343 lines
44 KiB
XML
343 lines
44 KiB
XML
<document id="6A0FEDB6CE6E35F1AEB89C3A8F23C3BF" ID-CLB-Dataset="145796" ID-DOI="10.11646/zootaxa.5188.1.2" ID-GBIF-Dataset="3355f15e-548c-438e-a3bb-d898a59dff7e" ID-ISSN="1175-5326" ID-Zenodo-Dep="7087337" ID-ZooBank="A9418E18-930F-42F9-AE89-408EDC27470B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1663360990953" checkinUser="plazi" docAuthor="Pham, Phong Huy" docDate="2022" docId="0878879BFFC6FF93FF255195FD56268C" docLanguage="en" docName="zootaxa.5188.1.2.pdf" docOrigin="Zootaxa 5188 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dasyproctus vietnamensis Pham 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="53" masterDocId="F441FFE3FFC1FF99FFB25714FF8F2409" masterDocTitle="Taxonomic studies on Dasyproctus Lepeletier & Brullé (Hymenoptera: Crabronidae) from North Vietnam with description of two new species" masterLastPageNumber="54" masterPageNumber="43" pageNumber="50" updateTime="1699381742709" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="BBEEA4F8A15190905606EBEFEE358717">
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<mods:title id="C868221BC8D0A460430777C9DAE552F7">Taxonomic studies on Dasyproctus Lepeletier & Brullé (Hymenoptera: Crabronidae) from North Vietnam with description of two new species</mods:title>
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<mods:name id="C7210B9EBE750995F8461D413D412451" type="personal">
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<mods:namePart id="722B63DD872D8003EAAF3D667E6BD0A5">Pham, Phong Huy</mods:namePart>
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<mods:nameIdentifier id="B151A4A357D275F544D97BCFE27A1BEC" type="ORCID">0000-0001-5350-3865</mods:nameIdentifier>
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<mods:affiliation id="FA2A25728864295EAAE44C3B4CE68A79">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam. phong. wasp @ gmail. com; https: // orcid. org / 0000 - 0001 - 5350 - 3865</mods:affiliation>
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<mods:nameIdentifier id="93DE804C9247CB6BA4C11C1D46C5788C" type="email">phong.wasp@gmail.com</mods:nameIdentifier>
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<mods:title id="E03FDB1C08A4E33D795ABCEEFC5FE4A8">Zootaxa</mods:title>
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<mods:part id="C4549EBF04A3B6919E12F5E076D39229">
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<mods:date id="05CBDEBD2D5D9C177C2743B6774FB681">2022</mods:date>
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<mods:number id="3A2209DCCF96E2CEA771CDF563C992C8">2022-09-15</mods:number>
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<mods:number id="7A731093457F2A7B75B0D187B985D7B5">5188</mods:number>
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<treatment id="0878879BFFC6FF93FF255195FD56268C" ID-DOI="http://doi.org/10.5281/zenodo.7087367" ID-GBIF-Taxon="200635745" ID-Zenodo-Dep="7087367" LSID="urn:lsid:plazi:treatment:0878879BFFC6FF93FF255195FD56268C" httpUri="http://treatment.plazi.org/id/0878879BFFC6FF93FF255195FD56268C" lastPageId="10" lastPageNumber="53" pageId="7" pageNumber="50">
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<heading id="DB2681E1FFC6FF9EFF255195FD242292" bold="true" box="[151,683,1665,1691]" fontSize="11" level="1" pageId="7" pageNumber="50" reason="1">
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<emphasis id="B2A5EA9FFFC6FF9EFF255195FD242292" bold="true" box="[151,683,1665,1691]" pageId="7" pageNumber="50">
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7.
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<taxonomicName id="47D14D0EFFC6FF9EFF075195FDCE2292" authority="Pham" authorityName="Pham" authorityYear="2022" box="[181,577,1665,1691]" class="Insecta" family="Crabronidae" genus="Dasyproctus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="vietnamensis" status="sp. nov.">
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<emphasis id="B2A5EA9FFFC6FF9EFF075195FE612292" bold="true" box="[181,494,1665,1691]" italics="true" pageId="7" pageNumber="50">Dasyproctus vietnamensis</emphasis>
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Pham
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,
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<taxonomicNameLabel id="A99657E4FFC6FF9EFDFC5195FD242292" box="[590,683,1665,1691]" pageId="7" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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<subSubSection id="C8CB6506FFC6FF9EFF2551B1FEA822B6" box="[151,295,1701,1727]" pageId="7" pageNumber="50" type="description">
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<paragraph id="806E368DFFC6FF9EFF2551B1FEA822B6" blockId="7.[151,683,1665,1727]" box="[151,295,1701,1727]" pageId="7" pageNumber="50">
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(
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<figureCitation id="18EA2A08FFC6FF9EFF2D51B1FEAF22B6" box="[159,288,1701,1727]" captionStart-0="FIGURES 49–54" captionStart-1="FIGURES 55–61" captionStartId-0="9.[152,270,1818,1842]" captionStartId-1="10.[152,270,1346,1370]" captionTargetBox-0="[152,1425,1172,1795]" captionTargetBox-1="[151,1428,669,1323]" captionTargetId-0="figure-468@9.[152,1436,1172,1795]" captionTargetId-1="figure-212@10.[151,1436,669,1323]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 49–54. Dasyproctus vietnamensis Pham sp. nov. Holotype ♀. 49. Head, frontal view; 50. Mesosoma, lateral view; 51. Metasoma, dorsal view; 52. Head, dorsal view; 53. Mesosoma, dorsal view; 54. Habitus, lateral view." captionText-1="FIGURES 55–61. Dasyproctus vietnamensis Pham sp. nov. Paratype ♂. 55. Head, frontal view; 56. Mesosoma, dorsal view; 57. Metasoma, dorsal view; 58. Head, dorsal view; 59. Mesosoma, lateral view; 60. Genitalia, ventral view; 61. Habitus, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.7087365" figureDoi-1="http://doi.org/10.5281/zenodo.7087369" httpUri-0="https://zenodo.org/record/7087365/files/figure.png" httpUri-1="https://zenodo.org/record/7087369/files/figure.png" pageId="7" pageNumber="50">Figs 49–61</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C8CB6506FFC6FF9EFF2551F9FE7323B5" pageId="7" pageNumber="50" type="materials_examined">
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<emphasis id="B2A5EA9FFFC6FF9EFF2551F9FEF1230E" bold="true" box="[151,382,1773,1799]" pageId="7" pageNumber="50">Material examined.</emphasis>
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,
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, pinned (deposited in
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),
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<collectingCountry id="F8C6761DFFC6FF9EFCE251F9FC59230E" box="[848,982,1773,1799]" name="Vietnam" pageId="7" pageNumber="50">VIETNAM</collectingCountry>
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:
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:
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,
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,
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,
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<collectingMethod id="59904E9AFFC6FF9EFF255005FEA72325" box="[151,296,1809,1836]" pageId="7" pageNumber="50">Malaise trap</collectingMethod>
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,
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<collectorName id="2D24535BFFC6FF9EFE845005FD9C2322" box="[310,531,1809,1835]" pageId="7" pageNumber="50">Coll. Mai Phu Quy</collectorName>
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[
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.
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(
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):
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<collectingCountry id="F8C6761DFFC6FF9EFCE95005FC6E2322" box="[859,993,1809,1835]" name="Vietnam" pageId="7" pageNumber="50">VIETNAM</collectingCountry>
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:
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<collectingRegion id="4215F86FFFC6FF9EFC5F5005FBE62322" box="[1005,1129,1809,1835]" country="Vietnam" name="Vinh Phuc" pageId="7" pageNumber="50">Vinh Phuc</collectingRegion>
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:
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<materialsCitation id="30B93CD0FFC6FF9EFBC55005FB162359" ID-GBIF-Occurrence="3909206334" collectingDate="2018-06-02" collectionCode="IEBR" collectorName="Coll. Phong Huy Pham" country="Vietnam" county="Me Linh Station" location="Ngoc Thanh" municipality="Biodiversity" pageId="7" pageNumber="50" specimenCount="1" specimenCount-female="1" stateProvince="Vinh Phuc" typeStatus="paratype">
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,
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<collectingCounty id="690F4E01FFC6FF9EFB075006FAFE2322" box="[1205,1393,1809,1836]" pageId="7" pageNumber="50">Me Linh Station</collectingCounty>
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for
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<collectingMunicipality id="600AACF7FFC6FF9EFF255021FEAC2346" box="[151,291,1845,1871]" pageId="7" pageNumber="50">Biodiversity</collectingMunicipality>
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,
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,
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<location id="850E6056FFC6FF9EFE7A5021FDBA2359" LSID="urn:lsid:plazi:treatment:0878879BFFC6FF93FF255195FD56268C:850E6056FFC6FF9EFE7A5021FDBA2359" box="[456,565,1845,1872]" country="Vietnam" county="Me Linh Station" municipality="Biodiversity" name="Phuc Yen" pageId="7" pageNumber="50" stateProvince="Vinh Phuc">Phuc Yen</location>
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,
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,
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<collectorName id="2D24535BFFC6FF9EFC845021FBB32346" box="[822,1084,1845,1872]" pageId="7" pageNumber="50">Coll. Phong Huy Pham</collectorName>
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[
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<collectionCode id="E6C0AE48FFC6FF9EFBF85022FB1D2359" box="[1098,1170,1846,1872]" pageId="7" pageNumber="50">IEBR</collectionCode>
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]
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</materialsCitation>
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.
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<materialsCitation id="30B93CD0FFC6FF9EFB165022FC28237D" ID-GBIF-Occurrence="3909206336" collectingDate="2012-08-16" collectorName="Coll. Hoa Thi Dang" country="Vietnam" county="Bai" location="Long Bien" municipality="Song Hong" pageId="7" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Ha Noi" typeStatus="paratype">
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<collectingRegion id="4215F86FFFC6FF9EFB165022FB752346" box="[1188,1274,1845,1872]" country="Vietnam" name="Ha Noi" pageId="7" pageNumber="50">Ha Noi</collectingRegion>
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:
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<specimenCount id="96D7FD04FFC6FF9EFAB75021FABE2359" box="[1285,1329,1845,1872]" pageId="7" pageNumber="50" type="male">1 ♂</specimenCount>
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,
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<collectingCounty id="690F4E01FFC6FF9EFA8C5021FAEA2346" box="[1342,1381,1845,1871]" pageId="7" pageNumber="50">Bai</collectingCounty>
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giua
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<collectingMunicipality id="600AACF7FFC6FF9EFF25504DFE95237D" box="[151,282,1881,1908]" pageId="7" pageNumber="50">Song Hong</collectingMunicipality>
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,
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<location id="850E6056FFC6FF9EFE94504EFE2E237A" LSID="urn:lsid:plazi:treatment:0878879BFFC6FF93FF255195FD56268C:850E6056FFC6FF9EFE94504EFE2E237A" box="[294,417,1881,1908]" country="Vietnam" county="Bai" municipality="Song Hong" name="Long Bien" pageId="7" pageNumber="50" stateProvince="Ha Noi">Long Bien</location>
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,
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<date id="F46F104DFFC6FF9EFE1F504DFDB9237A" box="[429,566,1881,1907]" pageId="7" pageNumber="50" value="2012-08-16">
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<collectingDate id="E42BE9A5FFC6FF9EFE1F504DFDB9237A" box="[429,566,1881,1907]" pageId="7" pageNumber="50" value="2012-08-16">16.viii.2012</collectingDate>
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,
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<collectingMethod id="59904E9AFFC6FF9EFDF1504EFD39237D" box="[579,694,1882,1908]" pageId="7" pageNumber="50">sweep net</collectingMethod>
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,
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<collectorName id="2D24535BFFC6FF9EFD70504DFC28237D" box="[706,935,1881,1908]" pageId="7" pageNumber="50">Coll. Hoa Thi Dang</collectorName>
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</materialsCitation>
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;
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<materialsCitation id="30B93CD0FFC6FF9EFC01504DFD172391" ID-GBIF-Occurrence="3909206313" collectingDate="2002-07-05" collectingDateMax="2002-07-15" collectingDateMin="2002-07-05" collectionCode="IEBR" collectorName="Coll. Long Dang Khuat" country="Vietnam" location="Thach That" municipality="Hoa Lac" pageId="7" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Ha Noi" typeStatus="paratype">
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<specimenCount id="96D7FD04FFC6FF9EFC01504DFC6F237D" box="[947,992,1881,1908]" pageId="7" pageNumber="50" type="male">1 ♂</specimenCount>
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,
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<collectingMunicipality id="600AACF7FFC6FF9EFC5F504EFBDE237D" box="[1005,1105,1882,1908]" pageId="7" pageNumber="50">Hoa Lac</collectingMunicipality>
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,
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,
|
||
<date id="F46F104DFFC6FF9EFB5E504DFA17237D" box="[1260,1432,1881,1908]" pageId="7" pageNumber="50" value="2002-07-05" valueMax="2002-07-15" valueMin="2002-07-05">
|
||
<collectingDate id="E42BE9A5FFC6FF9EFB5E504DFA17237D" box="[1260,1432,1881,1908]" pageId="7" pageNumber="50" value="2002-07-05" valueMax="2002-07-15" valueMin="2002-07-05">05–15.vii.2002</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="59904E9AFFC6FF9EFF255069FEAA2391" box="[151,293,1917,1944]" pageId="7" pageNumber="50">Malaise trap</collectingMethod>
|
||
,
|
||
<collectorName id="2D24535BFFC6FF9EFE835069FDB5239E" box="[305,570,1917,1944]" pageId="7" pageNumber="50">Coll. Long Dang Khuat</collectorName>
|
||
[
|
||
<collectionCode id="E6C0AE48FFC6FF9EFDFB506AFD1E2391" box="[585,657,1918,1944]" pageId="7" pageNumber="50">IEBR</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="30B93CD0FFC6FF9EFD115069FE7723B5" ID-GBIF-Occurrence="3909206339" collectingDate="2017-06-23" collectionCode="IEBR" collectorName="Thung Den Tran & Coll. Phong Huy Pham" country="Vietnam" location="Trang An" pageId="7" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Ninh Binh" typeStatus="paratype">
|
||
<collectingRegion id="4215F86FFFC6FF9EFD115069FC92239E" box="[675,797,1917,1943]" country="Vietnam" name="Ninh Binh" pageId="7" pageNumber="50">Ninh Binh</collectingRegion>
|
||
:
|
||
<specimenCount id="96D7FD04FFC6FF9EFC9A5069FCDB2391" box="[808,852,1917,1944]" pageId="7" pageNumber="50" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectorName id="2D24535BFFC6FF9EFCD25069FB922391" box="[864,1053,1917,1944]" pageId="7" pageNumber="50">Thung Den Tran</collectorName>
|
||
,
|
||
<location id="850E6056FFC6FF9EFB95506AFB1B239E" LSID="urn:lsid:plazi:treatment:0878879BFFC6FF93FF255195FD56268C:850E6056FFC6FF9EFB95506AFB1B239E" box="[1063,1172,1917,1944]" country="Vietnam" name="Trang An" pageId="7" pageNumber="50" stateProvince="Ninh Binh">Trang An</location>
|
||
,
|
||
<date id="F46F104DFFC6FF9EFB125069FA96239E" box="[1184,1305,1917,1943]" pageId="7" pageNumber="50" value="2017-06-23">
|
||
<collectingDate id="E42BE9A5FFC6FF9EFB125069FA96239E" box="[1184,1305,1917,1943]" pageId="7" pageNumber="50" value="2017-06-23">23.vi.2017</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="59904E9AFFC6FF9EFA94506AFA172391" box="[1318,1432,1918,1944]" pageId="7" pageNumber="50">sweep net</collectingMethod>
|
||
,
|
||
<collectorName id="2D24535BFFC6FF9EFF2550B5FE1423B2" box="[151,411,1953,1980]" pageId="7" pageNumber="50">Coll. Phong Huy Pham</collectorName>
|
||
[
|
||
<collectionCode id="E6C0AE48FFC6FF9EFE1B50B6FE7E23B5" box="[425,497,1954,1980]" pageId="7" pageNumber="50">IEBR</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="D4AE6605FFC9FF91FF2A545AFB762782" ID-DOI="http://doi.org/10.5281/zenodo.7087363" ID-Zenodo-Dep="7087363" httpUri="https://zenodo.org/record/7087363/files/figure.png" pageId="8" pageNumber="51" startId="8.[152,270,846,870]" targetBox="[151,1431,181,822]" targetPageId="8">
|
||
<paragraph id="806E368DFFC9FF91FF2A545AFB762782" blockId="8.[151,1436,846,907]" pageId="8" pageNumber="51">
|
||
<emphasis id="B2A5EA9FFFC9FF91FF2A545AFED6276F" bold="true" box="[152,345,846,870]" pageId="8" pageNumber="51">FIGURES 42–48.</emphasis>
|
||
<taxonomicName id="47D14D0EFFC9FF91FEEC545BFD1A276E" authority="Leclercq" authorityName="Leclercq" authorityYear="1956" box="[350,661,847,871]" class="Insecta" family="Crabronidae" genus="Dasyproctus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="51" phylum="Arthropoda" rank="species" species="pentheri">
|
||
<emphasis id="B2A5EA9FFFC9FF91FEEC545BFDB9276E" box="[350,566,847,871]" italics="true" pageId="8" pageNumber="51">Dasyproctus pentheri</emphasis>
|
||
Leclercq
|
||
</taxonomicName>
|
||
♂. 42. Head, frontal view; 43.
