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<document ID-DOI="http://dx.doi.org/10.3897/jhr.82.65252" ID-GBIF-Dataset="142ee8b9-8eb6-46c6-842a-88af6c2798ee" ID-Pensoft-Pub="1314-2607-82-361" ID-Pensoft-UUID="85B70BB83E44587FAA6E7D6B26D82A6F" ID-Zenodo-Dep="4741754" ID-ZooBank="C2CA4E54C2FE42DB806A6770DB6B5832" ModsDocID="1314-2607-82-361" checkinTime="1619783719098" checkinUser="pensoft" docAuthor="Soliman, Ahmed Mostafa" docDate="2021" docId="4CEECDE3AEC550D18F8631CCA3B705C7" docLanguage="en" docName="JourHymenoptRes 82: 361-415" docOrigin="Journal of Hymenoptera Research 82" docPubDate="2021-04-29" docSource="http://dx.doi.org/10.3897/jhr.82.65252" docTitle="Pseudophotopsis mascatiana Invrea 1962" docType="treatment" docVersion="4" id="85B70BB83E44587FAA6E7D6B26D82A6F" lastPageNumber="361" masterDocId="85B70BB83E44587FAA6E7D6B26D82A6F" masterDocTitle="The velvet ant genus Pseudophotopsis Andre, 1896 (Hymenoptera, Mutillidae) in the Arabian Peninsula, with the description of two new species" masterLastPageNumber="415" masterPageNumber="361" pageNumber="361" updateTime="1678755765704" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The velvet ant genus Pseudophotopsis Andre, 1896 (Hymenoptera, Mutillidae) in the Arabian Peninsula, with the description of two new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Soliman, Ahmed Mostafa</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5284-713X</mods:nameIdentifier>
<mods:affiliation>Plant Protection Department, College of Food and Agriculture Sciences, King Saud University, P. O. BOX 2460, Riyadh 11451, Saudi Arabia &amp; Zoology Department, Faculty of Science (Boys), Al-Azhar University, P. O. Box 11884, Nasr City, Cairo, Egypt</mods:affiliation>
<mods:nameIdentifier type="email">ammsoliman@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Hymenoptera Research</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-04-29</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>82</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>361</mods:start>
<mods:end>415</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/jhr.82.65252</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.82.65252</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2607-82-361</mods:identifier>
<mods:identifier type="ZooBank">C2CA4E54C2FE42DB806A6770DB6B5832</mods:identifier>
<mods:identifier type="Pensoft-UUID">85B70BB83E44587FAA6E7D6B26D82A6F</mods:identifier>
<mods:identifier type="Zenodo-Dep">4741754</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="181758014" LSID="urn:lsid:plazi:treatment:4CEECDE3AEC550D18F8631CCA3B705C7" httpUri="http://treatment.plazi.org/id/4CEECDE3AEC550D18F8631CCA3B705C7" lastPageNumber="361" pageId="0" pageNumber="361">
<subSubSection pageId="0" pageNumber="361" type="nomenclature">
<paragraph pageId="0" pageNumber="361">
<taxonomicName LSID="4CEECDE3-AEC5-50D1-8F86-31CCA3B705C7" authority="Invrea, 1962" authorityName="Invrea" authorityYear="1962" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudophotopsis mascatiana" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="mascatiana">Pseudophotopsis mascatiana Invrea, 1962</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="361" type="description">
<paragraph pageId="0" pageNumber="361">
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Pseudophotopsis mascatiana Invrea (Male) A, B habitus, lateral and dorsal views, respectively C head, frontal view D head and antennal scape, dorsal view." figureDoi="10.3897/jhr.82.65252.figure15" httpUri="https://binary.pensoft.net/fig/537896" pageId="0" pageNumber="361">Figures 15</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Pseudophotopsis mascatiana Invrea (Male) A antenna B head and antennal scape, lateral view C, D mesosoma, lateral and dorsal views, respectively." figureDoi="10.3897/jhr.82.65252.figure16" httpUri="https://binary.pensoft.net/fig/537897" pageId="0" pageNumber="361">, 16</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Pseudophotopsis mascatiana Invrea (Male) A right fore wing B, C metasoma, dorsal and ventral views, respectively." figureDoi="10.3897/jhr.82.65252.figure17" httpUri="https://binary.pensoft.net/fig/537898" pageId="0" pageNumber="361">, 17</figureCitation>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Pseudophotopsis mascatiana Invrea (Female) A, B habitus, lateral and dorsal views, respectively." figureDoi="10.3897/jhr.82.65252.figure18" httpUri="https://binary.pensoft.net/fig/537899" pageId="0" pageNumber="361">, 18</figureCitation>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Pseudophotopsis mascatiana Invrea (Female) A-C head, frontal, dorsal and lateral views, respectively D mandible, fronto-lateral view." figureDoi="10.3897/jhr.82.65252.figure19" httpUri="https://binary.pensoft.net/fig/537900" pageId="0" pageNumber="361">, 19</figureCitation>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pseudophotopsis mascatiana Invrea (Female) A antenna (only part of scape) B, C mesosoma, dorsal and lateral views, respectively." figureDoi="10.3897/jhr.82.65252.figure20" httpUri="https://binary.pensoft.net/fig/537901" pageId="0" pageNumber="361">, 20</figureCitation>
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudophotopsis mascatiana Invrea (Female) A, B mid and hind tibiae and basitarsomeres, respectively (apices of mid and hind tibiae indicated) C, D metasoma, dorsal and ventral views, respectively." figureDoi="10.3897/jhr.82.65252.figure21" httpUri="https://binary.pensoft.net/fig/537902" pageId="0" pageNumber="361">, 21</figureCitation>
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudophotopsis mascatiana Invrea (Female) A T 4 - T 6, dorsal view B T 6 (after removing setae surrounding pygidial area)." figureDoi="10.3897/jhr.82.65252.figure22" httpUri="https://binary.pensoft.net/fig/537903" pageId="0" pageNumber="361">, 22</figureCitation>
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Male genitalia A, B Pseudophotopsis mascatiana Invrea, dorsal and ventral views, respectively C, D P. maura Bischoff, dorsal and ventral views, respectively." figureDoi="10.3897/jhr.82.65252.figure34" httpUri="https://binary.pensoft.net/fig/537915" pageId="0" pageNumber="361">; 34A, B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="361" type="reference_group">
<paragraph pageId="0" pageNumber="361">
<taxonomicName authorityName="Invrea" authorityYear="1962" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudophotopsis mascatiana" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="mascatiana">Pseudophotopsis mascatiana</taxonomicName>
Invrea, 1962: 324, (♂ holotype),
<normalizedToken originalValue="“Migiurtinia”">&quot;Migiurtinia&quot;</normalizedToken>
(Somalia).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="361" type="diagnosis">
<paragraph pageId="0" pageNumber="361">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="361">
<emphasis bold="true" pageId="0" pageNumber="361">Male</emphasis>
(Figs
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Pseudophotopsis mascatiana Invrea (Male) A, B habitus, lateral and dorsal views, respectively C head, frontal view D head and antennal scape, dorsal view." figureDoi="10.3897/jhr.82.65252.figure15" httpUri="https://binary.pensoft.net/fig/537896" pageId="0" pageNumber="361">15</figureCitation>
-
