176 lines
16 KiB
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176 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.983.52630" ID-GBIF-Dataset="d3b857f7-25c1-4b21-9d39-4510ab675fca" ID-PMC="PMC7655787" ID-Pensoft-Pub="1313-2970-983-1" ID-Pensoft-UUID="C890C7FD4B2D57A8B1A062305ED42D53" ID-PubMed="33244289" ID-ZooBank="7DED4CE2934C4539945F758930C927F9" ModsDocID="1313-2970-983-1" checkinTime="1604475085656" checkinUser="pensoft" docAuthor="Smith, Sarah M., Beaver, Roger A. & Cognato, Anthony I." docDate="2020" docId="FAE7E68167F5AFA5E27B0FFCD768ACBF" docLanguage="en" docName="ZooKeys 983: 1-442" docOrigin="ZooKeys 983" docSource="http://dx.doi.org/10.3897/zookeys.983.52630" docTitle="Amasa Lea 1894" docType="treatment" docVersion="4" id="C890C7FD4B2D57A8B1A062305ED42D53" lastPageNumber="1" masterDocId="C890C7FD4B2D57A8B1A062305ED42D53" masterDocTitle="A monograph of the Xyleborini (Coleoptera, Curculionidae, Scolytinae) of the Indochinese Peninsula (except Malaysia) and China" masterLastPageNumber="442" masterPageNumber="1" pageNumber="1" updateTime="1668169715782" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A monograph of the Xyleborini (Coleoptera, Curculionidae, Scolytinae) of the Indochinese Peninsula (except Malaysia) and China</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Smith, Sarah M.</mods:namePart>
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<mods:affiliation>Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-5173-3736</mods:nameIdentifier>
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<mods:nameIdentifier type="email">camptocerus@gmail.com</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Beaver, Roger A.</mods:namePart>
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<mods:affiliation>161 / 2 Mu 5, Soi Wat Pranon, T. Donkaew, A. Maerim, Chiangmai 50180, Thailand</mods:affiliation>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cognato, Anthony I.</mods:namePart>
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<mods:affiliation>Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>983</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>442</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.983.52630</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.983.52630</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-983-1</mods:identifier>
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<mods:identifier type="ZooBank">7DED4CE2934C4539945F758930C927F9</mods:identifier>
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<treatment ID-GBIF-Taxon="168754447" LSID="urn:lsid:plazi:treatment:FAE7E68167F5AFA5E27B0FFCD768ACBF" httpUri="http://treatment.plazi.org/id/FAE7E68167F5AFA5E27B0FFCD768ACBF" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authority="Lea, 1894" authorityName="Lea" authorityYear="1894" class="Insecta" family="Curculionidae" genus="Amasa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amasa" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Amasa Lea, 1894</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Lea" authorityYear="1894" class="Insecta" family="Curculionidae" genus="Amasa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amasa" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Amasa</taxonomicName>
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Lea, 1894: 322.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Eggers" authorityYear="1930" class="Insecta" family="Curculionidae" genus="Pseudoxyleborus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudoxyleborus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Pseudoxyleborus</taxonomicName>
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Eggers, 1930: 206. Synonymy:
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<bibRefCitation author="Wood, SL" journalOrPublisher="The Great Basin Naturalist" pageId="0" pageNumber="1" pagination="223 - 230" refId="B485" refString="Wood, SL, 1984. New generic synonymy and new genera of Scolytidae (Coleoptera). The Great Basin Naturalist 44: 223 - 230" title="New generic synonymy and new genera of Scolytidae (Coleoptera)." volume="44" year="1984">Wood 1984</bibRefCitation>
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: 223.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Wood" authorityYear="1980" class="Insecta" family="Curculionidae" genus="Anaxyleborus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anaxyleborus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Anaxyleborus</taxonomicName>
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Wood, 1980: 90. Synonymy:
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.4489" author="Wood, SL" journalOrPublisher="The Great Basin Naturalist" pageId="0" pageNumber="1" pagination="647 - 659" refId="B484" refString="Wood, SL, 1983. New synonymy and new species of American bark beetles (Coleoptera: Scolytidae), Part IX. The Great Basin Naturalist 43: 647 - 659, DOI: https://doi.org/10.5962/bhl.part.4489" title="New synonymy and new species of American bark beetles (Coleoptera: Scolytidae), Part IX." url="https://doi.org/10.5962/bhl.part.4489" volume="43" year="1983">Wood 1983</bibRefCitation>
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: 647.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type species">
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<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Lea" authorityYear="1893" class="Insecta" family="Curculionidae" genus="Amasa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amasa thoracica" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thoracica">
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<emphasis italics="true" pageId="0" pageNumber="1">Amasa thoracica</emphasis>
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</taxonomicName>
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Lea, 1894 =
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<taxonomicName authorityName="Erichson" authorityYear="1842" class="Insecta" family="Curculionidae" genus="Tomicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomicus truncatus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="truncatus">
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<emphasis italics="true" pageId="0" pageNumber="1">Tomicus truncatus</emphasis>
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</taxonomicName>
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Erichson, 1842; monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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2.5-5.0 mm, 2.11-3.4
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide.