|
||
<taxonomicName id="47D14D0EFFC9FF91FC7B545BFBB6276F" box="[969,1081,847,870]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="53" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
|
||
, lateral view; 44. Metasoma, dorsal view; 45. Head, dorsal view; 46. Metasoma, dorsal view; 47. Genitalia, ventral view; 48. Habitus, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C8CB6506FFC9FF91FF7554ACFCFE23F2" pageId="8" pageNumber="51" type="diagnosis">
|
||
<paragraph id="806E368DFFC9FF91FF7554ACFAEC2175" blockId="8.[151,1437,952,2043]" pageId="8" pageNumber="51">
|
||
<emphasis id="B2A5EA9FFFC9FF91FF7554ACFECD27DB" bold="true" box="[199,322,952,978]" pageId="8" pageNumber="51">Diagnosis.</emphasis>
|
||
The new species can be easily distinguished from its congeners by the following combination of characters: in females: head rectangular in frontal view; free margin of clypeus slightly indented medially; scape yellow with small black spot on inner lateral side; mandible yellow basally, reddish to black apically; IOC conspicuous; vertex with sparse, small punctures, and longitudinal, shallow pit near inner eye margin; anterior carina of pronotal collar laterally bending towards insertion of fore coxa; mesopleuron with yellow spot, scutellum with large yellow band medially; propodeum with Y-shaped furrow posteriorly; hind tibia yellow postero-dorsally except at apex, with hook-shaped carina postero-dorsally, and brown macula antero-dorsally; tarsomere 1 yellow basally; T1 immaculate; T2 with large, round depression at base; pygidial plate reddish brown, T5 and pygidial plate with brown-yellow setae. In males: IOC present; clypeus with median carina, free margin truncate medially; gena without striae; frons densely punctate; flagellomeres II‒IV cylindrical; mandible entirely black except small yellow spot at base; scutum distinctly sparsely punctate; axilla and two spots on scutellum yellow; T1 immaculate, longer than hind femur, and shorter than T2‒T4 combined, its sides not parallel; T2 with large, round depression at base; T2‒T5 largely, brightly yellow maculate; T6 with largely, brightly yellow band anteriorly. T2‒T7 impunctate.
|
||
</paragraph>
|
||
<paragraph id="806E368DFFC9FF91FF755291FCFE23F2" blockId="8.[151,1437,952,2043]" pageId="8" pageNumber="51">
|
||
The new species runs close to
|
||
<taxonomicName id="47D14D0EFFC9FF91FD955292FD3D21A9" authority="Ma" authorityName="Ma" authorityYear="1936" box="[551,690,1414,1440]" class="Insecta" family="Crabronidae" genus="Dasyproctus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="51" phylum="Arthropoda" rank="species" species="jungi">
|
||
<emphasis id="B2A5EA9FFFC9FF91FD955292FD0A2196" box="[551,645,1414,1439]" italics="true" pageId="8" pageNumber="51">D. jungi</emphasis>
|
||
Ma
|
||
</taxonomicName>
|
||
in the key to Asian and Oceanic
|
||
<taxonomicName id="47D14D0EFFC9FF91FB815292FAF121A9" authority="(Leclercq 2015)" baseAuthorityName="Leclercq" baseAuthorityYear="2015" box="[1075,1406,1413,1440]" class="Insecta" family="Crabronidae" genus="Dasyproctus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="51" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2A5EA9FFFC9FF91FB815292FB4F2196" box="[1075,1216,1414,1439]" italics="true" pageId="8" pageNumber="51">Dasyproctus</emphasis>
|
||
(
|
||
<bibRefCitation id="E4404B7CFFC9FF91FB625291FAF921A9" author="Leclercq, J." box="[1232,1398,1413,1440]" pageId="8" pageNumber="51" pagination="41 - 89" refId="ref8557" refString="Leclercq, J. (2015) Hymenopteres Crabroniens d'Afrique, d'Asie et d'Oceanie du genre Dasyproctus Lepeletier & Brulle 1835 (Hymenoptera: Crabronidae: Crabroninae). Entomologie Faunistique, 68, 41 - 89." type="journal article" year="2015">Leclercq 2015</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in having, in females: vertex with sparse, small punctures, and longitudinal pit near inner eye margin; gena sparsely punctate, with dense short silvery setae; mandible yellow basally; anterior carina of pronotal collar laterally bending towards insertion of fore coxa, medially parallel to anterior margin of scutum; scutum with dense, small punctures; pronotal collar distinctly notched medially; metanotum with longitudinal striae mixed with small punctures; tegula testaceous; propodeum with coarse, oblique rugae dorsally, arched striae laterally, Y-shaped furrow on posterior aspect; T1 immaculate; T2 with large, round depression at base; T2‒T4 with largely, brightly yellow maculae. In males: clypeal with free margin truncate medially; mandible sometimes with small yellow spot at base; IOC distinct; gena sparsely punctate; flagellomeres II‒IV cylindrical; pedicel and flagellum black; scutum with sparse, small punctures; dorsal surface of fore tibia yellow; T1 longer than hind femur; T2 with large, round depression at base. However, it differs from the latter in having, in females: head rectangular; flagellum I brownish; mandible reddish to black apically; vertex with longitudinal, shallow pit near inner eye margin; hind tibia yellow postero-dorsally except apex, with hook-shaped carina postero-dorsally, and brown macula antero-dorsally; all tarsi brown except tarsomere 1 yellow basally; T2 with shallow depression basally; pygidial plate reddish brown; T5 and pygidial plate with brown-yellow setae. In males: mandible entirely black excepting small yellow spot at base; scape except dorsal surface, pronotal collar, pronotal lobe, dorsal surface of fore tibia yellow, yellow band on dorsal surface of mid and hind tibiae medially; petiole dorsally 2.5 × as long as wide; T2 with shallow depression basally; T5 with brightly yellow maculae, T6 with largely, brightly yellow band anteriorly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C8CB6506FFC8FF93FF755782FDD42634" lastPageId="10" lastPageNumber="53" pageId="9" pageNumber="52" type="description">
|
||
<paragraph id="806E368DFFC8FF90FF755782FAB624B8" blockId="9.[151,1437,150,1149]" box="[199,1337,150,177]" pageId="9" pageNumber="52">
|
||
<emphasis id="B2A5EA9FFFC8FF90FF755782FE3824B9" bold="true" box="[199,439,150,176]" pageId="9" pageNumber="52">Description. Female</emphasis>
|
||
,
|
||
<typeStatus id="5F6A882FFFC8FF90FE775782FDA824B9" box="[453,551,150,176]" pageId="9" pageNumber="52" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="18EA2A08FFC8FF90FD875782FD3524B8" box="[565,698,150,177]" captionStart="FIGURES 49–54" captionStartId="9.[152,270,1818,1842]" captionTargetBox="[152,1425,1172,1795]" captionTargetId="figure-468@9.[152,1436,1172,1795]" captionTargetPageId="9" captionText="FIGURES 49–54. Dasyproctus vietnamensis Pham sp. nov. Holotype ♀. 49. Head, frontal view; 50. Mesosoma, lateral view; 51. Metasoma, dorsal view; 52. Head, dorsal view; 53. Mesosoma, dorsal view; 54. Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7087365" httpUri="https://zenodo.org/record/7087365/files/figure.png" pageId="9" pageNumber="52">Figs 49‒54</figureCitation>
|
||
): Body length 8.5‒9.0 mm, forewing length 6.0‒
|
||
<quantity id="47299B68FFC8FF90FB6F5782FABA24B8" box="[1245,1333,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="9" pageNumber="52" unit="mm" value="6.3">6.3 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="806E368DFFC8FF90FF7557AEFE712581" blockId="9.[151,1437,150,1149]" pageId="9" pageNumber="52">
|
||
<emphasis id="B2A5EA9FFFC8FF90FF7557AEFE9224DC" bold="true" box="[199,285,186,213]" italics="true" pageId="9" pageNumber="52">Colour.</emphasis>
|
||
Body black with following yellow parts: scape except small black spot on inner lateral side; mandible except reddish to black apically; pronotal collar; pronotal lobe; large spot on mesopleuron; axilla; large median band on scutellum; brightly yellow bimaculae on T2‒T5; small apical spot on fore femur; large ventral band on mid femur; ventral surface of fore and mid tibiae; hind tibia except apical part and large spot antero-dorsally; tarsomere 1 basally. Brown to black-brown are: flagellum I, tegula, pedicel, pygidial plate, ventral surface of all tibiae, and all tarsi except tarsomere 1 basally.
|
||
</paragraph>
|
||
<paragraph id="806E368DFFC8FF90FF755686FB0E26C4" blockId="9.[151,1437,150,1149]" pageId="9" pageNumber="52">
|
||
<emphasis id="B2A5EA9FFFC8FF90FF755686FE8A25A4" bold="true" box="[199,261,402,429]" italics="true" pageId="9" pageNumber="52">Head</emphasis>
|
||
(
|
||
<figureCitation id="18EA2A08FFC8FF90FEAA5686FE3525A4" box="[280,442,402,429]" captionStart="FIGURES 49–54" captionStartId="9.[152,270,1818,1842]" captionTargetBox="[152,1425,1172,1795]" captionTargetId="figure-468@9.[152,1436,1172,1795]" captionTargetPageId="9" captionText="FIGURES 49–54. Dasyproctus vietnamensis Pham sp. nov. Holotype ♀. 49. Head, frontal view; 50. Mesosoma, lateral view; 51. Metasoma, dorsal view; 52. Head, dorsal view; 53. Mesosoma, dorsal view; 54. Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7087365" httpUri="https://zenodo.org/record/7087365/files/figure.png" pageId="9" pageNumber="52">Figs 49 & 52</figureCitation>
|
||
): In frontal view nearly rectangular, about 1.5 × as long as wide; mandible tridentate apically, with small tooth on inner margin; clypeus covered with dense silvery setae, slightly beveled, free margin slightly indented medially, blunt tooth on each side of free margin, longitudinal median carina reaching free margin; antennal sockets contiguous with one another and with inner eye margin; scape with two longitudinal, parallel carinae ventrally, length of scape 2.2 × distance between eyes at base of antennal socket; flagellum covered with thick short setae, flagellomere I as long as II, relative lengths of scape: pedicel: flagellomere I = 30: 5: 9; scapal basin with thick, adpressed silvery setae; IOC conspicuous; frons without furrow above IOC, densely punctate; OOD 1.18 × POD; vertex with sparse, small punctures, and well-imprinted supra-orbital fovea; frons and vertex covered with short brownish setae; gena sparsely punctate, with dense short silvery setae.
|
||
</paragraph>
|
||
<paragraph id="806E368DFFC8FF90FF7555C3FB9F27C1" blockId="9.[151,1437,150,1149]" pageId="9" pageNumber="52">
|
||
<taxonomicName id="47D14D0EFFC8FF90FF7555C3FECD26F8" box="[199,322,727,753]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2A5EA9FFFC8FF90FF7555C3FECD26F8" bold="true" box="[199,322,727,753]" italics="true" pageId="9" pageNumber="52">Mesosoma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="18EA2A08FFC8FF90FEE055C2FE6326F8" box="[338,492,726,753]" captionStart="FIGURES 49–54" captionStartId="9.[152,270,1818,1842]" captionTargetBox="[152,1425,1172,1795]" captionTargetId="figure-468@9.[152,1436,1172,1795]" captionTargetPageId="9" captionText="FIGURES 49–54. Dasyproctus vietnamensis Pham sp. nov. Holotype ♀. 49. Head, frontal view; 50. Mesosoma, lateral view; 51. Metasoma, dorsal view; 52. Head, dorsal view; 53. Mesosoma, dorsal view; 54. Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7087365" httpUri="https://zenodo.org/record/7087365/files/figure.png" pageId="9" pageNumber="52">Figs 50 & 53</figureCitation>
|
||
): Anterior carina of pronotal collar laterally bending towards insertion of fore coxa, medially parallel to anterior margin of scutum; pronotum conspicuously striate laterally; pronotal collar distinctly notched medially; scutum with dense, small punctures, and a series of short rugae posteriorly; scutellum and mesopleuron with sparse, fine, small punctures; metanotum with longitudinal striae mixed with small punctures; mesopleuron matt; metapleuron shining, with conspicuous, oblique rugae; propodeum with coarse, oblique rugae dorsally, arched striae laterally, Y-shaped furrow on posterior aspect.
|
||
<taxonomicName id="47D14D0EFFC8FF90FC28549FFB9C27AC" box="[922,1043,907,933]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="53" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
|
||
with white setae, sparse on scutum and denser on propodeum; hind tibia with hook-shaped carina postero-dorsally.
|
||
</paragraph>
|
||
<paragraph id="806E368DFFC8FF90FF7554C7FAA52074" blockId="9.[151,1437,150,1149]" pageId="9" pageNumber="52">
|
||
<emphasis id="B2A5EA9FFFC8FF90FF7554C7FEB027E4" bold="true" box="[199,319,979,1005]" italics="true" pageId="9" pageNumber="52">Metasoma</emphasis>
|
||
(
|
||
<figureCitation id="18EA2A08FFC8FF90FEE254C6FE6027E4" box="[336,495,978,1005]" captionStart="FIGURES 49–54" captionStartId="9.[152,270,1818,1842]" captionTargetBox="[152,1425,1172,1795]" captionTargetId="figure-468@9.[152,1436,1172,1795]" captionTargetPageId="9" captionText="FIGURES 49–54. Dasyproctus vietnamensis Pham sp. nov. Holotype ♀. 49. Head, frontal view; 50. Mesosoma, lateral view; 51. Metasoma, dorsal view; 52. Head, dorsal view; 53. Mesosoma, dorsal view; 54. Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7087365" httpUri="https://zenodo.org/record/7087365/files/figure.png" pageId="9" pageNumber="52">Figs 51 & 54</figureCitation>
|
||
): petiolate; T1 immaculate, with apical width about 2.6 × its basal width dorsally, length of T1 equal to that of hind femur, 1.4 × that of T2‒T4 combined; T2 with large, round depression at base; T1‒T5 with sparse, fine, small punctures; T2‒T5 with large maculae, these on T2 smaller than those on T3 and T4, these on T3 equal to those on T4; pygidial plate with coarse, dense punctures, conspicuously narrowed and gutterlike apically. Metasoma with whitish brown setae except T5 and pygidial plate with brown-yellow setae.
|
||
</paragraph>
|
||
<caption id="D4AE6605FFC8FF90FF2A500EFB4C235E" ID-DOI="http://doi.org/10.5281/zenodo.7087365" ID-Zenodo-Dep="7087365" httpUri="https://zenodo.org/record/7087365/files/figure.png" pageId="9" pageNumber="52" startId="9.[152,270,1818,1842]" targetBox="[152,1425,1172,1795]" targetPageId="9">
|
||
<paragraph id="806E368DFFC8FF90FF2A500EFB4C235E" blockId="9.[151,1436,1818,1879]" pageId="9" pageNumber="52">
|
||
<emphasis id="B2A5EA9FFFC8FF90FF2A500EFED4233A" bold="true" box="[152,347,1818,1843]" pageId="9" pageNumber="52">FIGURES 49–54.</emphasis>
|
||
<taxonomicName id="47D14D0EFFC8FF90FED3500FFD26233A" authority="Pham" authorityName="Pham" authorityYear="2022" box="[353,681,1819,1843]" class="Insecta" family="Crabronidae" genus="Dasyproctus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="52" phylum="Arthropoda" rank="species" species="vietnamensis" status="sp. nov.">
|
||
<emphasis id="B2A5EA9FFFC8FF90FED3500FFDE6233B" box="[353,617,1819,1842]" italics="true" pageId="9" pageNumber="52">Dasyproctus vietnamensis</emphasis>
|
||
Pham
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A99657E4FFC8FF90FD1D500FFC8D233B" box="[687,770,1819,1843]" pageId="9" pageNumber="52" rank="species">
|
||
<emphasis id="B2A5EA9FFFC8FF90FD1D500FFD74233A" bold="true" box="[687,763,1819,1843]" pageId="9" pageNumber="52">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
Holotype ♀. 49. Head, frontal view; 50.
|
||
<taxonomicName id="47D14D0EFFC8FF90FB16500FFA9B233B" box="[1188,1300,1819,1842]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="53" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
|
||
, lateral view; 51. Metasoma, dorsal view; 52. Head, dorsal view; 53.
|
||
<taxonomicName id="47D14D0EFFC8FF90FD75502BFCB8235F" box="[711,823,1855,1878]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="53" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
|
||
, dorsal view; 54. Habitus, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="806E368DFFC8FF90FF755091FB0B2396" blockId="9.[199,1436,1925,2023]" box="[199,1156,1925,1952]" pageId="9" pageNumber="52">
|
||
<emphasis id="B2A5EA9FFFC8FF90FF755091FE8B2396" bold="true" box="[199,260,1925,1951]" pageId="9" pageNumber="52">
|
||
<collectingRegion id="4215F86FFFC8FF90FF755091FE8B2396" box="[199,260,1925,1951]" country="Maldives" name="Maale" pageId="9" pageNumber="52">Male</collectingRegion>
|
||
</emphasis>
|
||
,
|
||
<typeStatus id="5F6A882FFFC8FF90FEA05091FEFD2396" box="[274,370,1925,1951]" pageId="9" pageNumber="52" type="paratype">paratype</typeStatus>
|
||
(
|
||
<figureCitation id="18EA2A08FFC8FF90FE335091FD8B23A9" box="[385,516,1925,1952]" captionStart="FIGURES 55–61" captionStartId="10.[152,270,1346,1370]" captionTargetBox="[151,1428,669,1323]" captionTargetId="figure-212@10.[151,1436,669,1323]" captionTargetPageId="10" captionText="FIGURES 55–61. Dasyproctus vietnamensis Pham sp. nov. Paratype ♂. 55. Head, frontal view; 56. Mesosoma, dorsal view; 57. Metasoma, dorsal view; 58. Head, dorsal view; 59. Mesosoma, lateral view; 60. Genitalia, ventral view; 61. Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7087369" httpUri="https://zenodo.org/record/7087369/files/figure.png" pageId="9" pageNumber="52">Figs 55‒61</figureCitation>
|
||
): Body length 6.5‒7.0 mm, forewing length 4.5‒5.0 mm.
|
||
</paragraph>
|
||
<paragraph id="806E368DFFC8FF90FF7550BDFD5723CA" blockId="9.[199,1436,1925,2023]" box="[199,728,1961,1987]" pageId="9" pageNumber="52">Structure as in female, but differing as follows:</paragraph>
|
||
<paragraph id="806E368DFFC8FF93FF7550D8FDAA2514" blockId="9.[199,1436,1925,2023]" lastBlockId="10.[151,1437,151,645]" lastPageId="10" lastPageNumber="53" pageId="9" pageNumber="52">
|
||
<emphasis id="B2A5EA9FFFC8FF90FF7550D8FE9723EE" bold="true" box="[199,280,1996,2023]" italics="true" pageId="9" pageNumber="52">Colour</emphasis>
|
||
. Scape yellow except inner lateral sides; mandible black, sometimes with small yellow spot at base; scutellum black except small yellow spot antero-laterally; mesopleuron sometimes with small yellow spot; small yellow spot on fore femur apically; dorsal surface of fore tibia yellow; yellow band on dorsal surface of mid and hind tibiae medially; large spot on dorsal surface of hind tibia; tegula testaceous; ventral surface of all tibiae blackish brown; T7 and all tarsi dark brown.
|
||
</paragraph>
|
||
<paragraph id="806E368DFFCBFF93FF755632FB02256C" blockId="10.[151,1437,151,645]" pageId="10" pageNumber="53">
|
||
<emphasis id="B2A5EA9FFFCBFF93FF755632FE8A2548" bold="true" box="[199,261,294,321]" italics="true" pageId="10" pageNumber="53">Head</emphasis>
|
||
(
|
||
<figureCitation id="18EA2A08FFCBFF93FEA75633FE3E2548" box="[277,433,294,321]" captionStart="FIGURES 55–61" captionStartId="10.[152,270,1346,1370]" captionTargetBox="[151,1428,669,1323]" captionTargetId="figure-212@10.[151,1436,669,1323]" captionTargetPageId="10" captionText="FIGURES 55–61. Dasyproctus vietnamensis Pham sp. nov. Paratype ♂. 55. Head, frontal view; 56. Mesosoma, dorsal view; 57. Metasoma, dorsal view; 58. Head, dorsal view; 59. Mesosoma, lateral view; 60. Genitalia, ventral view; 61. Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7087369" httpUri="https://zenodo.org/record/7087369/files/figure.png" pageId="10" pageNumber="53">Figs 55 & 58</figureCitation>
|
||
): clypeus with free margin truncate medially; mandible bidentate apically; flagellomeres II‒IV cylindrical; frons densely, largely punctate, with shallow, narrow furrow above IOC.
|
||
</paragraph>
|
||
<paragraph id="806E368DFFCBFF93FF75567BFE4A25D8" blockId="10.[151,1437,151,645]" pageId="10" pageNumber="53">
|
||
<taxonomicName id="47D14D0EFFCBFF93FF75567BFECD2580" box="[199,322,367,393]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="53" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2A5EA9FFFCBFF93FF75567BFECD2580" bold="true" box="[199,322,367,393]" italics="true" pageId="10" pageNumber="53">Mesosoma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="18EA2A08FFCBFF93FEE2567BFE682580" box="[336,487,366,393]" captionStart="FIGURES 55–61" captionStartId="10.[152,270,1346,1370]" captionTargetBox="[151,1428,669,1323]" captionTargetId="figure-212@10.[151,1436,669,1323]" captionTargetPageId="10" captionText="FIGURES 55–61. Dasyproctus vietnamensis Pham sp. nov. Paratype ♂. 55. Head, frontal view; 56. Mesosoma, dorsal view; 57. Metasoma, dorsal view; 58. Head, dorsal view; 59. Mesosoma, lateral view; 60. Genitalia, ventral view; 61. Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7087369" httpUri="https://zenodo.org/record/7087369/files/figure.png" pageId="10" pageNumber="53">Figs 56 & 59</figureCitation>
|
||
): Anterior carina of pronotal collar laterally curving towards pronotal lobe; pronotum smooth anteriorly; scutum with sparse, small punctures; scutellum with conspicuous rugae; propodeal dorsum coarsely, largely reticulate.
|
||
</paragraph>
|
||
<paragraph id="806E368DFFCBFF93FF7556CFFDD42634" blockId="10.[151,1437,151,645]" pageId="10" pageNumber="53">
|
||
<emphasis id="B2A5EA9FFFCBFF93FF7556CFFEB025FC" bold="true" box="[199,319,475,501]" italics="true" pageId="10" pageNumber="53">Metasoma</emphasis>
|
||
(
|
||
<figureCitation id="18EA2A08FFCBFF93FEE256CFFE6125FC" box="[336,494,474,501]" captionStart="FIGURES 55–61" captionStartId="10.[152,270,1346,1370]" captionTargetBox="[151,1428,669,1323]" captionTargetId="figure-212@10.[151,1436,669,1323]" captionTargetPageId="10" captionText="FIGURES 55–61. Dasyproctus vietnamensis Pham sp. nov. Paratype ♂. 55. Head, frontal view; 56. Mesosoma, dorsal view; 57. Metasoma, dorsal view; 58. Head, dorsal view; 59. Mesosoma, lateral view; 60. Genitalia, ventral view; 61. Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7087369" httpUri="https://zenodo.org/record/7087369/files/figure.png" pageId="10" pageNumber="53">Figs 57 & 61</figureCitation>
|
||
): petiolate; T1 longer than hind femur, shorter than T2‒T4 combined, its sides not parallel, 2.5 × as long as wide dorsally; T2‒T5 largely maculate, T6 with large yellow band anteriorly. T2‒T7 impunctate.
|
||
<collectingRegion id="4215F86FFFCBFF93FE905537FED32634" box="[290,348,547,573]" country="Maldives" name="Maale" pageId="10" pageNumber="53">Male</collectingRegion>
|
||
genitalia as in
|
||
<figureCitation id="18EA2A08FFCBFF93FDB75537FDD82634" box="[517,599,547,573]" captionStart="FIGURES 55–61" captionStartId="10.[152,270,1346,1370]" captionTargetBox="[151,1428,669,1323]" captionTargetId="figure-212@10.[151,1436,669,1323]" captionTargetPageId="10" captionText="FIGURES 55–61. Dasyproctus vietnamensis Pham sp. nov. Paratype ♂. 55. Head, frontal view; 56. Mesosoma, dorsal view; 57. Metasoma, dorsal view; 58. Head, dorsal view; 59. Mesosoma, lateral view; 60. Genitalia, ventral view; 61. Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7087369" httpUri="https://zenodo.org/record/7087369/files/figure.png" pageId="10" pageNumber="53">Fig. 60</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C8CB6506FFCBFF93FF755553FB332668" box="[199,1212,583,609]" pageId="10" pageNumber="53" type="etymology">
|
||
<paragraph id="806E368DFFCBFF93FF755553FB332668" blockId="10.[151,1437,151,645]" box="[199,1212,583,609]" pageId="10" pageNumber="53">
|
||
<emphasis id="B2A5EA9FFFCBFF93FF755553FEC72668" bold="true" box="[199,328,583,609]" pageId="10" pageNumber="53">Etymology</emphasis>
|
||
. The specific name of this new species refers to its country of origin,
|
||
<collectingCountry id="F8C6761DFFCBFF93FBE75553FB372668" box="[1109,1208,583,609]" name="Vietnam" pageId="10" pageNumber="53">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C8CB6506FFCBFF93FF75557EFD56268C" box="[199,729,618,645]" pageId="10" pageNumber="53" type="distribution">
|
||
<paragraph id="806E368DFFCBFF93FF75557EFD56268C" blockId="10.[151,1437,151,645]" box="[199,729,618,645]" pageId="10" pageNumber="53">
|
||
<emphasis id="B2A5EA9FFFCBFF93FF75557EFEEE268D" bold="true" box="[199,353,618,644]" pageId="10" pageNumber="53">Distribution.</emphasis>
|
||
<collectingCountry id="F8C6761DFFCBFF93FED5557FFE45268C" box="[359,458,619,645]" name="Vietnam" pageId="10" pageNumber="53">Vietnam</collectingCountry>
|
||
:
|
||
<collectingRegion id="4215F86FFFCBFF93FE67557FFDA8268C" box="[469,551,619,645]" country="Vietnam" name="Ha Noi" pageId="10" pageNumber="53">Ha Noi</collectingRegion>
|
||
and
|
||
<collectingRegion id="4215F86FFFCBFF93FDEF557FFD59268C" box="[605,726,619,645]" country="Vietnam" name="Vinh Phuc" pageId="10" pageNumber="53">Vinh Phuc</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |