaire, 1891</bibRefCitation>
)
</taxonomicName>
, paralectotype (MNHN).
<emphasis id="B9680A47A63D0622FE26FBB6FE0EE008" bold="true" box="[477,492,1055,1081]" pageId="8" pageNumber="9">2</emphasis>
. Parameres in dorsal (A), ventral (B) and lateral (C) views.
<emphasis id="B9680A47A63D0622FB52FBB6FB5AE008" bold="true" box="[1193,1208,1055,1081]" pageId="8" pageNumber="9">3</emphasis>
. Endophallus in lateral view.
<emphasis id="B9680A47A63D0622FEA8FBEAFE80E06C" bold="true" box="[339,354,1091,1117]" pageId="8" pageNumber="9">4</emphasis>
. Female genitalia of
<taxonomicName id="4C1CADD6A63D0622FDA2FBEDFC55E06C" authority="(Hope, 1831)" authorityName="CH. Bu Gao &amp; M. Coca-Abia" baseAuthorityName="Hope" baseAuthorityYear="1831" box="[601,951,1091,1117]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="B9680A47A63D0622FDA2FBEDFCF3E06C" box="[601,785,1091,1117]" italics="true" pageId="8" pageNumber="9">M. abdominalis</emphasis>
(
<bibRefCitation id="EF8DABA4A63D0622FCDBFBEAFC4CE06C" author="Hope F. W." box="[800,942,1091,1117]" pageId="8" pageNumber="9" pagination="21 - 32" refId="ref44170" refString="Hope F. W. 1831 Synopsis of the new species of Nepal insects in the collection of Major-General Hardwicke. In: Gray J. E. (ed.) The Zoological Miscellany Vol. 1: 21 - 32. Treuttel, Wurtz and Co., London. https: // doi. org / 10.5962 / bhl. title. 113722" type="journal article" year="1831">Hope, 1831</bibRefCitation>
)
</taxonomicName>
, lectotype (BMNH), in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm.
</paragraph>
</caption>
<caption id="DF6386DDA63D0622FF46F8D3FA95E3ED" ID-DOI="http://doi.org/10.5281/zenodo.4770301" ID-Zenodo-Dep="4770301" httpUri="https://zenodo.org/record/4770301/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,241,1914,1940]" targetBox="[277,1310,1238,1870]" targetPageId="8">
<paragraph id="8BA3D655A63D0622FF46F8D3FA95E3ED" blockId="8.[189,1399,1914,2012]" pageId="8" pageNumber="9">
<emphasis id="B9680A47A63D0622FF46F8D3FECDE3A4" bold="true" box="[189,303,1914,1941]" pageId="8" pageNumber="9">Figs 5–7.</emphasis>
Morphotype II “
<emphasis id="B9680A47A63D0622FDF9F8D2FD84E3A4" box="[514,614,1915,1941]" italics="true" pageId="8" pageNumber="9">Gressitti</emphasis>
”.
<emphasis id="B9680A47A63D0622FD7DF8D2FD50E3A4" bold="true" box="[646,690,1915,1941]" pageId="8" pageNumber="9">5–6</emphasis>
. Male genitalia of
<taxonomicName id="4C1CADD6A63D0622FC64F8D2FAE0E3A4" authority="(Frey, 1971)" baseAuthorityName="Frey" baseAuthorityYear="1971" box="[927,1282,1915,1941]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="herteli">
<emphasis id="B9680A47A63D0622FC64F8D2FB82E3A4" box="[927,1120,1915,1941]" italics="true" pageId="8" pageNumber="9">Miridiba herteli</emphasis>
(
<bibRefCitation id="EF8DABA4A63D0622FB88F8D2FB1BE3A4" author="Frey G." box="[1139,1273,1915,1941]" pageId="8" pageNumber="9" pagination="206 - 225" refId="ref43999" refString="Frey G. 1971. Bestimmungstabelle der Indischen und Ceylonesischen arten der Gattung Holotrichia Hope. Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 22: 206 - 225." type="journal article" year="1971">Frey, 1971</bibRefCitation>
)
</taxonomicName>
, paratype (NHMB).
<emphasis id="B9680A47A63D0622FEC3F837FEA5E389" bold="true" box="[312,327,1950,1976]" pageId="8" pageNumber="9">5</emphasis>
. Parameres in lateral view.
<emphasis id="B9680A47A63D0622FD75F837FD7EE389" bold="true" box="[654,668,1950,1976]" pageId="8" pageNumber="9">6</emphasis>
. Endophallus and temones in dorsal view.
<emphasis id="B9680A47A63D0622FB6DF837FB46E389" bold="true" box="[1174,1188,1950,1976]" pageId="8" pageNumber="9">7</emphasis>
. Female genitalia of
<taxonomicName id="4C1CADD6A63D0622FF27F86BFE1AE3EA" authority="(Frey, 1970)" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Frey" baseAuthorityYear="1970" box="[220,504,1985,2011]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="gressitti">
<emphasis id="B9680A47A63D0622FF27F86BFEBDE3EA" box="[220,351,1986,2011]" italics="true" pageId="8" pageNumber="9">M. gressitti</emphasis>
(
<bibRefCitation id="EF8DABA4A63D0622FE95F868FE12E3EA" author="Frey G." box="[366,496,1985,2011]" pageId="8" pageNumber="9" pagination="244 - 265" refId="ref43949" refString="Frey G. 1970. Bestimmungstabelle der Holotrichia - Arten Hinterindiens (Vietnam, Laos, Thailand, Kambodscha, Burma, Malaya) (Col., Scarab, Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 21: 244 - 265." type="journal article" year="1970">Frey, 1970</bibRefCitation>
)
</taxonomicName>
, paratype (NHMB) in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BA3D655A63C0623FF46FE3DFE08E59E" blockId="9.[189,490,404,431]" box="[189,490,404,431]" pageId="9" pageNumber="10">
<emphasis id="B9680A47A63C0623FF46FE3DFE6EE59F" bold="true" box="[189,396,404,430]" pageId="9" pageNumber="10">Female genitalia</emphasis>
(
<figureCitation id="1327CAD0A63C0623FE60FE3DFE03E59E" box="[411,481,404,431]" captionStart="Figs 5–7" captionStartId="8.[189,241,1914,1940]" captionTargetBox="[277,1310,1238,1870]" captionTargetId="figure-79@8.[264,1323,1229,1878]" captionTargetPageId="8" captionText="Figs 5–7. Morphotype II “Gressitti”. 5–6. Male genitalia of Miridiba herteli (Frey, 1971), paratype (NHMB). 5. Parameres in lateral view. 6. Endophallus and temones in dorsal view. 7. Female genitalia of M. gressitti (Frey, 1970), paratype (NHMB) in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770301" httpUri="https://zenodo.org/record/4770301/files/figure.png" pageId="9" pageNumber="10">Fig. 7</figureCitation>
)
</paragraph>
<paragraph id="8BA3D655A63C0623FF46FE6DFBA3E67B" blockId="9.[189,1401,452,587]" pageId="9" pageNumber="10">Anal fold with all vestigial tergites similar in size; gonopore fold with a pair of elongate plates with sensillae at posterior edge; vestigial sternites weakly developed with a few sensillae. Median oviduct with wrinkled and hardened epithelium and two rectangular plates at distal dorsal wall. Pouch of bursa copulatrix striated with punctate sensillae. Spermatheca with short peduncle.</paragraph>
<paragraph id="8BA3D655A63C0623FDBEFDDAFC0DE6BC" blockId="9.[581,1007,626,689]" box="[581,1007,626,653]" pageId="9" pageNumber="10">
<heading id="D0EB6139A63C0623FDBEFDDAFC0DE6BC" box="[581,1007,626,653]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="10" reason="2">
<emphasis id="B9680A47A63C0623FDBEFDDAFC0DE6BC" bold="true" box="[581,1007,626,653]" pageId="9" pageNumber="10">
Morphotype III “
<emphasis id="B9680A47A63C0623FCDAFDDBFC02E6BC" bold="true" box="[801,992,626,653]" italics="true" pageId="9" pageNumber="10">Leucophthalma</emphasis>
”
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A63C0623FD0CFD3EFCDEE680" blockId="9.[581,1007,626,689]" box="[759,828,663,689]" pageId="9" pageNumber="10">
<figureCitation id="1327CAD0A63C0623FD0CFD3EFCDEE680" box="[759,828,663,689]" captionStart="Fig" captionStartId="10.[189,232,720,746]" captionTargetBox="[259,1335,326,633]" captionTargetId="figure-17@10.[238,1349,305,646]" captionTargetPageId="10" captionText="Fig. 8. Morphotype III “Leucophthalma”. Male genitalia of M. leucophthalma (Wiedemann, 1819) (non-type, MFNB). Parameres in dorsal (A) and lateral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770305" httpUri="https://zenodo.org/record/4770305/files/figure.png" pageId="9" pageNumber="10">Fig. 8</figureCitation>
</paragraph>
<paragraph id="8BA3D655A63C0623FF46FD73FE2DE6C5" blockId="9.[189,463,729,756]" box="[189,463,729,756]" pageId="9" pageNumber="10">
<emphasis id="B9680A47A63C0623FF46FD73FE92E6C2" bold="true" box="[189,368,729,756]" pageId="9" pageNumber="10">Male genitalia</emphasis>
(
<figureCitation id="1327CAD0A63C0623FE84FD73FE27E6C5" box="[383,453,730,756]" captionStart="Fig" captionStartId="10.[189,232,720,746]" captionTargetBox="[259,1335,326,633]" captionTargetId="figure-17@10.[238,1349,305,646]" captionTargetPageId="10" captionText="Fig. 8. Morphotype III “Leucophthalma”. Male genitalia of M. leucophthalma (Wiedemann, 1819) (non-type, MFNB). Parameres in dorsal (A) and lateral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770305" httpUri="https://zenodo.org/record/4770305/files/figure.png" pageId="9" pageNumber="10">Fig. 8</figureCitation>
)
</paragraph>
<paragraph id="8BA3D655A63C0623FF46FCA3FC7FE785" blockId="9.[188,1399,777,949]" pageId="9" pageNumber="10">
Parameres (
<bibRefCitation id="EF8DABA4A63C0623FEBDFCA0FDECE712" author="Coca-Abia M. M." box="[326,526,777,803]" pageId="9" pageNumber="10" pagination="673 - 685" refId="ref43499" refString="Coca-Abia M. M. 2008. Revision of the genus Miridiba Reitter, 1901 (Coleoptera, Scarabaeidae, Melolonthinae). Zoological Science 25 (6): 673 - 685. https: // doi. org / 10.2108 / zsj. 25.673" type="journal article" year="2008">Coca-Abia 2008</bibRefCitation>
: fig. 17) consisting of dorsal and ventral branches separated in ventral and lateral views; dorsal branches only separated in distal end. Two ventral branches longer than dorsal ones, strong and curved downward. Collum absent. Endophallus with an area of soft setae, hair-like, at distal end, in some species spines arranged helicoidally around internal walls of sac. Temones laid at proximal half of endophallus, reduced; apophysis fused at proximal end.
</paragraph>
<paragraph id="8BA3D655A63C0623FF46FC74FE6EE7C6" blockId="9.[189,396,989,1015]" box="[189,396,989,1015]" pageId="9" pageNumber="10">
<heading id="D0EB6139A63C0623FF46FC74FE6EE7C6" bold="true" box="[189,396,989,1015]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B9680A47A63C0623FF46FC74FE6EE7C6" bold="true" box="[189,396,989,1015]" pageId="9" pageNumber="10">Female genitalia</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A63C0623FF46FBA5FC80E0A2" blockId="9.[189,1399,1036,1172]" pageId="9" pageNumber="10">Anal fold with all vestigial tergites similar in size; gonopore fold with plates varying in shapes with sensillae at posterior margin; vestigial sternites weakly developed with or without sensillae. Median oviduct strongly sclerotized. Peduncle of bursa copulatrix very long. Spermathecal reduced, gland strongly developed with caecum longer than tubular duct.</paragraph>
<paragraph id="8BA3D655A63C0623FD90FB15FC2BE0E4" blockId="9.[619,969,1211,1274]" box="[619,969,1211,1238]" pageId="9" pageNumber="10">
<heading id="D0EB6139A63C0623FD90FB15FC2BE0E4" box="[619,969,1211,1238]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="10" reason="2">
<emphasis id="B9680A47A63C0623FD90FB15FC2BE0E4" bold="true" box="[619,969,1211,1238]" pageId="9" pageNumber="10">
Morphotype IV “
<emphasis id="B9680A47A63C0623FCBEFB12FC58E0E7" bold="true" box="[837,954,1211,1238]" italics="true" pageId="9" pageNumber="10">Bidentata</emphasis>
”
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A63C0623FD1BFB49FCB1E0CB" blockId="9.[619,969,1211,1274]" box="[736,851,1248,1274]" pageId="9" pageNumber="10">
<figureCitation id="1327CAD0A63C0623FD1BFB49FCB1E0CB" box="[736,851,1248,1274]" captionStart="Figs 9–11" captionStartId="10.[189,241,1884,1910]" captionTargetBox="[277,1310,1002,1836]" captionTargetId="figure-47@10.[264,1323,988,1849]" captionTargetPageId="10" captionText="Figs 9–11. Morphotype IV “Bidentata”. 9–10. Male genitalia of M. behrensi Brenske, 1892, lectotype (MFNB). 9. Parameres in dorsal (A), ventral (B) and lateral (C) views. 10. Endophallus and temones in ventral view. 11. Female genitalia of M. saigonensis (Moser, 1912),?syntype (MFNB) in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770308" httpUri="https://zenodo.org/record/4770308/files/figure.png" pageId="9" pageNumber="10">Figs 9–11</figureCitation>
</paragraph>
<paragraph id="8BA3D655A63C0623FF46FA8AFE92E10D" blockId="9.[189,368,1314,1341]" box="[189,368,1314,1341]" pageId="9" pageNumber="10">
<heading id="D0EB6139A63C0623FF46FA8AFE92E10D" bold="true" box="[189,368,1314,1341]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B9680A47A63C0623FF46FA8AFE92E10D" bold="true" box="[189,368,1314,1341]" pageId="9" pageNumber="10">Male genitalia</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A63C0623FF46FAFAFCA7E277" blockId="9.[186,1401,1362,1607]" pageId="9" pageNumber="10">
Parameres (
<figureCitation id="1327CAD0A63C0623FEB1FAFBFE70E15D" box="[330,402,1362,1388]" captionStart="Figs 9–11" captionStartId="10.[189,241,1884,1910]" captionTargetBox="[277,1310,1002,1836]" captionTargetId="figure-47@10.[264,1323,988,1849]" captionTargetPageId="10" captionText="Figs 9–11. Morphotype IV “Bidentata”. 9–10. Male genitalia of M. behrensi Brenske, 1892, lectotype (MFNB). 9. Parameres in dorsal (A), ventral (B) and lateral (C) views. 10. Endophallus and temones in ventral view. 11. Female genitalia of M. saigonensis (Moser, 1912),?syntype (MFNB) in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770308" httpUri="https://zenodo.org/record/4770308/files/figure.png" pageId="9" pageNumber="10">Fig. 9</figureCitation>
) consisting of one dorsal and two ventral branches separated in lateral and ventral views. Presumably, dorsal branch resulted of two branches coalescence, showing a central sutural scar toward apical ostium varying in shapes and designs at distal end. Ventral branches bilaterally symmetrical, outer edge curved and elevated or straight; inner edge straight and converge at distal end. Collum absent. Endophallus (
<figureCitation id="1327CAD0A63C0623FE9BFA4AFE54E1CC" box="[352,438,1507,1533]" captionStart="Figs 9–11" captionStartId="10.[189,241,1884,1910]" captionTargetBox="[277,1310,1002,1836]" captionTargetId="figure-47@10.[264,1323,988,1849]" captionTargetPageId="10" captionText="Figs 9–11. Morphotype IV “Bidentata”. 9–10. Male genitalia of M. behrensi Brenske, 1892, lectotype (MFNB). 9. Parameres in dorsal (A), ventral (B) and lateral (C) views. 10. Endophallus and temones in ventral view. 11. Female genitalia of M. saigonensis (Moser, 1912),?syntype (MFNB) in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770308" httpUri="https://zenodo.org/record/4770308/files/figure.png" pageId="9" pageNumber="10">Fig. 10</figureCitation>
) with a few spines and, in some species, raspulae with dense sensillae. Temones laid at proximal half of endophallus, apophysis joined at proximal end, some species with distal ends surrounding endophallus partially (T-shaped) (
<figureCitation id="1327CAD0A63C0623FD1BF985FCDAE277" box="[736,824,1580,1606]" captionStart="Figs 9–11" captionStartId="10.[189,241,1884,1910]" captionTargetBox="[277,1310,1002,1836]" captionTargetId="figure-47@10.[264,1323,988,1849]" captionTargetPageId="10" captionText="Figs 9–11. Morphotype IV “Bidentata”. 9–10. Male genitalia of M. behrensi Brenske, 1892, lectotype (MFNB). 9. Parameres in dorsal (A), ventral (B) and lateral (C) views. 10. Endophallus and temones in ventral view. 11. Female genitalia of M. saigonensis (Moser, 1912),?syntype (MFNB) in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770308" httpUri="https://zenodo.org/record/4770308/files/figure.png" pageId="9" pageNumber="10">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="8BA3D655A63C0623FF46F9C7FE1AE2B9" blockId="9.[189,504,1646,1672]" box="[189,504,1646,1672]" pageId="9" pageNumber="10">
<emphasis id="B9680A47A63C0623FF46F9C7FE6EE2B9" bold="true" box="[189,396,1646,1672]" pageId="9" pageNumber="10">Female genitalia</emphasis>
(
<figureCitation id="1327CAD0A63C0623FE60F9C7FE0DE2B9" box="[411,495,1646,1672]" captionStart="Figs 9–11" captionStartId="10.[189,241,1884,1910]" captionTargetBox="[277,1310,1002,1836]" captionTargetId="figure-47@10.[264,1323,988,1849]" captionTargetPageId="10" captionText="Figs 9–11. Morphotype IV “Bidentata”. 9–10. Male genitalia of M. behrensi Brenske, 1892, lectotype (MFNB). 9. Parameres in dorsal (A), ventral (B) and lateral (C) views. 10. Endophallus and temones in ventral view. 11. Female genitalia of M. saigonensis (Moser, 1912),?syntype (MFNB) in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770308" httpUri="https://zenodo.org/record/4770308/files/figure.png" pageId="9" pageNumber="10">Fig. 11</figureCitation>
)
</paragraph>
<paragraph id="8BA3D655A63C0623FF46F937FE12E378" blockId="9.[188,1399,1694,1865]" pageId="9" pageNumber="10">Anal fold with all vestigial tergites similar in size; gonopore fold with a pair of plates, without sensillae, in rectangular-elongate shape extending into median oviduct dorsal wall; vestigial sternites weakly developed with a few sensillae. Median oviduct with wrinkled and/or hardened epithelium sclerotized to a greater or lesser extent. Bursa copulatrix with long peduncle. Spermatheca with short peduncle, gland with marked pouch.</paragraph>
<caption id="DF6386DDA63F0620FF46FD79FB26E73F" ID-DOI="http://doi.org/10.5281/zenodo.4770305" ID-Zenodo-Dep="4770305" httpUri="https://zenodo.org/record/4770305/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,720,746]" targetBox="[259,1335,326,633]" targetPageId="10">
<paragraph id="8BA3D655A63F0620FF46FD79FB26E73F" blockId="10.[189,1398,720,782]" pageId="10" pageNumber="11">
<emphasis id="B9680A47A63F0620FF46FD79FEEDE6DB" bold="true" box="[189,271,720,746]" pageId="10" pageNumber="11">Fig. 8.</emphasis>
Morphotype III “
<emphasis id="B9680A47A63F0620FE15FD79FD47E6DB" box="[494,677,720,746]" italics="true" pageId="10" pageNumber="11">Leucophthalma</emphasis>
”. Male genitalia of
<taxonomicName id="4C1CADD6A63F0620FC59FD78FA94E6DB" authority="(Wiedemann, 1819)" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[930,1398,720,746]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="leucophthalma">
<emphasis id="B9680A47A63F0620FC59FD78FB9EE6DB" box="[930,1148,720,746]" italics="true" pageId="10" pageNumber="11">M. leucophthalma</emphasis>
(
<bibRefCitation id="EF8DABA4A63F0620FB6BFD79FA8CE6DB" author="Wiedemann C. R. W." box="[1168,1390,720,746]" pageId="10" pageNumber="11" pagination="157 - 183" refId="ref46073" refString="Wiedemann C. R. W. 1819. VI. Neue Kafer aus Bengalen und Java; befchrieben vom Herausgeber. Zoologisches Magazin, Kiel 1 (3): 157 - 183." type="journal article" year="1819">Wiedemann, 1819</bibRefCitation>
)
</taxonomicName>
(non-type, MFNB). Parameres in dorsal (A) and lateral (B) views. Scale bars = 1.0 mm.
</paragraph>
</caption>
<caption id="DF6386DDA63F0620FF46F8F5FD6DE3D0" ID-DOI="http://doi.org/10.5281/zenodo.4770308" ID-Zenodo-Dep="4770308" httpUri="https://zenodo.org/record/4770308/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,241,1884,1910]" targetBox="[277,1310,1002,1836]" targetPageId="10">
<paragraph id="8BA3D655A63F0620FF46F8F5FD6DE3D0" blockId="10.[188,1399,1884,2017]" pageId="10" pageNumber="11">
<emphasis id="B9680A47A63F0620FF46F8F5FEDBE347" bold="true" box="[189,313,1884,1910]" pageId="10" pageNumber="11">Figs 9–11.</emphasis>
Morphotype IV “
<emphasis id="B9680A47A63F0620FDEBF8F5FD60E347" box="[528,642,1884,1910]" italics="true" pageId="10" pageNumber="11">Bidentata</emphasis>
”.
<emphasis id="B9680A47A63F0620FD64F8F5FD38E347" bold="true" box="[671,730,1884,1910]" pageId="10" pageNumber="11">9–10</emphasis>
. Male genitalia of
<taxonomicName id="4C1CADD6A63F0620FC46F8F4FB1CE346" authority="Brenske, 1892" authorityName="Brenske" authorityYear="1892" box="[957,1278,1884,1911]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="behrensi">
<emphasis id="B9680A47A63F0620FC46F8F4FBABE347" box="[957,1097,1884,1910]" italics="true" pageId="10" pageNumber="11">M. behrensi</emphasis>
<bibRefCitation id="EF8DABA4A63F0620FBAAF8F5FB1CE346" author="Brenske E." box="[1105,1278,1884,1911]" pageId="10" pageNumber="11" pagination="159 - 192" refId="ref43256" refString="Brenske E. 1892. Neue arten der Coleopteren-Gattung Holotrichia (Lachnosterna). Berliner Entomologische Zeitschrift 37 (2): 159 - 192." type="journal article" year="1892">Brenske, 1892</bibRefCitation>
</taxonomicName>
, lectotype (MFNB).
<emphasis id="B9680A47A63F0620FEC9F8D6FEA3E3A8" bold="true" box="[306,321,1919,1945]" pageId="10" pageNumber="11">9</emphasis>
. Parameres in dorsal (A), ventral (B) and lateral (C) views.
<emphasis id="B9680A47A63F0620FC04F8D6FBFEE3A8" bold="true" box="[1023,1052,1919,1945]" pageId="10" pageNumber="11">10</emphasis>
. Endophallus and temones in ventral view.
<emphasis id="B9680A47A63F0620FEA2F80AFE97E38C" bold="true" box="[345,373,1955,1981]" pageId="10" pageNumber="11">11</emphasis>
. Female genitalia of
<taxonomicName id="4C1CADD6A63F0620FD90F80DFC28E38C" authority="(Moser, 1912)" baseAuthorityName="Moser" baseAuthorityYear="1912" box="[619,970,1955,1981]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="saigonensis">
<emphasis id="B9680A47A63F0620FD90F80DFCF8E38C" box="[619,794,1956,1981]" italics="true" pageId="10" pageNumber="11">M. saigonensis</emphasis>
(
<bibRefCitation id="EF8DABA4A63F0620FCD2F80AFC20E38C" author="Moser J." box="[809,962,1955,1981]" pageId="10" pageNumber="11" pagination="420 - 449" refId="ref45321" refString="Moser J. 1912. Neue arten der Melolonthiden-Gattungen Holotrichia und Pentelia. Annales de la Societe entomologique de Belgique 56: 420 - 449." type="journal article" year="1912">Moser, 1912</bibRefCitation>
)
</taxonomicName>
,?syntype (MFNB) in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BA3D655A63E0621FD82FEA0FC59E513" blockId="11.[633,955,264,327]" box="[633,955,264,291]" pageId="11" pageNumber="12">
<heading id="D0EB6139A63E0621FD82FEA0FC59E513" box="[633,955,264,291]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
<emphasis id="B9680A47A63E0621FD82FEA0FC59E513" bold="true" box="[633,955,264,291]" pageId="11" pageNumber="12">
Morphotype V “
<emphasis id="B9680A47A63E0621FCBCFEA1FC4EE513" bold="true" box="[839,940,264,290]" italics="true" pageId="11" pageNumber="12">Sinensis</emphasis>
”
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A63E0621FD23FE84FCB9E576" blockId="11.[633,955,264,327]" box="[728,859,301,327]" pageId="11" pageNumber="12">
<figureCitation id="1327CAD0A63E0621FD23FE84FCB9E576" box="[728,859,301,327]" captionStart="Figs 12–14" captionStartId="12.[189,241,954,980]" captionTargetBox="[283,1310,296,918]" captionTargetId="figure-18@12.[264,1323,283,920]" captionTargetPageId="12" captionText="Figs 12–14. Morphotype V “Sinensis”. 12–13. Male genitalia of M. sinensis (Hope, 1842), lectotype (BMNH). 12. Parameres in dorsal (A), ventral (B) and lateral (C) views. 13. Endophallus and temones in ventral view. 14. Female genitalia of M. dalatensis Frey, 1970, holotype (NHMB) in lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770312" httpUri="https://zenodo.org/record/4770312/files/figure.png" pageId="11" pageNumber="12">Figs 12–14</figureCitation>
</paragraph>
<paragraph id="8BA3D655A63E0621FF46FEC6FE92E5B8" blockId="11.[189,368,367,393]" box="[189,368,367,393]" pageId="11" pageNumber="12">
<heading id="D0EB6139A63E0621FF46FEC6FE92E5B8" bold="true" box="[189,368,367,393]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B9680A47A63E0621FF46FEC6FE92E5B8" bold="true" box="[189,368,367,393]" pageId="11" pageNumber="12">Male genitalia</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A63E0621FF46FE37FD7CE69F" blockId="11.[189,1400,414,686]" pageId="11" pageNumber="12">
Parameres (
<figureCitation id="1327CAD0A63E0621FEB2FE37FE7DE589" box="[329,415,414,440]" captionStart="Figs 12–14" captionStartId="12.[189,241,954,980]" captionTargetBox="[283,1310,296,918]" captionTargetId="figure-18@12.[264,1323,283,920]" captionTargetPageId="12" captionText="Figs 12–14. Morphotype V “Sinensis”. 12–13. Male genitalia of M. sinensis (Hope, 1842), lectotype (BMNH). 12. Parameres in dorsal (A), ventral (B) and lateral (C) views. 13. Endophallus and temones in ventral view. 14. Female genitalia of M. dalatensis Frey, 1970, holotype (NHMB) in lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770312" httpUri="https://zenodo.org/record/4770312/files/figure.png" pageId="11" pageNumber="12">Fig. 12</figureCitation>
) consisting of two dorsal and two ventral branches separated in dorsal and ventral views and fused laterally. Lateral scars, presumably, formed by the fusion of dorsal and ventral branches, or joined totally or partially by a lateral membranous area. Dorsal branches with symmetrical or asymmetrical design; collum undefined in lateral view, with a membranous area in dorsal view. Ventral branches bilaterally symmetrical, straight and convergent at distal end. Phallobase shorter than paramere. Endophallus (
<figureCitation id="1327CAD0A63E0621FE1BFDE7FDD5E659" box="[480,567,590,616]" captionStart="Figs 12–14" captionStartId="12.[189,241,954,980]" captionTargetBox="[283,1310,296,918]" captionTargetId="figure-18@12.[264,1323,283,920]" captionTargetPageId="12" captionText="Figs 12–14. Morphotype V “Sinensis”. 12–13. Male genitalia of M. sinensis (Hope, 1842), lectotype (BMNH). 12. Parameres in dorsal (A), ventral (B) and lateral (C) views. 13. Endophallus and temones in ventral view. 14. Female genitalia of M. dalatensis Frey, 1970, holotype (NHMB) in lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770312" httpUri="https://zenodo.org/record/4770312/files/figure.png" pageId="11" pageNumber="12">Fig. 13</figureCitation>
) with a dense area of small spines (raspulae), stronger towards distal end, or raspulae absent, only acicular soft setae. Temones laid at proximal half of endophallus, slender, apophysis separated each other (
<figureCitation id="1327CAD0A63E0621FDC2FD3DFD72E69F" box="[569,656,660,686]" captionStart="Figs 12–14" captionStartId="12.[189,241,954,980]" captionTargetBox="[283,1310,296,918]" captionTargetId="figure-18@12.[264,1323,283,920]" captionTargetPageId="12" captionText="Figs 12–14. Morphotype V “Sinensis”. 12–13. Male genitalia of M. sinensis (Hope, 1842), lectotype (BMNH). 12. Parameres in dorsal (A), ventral (B) and lateral (C) views. 13. Endophallus and temones in ventral view. 14. Female genitalia of M. dalatensis Frey, 1970, holotype (NHMB) in lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770312" httpUri="https://zenodo.org/record/4770312/files/figure.png" pageId="11" pageNumber="12">Fig. 13</figureCitation>
).
</paragraph>
<paragraph id="8BA3D655A63E0621FF46FD7FFE1BE6C1" blockId="11.[189,505,725,752]" box="[189,505,725,752]" pageId="11" pageNumber="12">
<emphasis id="B9680A47A63E0621FF46FD7FFE6EE6DE" bold="true" box="[189,396,725,752]" pageId="11" pageNumber="12">Female genitalia</emphasis>
(
<figureCitation id="1327CAD0A63E0621FE60FD7FFE12E6C1" box="[411,496,726,752]" captionStart="Figs 12–14" captionStartId="12.[189,241,954,980]" captionTargetBox="[283,1310,296,918]" captionTargetId="figure-18@12.[264,1323,283,920]" captionTargetPageId="12" captionText="Figs 12–14. Morphotype V “Sinensis”. 12–13. Male genitalia of M. sinensis (Hope, 1842), lectotype (BMNH). 12. Parameres in dorsal (A), ventral (B) and lateral (C) views. 13. Endophallus and temones in ventral view. 14. Female genitalia of M. dalatensis Frey, 1970, holotype (NHMB) in lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770312" httpUri="https://zenodo.org/record/4770312/files/figure.png" pageId="11" pageNumber="12">Fig. 14</figureCitation>
)
</paragraph>
<paragraph id="8BA3D655A63E0621FF46FCADFED4E79A" blockId="11.[189,1400,772,939]" pageId="11" pageNumber="12">Anal fold with all vestigial tergites similar in size; gonopore fold with a pair of short sensory elongate plates, with sensilla at posterior edge; vestigial sternites weakly developed without apparent sensillae. Median oviduct with wrinkled and hardened epithelium sclerotized to a greater or lesser extent. Bursa copulatrix with short peduncle, pouch arranged transversely. Spermatheca with short peduncle, gland very long.</paragraph>
<paragraph id="8BA3D655A63E0621FD99FC7AFC30E7DD" blockId="11.[610,978,978,1040]" box="[610,978,978,1005]" pageId="11" pageNumber="12">
<heading id="D0EB6139A63E0621FD99FC7AFC30E7DD" box="[610,978,978,1005]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
<emphasis id="B9680A47A63E0621FD99FC7AFC30E7DD" bold="true" box="[610,978,978,1005]" pageId="11" pageNumber="12">
Morphotype VI “
<emphasis id="B9680A47A63E0621FCC7FC7BFC21E7DD" bold="true" box="[828,963,978,1004]" italics="true" pageId="11" pageNumber="12">Borneensis</emphasis>
”
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A63E0621FD23FC5FFCB9E021" blockId="11.[610,978,978,1040]" box="[728,859,1013,1040]" pageId="11" pageNumber="12">
<figureCitation id="1327CAD0A63E0621FD23FC5FFCB9E021" box="[728,859,1013,1040]" captionStart="Figs 15–17" captionStartId="12.[189,241,1860,1886]" captionTargetBox="[277,1313,1218,1772]" captionTargetId="figure-19@12.[264,1323,1202,1785]" captionTargetPageId="12" captionText="Figs 15–17. Morphotype VI “Borneensis”. 15. Parameres of M. borneensis (Moser, 1918), lectotype (BMNH) in dorsal (A) and lateral (B) views. 16. Endophallus and temones of M. coxalis (Arrow, 1944), lectotype (BMNH) in dorsal view. 17. Female genitalia of M. coxalis, paralectotype (BMNH) in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770314" httpUri="https://zenodo.org/record/4770314/files/figure.png" pageId="11" pageNumber="12">Figs 15–17</figureCitation>
</paragraph>
<paragraph id="8BA3D655A63E0621FF46FB9EFE92E060" blockId="11.[189,368,1079,1105]" box="[189,368,1079,1105]" pageId="11" pageNumber="12">
<heading id="D0EB6139A63E0621FF46FB9EFE92E060" bold="true" box="[189,368,1079,1105]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B9680A47A63E0621FF46FB9EFE92E060" bold="true" box="[189,368,1079,1105]" pageId="11" pageNumber="12">Male genitalia</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A63E0621FF46FBCFFCFEE165" blockId="11.[187,1401,1125,1364]" pageId="11" pageNumber="12">
Parameres (
<figureCitation id="1327CAD0A63E0621FEB2FBCCFE42E0B1" box="[329,416,1125,1152]" captionStart="Figs 15–17" captionStartId="12.[189,241,1860,1886]" captionTargetBox="[277,1313,1218,1772]" captionTargetId="figure-19@12.[264,1323,1202,1785]" captionTargetPageId="12" captionText="Figs 15–17. Morphotype VI “Borneensis”. 15. Parameres of M. borneensis (Moser, 1918), lectotype (BMNH) in dorsal (A) and lateral (B) views. 16. Endophallus and temones of M. coxalis (Arrow, 1944), lectotype (BMNH) in dorsal view. 17. Female genitalia of M. coxalis, paralectotype (BMNH) in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770314" httpUri="https://zenodo.org/record/4770314/files/figure.png" pageId="11" pageNumber="12">Fig. 15</figureCitation>
) consisting of two dorsal and two ventral branches separated in dorsal and lateral views. Dorsal branches varying in shapes, reduced and shorter than ventral ones. Ventral branches joined by a dorsal membrane and fused ventrally forming up a tubular structure through which endophallus passes; apices of ventral branches separate. Collum absent. Parameres and phallobase joined by a wide dorsal membrane varying in shapes. Endophallus (
<figureCitation id="1327CAD0A63E0621FCE9FB5BFC8AE13C" box="[786,872,1266,1293]" captionStart="Figs 15–17" captionStartId="12.[189,241,1860,1886]" captionTargetBox="[277,1313,1218,1772]" captionTargetId="figure-19@12.[264,1323,1202,1785]" captionTargetPageId="12" captionText="Figs 15–17. Morphotype VI “Borneensis”. 15. Parameres of M. borneensis (Moser, 1918), lectotype (BMNH) in dorsal (A) and lateral (B) views. 16. Endophallus and temones of M. coxalis (Arrow, 1944), lectotype (BMNH) in dorsal view. 17. Female genitalia of M. coxalis, paralectotype (BMNH) in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770314" httpUri="https://zenodo.org/record/4770314/files/figure.png" pageId="11" pageNumber="12">Fig. 16</figureCitation>
) almost empty, with few spines at middle of sac and inconspicuous sensillae. Temones laid at proximal half of endophallus, apophysis joined, distal ends surrounding endophallus completely (
<figureCitation id="1327CAD0A63E0621FD4CFA90FCEDE165" box="[695,783,1337,1364]" captionStart="Figs 15–17" captionStartId="12.[189,241,1860,1886]" captionTargetBox="[277,1313,1218,1772]" captionTargetId="figure-19@12.[264,1323,1202,1785]" captionTargetPageId="12" captionText="Figs 15–17. Morphotype VI “Borneensis”. 15. Parameres of M. borneensis (Moser, 1918), lectotype (BMNH) in dorsal (A) and lateral (B) views. 16. Endophallus and temones of M. coxalis (Arrow, 1944), lectotype (BMNH) in dorsal view. 17. Female genitalia of M. coxalis, paralectotype (BMNH) in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770314" httpUri="https://zenodo.org/record/4770314/files/figure.png" pageId="11" pageNumber="12">Fig. 16</figureCitation>
).
</paragraph>
<paragraph id="8BA3D655A63E0621FF46FAD3FE1BE1A5" blockId="11.[188,1399,1402,1612]" box="[189,505,1402,1428]" pageId="11" pageNumber="12">
<emphasis id="B9680A47A63E0621FF46FAD3FE6EE1A5" bold="true" box="[189,396,1402,1428]" pageId="11" pageNumber="12">Female genitalia</emphasis>
(
<figureCitation id="1327CAD0A63E0621FE60FAD3FE12E1A5" box="[411,496,1402,1428]" captionStart="Figs 15–17" captionStartId="12.[189,241,1860,1886]" captionTargetBox="[277,1313,1218,1772]" captionTargetId="figure-19@12.[264,1323,1202,1785]" captionTargetPageId="12" captionText="Figs 15–17. Morphotype VI “Borneensis”. 15. Parameres of M. borneensis (Moser, 1918), lectotype (BMNH) in dorsal (A) and lateral (B) views. 16. Endophallus and temones of M. coxalis (Arrow, 1944), lectotype (BMNH) in dorsal view. 17. Female genitalia of M. coxalis, paralectotype (BMNH) in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770314" httpUri="https://zenodo.org/record/4770314/files/figure.png" pageId="11" pageNumber="12">Fig. 17</figureCitation>
)
</paragraph>
<paragraph id="8BA3D655A63E0621FF46FA0CFD5EE27D" blockId="11.[188,1399,1402,1612]" pageId="11" pageNumber="12">Anal fold with all vestigial tergites similar in size; gonopore fold with a pair of conspicuous plates varying in shapes without sensillae, extending into median oviduct dorsal wall; vestigial sternites absent; ventral wall of vulva with a gently sclerotized area with punctate sensillae. Median oviduct with wrinkled and/or hardened epithelium sclerotized to a greater or lesser extent. Bursa copulatrix with short peduncle. Spermatheca with long peduncle.</paragraph>
<paragraph id="8BA3D655A63E0621FDACF9DAFC3FE2BC" blockId="11.[599,989,1651,1713]" box="[599,989,1651,1678]" pageId="11" pageNumber="12">
<heading id="D0EB6139A63E0621FDACF9DAFC3FE2BC" box="[599,989,1651,1678]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
<emphasis id="B9680A47A63E0621FDACF9DAFC3FE2BC" bold="true" box="[599,989,1651,1678]" pageId="11" pageNumber="12">
Morphotype VII “
<emphasis id="B9680A47A63E0621FCC7F9DAFC2FE2BF" bold="true" box="[828,973,1651,1678]" italics="true" pageId="11" pageNumber="12">Rugaticollis</emphasis>
”
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A63E0621FD23F93FFCB9E280" blockId="11.[599,989,1651,1713]" box="[728,859,1686,1713]" pageId="11" pageNumber="12">
<figureCitation id="1327CAD0A63E0621FD23F93FFCB9E280" box="[728,859,1686,1713]" captionStart="Figs 18–19" captionStartId="13.[189,241,983,1009]" captionTargetBox="[281,1310,319,842]" captionTargetId="figure-187@13.[264,1323,299,855]" captionTargetPageId="13" captionText="Figs 18–19. Morphotype VII “Rugaticollis”. Male genitalia of M. rugaticollis (Moser, 1913), lectotype (MFNB). 18. Parameres in dorsal (A), ventral (B) and lateral (C) views. 19. Endophallus and temones in lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770320" httpUri="https://zenodo.org/record/4770320/files/figure.png" pageId="11" pageNumber="12">Figs 18–19</figureCitation>
</paragraph>
<paragraph id="8BA3D655A63E0621FF46F971FE92E2C3" blockId="11.[189,368,1752,1778]" box="[189,368,1752,1778]" pageId="11" pageNumber="12">
<heading id="D0EB6139A63E0621FF46F971FE92E2C3" bold="true" box="[189,368,1752,1778]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B9680A47A63E0621FF46F971FE92E2C3" bold="true" box="[189,368,1752,1778]" pageId="11" pageNumber="12">Male genitalia</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A63E0621FF46F8AEFA94E39C" blockId="11.[186,1400,1798,1966]" pageId="11" pageNumber="12">
Parameres (
<figureCitation id="1327CAD0A63E0621FEBEF8AFFE75E310" box="[325,407,1798,1825]" captionStart="Figs 18–19" captionStartId="13.[189,241,983,1009]" captionTargetBox="[281,1310,319,842]" captionTargetId="figure-187@13.[264,1323,299,855]" captionTargetPageId="13" captionText="Figs 18–19. Morphotype VII “Rugaticollis”. Male genitalia of M. rugaticollis (Moser, 1913), lectotype (MFNB). 18. Parameres in dorsal (A), ventral (B) and lateral (C) views. 19. Endophallus and temones in lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770320" httpUri="https://zenodo.org/record/4770320/files/figure.png" pageId="11" pageNumber="12">Fig. 18</figureCitation>
) consisting of one dorsal and two ventral branches. Presumably, dorsal branch resulted of two branches coalescence, showing a central sutural scar opened toward apical ostium. Dorsal branch thick and wide, gently narrowing toward apex in dorsal view; with a notch varying in shapes at distal end in lateral and ventral views. Collum undefined in lateral view and with a triangular-shaped area less sclerotized at proximal half. Ventral branches longer and thinner than dorsal ones, with apices gently
</paragraph>
<caption id="DF6386DDA6390626FF46FC13FE57E00E" ID-DOI="http://doi.org/10.5281/zenodo.4770312" ID-Zenodo-Dep="4770312" httpUri="https://zenodo.org/record/4770312/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,241,954,980]" targetBox="[283,1310,296,918]" targetPageId="12">
<paragraph id="8BA3D655A6390626FF46FC13FE57E00E" blockId="12.[189,1399,954,1087]" pageId="12" pageNumber="13">
<emphasis id="B9680A47A6390626FF46FC13FEA9E7E5" bold="true" box="[189,331,954,980]" pageId="12" pageNumber="13">Figs 12–14.</emphasis>
Morphotype V “
<emphasis id="B9680A47A6390626FDE0FC12FD99E7E4" box="[539,635,955,981]" italics="true" pageId="12" pageNumber="13">Sinensis</emphasis>
”.
<emphasis id="B9680A47A6390626FD62FC13FD01E7E5" bold="true" box="[665,739,954,980]" pageId="12" pageNumber="13">12–13</emphasis>
. Male genitalia of
<taxonomicName id="4C1CADD6A6390626FC36FC12FB1FE7E4" authority="(Hope, 1842)" baseAuthorityName="Hope" baseAuthorityYear="1842" box="[973,1277,955,981]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B9680A47A6390626FC36FC12FBB6E7E5" box="[973,1108,955,980]" italics="true" pageId="12" pageNumber="13">M. sinensis</emphasis>
(
<bibRefCitation id="EF8DABA4A6390626FB98FC12FB16E7E4" author="Hope F. W." box="[1123,1268,955,981]" pageId="12" pageNumber="13" pagination="59 - 66" refId="ref44236" refString="Hope F. W. 1842. Descriptions of the Coleopterous insects sent to England by Dr. Cantor from Chusan and Canton, with observations on the Entomology of China. Proceedings of the Entomological Society of London 1841: 59 - 66." type="journal article" year="1842">Hope, 1842</bibRefCitation>
)
</taxonomicName>
, lectotype (BMNH).
<emphasis id="B9680A47A6390626FEC3FC77FEB7E7C9" bold="true" box="[312,341,990,1016]" pageId="12" pageNumber="13">12</emphasis>
. Parameres in dorsal (A), ventral (B) and lateral (C) views.
<emphasis id="B9680A47A6390626FBE7FC77FBDBE7C9" bold="true" box="[1052,1081,990,1016]" pageId="12" pageNumber="13">13</emphasis>
. Endophallus and temones in ventral view.
<emphasis id="B9680A47A6390626FE79FBA8FE7DE02A" bold="true" box="[386,415,1025,1051]" pageId="12" pageNumber="13">14</emphasis>
. Female genitalia of
<taxonomicName id="4C1CADD6A6390626FD59FBABFC2DE02A" authority="Frey, 1970" authorityName="Frey" authorityYear="1970" box="[674,975,1025,1052]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dalatensis">
<emphasis id="B9680A47A6390626FD59FBABFCA1E02A" box="[674,835,1025,1051]" italics="true" pageId="12" pageNumber="13">M. dalatensis</emphasis>
<bibRefCitation id="EF8DABA4A6390626FCB6FBABFC2DE02A" author="Frey G." box="[845,975,1025,1052]" pageId="12" pageNumber="13" pagination="244 - 265" refId="ref43949" refString="Frey G. 1970. Bestimmungstabelle der Holotrichia - Arten Hinterindiens (Vietnam, Laos, Thailand, Kambodscha, Burma, Malaya) (Col., Scarab, Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 21: 244 - 265." type="journal article" year="1970">Frey, 1970</bibRefCitation>
</taxonomicName>
, holotype (NHMB) in lateral view. Scale bars = 1.0 mm.
</paragraph>
</caption>
<caption id="DF6386DDA6390626FF46F8EDFD10E3F9" ID-DOI="http://doi.org/10.5281/zenodo.4770314" ID-Zenodo-Dep="4770314" httpUri="https://zenodo.org/record/4770314/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,241,1860,1886]" targetBox="[277,1313,1218,1772]" targetPageId="12">
<paragraph id="8BA3D655A6390626FF46F8EDFD10E3F9" blockId="12.[188,1399,1860,1992]" pageId="12" pageNumber="13">
<emphasis id="B9680A47A6390626FF46F8EDFEA9E36F" bold="true" box="[189,331,1860,1886]" pageId="12" pageNumber="13">Figs 15–17.</emphasis>
Morphotype VI “
<emphasis id="B9680A47A6390626FDDCF8ECFD4BE36F" box="[551,681,1861,1886]" italics="true" pageId="12" pageNumber="13">Borneensis</emphasis>
”.
<emphasis id="B9680A47A6390626FD3CF8EDFD06E36F" bold="true" box="[711,740,1860,1886]" pageId="12" pageNumber="13">15</emphasis>
. Parameres of
<taxonomicName id="4C1CADD6A6390626FC66F8ECFB1EE36F" authority="(Moser, 1918)" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Moser" baseAuthorityYear="1918" box="[925,1276,1860,1886]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis id="B9680A47A6390626FC66F8ECFBA4E36F" box="[925,1094,1860,1886]" italics="true" pageId="12" pageNumber="13">M. borneensis</emphasis>
(
<bibRefCitation id="EF8DABA4A6390626FBA3F8EDFB11E36F" author="Moser J." box="[1112,1267,1860,1886]" pageId="12" pageNumber="13" pagination="297 - 349" refId="ref45453" refString="Moser J. 1918. Beitrag zur Kenntnis der Melolonthiden. (Col.) VIII. Stettiner entomologische Zeitung 79: 297 - 349." type="journal article" year="1918">Moser, 1918</bibRefCitation>
)
</taxonomicName>
, lectotype (BMNH) in dorsal (A) and lateral (B) views.
<emphasis id="B9680A47A6390626FD30F8CEFD0AE3B0" bold="true" box="[715,744,1895,1921]" pageId="12" pageNumber="13">16</emphasis>
. Endophallus and temones of
<taxonomicName id="4C1CADD6A6390626FBBCF8C1FA90E3B0" authority="(Arrow, 1944)" authorityName="CH. Bu Gao &amp; M. Coca-Abia" authorityYear="2021" baseAuthorityName="Arrow" baseAuthorityYear="1944" box="[1095,1394,1895,1921]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="coxalis">
<emphasis id="B9680A47A6390626FBBCF8C1FB23E3B0" box="[1095,1217,1895,1921]" italics="true" pageId="12" pageNumber="13">M. coxalis</emphasis>
(
<bibRefCitation id="EF8DABA4A6390626FB2BF8CEFA8BE3B0" author="Arrow G. J." box="[1232,1385,1895,1921]" pageId="12" pageNumber="13" pagination="631 - 648" refId="ref43013" refString="Arrow G. J. 1944. LV. - Systematic notes on Melolonthine beetles belonging to Holotrichia and related genera. The Annals and Magazine of Natural History 11 (82): 631 - 648. https: // doi. org / 10.1080 / 00222934408527462" type="journal article" year="1944">Arrow, 1944</bibRefCitation>
)
</taxonomicName>
, lectotype (BMNH) in dorsal view.
<emphasis id="B9680A47A6390626FDACF823FD96E395" bold="true" box="[599,628,1930,1956]" pageId="12" pageNumber="13">17</emphasis>
. Female genitalia of
<taxonomicName id="4C1CADD6A6390626FC95F822FC0BE395" authorityName="CH. Bu Gao &amp; M. Coca-Abia" authorityYear="2021" baseAuthorityName="Arrow" baseAuthorityYear="1944" box="[878,1001,1930,1956]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="coxalis">
<emphasis id="B9680A47A6390626FC95F822FC0BE395" box="[878,1001,1930,1956]" italics="true" pageId="12" pageNumber="13">M. coxalis</emphasis>
</taxonomicName>
, paralectotype (BMNH) in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm.
</paragraph>
</caption>
<caption id="DF6386DDA6380627FF46FC7EFD89E008" ID-DOI="http://doi.org/10.5281/zenodo.4770320" ID-Zenodo-Dep="4770320" httpUri="https://zenodo.org/record/4770320/files/figure.png" pageId="13" pageNumber="14" startId="13.[189,241,983,1009]" targetBox="[281,1310,319,842]" targetPageId="13">
<paragraph id="8BA3D655A6380627FF46FC7EFD89E008" blockId="13.[189,1399,983,1081]" pageId="13" pageNumber="14">
<emphasis id="B9680A47A6380627FF46FC7EFEABE7C3" bold="true" box="[189,329,983,1010]" pageId="13" pageNumber="14">Figs 18–19.</emphasis>
Morphotype VII “
<emphasis id="B9680A47A6380627FDDDFC71FD56E7C3" box="[550,692,984,1010]" italics="true" pageId="13" pageNumber="14">Rugaticollis</emphasis>
”. Male genitalia of
<taxonomicName id="4C1CADD6A6380627FC5BFC70FB1DE7C3" authority="(Moser, 1913)" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[928,1279,984,1010]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="rugaticollis">
<emphasis id="B9680A47A6380627FC5BFC70FBACE7C3" box="[928,1102,984,1010]" italics="true" pageId="13" pageNumber="14">M. rugaticollis</emphasis>
(Moser, 1913)
</taxonomicName>
, lectotype (MFNB).
<emphasis id="B9680A47A6380627FEC8FC52FEB2E024" bold="true" box="[307,336,1019,1045]" pageId="13" pageNumber="14">18</emphasis>
. Parameres in dorsal (A), ventral (B) and lateral (C) views.
<emphasis id="B9680A47A6380627FBE0FC52FBD5E024" bold="true" box="[1051,1079,1019,1045]" pageId="13" pageNumber="14">19</emphasis>
. Endophallus and temones in lateral view. Scale bars = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BA3D655A6380627FF46FB15FC02E0CB" blockId="13.[189,1398,1212,1274]" pageId="13" pageNumber="14">
raised. Endophallus (
<figureCitation id="1327CAD0A6380627FE4FFB15FDEBE0E7" box="[436,521,1212,1238]" captionStart="Figs 18–19" captionStartId="13.[189,241,983,1009]" captionTargetBox="[281,1310,319,842]" captionTargetId="figure-187@13.[264,1323,299,855]" captionTargetPageId="13" captionText="Figs 18–19. Morphotype VII “Rugaticollis”. Male genitalia of M. rugaticollis (Moser, 1913), lectotype (MFNB). 18. Parameres in dorsal (A), ventral (B) and lateral (C) views. 19. Endophallus and temones in lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770320" httpUri="https://zenodo.org/record/4770320/files/figure.png" pageId="13" pageNumber="14">Fig. 19</figureCitation>
) with a sacculus bearing several short spines at distal half. Temones laid at distal end of endophallus, apophysis slender and separated (
<figureCitation id="1327CAD0A6380627FC80FB76FC30E0CB" box="[891,978,1247,1274]" captionStart="Figs 18–19" captionStartId="13.[189,241,983,1009]" captionTargetBox="[281,1310,319,842]" captionTargetId="figure-187@13.[264,1323,299,855]" captionTargetPageId="13" captionText="Figs 18–19. Morphotype VII “Rugaticollis”. Male genitalia of M. rugaticollis (Moser, 1913), lectotype (MFNB). 18. Parameres in dorsal (A), ventral (B) and lateral (C) views. 19. Endophallus and temones in lateral view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770320" httpUri="https://zenodo.org/record/4770320/files/figure.png" pageId="13" pageNumber="14">Fig. 19</figureCitation>
).
</paragraph>
<paragraph id="8BA3D655A6380627FF46FA88FE6EE10A" blockId="13.[189,396,1313,1339]" box="[189,396,1313,1339]" pageId="13" pageNumber="14">
<heading id="D0EB6139A6380627FF46FA88FE6EE10A" bold="true" box="[189,396,1313,1339]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B9680A47A6380627FF46FA88FE6EE10A" bold="true" box="[189,396,1313,1339]" pageId="13" pageNumber="14">Female genitalia</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6380627FF46FAE6FEDBE158" blockId="13.[189,313,1359,1385]" box="[189,313,1359,1385]" pageId="13" pageNumber="14">Unknown.</paragraph>
<paragraph id="8BA3D655A6380627FD97FA38FC25E19A" blockId="13.[620,967,1425,1486]" box="[620,967,1425,1451]" pageId="13" pageNumber="14">
<heading id="D0EB6139A6380627FD97FA38FC25E19A" box="[620,967,1425,1451]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="14" reason="2">
<emphasis id="B9680A47A6380627FD97FA38FC25E19A" bold="true" box="[620,967,1425,1451]" pageId="13" pageNumber="14">
Morphotype VIII “
<emphasis id="B9680A47A6380627FCA6FA38FC5AE19A" bold="true" box="[861,952,1425,1451]" italics="true" pageId="13" pageNumber="14">Scutata</emphasis>
”
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6380627FD23FA1DFCB9E1FF" blockId="13.[620,967,1425,1486]" box="[728,859,1460,1486]" pageId="13" pageNumber="14">
<figureCitation id="1327CAD0A6380627FD23FA1DFCB9E1FF" box="[728,859,1460,1486]" captionStart="Figs 20–22" captionStartId="14.[189,241,1251,1277]" captionTargetBox="[291,1324,285,1192]" captionTargetId="figure-142@14.[278,1337,265,1205]" captionTargetPageId="14" captionText="Figs 20–22. Morphotype VIII “Scutata”. 20. Male genitalia of M. scutata (Reitter, 1902), lectotype (MFNB), parameres in dorsal (A), ventral (B) and lateral (C) views. 21. Endophallus and temones of M. lassallei Keith, 2010, holotype (DKC) in lateral view. 22. Female genitalia of M. scutata, non-type (MFNB) in dorsal view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770322" httpUri="https://zenodo.org/record/4770322/files/figure.png" pageId="13" pageNumber="14">Figs 20–22</figureCitation>
</paragraph>
<paragraph id="8BA3D655A6380627FF46FA5FFE92E23E" blockId="13.[189,368,1525,1552]" box="[189,368,1525,1552]" pageId="13" pageNumber="14">
<heading id="D0EB6139A6380627FF46FA5FFE92E23E" bold="true" box="[189,368,1525,1552]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B9680A47A6380627FF46FA5FFE92E23E" bold="true" box="[189,368,1525,1552]" pageId="13" pageNumber="14">Male genitalia</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6380627FF46F98CFE52E2DF" blockId="13.[188,1399,1572,1774]" pageId="13" pageNumber="14">
Parameres (
<figureCitation id="1327CAD0A6380627FEBCF98DFE7EE20F" box="[327,412,1572,1598]" captionStart="Figs 20–22" captionStartId="14.[189,241,1251,1277]" captionTargetBox="[291,1324,285,1192]" captionTargetId="figure-142@14.[278,1337,265,1205]" captionTargetPageId="14" captionText="Figs 20–22. Morphotype VIII “Scutata”. 20. Male genitalia of M. scutata (Reitter, 1902), lectotype (MFNB), parameres in dorsal (A), ventral (B) and lateral (C) views. 21. Endophallus and temones of M. lassallei Keith, 2010, holotype (DKC) in lateral view. 22. Female genitalia of M. scutata, non-type (MFNB) in dorsal view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770322" httpUri="https://zenodo.org/record/4770322/files/figure.png" pageId="13" pageNumber="14">Fig. 20</figureCitation>
) consisting of one dorsal branch semi-tubular with proximal half obturated ventrally by a strong membrane; sutural scar absent; apex varying in shapes. Collum absent. Ventral branches thin, fused to dorsal one, emerging from edge of ventral obturating membrane toward distal end. Endophallus (
<figureCitation id="1327CAD0A6380627FE99F927FE58E299" box="[354,442,1678,1704]" captionStart="Figs 20–22" captionStartId="14.[189,241,1251,1277]" captionTargetBox="[291,1324,285,1192]" captionTargetId="figure-142@14.[278,1337,265,1205]" captionTargetPageId="14" captionText="Figs 20–22. Morphotype VIII “Scutata”. 20. Male genitalia of M. scutata (Reitter, 1902), lectotype (MFNB), parameres in dorsal (A), ventral (B) and lateral (C) views. 21. Endophallus and temones of M. lassallei Keith, 2010, holotype (DKC) in lateral view. 22. Female genitalia of M. scutata, non-type (MFNB) in dorsal view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770322" httpUri="https://zenodo.org/record/4770322/files/figure.png" pageId="13" pageNumber="14">Fig. 21</figureCitation>
) with an area of conspicuous acicular setae (raspulae) and spines at distal end; a few spines isolated at middle of sac. Temones (
<figureCitation id="1327CAD0A6380627FCEEF918FC8FE2FA" box="[789,877,1713,1739]" captionStart="Figs 20–22" captionStartId="14.[189,241,1251,1277]" captionTargetBox="[291,1324,285,1192]" captionTargetId="figure-142@14.[278,1337,265,1205]" captionTargetPageId="14" captionText="Figs 20–22. Morphotype VIII “Scutata”. 20. Male genitalia of M. scutata (Reitter, 1902), lectotype (MFNB), parameres in dorsal (A), ventral (B) and lateral (C) views. 21. Endophallus and temones of M. lassallei Keith, 2010, holotype (DKC) in lateral view. 22. Female genitalia of M. scutata, non-type (MFNB) in dorsal view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770322" httpUri="https://zenodo.org/record/4770322/files/figure.png" pageId="13" pageNumber="14">Fig. 21</figureCitation>
) laid at distal part of endophallus, slender; apophysis separated.
</paragraph>
<paragraph id="8BA3D655A6380627FF46F8BFFE1BE301" blockId="13.[189,505,1813,1840]" box="[189,505,1813,1840]" pageId="13" pageNumber="14">
<emphasis id="B9680A47A6380627FF46F8BFFE6EE31E" bold="true" box="[189,396,1813,1840]" pageId="13" pageNumber="14">Female genitalia</emphasis>
(
<figureCitation id="1327CAD0A6380627FE60F8BCFE13E301" box="[411,497,1813,1840]" captionStart="Figs 20–22" captionStartId="14.[189,241,1251,1277]" captionTargetBox="[291,1324,285,1192]" captionTargetId="figure-142@14.[278,1337,265,1205]" captionTargetPageId="14" captionText="Figs 20–22. Morphotype VIII “Scutata”. 20. Male genitalia of M. scutata (Reitter, 1902), lectotype (MFNB), parameres in dorsal (A), ventral (B) and lateral (C) views. 21. Endophallus and temones of M. lassallei Keith, 2010, holotype (DKC) in lateral view. 22. Female genitalia of M. scutata, non-type (MFNB) in dorsal view. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770322" httpUri="https://zenodo.org/record/4770322/files/figure.png" pageId="13" pageNumber="14">Fig. 22</figureCitation>
)
</paragraph>
<paragraph id="8BA3D655A6380627FF46F8EDFC8DE3F9" blockId="13.[188,1399,1860,1992]" pageId="13" pageNumber="14">Anal fold with all vestigial tergites similar in size; gonopore fold with a pair of sensory plates in ovaltransverse shaped, plates with sensilla along internal margin; vestigial sternites weakly developed with a few sensillae. Median oviduct slightly developed with membranous epithelium. Bursa copulatrix with part of peduncle striated. Spermatheca with long peduncle.</paragraph>
<caption id="DF6386DDA63B0624FF46FB4AFD35E159" ID-DOI="http://doi.org/10.5281/zenodo.4770322" ID-Zenodo-Dep="4770322" httpUri="https://zenodo.org/record/4770322/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,241,1251,1277]" targetBox="[291,1324,285,1192]" targetPageId="14">
<paragraph id="8BA3D655A63B0624FF46FB4AFD35E159" blockId="14.[189,1399,1251,1384]" pageId="14" pageNumber="15">
<emphasis id="B9680A47A63B0624FF46FB4AFEAEE0CC" bold="true" box="[189,332,1251,1277]" pageId="14" pageNumber="15">Figs 20–22.</emphasis>
Morphotype VIII “
<emphasis id="B9680A47A63B0624FDC5FB4DFD74E0CF" box="[574,662,1252,1278]" italics="true" pageId="14" pageNumber="15">Scutata</emphasis>
”.
<emphasis id="B9680A47A63B0624FD4EFB4AFD30E0CC" bold="true" box="[693,722,1251,1277]" pageId="14" pageNumber="15">20</emphasis>
. Male genitalia of
<taxonomicName id="4C1CADD6A63B0624FC44FB4DFB1EE0CF" authority="(Reitter, 1902)" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Reitter" baseAuthorityYear="1902" box="[959,1276,1251,1278]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A63B0624FC44FB4DFBDDE0CC" box="[959,1087,1252,1277]" italics="true" pageId="14" pageNumber="15">M. scutata</emphasis>
(
<bibRefCitation id="EF8DABA4A63B0624FBAAFB4AFB11E0CF" author="Reitter E." box="[1105,1267,1251,1278]" pageId="14" pageNumber="15" pagination="93 - 303" refId="ref45541" refString="Reitter E. 1902. Bestimmungs-Tabelle der Melolonthidae aus der europaischen Fauna und den angrenzenden Landern, enthaltend die Gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn 40: 93 - 303." type="journal article" year="1902">Reitter, 1902</bibRefCitation>
)
</taxonomicName>
, lectotype (MFNB), parameres in dorsal (A), ventral (B) and lateral (C) views.
<emphasis id="B9680A47A63B0624FC0DFAAFFBF1E111" bold="true" box="[1014,1043,1286,1312]" pageId="14" pageNumber="15">21</emphasis>
. Endophallus and temones of
<taxonomicName id="4C1CADD6A63B0624FF44FA82FE3FE175" authority="Keith, 2010" authorityName="Keith" authorityYear="2010" box="[191,477,1322,1348]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="lassallei">
<emphasis id="B9680A47A63B0624FF44FA82FEABE175" box="[191,329,1322,1348]" italics="true" pageId="14" pageNumber="15">M. lassallei</emphasis>
Keith, 2010
</taxonomicName>
, holotype (DKC) in lateral view.
<emphasis id="B9680A47A63B0624FC91FA83FC65E175" bold="true" box="[874,903,1322,1348]" pageId="14" pageNumber="15">22</emphasis>
. Female genitalia of
<taxonomicName id="4C1CADD6A63B0624FB78FA82FB1DE175" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Reitter" baseAuthorityYear="1902" box="[1155,1279,1323,1348]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="B9680A47A63B0624FB78FA82FB1DE175" box="[1155,1279,1323,1348]" italics="true" pageId="14" pageNumber="15">M. scutata</emphasis>
</taxonomicName>
, non-type (MFNB) in dorsal view. Scale bars = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BA3D655A63B0624FDA3FA44FC3EE237" blockId="14.[600,988,1516,1578]" box="[600,988,1516,1543]" pageId="14" pageNumber="15">
<heading id="D0EB6139A63B0624FDA3FA44FC3EE237" box="[600,988,1516,1543]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="15" reason="2">
<emphasis id="B9680A47A63B0624FDA3FA44FC3EE237" bold="true" box="[600,988,1516,1543]" pageId="14" pageNumber="15">
Morphotype IX “
<emphasis id="B9680A47A63B0624FCC9FA45FC2FE236" bold="true" box="[818,973,1516,1543]" italics="true" pageId="14" pageNumber="15">Ciliatipennis</emphasis>
”
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A63B0624FD0BF9B9FCA1E21B" blockId="14.[600,988,1516,1578]" box="[752,835,1552,1578]" pageId="14" pageNumber="15">
<figureCitation id="1327CAD0A63B0624FD0BF9B9FCA1E21B" box="[752,835,1552,1578]" captionStart="Fig" captionStartId="15.[191,234,940,966]" captionTargetBox="[279,1310,277,894]" captionTargetId="figure-236@15.[264,1323,262,907]" captionTargetPageId="15" captionText="Fig. 23. Morphotype IX “Ciliatipennis”. Male genitalia of M. ciliatipennis (Moser, 1903), lectotype (MFNB). Parameres in dorsal (A), ventral (B) and lateral (C) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770326" httpUri="https://zenodo.org/record/4770326/files/figure.png" pageId="14" pageNumber="15">Fig. 23</figureCitation>
</paragraph>
<paragraph id="8BA3D655A63B0624FF46F9FBFE92E25A" blockId="14.[189,368,1617,1644]" box="[189,368,1617,1644]" pageId="14" pageNumber="15">
<heading id="D0EB6139A63B0624FF46F9FBFE92E25A" bold="true" box="[189,368,1617,1644]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B9680A47A63B0624FF46F9FBFE92E25A" bold="true" box="[189,368,1617,1644]" pageId="14" pageNumber="15">Male genitalia</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A63B0624FF46F929FC83E35F" blockId="14.[188,1399,1664,1902]" pageId="14" pageNumber="15">
Parameres (
<figureCitation id="1327CAD0A63B0624FEB3F929FE7CE2AB" box="[328,414,1664,1690]" captionStart="Fig" captionStartId="15.[191,234,940,966]" captionTargetBox="[279,1310,277,894]" captionTargetId="figure-236@15.[264,1323,262,907]" captionTargetPageId="15" captionText="Fig. 23. Morphotype IX “Ciliatipennis”. Male genitalia of M. ciliatipennis (Moser, 1903), lectotype (MFNB). Parameres in dorsal (A), ventral (B) and lateral (C) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770326" httpUri="https://zenodo.org/record/4770326/files/figure.png" pageId="14" pageNumber="15">Fig. 23</figureCitation>
) consisting of one strong dorsal tubular complex divided into two parts, both parts with two branches at distal end; upper part with branches (
<figureCitation id="1327CAD0A63B0624FC6BF90AFC1DE28C" box="[912,1023,1699,1725]" captionStart="Fig" captionStartId="15.[191,234,940,966]" captionTargetBox="[279,1310,277,894]" captionTargetId="figure-236@15.[264,1323,262,907]" captionTargetPageId="15" captionText="Fig. 23. Morphotype IX “Ciliatipennis”. Male genitalia of M. ciliatipennis (Moser, 1903), lectotype (MFNB). Parameres in dorsal (A), ventral (B) and lateral (C) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770326" httpUri="https://zenodo.org/record/4770326/files/figure.png" pageId="14" pageNumber="15">Fig. 23A</figureCitation>
) varying in shapes and length; branches of lower part (
<figureCitation id="1327CAD0A63B0624FE2AF96FFDDBE2D1" box="[465,569,1734,1760]" captionStart="Fig" captionStartId="15.[191,234,940,966]" captionTargetBox="[279,1310,277,894]" captionTargetId="figure-236@15.[264,1323,262,907]" captionTargetPageId="15" captionText="Fig. 23. Morphotype IX “Ciliatipennis”. Male genitalia of M. ciliatipennis (Moser, 1903), lectotype (MFNB). Parameres in dorsal (A), ventral (B) and lateral (C) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770326" httpUri="https://zenodo.org/record/4770326/files/figure.png" pageId="14" pageNumber="15">Fig. 23B</figureCitation>
) shorter than those of dorsal part, convergent at distal end with varying shapes tips. In lateral view, branches of dorsal complex form a concavity (
<figureCitation id="1327CAD0A63B0624FBDCF940FB72E335" box="[1063,1168,1769,1796]" captionStart="Fig" captionStartId="15.[191,234,940,966]" captionTargetBox="[279,1310,277,894]" captionTargetId="figure-236@15.[264,1323,262,907]" captionTargetPageId="15" captionText="Fig. 23. Morphotype IX “Ciliatipennis”. Male genitalia of M. ciliatipennis (Moser, 1903), lectotype (MFNB). Parameres in dorsal (A), ventral (B) and lateral (C) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770326" httpUri="https://zenodo.org/record/4770326/files/figure.png" pageId="14" pageNumber="15">Fig. 23C</figureCitation>
) varying in shapes. Collum absent. Ventral branches reduced, fused laterally to dorsal tubular complex with a weak lateral scar. Endophallus with short spines and pointed sensilla at middle (
<bibRefCitation id="EF8DABA4A63B0624FC1AF899FB42E37B" author="Gao C. - B. &amp; Bai M. &amp; Fang H. &amp; Yu Z. - G." box="[993,1184,1840,1866]" pageId="14" pageNumber="15" pagination="1 - 20" refId="ref44032" refString="Gao C. - B., Bai M., Fang H. &amp; Yu Z. - G. 2018. Four new species of the genus Miridiba Reitter (Coleoptera: Scarabaeidae: Melolonthinae) from China. Zootaxa 4527 (1): 1 - 20. https: // doi. org / 10.11646 / zootaxa. 4527.1.1" type="journal article" year="2018">
Gao
<emphasis id="B9680A47A63B0624FBE5F898FBB3E37B" box="[1054,1105,1840,1866]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. 2018
</bibRefCitation>
: fig. 3j). Temones slender and apophysis separated (
<bibRefCitation id="EF8DABA4A63B0624FDBDF8FAFCE2E35C" author="Gao C. - B. &amp; Bai M. &amp; Fang H. &amp; Yu Z. - G." box="[582,768,1875,1902]" pageId="14" pageNumber="15" pagination="1 - 20" refId="ref44032" refString="Gao C. - B., Bai M., Fang H. &amp; Yu Z. - G. 2018. Four new species of the genus Miridiba Reitter (Coleoptera: Scarabaeidae: Melolonthinae) from China. Zootaxa 4527 (1): 1 - 20. https: // doi. org / 10.11646 / zootaxa. 4527.1.1" type="journal article" year="2018">
Gao
<emphasis id="B9680A47A63B0624FD84F8FDFD51E35C" box="[639,691,1875,1901]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. 2018
</bibRefCitation>
: fig. 3j).
</paragraph>
<paragraph id="8BA3D655A63B0624FF46F83CFE6EE39F" blockId="14.[189,396,1940,1967]" box="[189,396,1940,1967]" pageId="14" pageNumber="15">
<heading id="D0EB6139A63B0624FF46F83CFE6EE39F" bold="true" box="[189,396,1940,1967]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B9680A47A63B0624FF46F83CFE6EE39F" bold="true" box="[189,396,1940,1967]" pageId="14" pageNumber="15">Female genitalia</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A63B0624FF46F86DFEAFE3EC" blockId="14.[189,333,1987,2014]" box="[189,333,1987,2014]" pageId="14" pageNumber="15">Not studied.</paragraph>
<caption id="DF6386DDA63A0625FF44FC05FB3EE7DB" ID-DOI="http://doi.org/10.5281/zenodo.4770326" ID-Zenodo-Dep="4770326" httpUri="https://zenodo.org/record/4770326/files/figure.png" pageId="15" pageNumber="16" startId="15.[191,234,940,966]" targetBox="[279,1310,277,894]" targetPageId="15">
<paragraph id="8BA3D655A63A0625FF44FC05FB3EE7DB" blockId="15.[189,1399,940,1002]" pageId="15" pageNumber="16">
<emphasis id="B9680A47A63A0625FF44FC05FEFDE7F6" bold="true" box="[191,287,940,967]" pageId="15" pageNumber="16">Fig. 23.</emphasis>
Morphotype IX “
<emphasis id="B9680A47A63A0625FE02FC04FD73E7F6" box="[505,657,941,967]" italics="true" pageId="15" pageNumber="16">Ciliatipennis</emphasis>
”. Male genitalia of
<taxonomicName id="4C1CADD6A63A0625FC72FC07FB1EE7F6" authority="(Moser, 1903)" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Moser" baseAuthorityYear="1903" box="[905,1276,941,967]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ciliatipennis">
<emphasis id="B9680A47A63A0625FC72FC07FBA6E7F6" box="[905,1092,941,967]" italics="true" pageId="15" pageNumber="16">M. ciliatipennis</emphasis>
(Moser, 1903)
</taxonomicName>
, lectotype (MFNB). Parameres in dorsal (A), ventral (B) and lateral (C) views. Scale bars = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BA3D655A63A0625FF46FB8BFCA7E00E" blockId="15.[189,837,1058,1087]" box="[189,837,1058,1087]" pageId="15" pageNumber="16">
<emphasis id="B9680A47A63A0625FF46FB8BFCA7E00E" bold="true" box="[189,837,1058,1087]" italics="true" pageId="15" pageNumber="16">
Species included in Morphotype I “
<taxonomicName id="4C1CADD6A63A0625FD74FB8BFCD5E00E" box="[655,823,1058,1087]" class="Insecta" family="Tachinidae" genus="Trichophora" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Trichophora</taxonomicName>
”
</emphasis>
</paragraph>
<paragraph id="8BA3D655A63A0625FF46FBFFFCE1E086" blockId="15.[189,1399,1110,1207]" pageId="15" pageNumber="16">The group is composed of twenty-nine species, which are characterized by antenna 9-segmented, head and pronotal surface densely punctate (distance between punctures smaller than diameter of puncture) and male and female genitalia as specified above.</paragraph>
</subSubSection>
</treatment>
</document>
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