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Pseudophotopsis mascatiana Invrea (Male) A right fore wing B, C metasoma, dorsal and ventral views, respectively." figureDoi="10.3897/jhr.82.65252.figure17" httpUri="https://binary.pensoft.net/fig/537898" pageId="0" pageNumber="361">17</figureCitation>
;
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Male genitalia A, B Pseudophotopsis mascatiana Invrea, dorsal and ventral views, respectively C, D P. maura Bischoff, dorsal and ventral views, respectively." figureDoi="10.3897/jhr.82.65252.figure34" httpUri="https://binary.pensoft.net/fig/537915" pageId="0" pageNumber="361">34A, B</figureCitation>
). Body length 8.8-16 mm. Metasomal segments darkened, at most 7th segment yellowish-brown to red (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Pseudophotopsis mascatiana Invrea (Male) A right fore wing B, C metasoma, dorsal and ventral views, respectively." figureDoi="10.3897/jhr.82.65252.figure17" httpUri="https://binary.pensoft.net/fig/537898" pageId="0" pageNumber="361">17B, C</figureCitation>
); fore wing slightly infuscate on distal half, with brown veins (Sc+R and pterostigma darker) (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Pseudophotopsis mascatiana Invrea (Male) A right fore wing B, C metasoma, dorsal and ventral views, respectively." figureDoi="10.3897/jhr.82.65252.figure17" httpUri="https://binary.pensoft.net/fig/537898" pageId="0" pageNumber="361">17A</figureCitation>
); antennal flagellomeres more or less light brown (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Pseudophotopsis mascatiana Invrea (Male) A antenna B head and antennal scape, lateral view C, D mesosoma, lateral and dorsal views, respectively." figureDoi="10.3897/jhr.82.65252.figure16" httpUri="https://binary.pensoft.net/fig/537897" pageId="0" pageNumber="361">16A</figureCitation>
). Distance between apex of lower mandibular tooth and upper mandibular ridge less than mandibular height at base (about 0.85
<normalizedToken originalValue="×">x</normalizedToken>
) (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Pseudophotopsis mascatiana Invrea (Male) A antenna B head and antennal scape, lateral view C, D mesosoma, lateral and dorsal views, respectively." figureDoi="10.3897/jhr.82.65252.figure16" httpUri="https://binary.pensoft.net/fig/537897" pageId="0" pageNumber="361">16B</figureCitation>
); POD 1.3-1.6
<normalizedToken originalValue="×">x</normalizedToken>
OOD (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Pseudophotopsis mascatiana Invrea (Male) A, B habitus, lateral and dorsal views, respectively C head, frontal view D head and antennal scape, dorsal view." figureDoi="10.3897/jhr.82.65252.figure15" httpUri="https://binary.pensoft.net/fig/537896" pageId="0" pageNumber="361">15D</figureCitation>
); metanotal horns well developed, distinctly longer than wide (about 1.5
<normalizedToken originalValue="×">x</normalizedToken>
) (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Pseudophotopsis mascatiana Invrea (Male) A antenna B head and antennal scape, lateral view C, D mesosoma, lateral and dorsal views, respectively." figureDoi="10.3897/jhr.82.65252.figure16" httpUri="https://binary.pensoft.net/fig/537897" pageId="0" pageNumber="361">16C</figureCitation>
); T1 distinctly long, about 1.45-1.65
<normalizedToken originalValue="×">x</normalizedToken>
as long as maximal width (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Pseudophotopsis mascatiana Invrea (Male) A, B habitus, lateral and dorsal views, respectively C head, frontal view D head and antennal scape, dorsal view." figureDoi="10.3897/jhr.82.65252.figure15" httpUri="https://binary.pensoft.net/fig/537896" pageId="0" pageNumber="361">15B</figureCitation>
); lateral felt line on T2 about 0.6-0.75
<normalizedToken originalValue="×">x</normalizedToken>
as long as distance between felt line and posterior tergal margin; S2 with extremely small median basal tubercle (mostly absent) (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Pseudophotopsis mascatiana Invrea (Male) A right fore wing B, C metasoma, dorsal and ventral views, respectively." figureDoi="10.3897/jhr.82.65252.figure17" httpUri="https://binary.pensoft.net/fig/537898" pageId="0" pageNumber="361">17C</figureCitation>
); parapenial lobe slightly oblique inward, with subrounded apex (Fig.
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Male genitalia A, B Pseudophotopsis mascatiana Invrea, dorsal and ventral views, respectively C, D P. maura Bischoff, dorsal and ventral views, respectively." figureDoi="10.3897/jhr.82.65252.figure34" httpUri="https://binary.pensoft.net/fig/537915" pageId="0" pageNumber="361">34A</figureCitation>
); genital ventral lobes at most oblique inward, with outer margin obtusely angulate (Fig.
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Male genitalia A, B Pseudophotopsis mascatiana Invrea, dorsal and ventral views, respectively C, D P. maura Bischoff, dorsal and ventral views, respectively." figureDoi="10.3897/jhr.82.65252.figure34" httpUri="https://binary.pensoft.net/fig/537915" pageId="0" pageNumber="361">34B</figureCitation>
).
</paragraph>
<caption ID-Zenodo-Dep="4741768" doi="10.3897/jhr.82.65252.figure15" httpUri="https://binary.pensoft.net/fig/537896" pageId="0" pageNumber="361" start="Figure 15" startId="F15">
<paragraph pageId="0" pageNumber="361">
<emphasis bold="true" pageId="0" pageNumber="361">Figure 15.</emphasis>
<taxonomicName authorityName="Invrea" authorityYear="1962" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudophotopsis mascatiana" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="mascatiana">
<emphasis italics="true" pageId="0" pageNumber="361">Pseudophotopsis mascatiana</emphasis>
</taxonomicName>
Invrea (Male)
<emphasis bold="true" pageId="0" pageNumber="361">A, B</emphasis>
habitus, lateral and dorsal views, respectively
<emphasis bold="true" pageId="0" pageNumber="361">C</emphasis>
head, frontal view
<emphasis bold="true" pageId="0" pageNumber="361">D</emphasis>
head and antennal scape, dorsal view.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="4741770" doi="10.3897/jhr.82.65252.figure16" httpUri="https://binary.pensoft.net/fig/537897" pageId="0" pageNumber="361" start="Figure 16" startId="F16">
<paragraph pageId="0" pageNumber="361">
<emphasis bold="true" pageId="0" pageNumber="361">Figure 16.</emphasis>
<taxonomicName authorityName="Invrea" authorityYear="1962" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudophotopsis mascatiana" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="mascatiana">
<emphasis italics="true" pageId="0" pageNumber="361">Pseudophotopsis mascatiana</emphasis>
</taxonomicName>
Invrea (Male)
<emphasis bold="true" pageId="0" pageNumber="361">A</emphasis>
antenna
<emphasis bold="true" pageId="0" pageNumber="361">B</emphasis>
head and antennal scape, lateral view
<emphasis bold="true" pageId="0" pageNumber="361">C, D</emphasis>
mesosoma, lateral and dorsal views, respectively.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="4741772" doi="10.3897/jhr.82.65252.figure17" httpUri="https://binary.pensoft.net/fig/537898" pageId="0" pageNumber="361" start="Figure 17" startId="F17">
<paragraph pageId="0" pageNumber="361">
<emphasis bold="true" pageId="0" pageNumber="361">Figure 17.</emphasis>
<taxonomicName authorityName="Invrea" authorityYear="1962" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudophotopsis mascatiana" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="mascatiana">
<emphasis italics="true" pageId="0" pageNumber="361">Pseudophotopsis mascatiana</emphasis>
</taxonomicName>
Invrea (Male)
<emphasis bold="true" pageId="0" pageNumber="361">A</emphasis>
right fore wing
<emphasis bold="true" pageId="0" pageNumber="361">B, C</emphasis>
metasoma, dorsal and ventral views, respectively.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="361">
<emphasis bold="true" pageId="0" pageNumber="361">Female</emphasis>
(Figs
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Pseudophotopsis mascatiana Invrea (Female) A, B habitus, lateral and dorsal views, respectively." figureDoi="10.3897/jhr.82.65252.figure18" httpUri="https://binary.pensoft.net/fig/537899" pageId="0" pageNumber="361">18</figureCitation>
-
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudophotopsis mascatiana Invrea (Female) A T 4 - T 6, dorsal view B T 6 (after removing setae surrounding pygidial area)." figureDoi="10.3897/jhr.82.65252.figure22" httpUri="https://binary.pensoft.net/fig/537903" pageId="0" pageNumber="361">22</figureCitation>
). Body length 10.8-16.5 mm. Anterior ocellus absent, only a pair of poorly developed posterior ocelli present (Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Pseudophotopsis mascatiana Invrea (Female) A-C head, frontal, dorsal and lateral views, respectively D mandible, fronto-lateral view." figureDoi="10.3897/jhr.82.65252.figure19" httpUri="https://binary.pensoft.net/fig/537900" pageId="0" pageNumber="361">19B</figureCitation>
); distance between apex of mandibular lower tooth and upper mandibular ridge 0.86-0.88
<normalizedToken originalValue="×">x</normalizedToken>
as long as mandibular height at the base (Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Pseudophotopsis mascatiana Invrea (Female) A-C head, frontal, dorsal and lateral views, respectively D mandible, fronto-lateral view." figureDoi="10.3897/jhr.82.65252.figure19" httpUri="https://binary.pensoft.net/fig/537900" pageId="0" pageNumber="361">19D</figureCitation>
); F1 about 1.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as F2 (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pseudophotopsis mascatiana Invrea (Female) A antenna (only part of scape) B, C mesosoma, dorsal and lateral views, respectively." figureDoi="10.3897/jhr.82.65252.figure20" httpUri="https://binary.pensoft.net/fig/537901" pageId="0" pageNumber="361">20A</figureCitation>
); mesosomal dorsum regularly punctate, with interspaces between punctures ridged but not tuberculate (smooth), clothed with long erect to suberect golden setae (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pseudophotopsis mascatiana Invrea (Female) A antenna (only part of scape) B, C mesosoma, dorsal and lateral views, respectively." figureDoi="10.3897/jhr.82.65252.figure20" httpUri="https://binary.pensoft.net/fig/537901" pageId="0" pageNumber="361">20B, C</figureCitation>
); pronotal dorsal face (excluding anterior flange) gently declivitous antero-medially, with convex anterior border (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pseudophotopsis mascatiana Invrea (Female) A antenna (only part of scape) B, C mesosoma, dorsal and lateral views, respectively." figureDoi="10.3897/jhr.82.65252.figure20" httpUri="https://binary.pensoft.net/fig/537901" pageId="0" pageNumber="361">20B</figureCitation>
); propodeal dorsal face gently declivitous (rounded) posteriorly, 0.70-0.77
<normalizedToken originalValue="×">x</normalizedToken>
as long as maximal width (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pseudophotopsis mascatiana Invrea (Female) A antenna (only part of scape) B, C mesosoma, dorsal and lateral views, respectively." figureDoi="10.3897/jhr.82.65252.figure20" httpUri="https://binary.pensoft.net/fig/537901" pageId="0" pageNumber="361">20B</figureCitation>
); meso- and metapleura mostly smooth (with scattered fine punctures) (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pseudophotopsis mascatiana Invrea (Female) A antenna (only part of scape) B, C mesosoma, dorsal and lateral views, respectively." figureDoi="10.3897/jhr.82.65252.figure20" httpUri="https://binary.pensoft.net/fig/537901" pageId="0" pageNumber="361">20C</figureCitation>
); mid and hind tibiae without prolongation at their apices (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudophotopsis mascatiana Invrea (Female) A, B mid and hind tibiae and basitarsomeres, respectively (apices of mid and hind tibiae indicated) C, D metasoma, dorsal and ventral views, respectively." figureDoi="10.3897/jhr.82.65252.figure21" httpUri="https://binary.pensoft.net/fig/537902" pageId="0" pageNumber="361">21A, B</figureCitation>
); hind tibia with narrow shallow groove extending along inner face (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudophotopsis mascatiana Invrea (Female) A, B mid and hind tibiae and basitarsomeres, respectively (apices of mid and hind tibiae indicated) C, D metasoma, dorsal and ventral views, respectively." figureDoi="10.3897/jhr.82.65252.figure21" httpUri="https://binary.pensoft.net/fig/537902" pageId="0" pageNumber="361">21B</figureCitation>
); T1-T5 and S2-S5 with apical fringe of compact golden setae (Figs
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Pseudophotopsis mascatiana Invrea (Female) A, B habitus, lateral and dorsal views, respectively." figureDoi="10.3897/jhr.82.65252.figure18" httpUri="https://binary.pensoft.net/fig/537899" pageId="0" pageNumber="361">18A, B</figureCitation>
;
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudophotopsis mascatiana Invrea (Female) A, B mid and hind tibiae and basitarsomeres, respectively (apices of mid and hind tibiae indicated) C, D metasoma, dorsal and ventral views, respectively." figureDoi="10.3897/jhr.82.65252.figure21" httpUri="https://binary.pensoft.net/fig/537902" pageId="0" pageNumber="361">21C, D</figureCitation>
); T2 densely punctate, punctures 1-2 diameters apart (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudophotopsis mascatiana Invrea (Female) A, B mid and hind tibiae and basitarsomeres, respectively (apices of mid and hind tibiae indicated) C, D metasoma, dorsal and ventral views, respectively." figureDoi="10.3897/jhr.82.65252.figure21" httpUri="https://binary.pensoft.net/fig/537902" pageId="0" pageNumber="361">21C</figureCitation>
); T6 with bell-shaped, finely wrinkled pygidial area (Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudophotopsis mascatiana Invrea (Female) A T 4 - T 6, dorsal view B T 6 (after removing setae surrounding pygidial area)." figureDoi="10.3897/jhr.82.65252.figure22" httpUri="https://binary.pensoft.net/fig/537903" pageId="0" pageNumber="361">22B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="361" type="description">
<paragraph pageId="0" pageNumber="361">Description of female</paragraph>
<paragraph pageId="0" pageNumber="361">
<emphasis bold="true" pageId="0" pageNumber="361">
(hitherto unknown) allotype (Figs
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Pseudophotopsis mascatiana Invrea (Female) A, B habitus, lateral and dorsal views, respectively." figureDoi="10.3897/jhr.82.65252.figure18" httpUri="https://binary.pensoft.net/fig/537899" pageId="0" pageNumber="361">18</figureCitation>
-
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudophotopsis mascatiana Invrea (Female) A T 4 - T 6, dorsal view B T 6 (after removing setae surrounding pygidial area)." figureDoi="10.3897/jhr.82.65252.figure22" httpUri="https://binary.pensoft.net/fig/537903" pageId="0" pageNumber="361">22</figureCitation>
).
</emphasis>
Body length 10.8 mm.
<emphasis bold="true" italics="true" pageId="0" pageNumber="361">Color</emphasis>
(Figs
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Pseudophotopsis mascatiana Invrea (Female) A, B habitus, lateral and dorsal views, respectively." figureDoi="10.3897/jhr.82.65252.figure18" httpUri="https://binary.pensoft.net/fig/537899" pageId="0" pageNumber="361">18A, B</figureCitation>
;
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Pseudophotopsis mascatiana Invrea (Female) A-C head, frontal, dorsal and lateral views, respectively D mandible, fronto-lateral view." figureDoi="10.3897/jhr.82.65252.figure19" httpUri="https://binary.pensoft.net/fig/537900" pageId="0" pageNumber="361">19A, D</figureCitation>
;
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pseudophotopsis mascatiana Invrea (Female) A antenna (only part of scape) B, C mesosoma, dorsal and lateral views, respectively." figureDoi="10.3897/jhr.82.65252.figure20" httpUri="https://binary.pensoft.net/fig/537901" pageId="0" pageNumber="361">20A</figureCitation>
;
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudophotopsis mascatiana Invrea (Female) A, B mid and hind tibiae and basitarsomeres, respectively (apices of mid and hind tibiae indicated) C, D metasoma, dorsal and ventral views, respectively." figureDoi="10.3897/jhr.82.65252.figure21" httpUri="https://binary.pensoft.net/fig/537902" pageId="0" pageNumber="361">21D</figureCitation>
). Head (including antennal scape, pedicel and F1 and basal third of mandible) and mesosoma (including legs) dark red (mid and hind legs darker); remaining flagellomeres light brown, F11 yellowish apically; mandible black apically; metasomal terga black (except blackish-red T6), sterna blackish-red to black.
</paragraph>
<caption ID-Zenodo-Dep="4741774" doi="10.3897/jhr.82.65252.figure18" httpUri="https://binary.pensoft.net/fig/537899" pageId="0" pageNumber="361" start="Figure 18" startId="F18">
<paragraph pageId="0" pageNumber="361">
<emphasis bold="true" pageId="0" pageNumber="361">Figure 18.</emphasis>
<taxonomicName authorityName="Invrea" authorityYear="1962" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudophotopsis mascatiana" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="mascatiana">
<emphasis italics="true" pageId="0" pageNumber="361">Pseudophotopsis mascatiana</emphasis>
</taxonomicName>
Invrea (Female)
<emphasis bold="true" pageId="0" pageNumber="361">A, B</emphasis>
habitus, lateral and dorsal views, respectively.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="361">
<emphasis bold="true" italics="true" pageId="0" pageNumber="361">Setation</emphasis>
(Figs
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Pseudophotopsis mascatiana Invrea (Female) A, B habitus, lateral and dorsal views, respectively." figureDoi="10.3897/jhr.82.65252.figure18" httpUri="https://binary.pensoft.net/fig/537899" pageId="0" pageNumber="361">18A, B</figureCitation>
;
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Pseudophotopsis mascatiana Invrea (Female) A-C head, frontal, dorsal and lateral views, respectively D mandible, fronto-lateral view." figureDoi="10.3897/jhr.82.65252.figure19" httpUri="https://binary.pensoft.net/fig/537900" pageId="0" pageNumber="361">19A-D</figureCitation>
;
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pseudophotopsis mascatiana Invrea (Female) A antenna (only part of scape) B, C mesosoma, dorsal and lateral views, respectively." figureDoi="10.3897/jhr.82.65252.figure20" httpUri="https://binary.pensoft.net/fig/537901" pageId="0" pageNumber="361">20C</figureCitation>
;
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudophotopsis mascatiana Invrea (Female) A, B mid and hind tibiae and basitarsomeres, respectively (apices of mid and hind tibiae indicated) C, D metasoma, dorsal and ventral views, respectively." figureDoi="10.3897/jhr.82.65252.figure21" httpUri="https://binary.pensoft.net/fig/537902" pageId="0" pageNumber="361">21C, D</figureCitation>
;
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudophotopsis mascatiana Invrea (Female) A T 4 - T 6, dorsal view B T 6 (after removing setae surrounding pygidial area)." figureDoi="10.3897/jhr.82.65252.figure22" httpUri="https://binary.pensoft.net/fig/537903" pageId="0" pageNumber="361">22A</figureCitation>
). Head (including scape, pedicel and F1 of antenna and basal third of mandible), pronotal lateral face, meso- and metapleura, legs, T1 and S1-S2 clothed with long erect white setae; mesosomal dorsum and T2-T5 with long erect to suberect golden setae; T1-T5 and S2-S5 with apical fringe of extremely dense (compact) golden setae, paler on sterna.
</paragraph>
<caption ID-Zenodo-Dep="4741776" doi="10.3897/jhr.82.65252.figure19" httpUri="https://binary.pensoft.net/fig/537900" pageId="0" pageNumber="361" start="Figure 19" startId="F19">
<paragraph pageId="0" pageNumber="361">
<emphasis bold="true" pageId="0" pageNumber="361">Figure 19.</emphasis>
<taxonomicName authorityName="Invrea" authorityYear="1962" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudophotopsis mascatiana" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="mascatiana">
<emphasis italics="true" pageId="0" pageNumber="361">Pseudophotopsis mascatiana</emphasis>
</taxonomicName>
Invrea (Female)
<emphasis bold="true" pageId="0" pageNumber="361">A-C</emphasis>
head, frontal, dorsal and lateral views, respectively
<emphasis bold="true" pageId="0" pageNumber="361">D</emphasis>
mandible, fronto-lateral view.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="361">
<emphasis bold="true" italics="true" pageId="0" pageNumber="361">Head</emphasis>
(Figs
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Pseudophotopsis mascatiana Invrea (Female) A, B habitus, lateral and dorsal views, respectively." figureDoi="10.3897/jhr.82.65252.figure18" httpUri="https://binary.pensoft.net/fig/537899" pageId="0" pageNumber="361">18B</figureCitation>
;
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Pseudophotopsis mascatiana Invrea (Female) A-C head, frontal, dorsal and lateral views, respectively D mandible, fronto-lateral view." figureDoi="10.3897/jhr.82.65252.figure19" httpUri="https://binary.pensoft.net/fig/537900" pageId="0" pageNumber="361">19A-D</figureCitation>
;
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pseudophotopsis mascatiana Invrea (Female) A antenna (only part of scape) B, C mesosoma, dorsal and lateral views, respectively." figureDoi="10.3897/jhr.82.65252.figure20" httpUri="https://binary.pensoft.net/fig/537901" pageId="0" pageNumber="361">20A</figureCitation>
). In dorsal view, subquadrangular, scarcely narrower than pronotum, convergent behind eyes, with broadly rounded temple; head height slightly greater than width; only a pair of poorly developed posterior ocelli present, anterior ocellus absent; frons, vertex, and gena densely setiferous foveate, interspaces between foveae thick (ridged) without tubercles; eye rather oval-shaped, widely separated from mandibular articulation (malar space about 0.85
<normalizedToken originalValue="×">x</normalizedToken>
mandibular basal height); scrobal carina well developed, polished and thick; clypeal longitudinal median carina ridged, free margin with a pair of small tubercles; mandible with strong subbasal lower tooth, distance between apex of tooth and upper mandibular ridge about 0.86
<normalizedToken originalValue="×">x</normalizedToken>
as long as mandibular height at base, subapically with distinctly small inner tooth. F1 1.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as maximal width, 1.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as F2, hardly longer than F3 (1.1
<normalizedToken originalValue="×">x</normalizedToken>
).
</paragraph>
<paragraph pageId="0" pageNumber="361">
<emphasis bold="true" italics="true" pageId="0" pageNumber="361">Mesosoma</emphasis>
(Figs
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pseudophotopsis mascatiana Invrea (Female) A antenna (only part of scape) B, C mesosoma, dorsal and lateral views, respectively." figureDoi="10.3897/jhr.82.65252.figure20" httpUri="https://binary.pensoft.net/fig/537901" pageId="0" pageNumber="361">20B, C</figureCitation>
;
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudophotopsis mascatiana Invrea (Female) A, B mid and hind tibiae and basitarsomeres, respectively (apices of mid and hind tibiae indicated) C, D metasoma, dorsal and ventral views, respectively." figureDoi="10.3897/jhr.82.65252.figure21" httpUri="https://binary.pensoft.net/fig/537902" pageId="0" pageNumber="361">21A, B</figureCitation>
). Dorsum regularly setiferous punctate, with interspaces between punctures ridged and polished, not tuberculate; pronotal dorsal face slightly wider than propodeum (1.15
<normalizedToken originalValue="×">x</normalizedToken>
), rather gently declivitous anteromedially, with slightly convex anterior border and broadly rounded humeral angles; pronotal lateral face foveate-reticulate; promesonotal suture gently arched; mesonotum suddenly contracted behind pronotum (about 0.75
<normalizedToken originalValue="×">x</normalizedToken>
as wide as pronotum), with posterior suture nearly straight; propodeal dorsal face about 0.77
<normalizedToken originalValue="×">x</normalizedToken>
as long as maximal width, subparallel at sides, gently declivitous (rounded) posteriorly; propodeal posterior face foveate-reticulate (interspaces between foveae not tuberculate); meso- and metapleura shiny, mostly smooth (with scattered fine punctures), mesepisternum with vertical row of close foveae on ventral half of strong mesopleural ridge; mid and hind tibiae with a double row of 5-7 strong spines on outer faces, both tibiae without prolongation at their apices; hind tibia with longitudinal narrow and shallow groove extending along inner face.
</paragraph>
<caption ID-Zenodo-Dep="4741780" doi="10.3897/jhr.82.65252.figure20" httpUri="https://binary.pensoft.net/fig/537901" pageId="0" pageNumber="361" start="Figure 20" startId="F20">
<paragraph pageId="0" pageNumber="361">
<emphasis bold="true" pageId="0" pageNumber="361">Figure 20.</emphasis>
<taxonomicName authorityName="Invrea" authorityYear="1962" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudophotopsis mascatiana" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="mascatiana">
<emphasis italics="true" pageId="0" pageNumber="361">Pseudophotopsis mascatiana</emphasis>
</taxonomicName>
Invrea (Female)
<emphasis bold="true" pageId="0" pageNumber="361">A</emphasis>
antenna (only part of scape)
<emphasis bold="true" pageId="0" pageNumber="361">B, C</emphasis>
mesosoma, dorsal and lateral views, respectively.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="361">
<emphasis bold="true" italics="true" pageId="0" pageNumber="361">Metasoma</emphasis>
(Figs
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudophotopsis mascatiana Invrea (Female) A, B mid and hind tibiae and basitarsomeres, respectively (apices of mid and hind tibiae indicated) C, D metasoma, dorsal and ventral views, respectively." figureDoi="10.3897/jhr.82.65252.figure21" httpUri="https://binary.pensoft.net/fig/537902" pageId="0" pageNumber="361">21C, D</figureCitation>
;
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudophotopsis mascatiana Invrea (Female) A T 4 - T 6, dorsal view B T 6 (after removing setae surrounding pygidial area)." figureDoi="10.3897/jhr.82.65252.figure22" httpUri="https://binary.pensoft.net/fig/537903" pageId="0" pageNumber="361">22A, B</figureCitation>
). Sessile; T1 without distinct dorsal face (sharply sloping forward), wider than long (1.3
<normalizedToken originalValue="×">x</normalizedToken>
), densely finely punctate apically (punctures 1-2 diameters apart), sparsely punctate at the base; T2 densely punctate throughout (punctures larger than those on T1, 1-2 diameters apart); exposed parts of T3-T5 finely punctate; T6 nearly entirely covered with long golden setae arising from apical fringe of T5 and laterally on T6 (removal of setae shows finely wrinkled, bell-shaped pygidial area of T6); S1 with strongly elevated inverted Y-shaped carina; S2 densely punctate, basally with large median tubercle.
</paragraph>
<caption ID-Zenodo-Dep="4741784" doi="10.3897/jhr.82.65252.figure21" httpUri="https://binary.pensoft.net/fig/537902" pageId="0" pageNumber="361" start="Figure 21" startId="F21">
<paragraph pageId="0" pageNumber="361">
<emphasis bold="true" pageId="0" pageNumber="361">Figure 21.</emphasis>
<taxonomicName authorityName="Invrea" authorityYear="1962" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudophotopsis mascatiana" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="mascatiana">
<emphasis italics="true" pageId="0" pageNumber="361">Pseudophotopsis mascatiana</emphasis>
</taxonomicName>
Invrea (Female)
<emphasis bold="true" pageId="0" pageNumber="361">A, B</emphasis>
mid and hind tibiae and basitarsomeres, respectively (apices of mid and hind tibiae indicated)
<emphasis bold="true" pageId="0" pageNumber="361">C, D</emphasis>
metasoma, dorsal and ventral views, respectively.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="4741786" doi="10.3897/jhr.82.65252.figure22" httpUri="https://binary.pensoft.net/fig/537903" pageId="0" pageNumber="361" start="Figure 22" startId="F22">
<paragraph pageId="0" pageNumber="361">
<emphasis bold="true" pageId="0" pageNumber="361">Figure 22.</emphasis>
<taxonomicName authorityName="Invrea" authorityYear="1962" class="Insecta" family="Mutillidae" genus="Pseudophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudophotopsis mascatiana" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="mascatiana">
<emphasis italics="true" pageId="0" pageNumber="361">Pseudophotopsis mascatiana</emphasis>
</taxonomicName>
Invrea (Female)
<emphasis bold="true" pageId="0" pageNumber="361">A</emphasis>
T4-T6, dorsal view
<emphasis bold="true" pageId="0" pageNumber="361">B</emphasis>
T6 (after removing setae surrounding pygidial area).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="361" type="materials_examined">
<paragraph pageId="0" pageNumber="361">Material examined</paragraph>
<paragraph pageId="0" pageNumber="361">
<emphasis bold="true" pageId="0" pageNumber="361">
(
<specimenCount type="male">174 males</specimenCount>
,
<specimenCount type="female">11 females</specimenCount>
).
</emphasis>
<specimenCount type="female">1♀</specimenCount>
(WM2),
<date value="2019-04-11">11.iv.2019</date>
, HP, leg. AMS [KSMA];
<specimenCount type="male">1♂</specimenCount>
(JB),
<date value="1983-09-10">10.ix.1983</date>
, leg. Buttiker W. [NMB];
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" location="Al-Wesam" specimenCount="1" specimenCount-male="1">
<specimenCount type="male">1♂</specimenCount>
(
<location LSID="urn:lsid:plazi:treatment:4CEECDE3AEC550D18F8631CCA3B705C7:61B14CC8CFFB8CFC79BEF132AED46649" name="Al-Wesam">Al-Wesam</location>
)
</materialsCitation>
,
<date value="2010-10-11">11.x.2010</date>
, leg. AlDhafer et al. [
<typeStatus>allotype</typeStatus>
KSMA];
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" location="Al-Shafa" specimenCount="2" specimenCount-male="2">
<specimenCount type="male">2♂</specimenCount>
(
<location LSID="urn:lsid:plazi:treatment:4CEECDE3AEC550D18F8631CCA3B705C7:E53D8355B7D4120BFFDD498B3683EABC" name="Al-Shafa">Al-Shafa</location>
)
</materialsCitation>
,
<date value="2010-10-12">12.x.2010</date>
, LT, leg. Al-Dhafer et al. [KSMA];
<specimenCount type="female">1♀</specimenCount>
(GR2),
<specimenCount type="female">1♀</specimenCount>
(GR3),
<date value="2014-04-08">8.iv.2014</date>
, PT, leg. Al-Dhafer et al. [KSMA];
<specimenCount type="male">2♂</specimenCount>
(SH6),
<specimenCount type="male">1♂</specimenCount>
(SH7),
<date value="2014-04-21">21.iv.2014</date>
, LT, leg. Al-Dhafer et al. [KSMA];
<specimenCount type="male">6♂</specimenCount>
(GR1),
<date value="2014-04-24">24.iv.2014</date>
, LT, leg. Al-Dhafer et al. [KSMA];
<specimenCount type="male">8♂</specimenCount>
(SH2),
<specimenCount type="male">2♂</specimenCount>
(SH5),
<specimenCount type="male">2♂</specimenCount>
(SH6),
<specimenCount type="male">6♂</specimenCount>
(SH7),
<date value="2014-06-03">3.vi.2014</date>
, LT, leg. Al-Dhafer et al. [KSMA];
<specimenCount type="male">18♂</specimenCount>
(GR1),
<specimenCount type="male">12♂</specimenCount>
(GR2),
<specimenCount type="male">3♂</specimenCount>
(GR3),
<date value="2014-06-08">8.vi.2014</date>
, LT, leg. Al-Dhafer et al. [KSMA];
<specimenCount type="male">4♂</specimenCount>
(GR1),
<specimenCount type="male">1♂</specimenCount>
(GR2),
<specimenCount type="male">1♂</specimenCount>
(GR6),
<date value="2014-08-26">26.viii.2014</date>
, LT, leg. Al-Dhafer et al. [KSMA];
<specimenCount type="male">2♂</specimenCount>
(GR1),
<specimenCount type="male">1♂</specimenCount>
(GR3),
<date value="2014-10-20">20.x.2014</date>
, LT, leg. Al-Dhafer et al. [KSMA];
<specimenCount type="male">1♂</specimenCount>
(WK),
<date value="2014-10-30">30.x.2014</date>
, LT, leg. Al-Harbi M. &amp; Rasool I. [KSMA];
<specimenCount type="female">1♀</specimenCount>
(SH7),
<date value="2015-01-26">26.i.2015</date>
, PT, leg. Al-Dhafer et al. [KSMA];
<specimenCount type="male">1♂</specimenCount>
(SH2),
<date value="2015-01-27">27.i.2015</date>
, LT, leg. Al-Dhafer et al. [KSMA];
<specimenCount type="female">2♀</specimenCount>
(SH3),
<specimenCount type="female">1♀</specimenCount>
(SH8),
<date value="2015-03-02">2.iii.2015</date>
, PT, leg. ElTorkey et al. [KSMA];
<specimenCount type="female">1♀</specimenCount>
(SH8),
<date value="2015-05-03">3.v.2015</date>
, PT, leg. Al-Dhafer et al. [KSMA];
<specimenCount type="male">1♂</specimenCount>
(SH3),
<specimenCount type="male">4♂</specimenCount>
(SH5),
<date value="2015-05-05">5.v.2015</date>
, LT, leg. Al-Dhafer et al. [KSMA];
<specimenCount type="male">1♂</specimenCount>
(GR2),
<date value="2015-05-07">7.v.2015</date>
, LT, leg. Al-Dhafer et al. [KSMA];
<specimenCount type="male">5♂</specimenCount>
, (WY2),
<date value="2015-05-22">22.v.2015</date>
, LT, leg. AMS [KSMA];
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" location="Al-Soudah" specimenCount="1" specimenCount-male="1">
<specimenCount type="male">1♂</specimenCount>
(
<location LSID="urn:lsid:plazi:treatment:4CEECDE3AEC550D18F8631CCA3B705C7:A2BF5666A1C78DA0290162F7208FD513" name="Al-Soudah">Al-Soudah</location>
)
</materialsCitation>
,
<date value="2015-05-23">23.v.2015</date>
, LT, leg. AMS [KSMA];
<specimenCount type="male">3♂</specimenCount>
(SH2),
<specimenCount type="male">2♂</specimenCount>
(SH5),
<specimenCount type="male">1♂</specimenCount>
(SH6),
<specimenCount type="male">1♂</specimenCount>
(SH7),
<specimenCount type="male">2♂</specimenCount>
(SH8),
<date value="2015-07-27">27.vii.2015</date>
, LT, leg. Al-Dhafer et al. [KSMA];
<specimenCount type="male">5♂</specimenCount>
&amp;
<specimenCount type="female">1♀</specimenCount>
(GR1),
<specimenCount type="male">1♂</specimenCount>
(GR2),
<specimenCount type="male">4♂</specimenCount>
&amp;
<specimenCount type="female">1♀</specimenCount>
(GR3),
<specimenCount type="male">10♂</specimenCount>
(GR4),
<specimenCount type="male">1♂</specimenCount>
(GR5),
<date value="2015-07-31">31.vii.2015</date>
, LT for males PT for females, leg. Al-Dhafer et al. [KSMA];
<specimenCount type="male">1♂</specimenCount>
(SH3),
<specimenCount type="male">1♂</specimenCount>
(SH6),
<specimenCount type="male">1♂</specimenCount>
(SH7),
<specimenCount type="male">1♂</specimenCount>
(SH8),
<date value="2015-09-02">2.ix.2015</date>
, LT, leg. Al-Dhafer et al. [KSMA];
<specimenCount type="male">2♂</specimenCount>
(GR1),
<specimenCount type="male">1♂</specimenCount>
(GR2),
<specimenCount type="male">1♂</specimenCount>
(GR3),
<specimenCount type="male">1♂</specimenCount>
(GR4),
<date value="2015-09-05">5.ix.2015</date>
, LT, leg. Al-Dhafer et al. [KSMA];
<specimenCount type="male">5♂</specimenCount>
(SH1),
<specimenCount type="male">3♂</specimenCount>
(SH2),
<specimenCount type="male">4♂</specimenCount>
(SH5),
<specimenCount type="male">1♂</specimenCount>
(SH8),
<date value="2015-11-14">14.xi.2015</date>
, LT, leg. Al-Dhafer et al. [KSMA];
<specimenCount type="male">1♂</specimenCount>
(WN1),
<date value="2015-11-15">15.xi.2015</date>
, LT, leg. AMS [KSMA];
<specimenCount type="male">1♂</specimenCount>
(GR1),
<specimenCount type="male">3♂</specimenCount>
(GR2),
<specimenCount type="male">1♂</specimenCount>
(GR4),
<date value="2015-11-18">18.xi.2015</date>
, LT, leg. Al-Dhafer et al. [KSMA];
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" location="Wadi Mashwas" specimenCount="1" specimenCount-male="1">
<specimenCount type="male">1♂</specimenCount>
(
<location LSID="urn:lsid:plazi:treatment:4CEECDE3AEC550D18F8631CCA3B705C7:5DB30C0F7284DCC5DF4425BBF7BC3C7D" name="Wadi Mashwas">Wadi Mashwas</location>
)
</materialsCitation>
,
<date value="2015-11-20">20.xi.2015</date>
, LT, leg. AMS [KSMA];
<specimenCount type="male">3♂</specimenCount>
(WM1),
<date value="2016-04-16">16.iv.2016</date>
, LT, leg. AMS [KSMA];
<specimenCount type="male">1♂</specimenCount>
(WBD),
<date value="2016-10-19">19.x.2016</date>
, LT, leg. AMS [KSMA];
<specimenCount type="male">1♂</specimenCount>
&amp;
<specimenCount type="female">1♀</specimenCount>
(WM2),
<date value="2019-04-11">11.iv.2019</date>
, LT for
<specimenCount>♂</specimenCount>
and HP for
<specimenCount>♀</specimenCount>
, leg. AMS [KSMA];
<specimenCount type="male">1♂</specimenCount>
(WK),
<date value="2019-04-12">12.iv.2019</date>
, LT, leg. AMS [KSMA].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="361" type="remarks">
<paragraph pageId="0" pageNumber="361">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="361">
Prior to the present study,
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. mascatiana" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="mascatiana">
<emphasis italics="true" pageId="0" pageNumber="361">P. mascatiana</emphasis>
</taxonomicName>
was known only from males and restricted in its distribution to the Afrotropical region in Somalia and Yemen (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1226.1.1" author="Lelej, AS" journalOrPublisher="Zootaxa" pageId="0" pageNumber="361" pagination="1 - 50" refId="B29" refString="Lelej, AS, van Harten, A, 2006. A review of the Mutillidae (Hymenoptera) of Yemen. Zootaxa 1226 (1): 1 - 50, DOI: https://doi.org/10.11646/zootaxa.1226.1.1" title="A review of the Mutillidae (Hymenoptera) of Yemen." url="https://doi.org/10.11646/zootaxa.1226.1.1" volume="1226" year="2006">Lelej and van Harten 2006</bibRefCitation>
). Here this species is recorded from Saudi Arabia with discovery of the opposite sex (female) for the first time. The sex association is based on the overlapping distribution areas of males and females (frequently both sexes were collected from the same locality at the same time or within a short period) (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution map of Pseudophotopsis mascatiana Invrea, and P. subaurea sp. nov. in the Arabian Peninsula." figureDoi="10.3897/jhr.82.65252.figure2" httpUri="https://binary.pensoft.net/fig/537883" pageId="0" pageNumber="361">2</figureCitation>
), and the general color pattern in both sexes. On the other hand, there is another species,
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. aurea" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="aurea">
<emphasis italics="true" pageId="0" pageNumber="361">P. aurea</emphasis>
</taxonomicName>
, common to southwestern Saudi Arabia and known from both sexes; the females of these two species are similar in having T1-T5 and S2-S5 with apical fringes of compact golden setae; however,
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. mascatiana" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="mascatiana">
<emphasis italics="true" pageId="0" pageNumber="361">P. mascatiana</emphasis>
</taxonomicName>
differs from
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. aurea" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="aurea">
<emphasis italics="true" pageId="0" pageNumber="361">P. aurea</emphasis>
</taxonomicName>
in the following aspects: the head and mesosoma are dark red (Fig.
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Pseudophotopsis mascatiana Invrea (Female) A, B habitus, lateral and dorsal views, respectively." figureDoi="10.3897/jhr.82.65252.figure18" httpUri="https://binary.pensoft.net/fig/537899" pageId="0" pageNumber="361">18A, B</figureCitation>
) (they are dark brown with slight reddish tint in
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. aurea" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="aurea">
<emphasis italics="true" pageId="0" pageNumber="361">P. aurea</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pseudophotopsis aurea (Klug) (Female) A, B habitus, lateral and dorsal views, respectively." figureDoi="10.3897/jhr.82.65252.figure6" httpUri="https://binary.pensoft.net/fig/537887" pageId="0" pageNumber="361">6A, B</figureCitation>
)); the frons and vertex with smooth (not tuberculate) ridges between the foveae (Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Pseudophotopsis mascatiana Invrea (Female) A-C head, frontal, dorsal and lateral views, respectively D mandible, fronto-lateral view." figureDoi="10.3897/jhr.82.65252.figure19" httpUri="https://binary.pensoft.net/fig/537900" pageId="0" pageNumber="361">19A, B</figureCitation>
) (with coarsely tuberculate ridges between the foveae in
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. aurea" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="aurea">
<emphasis italics="true" pageId="0" pageNumber="361">P. aurea</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Pseudophotopsis aurea (Klug) (Female) A head, frontal view B head and antennal scape, dorsal view C antenna (with only part of scape) D mesosoma, dorsal view." figureDoi="10.3897/jhr.82.65252.figure7" httpUri="https://binary.pensoft.net/fig/537888" pageId="0" pageNumber="361">7A, B</figureCitation>
); the anterior ocellus is absent, only a pair of poorly developed posterior ocelli being present (Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Pseudophotopsis mascatiana Invrea (Female) A-C head, frontal, dorsal and lateral views, respectively D mandible, fronto-lateral view." figureDoi="10.3897/jhr.82.65252.figure19" httpUri="https://binary.pensoft.net/fig/537900" pageId="0" pageNumber="361">19A, B</figureCitation>
) (three well developed ocelli are present in
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. aurea" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="aurea">
<emphasis italics="true" pageId="0" pageNumber="361">P. aurea</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Pseudophotopsis aurea (Klug) (Female) A head, frontal view B head and antennal scape, dorsal view C antenna (with only part of scape) D mesosoma, dorsal view." figureDoi="10.3897/jhr.82.65252.figure7" httpUri="https://binary.pensoft.net/fig/537888" pageId="0" pageNumber="361">7A, B</figureCitation>
)); the mesosomal dorsum is clothed with golden setae, regularly punctate, with smooth (not tuberculate) ridges between the punctures (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pseudophotopsis mascatiana Invrea (Female) A antenna (only part of scape) B, C mesosoma, dorsal and lateral views, respectively." figureDoi="10.3897/jhr.82.65252.figure20" httpUri="https://binary.pensoft.net/fig/537901" pageId="0" pageNumber="361">20B, C</figureCitation>
) (in
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. aurea" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="aurea">
<emphasis italics="true" pageId="0" pageNumber="361">P. aurea</emphasis>
</taxonomicName>
the mesosomal dorsum clothed with pale (yellowish-white) setae, coarsely foveate, with ridges between foveae tuberculate (Figs
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Pseudophotopsis aurea (Klug) (Female) A head, frontal view B head and antennal scape, dorsal view C antenna (with only part of scape) D mesosoma, dorsal view." figureDoi="10.3897/jhr.82.65252.figure7" httpUri="https://binary.pensoft.net/fig/537888" pageId="0" pageNumber="361">7D</figureCitation>
;
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pseudophotopsis aurea (Klug) (Female) A head and mesosoma, lateral view (tubercles on propodeal dorsum indicated) B, C mid and hind tibiae and basitarsomeres, respectively (prolongation at the apices of mid and hind tibiae indicated) D metasoma, dorsal view." figureDoi="10.3897/jhr.82.65252.figure8" httpUri="https://binary.pensoft.net/fig/537889" pageId="0" pageNumber="361">8A</figureCitation>
)); the pronotal dorsal face gently declivitous antero-medially, with a convex anterior border (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pseudophotopsis mascatiana Invrea (Female) A antenna (only part of scape) B, C mesosoma, dorsal and lateral views, respectively." figureDoi="10.3897/jhr.82.65252.figure20" httpUri="https://binary.pensoft.net/fig/537901" pageId="0" pageNumber="361">20B</figureCitation>
) (sharply declivitous antero-medially, with a straight anterior border in
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. aurea" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="aurea">
<emphasis italics="true" pageId="0" pageNumber="361">P. aurea</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Pseudophotopsis aurea (Klug) (Female) A head, frontal view B head and antennal scape, dorsal view C antenna (with only part of scape) D mesosoma, dorsal view." figureDoi="10.3897/jhr.82.65252.figure7" httpUri="https://binary.pensoft.net/fig/537888" pageId="0" pageNumber="361">7D</figureCitation>
)); the propodeal dorsal face is subparallel at its sides, gently declivitous (rounded) posteriorly, and 0.70-0.77
<normalizedToken originalValue="×">x</normalizedToken>
as long as its maximal width (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pseudophotopsis mascatiana Invrea (Female) A antenna (only part of scape) B, C mesosoma, dorsal and lateral views, respectively." figureDoi="10.3897/jhr.82.65252.figure20" httpUri="https://binary.pensoft.net/fig/537901" pageId="0" pageNumber="361">20B, C</figureCitation>
) (in
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. aurea" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="aurea">
<emphasis italics="true" pageId="0" pageNumber="361">P. aurea</emphasis>
</taxonomicName>
the propodeal dorsal face is strongly expanded at the sides, sharply declivitous (truncate) posteriorly, and 0.50-0.55
<normalizedToken originalValue="×">x</normalizedToken>
as long as its maximal width (Figs
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Pseudophotopsis aurea (Klug) (Female) A head, frontal view B head and antennal scape, dorsal view C antenna (with only part of scape) D mesosoma, dorsal view." figureDoi="10.3897/jhr.82.65252.figure7" httpUri="https://binary.pensoft.net/fig/537888" pageId="0" pageNumber="361">7D</figureCitation>
;
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pseudophotopsis aurea (Klug) (Female) A head and mesosoma, lateral view (tubercles on propodeal dorsum indicated) B, C mid and hind tibiae and basitarsomeres, respectively (prolongation at the apices of mid and hind tibiae indicated) D metasoma, dorsal view." figureDoi="10.3897/jhr.82.65252.figure8" httpUri="https://binary.pensoft.net/fig/537889" pageId="0" pageNumber="361">8A</figureCitation>
)); meso- and metapleura are mostly smooth (Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pseudophotopsis mascatiana Invrea (Female) A antenna (only part of scape) B, C mesosoma, dorsal and lateral views, respectively." figureDoi="10.3897/jhr.82.65252.figure20" httpUri="https://binary.pensoft.net/fig/537901" pageId="0" pageNumber="361">20C</figureCitation>
) (sparsely punctate in
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. aurea" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="aurea">
<emphasis italics="true" pageId="0" pageNumber="361">P. aurea</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pseudophotopsis aurea (Klug) (Female) A head and mesosoma, lateral view (tubercles on propodeal dorsum indicated) B, C mid and hind tibiae and basitarsomeres, respectively (prolongation at the apices of mid and hind tibiae indicated) D metasoma, dorsal view." figureDoi="10.3897/jhr.82.65252.figure8" httpUri="https://binary.pensoft.net/fig/537889" pageId="0" pageNumber="361">8A</figureCitation>
)); the mid and hind tibiae lack prolongation at their apices, hind tibia with a groove extending along inner face (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pseudophotopsis mascatiana Invrea (Female) A, B mid and hind tibiae and basitarsomeres, respectively (apices of mid and hind tibiae indicated) C, D metasoma, dorsal and ventral views, respectively." figureDoi="10.3897/jhr.82.65252.figure21" httpUri="https://binary.pensoft.net/fig/537902" pageId="0" pageNumber="361">21A, B</figureCitation>
) (the mid and hind tibiae with strong prolongation at their apices, hind tibia without such groove on inner face in
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. aurea" order="Hymenoptera" pageId="0" pageNumber="361" phylum="Arthropoda" rank="species" species="aurea">
<emphasis italics="true" pageId="0" pageNumber="361">P. aurea</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pseudophotopsis aurea (Klug) (Female) A head and mesosoma, lateral view (tubercles on propodeal dorsum indicated) B, C mid and hind tibiae and basitarsomeres, respectively (prolongation at the apices of mid and hind tibiae indicated) D metasoma, dorsal view." figureDoi="10.3897/jhr.82.65252.figure8" httpUri="https://binary.pensoft.net/fig/537889" pageId="0" pageNumber="361">8B, C</figureCitation>
)); T6 with a bell-shaped, finely wrinkled pygidial area (Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pseudophotopsis mascatiana Invrea (Female) A T 4 - T 6, dorsal view B T 6 (after removing setae surrounding pygidial area)." figureDoi="10.3897/jhr.82.65252.figure22" httpUri="https://binary.pensoft.net/fig/537903" pageId="0" pageNumber="361">22B</figureCitation>
) (T6 with an oval-shaped pygidial area that is minutely rugose on the disc and obliquely striate laterally (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Pseudophotopsis aurea (Klug) (Female) A T 4 - T 6, dorsal view B T 6 (after removing setae surrounding pygidial area) C metasoma, ventral view (felt lines on T 2 and S 2 indicated)." figureDoi="10.3897/jhr.82.65252.figure9" httpUri="https://binary.pensoft.net/fig/537890" pageId="0" pageNumber="361">9A, B</figureCitation>
)).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="361" type="distribution">
<paragraph pageId="0" pageNumber="361">Distribution in the Arabian Peninsula.</paragraph>
<paragraph pageId="0" pageNumber="361">
<normalizedToken originalValue="Sana’a">Sana'a</normalizedToken>
, Dhamar, Amran,
<normalizedToken originalValue="Ta’izz">Ta'izz</normalizedToken>
, and Al-Hudaydah (Yemen) (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1226.1.1" author="Lelej, AS" journalOrPublisher="Zootaxa" pageId="0" pageNumber="361" pagination="1 - 50" refId="B29" refString="Lelej, AS, van Harten, A, 2006. A review of the Mutillidae (Hymenoptera) of Yemen. Zootaxa 1226 (1): 1 - 50, DOI: https://doi.org/10.11646/zootaxa.1226.1.1" title="A review of the Mutillidae (Hymenoptera) of Yemen." url="https://doi.org/10.11646/zootaxa.1226.1.1" volume="1226" year="2006">Lelej and van Harten 2006</bibRefCitation>
); Asir, Al-Baha, and Makkah regions (southwestern Saudi Arabia) (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution map of Pseudophotopsis mascatiana Invrea, and P. subaurea sp. nov. in the Arabian Peninsula." figureDoi="10.3897/jhr.82.65252.figure2" httpUri="https://binary.pensoft.net/fig/537883" pageId="0" pageNumber="361">2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="361" type="distribution">
<paragraph pageId="0" pageNumber="361">Global distribution.</paragraph>
<paragraph pageId="0" pageNumber="361">
Somalia, Yemen (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4754.1.15" author="Gadallah, NS" journalOrPublisher="Zootaxa" pageId="0" pageNumber="361" pagination="141 - 152" refId="B19" refString="Gadallah, NS, Lelej, AS, Brothers, DJ, 2020. Biodiversity of the aculeate wasps (Hymenoptera: Aculeata) of the Arabian Peninsula: Vespoidea, Mutillidae. Zootaxa 4754 (1): 141 - 152, DOI: https://doi.org/10.11646/zootaxa.4754.1.15" title="Biodiversity of the aculeate wasps (Hymenoptera: Aculeata) of the Arabian Peninsula: Vespoidea, Mutillidae." url="https://doi.org/10.11646/zootaxa.4754.1.15" volume="4754" year="2020">Gadallah et al. 2020</bibRefCitation>
); Saudi Arabia (new record).
</paragraph>
</subSubSection>
</treatment>
</document>
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