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<taxonomicName authorityName="Lea" authorityYear="1894" class="Insecta" family="Curculionidae" genus="Amasa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amasa" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Amasa</emphasis>
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</taxonomicName>
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is distinguished by the declivity truncate, margined with a circumdeclivital ring; antennal club flattened, types 4 or 5 (typically type 4), club sutures sinuate, two sutures visible on posterior face; protibiae typically slender, inflated and granulate on posterior face (rarely distinctly triangular or unarmed on posterior face); anterior margin of pronotum with a row of serrations; scutellum flat, flush with elytral surface; declivital face with three striae; procoxae contiguous or narrowly separated; and mycangial tufts absent.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="similar genera">
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<paragraph pageId="0" pageNumber="1">Similar genera.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Hagedorn" authorityYear="1912" class="Insecta" family="Curculionidae" genus="Cyclorhipidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclorhipidion" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Cyclorhipidion</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Browne" authorityYear="1961" class="Insecta" family="Curculionidae" genus="Pseudowebbia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudowebbia" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Pseudowebbia</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Hulcr & Cognato" authorityYear="2010" class="Insecta" family="Curculionidae" genus="Truncaudum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Truncaudum" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Truncaudum</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Hopkins" authorityYear="1915" class="Insecta" family="Curculionidae" genus="Webbia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Webbia" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Webbia</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Reitter" authorityYear="1913" class="Insecta" family="Curculionidae" genus="Xylosandrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xylosandrus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Xylosandrus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Distributed throughout Asia and Australasia, also occurring in Madagascar. One species has been introduced to Brazil, Chile and Uruguay (
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<bibRefCitation DOI="https://doi.org/10.1649/072.065.0419" author="Flechtmann, CAH" journalOrPublisher="The Coleopterists Bulletin" pageId="0" pageNumber="1" pagination="417 - 421" refId="B162" refString="Flechtmann, CAH, Cognato, AI, 2011. First report of Amasa truncata (Coleoptera: Curculionidae: Scolytinae) in Brazil. The Coleopterists Bulletin 65: 417 - 421, DOI: https://doi.org/10.1649/072.065.0419" title="First report of Amasa truncata (Coleoptera: Curculionidae: Scolytinae) in Brazil." url="https://doi.org/10.1649/072.065.0419" volume="65" year="2011">Flechtmann and Cognato 2011</bibRefCitation>
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;
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<bibRefCitation DOI="https://doi.org/10.1111/evo.13219" author="Gomez, D" journalOrPublisher="The Coleopterists Bulletin" pageId="0" pageNumber="1" pagination="825 - 826" refId="B174" refString="Gomez, D, Suarez, M, Martinez, G, 2017. Amasa truncata (Erichson) (Coleoptera: Curculionidae: Scolytinae): a new exotic ambrosia beetle in Uruguay. The Coleopterists Bulletin 71: 825 - 826, DOI: https://doi.org/10.1111/evo.13219" title="Amasa truncata (Erichson) (Coleoptera: Curculionidae: Scolytinae): a new exotic ambrosia beetle in Uruguay." url="https://doi.org/10.1111/evo.13219" volume="71" year="2017">
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<normalizedToken originalValue="Gómez">Gomez</normalizedToken>
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et al. 2017
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</bibRefCitation>
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;
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<bibRefCitation DOI="https://doi.org/10.3390/d10020040" author="Kirkendall, LR" journalOrPublisher="Diversity" pageId="0" pageNumber="1" pagination="1 - 40" refId="B230" refString="Kirkendall, LR, 2018. Invasive bark beetles (Coleoptera, Curculionidae, Scolytinae) in Chile and Argentina, including two species new for South America, and the correct identity of the Orthotomicus species in Chile and Argentina. Diversity 10: 1 - 40, DOI: https://doi.org/10.3390/d10020040" title="Invasive bark beetles (Coleoptera, Curculionidae, Scolytinae) in Chile and Argentina, including two species new for South America, and the correct identity of the Orthotomicus species in Chile and Argentina." url="https://doi.org/10.3390/d10020040" volume="10" year="2018">Kirkendall 2018</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="gallery system">
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<paragraph pageId="0" pageNumber="1">Gallery system.</paragraph>
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<paragraph pageId="0" pageNumber="1">This usually comprises a short radial tunnel leading to a single, large, flat brood chamber, extending in the longitudinal plane.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Lea" authorityYear="1894" class="Insecta" family="Curculionidae" genus="Amasa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amasa" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Amasa</emphasis>
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</taxonomicName>
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is easily confused with other species possessing truncate declivities in the genera listed above. Most species can be readily distinguished by the type 4 antennal club with sinuate sutures and the presence of only three striae on the declivital face.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Previous morphological studies of
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<taxonomicName authorityName="Lea" authorityYear="1894" class="Insecta" family="Curculionidae" genus="Amasa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amasa" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Amasa</emphasis>
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</taxonomicName>
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have suggested that species are very morphologically variable (
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<bibRefCitation author="Hulcr, J" journalOrPublisher="Thomas Say Publications in Entomology, Entomological Society of America, Lanham" pageId="0" pageNumber="1" refId="B205" refString="Hulcr, J, Cognato, AI, 2013. Xyleborini of New Guinea: A Taxonomic Monograph. Thomas Say Publications in Entomology, Entomological Society of America, Lanham" title="Xyleborini of New Guinea: A Taxonomic Monograph." year="2013">Hulcr and Cognato 2013</bibRefCitation>
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). As a result, many species were considered conspecific and part of a morphological continuum. Molecular data generated as part of this study has demonstrated that
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<taxonomicName authorityName="Lea" authorityYear="1894" class="Insecta" family="Curculionidae" genus="Amasa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amasa" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Amasa</emphasis>
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</taxonomicName>
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species are actually morphologically conserved even across broad ranges (
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<bibRefCitation DOI="https://doi.org/10.1649/0010-065X-74.1.37" author="Smith, SM" journalOrPublisher="The Coleopterists Bulletin" pageId="0" pageNumber="1" pagination="37 - 41" refId="B446" refString="Smith, SM, Beaver, RA, Cognato, AI, 2020. Taxonomic changes for Indo-Malayan ambrosia beetles (Coleoptera: Curculionidae: Scolytinae: Xyleborini). The Coleopterists Bulletin 74: 37 - 41, DOI: https://doi.org/10.1649/0010-065X-74.1.37" title="Taxonomic changes for Indo-Malayan ambrosia beetles (Coleoptera: Curculionidae: Scolytinae: Xyleborini)." url="https://doi.org/10.1649/0010-065X-74.1.37" volume="74" year="2020">Smith et al. 2020</bibRefCitation>
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).
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<taxonomicName authorityName="Lea" authorityYear="1894" class="Insecta" family="Curculionidae" genus="Amasa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amasa" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Amasa</emphasis>
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</taxonomicName>
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species outside our coverage area are thus in need of revision. Potentially much of the diversity is awaiting discovery.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |