255 lines
29 KiB
XML
255 lines
29 KiB
XML
<document id="6FFBD11D46BED92DE552A19E0087B714" ID-DOI="10.15298/rusentj.32.1.03" ID-ISSN="0132-8069" ID-Zenodo-Dep="10949096" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1712198792598" checkinUser="felipe" docAuthor="Fedorenko, D. N." docDate="2023" docId="2F4187C3FFCD16657984F98323A273D8" docLanguage="en" docName="RussEntomolJ.32.1.16-39.pdf" docOrigin="Russian Entomological Journal 32 (1)" docSource="http://dx.doi.org/10.15298/rusentj.32.1.03" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.1:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="1" docTitle="(Neohaptoderus) Tschitscherine 1898" docType="treatment" docVersion="5" lastPageNumber="23" masterDocId="D378FFBBFFCB166D7966FF9A2163720F" masterDocTitle="New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam" masterLastPageNumber="39" masterPageNumber="16" pageNumber="22" updateTime="1712734223118" updateUser="ExternalLinkService">
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<mods:titleInfo id="884F9CB92CA832F5857F3F6465BC915C">
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<mods:title id="F8BEA8AF6F00BD8701243FF2A9F95673">New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam</mods:title>
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<mods:namePart id="FDA04391E4993ED74F070CA95007D466">Fedorenko, D. N.</mods:namePart>
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<mods:title id="7D12901AD9B1BF48E95F935D108C8A04">Russian Entomological Journal</mods:title>
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<mods:part id="87D08C710DD1057EA1E98A15C1AE4ABA">
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<mods:date id="CA2A3952D407A0EA2A874ACBCAA51A66">2023</mods:date>
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<mods:number id="B741243BB7B43F3CEF705FD3209823F2">2023-03-31</mods:number>
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<mods:number id="3365C91DDA8DEE3D83C8F68D2F52DA77">32</mods:number>
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<mods:url id="BF6DC860550A8E3403068CE629BBD5D9">http://dx.doi.org/10.15298/rusentj.32.1.03</mods:url>
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<mods:identifier id="F3D46568CA84B4EABAFD30DDF5894E76" type="DOI">10.15298/rusentj.32.1.03</mods:identifier>
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<mods:identifier id="C3A8B7C8AF23A7A7EF0E416EC7B74E70" type="ISSN">0132-8069</mods:identifier>
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<subSubSection id="EFF2655EFFCD166B7984F9832382743E" box="[226,737,1560,1585]" pageId="6" pageNumber="22" type="nomenclature">
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<paragraph id="A75736D5FFCD166B7984F9832382743E" blockId="6.[226,737,1560,1585]" box="[226,737,1560,1585]" pageId="6" pageNumber="22">
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<heading id="FC1F81B9FFCD166B7984F9832382743E" box="[226,737,1560,1585]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="22" reason="3">
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Subgenus
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<taxonomicName id="60E84D56FFCD166B7834F9822382743E" ID-CoL="8MBT6" authority="Tschitscherine, 1898" authorityName="Tschitscherine" authorityYear="1898" box="[338,737,1560,1585]" kingdom="Animalia" pageId="6" pageNumber="22" rank="subGenus" subGenus="Neohaptoderus">
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<emphasis id="959CEAC7FFCD166B7834F982209A743F" bold="true" box="[338,505,1560,1584]" italics="true" pageId="6" pageNumber="22">Neohaptoderus</emphasis>
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Tschitschérine, 1898
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="EFF2655EFFCD166B7984F9D823127463" pageId="6" pageNumber="22" type="reference_group">
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||
<paragraph id="A75736D5FFCD166B7984F9D823127463" blockId="6.[189,775,1602,2066]" pageId="6" pageNumber="22">
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<bibRefCitation id="C3794B24FFCD166B7984F9D820B2745A" author="Tschitscherine T." box="[226,465,1602,1621]" pageId="6" pageNumber="22" pagination="1 - 224" refId="ref20688" refString="Tschitscherine T. 1898 a. Materiaux pour servir a l'etude des feroniens. IV // Horae Soc. Ent. Ross. T. 32. P. 1 - 224." type="journal article" year="1898">Tschitschérine, 1898a: 183</bibRefCitation>
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(
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<taxonomicName id="60E84D56FFCD166B7887F9D82345745B" authority="Leach, 1817" box="[481,550,1602,1620]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="22" phylum="Arthropoda" rank="subGenus" subGenus="Feronia">
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<emphasis id="959CEAC7FFCD166B7887F9D82345745B" box="[481,550,1602,1620]" italics="true" pageId="6" pageNumber="22">Feronia</emphasis>
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</taxonomicName>
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subg.);
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<bibRefCitation id="C3794B24FFCD166B7B15F9D82262745A" author="Sciaky R." box="[627,769,1602,1621]" pageId="6" pageNumber="22" pagination="1 - 19" refId="ref20291" refString="Sciaky R. 1994. Revision of Pterostichus subg. Morphohaptoderus Tschitscherine, 1898 with description of ten new species from China (Coleoptera: Carabidae) // Koleopt. Rundsch. Bd. 64. P. 1 - 19." type="book chapter" year="1994">Sciaky, 1994: 1</bibRefCitation>
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;
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<bibRefCitation id="C3794B24FFCD166B79DBF9C020CE7462" author="Sciaky R. & Wrase D. W." box="[189,429,1626,1645]" pageId="6" pageNumber="22" pagination="1087 - 1139" refId="ref20333" refString="Sciaky R., Wrase D. W. 1997. Twenty-nine new taxa of Pterostichinae from Shaanxi (Coleoptera, Carabidae) // Linzer Biol. Beitr. Bd. 29. H. 2. P. 1087 - 1139." type="journal article" year="1997">Sciaky, Wrase, 1997: 1091</bibRefCitation>
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;
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<bibRefCitation id="C3794B24FFCD166B78D1F9C0230E7463" author="Shi H. & Sciaky R. & Liang H. & Zhou H." box="[439,621,1625,1645]" pageId="6" pageNumber="22" pagination="101 - 135" refId="ref20376" refString="Shi H., Sciaky R., Liang H., Zhou H. 2013. A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions // Zootaxa. Vol. 3664. No. 2. P. 101 - 135." type="book chapter" year="2013">Shi et al., 2013: 105</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="EFF2655EFFCD166B7984F9E823797492" pageId="6" pageNumber="22" type="materials_examined">
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<paragraph id="A75736D5FFCD166B7984F9E823797492" blockId="6.[189,775,1602,2066]" pageId="6" pageNumber="22">
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<typeStatus id="78538877FFCD166B7984F9E8206D748A" box="[226,270,1650,1669]" pageId="6" pageNumber="22">Type</typeStatus>
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species:
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<taxonomicName id="60E84D56FFCD166B7805F9E823B5748A" authority="Tschitscherine, 1898" authorityName="Tschitscherine" authorityYear="1898" box="[355,726,1650,1669]" class="Insecta" family="Carabidae" genus="Feronia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="22" phylum="Arthropoda" rank="species" species="berezowskii">
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<emphasis id="959CEAC7FFCD166B7805F9E82375748A" box="[355,534,1650,1669]" italics="true" pageId="6" pageNumber="22">Feronia berezowskii</emphasis>
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Tschitschérine, 1898
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</taxonomicName>
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, subsequently designated by
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<bibRefCitation id="C3794B24FFCD166B78F0F91023747492" author="Lorenz W." box="[406,535,1674,1693]" pageId="6" pageNumber="22" refId="ref19861" refString="Lorenz W. 1998. Systematic list of extant ground beetles of the world (Insecta Coleoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 1 st ed. Tutzing. 502 pp." type="book" year="1998">Lorenz [1998]</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="EFF2655EFFCD166B7984F93820C97A1D" pageId="6" pageNumber="22" type="diagnosis">
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<paragraph id="A75736D5FFCD166B7984F938208E7558" blockId="6.[189,775,1602,2066]" pageId="6" pageNumber="22">
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DIAGNOSIS. A polytypic subgenus of
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<taxonomicName id="60E84D56FFCD166B7B3DF93B23B774B9" authorityName="Bonelli" authorityYear="1810" box="[603,724,1697,1718]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="22" phylum="Arthropoda" rank="genus">
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<emphasis id="959CEAC7FFCD166B7B3DF93B23B774B9" box="[603,724,1697,1718]" italics="true" pageId="6" pageNumber="22">Pterostichus</emphasis>
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</taxonomicName>
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from southern
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<collectingCountry id="DFFF7645FFCD166B7871F926203274DD" box="[279,337,1724,1746]" name="China" pageId="6" pageNumber="22">China</collectingCountry>
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to northern
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<collectingCountry id="DFFF7645FFCD166B78ABF926234074DD" box="[461,547,1724,1746]" name="Vietnam" pageId="6" pageNumber="22">Vietnam</collectingCountry>
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, defined chiefly by the combination of the body apterous, the metepisternum short,
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<emphasis id="959CEAC7FFCD166B79DBF96821B17508" box="[189,210,1778,1799]" italics="true" pageId="6" pageNumber="22">cx</emphasis>
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3 bisetose and
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<emphasis id="959CEAC7FFCD166B780FF968201A7508" box="[361,377,1778,1799]" italics="true" pageId="6" pageNumber="22">tr</emphasis>
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3 with seta. Most species also share the pronotum distinctly bisulcate on each side of base, the elytron with 1–2 discal setae (d2 and/or d3) adjoining stria 2, and the tarsomere 5 glabrous ventrally.
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</paragraph>
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<paragraph id="A75736D5FFCD166B7984F8C620C97A1D" blockId="6.[189,775,1602,2066]" pageId="6" pageNumber="22">
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Body setation otherwise complete: clypeus bisetose, two supraocular setae, two elytral apical setae in stria 7,
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<emphasis id="959CEAC7FFCD166B7B8FF8ED23997583" box="[745,762,1911,1932]" italics="true" pageId="6" pageNumber="22">fe</emphasis>
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1 posterior face trisetose,
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<emphasis id="959CEAC7FFCD166B78CDF80820DF75A8" box="[427,444,1938,1959]" italics="true" pageId="6" pageNumber="22">fe</emphasis>
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2 with two anteroventral setae in basal three fifths and two anterodorsal setae in apical two fifths,
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<emphasis id="959CEAC7FFCD166B799CF85D206875D3" box="[250,267,1991,2012]" italics="true" pageId="6" pageNumber="22">fe</emphasis>
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3 bisetose;
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<emphasis id="959CEAC7FFCD166B7810F85220E075D2" box="[374,387,1992,2013]" italics="true" pageId="6" pageNumber="22">ti</emphasis>
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3 toward apex with 1–3 lateral spiniform setae. Tarsomere 5 glabrous ventrally in more than 80% species of the subgenus.
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</paragraph>
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</subSubSection>
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<subSubSection id="EFF2655EFFCD166B7A34FEA125B07578" pageId="6" pageNumber="22" type="description">
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<paragraph id="A75736D5FFCD166B7A34FEA1247F7363" blockId="6.[812,1399,315,2071]" pageId="6" pageNumber="22">
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REDESCRIPTION (For recognition only). Body rather medium-sized, BL
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<quantity id="60109B30FFCD166B7A8EFECC252B7364" box="[1000,1096,342,364]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.6" metricValueMin="0.7" pageId="6" pageNumber="22" unit="mm" value="11.5" valueMax="16.0" valueMin="7.0">7–16 mm</quantity>
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. Dorsum shiny black.
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</paragraph>
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<paragraph id="A75736D5FFCD166B7A34FEEB22E4709E" blockId="6.[812,1399,315,2071]" pageId="6" pageNumber="22">Head medium-sized, with neck constriction distinct to imperceptible. Eyes more or less flattened and often rather small, with genae fairly long. Clypeus subsinuate apically, frontoclypeal suture fine and straight, imperceptible on sides. Frontal sulci smooth, usually moderately long, moderately deep and slightly diverging. Antennal scape with preapical seta, pedicel glabrous except for ventral seta and apical verticillate setae, antennomeres 4–11 pubescent. Mentum tooth bifid, submentum quadrisetose. Penultimate labial palpomere bisetose, apical labial and maxillary palpomeres subfusiform.</paragraph>
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<paragraph id="A75736D5FFCD166B7A34FD0C242F70F3" blockId="6.[812,1399,315,2071]" pageId="6" pageNumber="22">Pronotum cordate to quadrate or circular, bisetose on each side; these finely beaded. Base with two, well-impressed, basolateral sulci on each side, adjoining along base; inner sulcus separate from lateral groove, rarely absent.</paragraph>
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<paragraph id="A75736D5FFCD166B7A34FC9B24347193" blockId="6.[812,1399,315,2071]" pageId="6" pageNumber="22">
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Elytra rather wide at bases, with striae 1–7 or 1–6 inside humeral angle; humeri and humeral tooth distinct, parascutellar seta present, parascutellar striole short to missing. Interval 3 with 1–2 setae, d2 and/or d3; USS: 13–17, sometimes 20–21;
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<collectingCountry id="DFFF7645FFCD166B7AD3FCF122B7718F" box="[949,980,875,896]" name="United States of America" pageId="6" pageNumber="22">US</collectingCountry>
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arranged into three groups, anterior (5–7, mostly six,
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<collectingCountry id="DFFF7645FFCD166B7AF9FC1C22A27193" box="[927,961,902,924]" name="United States of America" pageId="6" pageNumber="22">US</collectingCountry>
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), intermediate (1–2), and posterior (8–9).
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</paragraph>
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<paragraph id="A75736D5FFCD166B7A34FC3B22E17658" blockId="6.[812,1399,315,2071]" pageId="6" pageNumber="22">
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Prosternal process with fine apical bead only (abraded in some older specimens); inclination wide, convex to flat, with margins edged or beaded, or not. Abdominal sternite VII bi- (
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<specimenCount id="B1EEFD5CFFCD166B7A52FC7422297609" box="[820,842,1006,1030]" count="1" pageId="6" pageNumber="22" type="male">♂</specimenCount>
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) or quadrisetose (
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<specimenCount id="B1EEFD5CFFCD166B7A9CFC7425647609" box="[1018,1031,1006,1030]" count="1" pageId="6" pageNumber="22" type="female">♀</specimenCount>
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) at apex, not or slightly modified into a shallow median concavity in male, sometimes with very fine median carina at bottom; rarely with a small median tubercle.
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</paragraph>
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<paragraph id="A75736D5FFCD166B7A34FBC6247B76CE" blockId="6.[812,1399,315,2071]" pageId="6" pageNumber="22">
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Legs strong;
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<emphasis id="959CEAC7FFCD166B7AB4FBC622BC767E" box="[978,991,1116,1137]" italics="true" pageId="6" pageNumber="22">ti</emphasis>
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1 with anterior face smooth and glabrous; tarsi carinate laterally and sulcate on each side of the carina. Protarsomeres 1–3 dilated, with ventral pad, in male, or strongly toothed at ventro-apical angles in female.
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</paragraph>
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<paragraph id="A75736D5FFCD166B7A34FB5C24127798" blockId="6.[812,1399,315,2071]" pageId="6" pageNumber="22">Aedeagus rather typical of the genus, median lobe in lateral view strongly curved to nearly geniculate; apex in dorsal view small triangular to trapezoidal. Right paramere mostly long, arcuate, narrow to very wide, sometimes hooked or moderately long, or short. Everted and inflated internal sac curved to the left and then ventrad and slightly basad; its inner curve mostly with an oblong, more or less sclerotized, concavity to receive a vesicle in form of an oblong sclerite.</paragraph>
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<paragraph id="A75736D5FFCD166B7A34FA0125B07578" blockId="6.[812,1399,315,2071]" pageId="6" pageNumber="22">
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Female tergite VIII fairly wide, rounded at apex; sternite VIII fairly wide, rounded on sides, evenly sclerotized (pigmented), with a small to indistinct, depigmented, latero-apical area; apical pubescence mostly long, dense, absent from sides, medial setae being longer; basolateral apodemes moderately wide, about half as wide at base as long, subtriangular or rectangular, sclerotized same as sternite proper. Tergite IX: laterotergite apically membranous, rounded, densely setose; gonosubcoxite with a row of dense ensiform setae along latero-apical margin. Gonocoxite crescent, with apex pointed, preapical nematiform seta, one inner (d, dorsal) and two outer (v, ventral) ensiform setae. This primary
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<typeStatus id="78538877FFCD166B7DFAF95B25A674D9" box="[1180,1221,1729,1750]" pageId="6" pageNumber="22">type</typeStatus>
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gonocoxite modified into subtriangular to elliptic one in most species, with apex rounded or blunt, and ensiform setae varying considerably between species, so that 1d+1v, 2d+2v, 2–3d+3–4v combinations occur. Reproductive tract as for
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<taxonomicName id="60E84D56FFCD166B7DDEF8B12413754F" authorityName="Tschitscherine" authorityYear="1898" box="[1208,1392,1835,1856]" pageId="6" pageNumber="17" rank="subGenus" subGenus="Morphohaptoderus">
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<emphasis id="959CEAC7FFCD166B7DDEF8B12413754F" box="[1208,1392,1835,1856]" italics="true" pageId="6" pageNumber="22">Morphohaptoderus</emphasis>
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</taxonomicName>
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, except for receptacle curved and seminal canal lacking basal sclerotization while varying much in length.
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</paragraph>
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</subSubSection>
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<subSubSection id="EFF2655EFFCD16657A34F8E123A273D8" lastPageId="8" lastPageNumber="24" pageId="6" pageNumber="22" type="discussion">
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<paragraph id="A75736D5FFCD166B7A34F8E125CB75A3" blockId="6.[812,1399,315,2071]" pageId="6" pageNumber="22">
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COMMENTS. This subgenus includes 17 described and many undescribed species from
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<collectingCountry id="DFFF7645FFCD166B7D01F80C25C775A3" box="[1127,1188,1942,1964]" name="China" pageId="6" pageNumber="22">China</collectingCountry>
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.
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</paragraph>
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<paragraph id="A75736D5FFCD166A7A34F82B218D73FE" blockId="6.[812,1399,315,2071]" lastBlockId="7.[189,775,312,657]" lastPageId="7" lastPageNumber="23" pageId="6" pageNumber="22">
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Some significant characters are polythetic within the subgenus. For instance, some species have the right paramere short (
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<emphasis id="959CEAC7FFCD166B7A0BF87D22E275F3" box="[877,897,2023,2044]" italics="true" pageId="6" pageNumber="22">vs</emphasis>
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. long in most of the others). Furthermore,
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<emphasis id="959CEAC7FFCD166B7C4BF87D245E75F3" box="[1325,1341,2023,2044]" italics="true" pageId="6" pageNumber="22">tr</emphasis>
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3 seta has been found to be missing in one species while polymorphic in another one,
|
||
<emphasis id="959CEAC7FFCC166A78E7FEA620FC735E" box="[385,415,316,337]" italics="true" pageId="7" pageNumber="23">i.e.</emphasis>
|
||
, either present (
|
||
<specimenCount id="B1EEFD5CFFCC166A7B5DFEA12319735F" box="[571,634,312,336]" count="2" pageId="7" pageNumber="23" type="male">2 ♂♂</specimenCount>
|
||
) or missing (
|
||
<specimenCount id="B1EEFD5CFFCC166A7B9CFEA121BB7364" count="2" pageId="7" pageNumber="23" type="female">2 ♀♀</specimenCount>
|
||
), or vestigial unilateral (
|
||
<specimenCount id="B1EEFD5CFFCC166A7886FEC9208E7364" box="[480,493,339,363]" count="1" pageId="7" pageNumber="23" type="female">♀</specimenCount>
|
||
). Inner apical setae of the abdominal sternite VII tend to disappear in females of at least two species. One of them is described below and
|
||
<emphasis id="959CEAC7FFCC166A7BA7FE1623B373AE" box="[705,720,396,417]" italics="true" pageId="7" pageNumber="23">P</emphasis>
|
||
. (
|
||
<emphasis id="959CEAC7FFCC166A7B8EFE16239B73AE" box="[744,760,396,417]" italics="true" pageId="7" pageNumber="23">N</emphasis>
|
||
.)
|
||
<emphasis id="959CEAC7FFCC166A79DBFE3C203573B4" box="[189,342,422,443]" italics="true" pageId="7" pageNumber="23">szetschuanensis</emphasis>
|
||
(
|
||
<bibRefCitation id="C3794B24FFCC166A7801FE3C235B73B3" author="Tschitscherine T." box="[359,568,422,444]" pageId="7" pageNumber="23" pagination="185 - 198" refId="ref20594" refString="Tschitscherine T. 1889. Insecta, a Cl. G. N. Potanin in China et in Mongolia novissime lecta. Insectes rapportes par Mr. Potanin de son voyage fait en 1884 - 85 - 86. VI. Genre Pterostichus // Horae Soc. Ent. Ross. T. 23. P. 185 - 198." type="journal article" year="1889">Tschitschérine, 1889</bibRefCitation>
|
||
) has been found to have 1+2 apical setae in one of
|
||
<specimenCount id="B1EEFD5CFFCC166A7888FE5B233973D8" box="[494,602,449,471]" count="2" pageId="7" pageNumber="23" type="female">two female</specimenCount>
|
||
specimens examined.
|
||
</paragraph>
|
||
<paragraph id="A75736D5FFCC16657984FE6C23A273D8" blockId="7.[189,775,312,657]" lastBlockId="8.[189,775,315,471]" lastPageId="8" lastPageNumber="24" pageId="7" pageNumber="23">
|
||
<bibRefCitation id="C3794B24FFCC166A7984FE6C20117004" author="Sciaky R." box="[226,370,501,524]" pageId="7" pageNumber="23" pagination="1 - 19" refId="ref20291" refString="Sciaky R. 1994. Revision of Pterostichus subg. Morphohaptoderus Tschitscherine, 1898 with description of ten new species from China (Coleoptera: Carabidae) // Koleopt. Rundsch. Bd. 64. P. 1 - 19." type="book chapter" year="1994">Sciaky [1994]</bibRefCitation>
|
||
supposed the closest relationships between
|
||
<taxonomicName id="60E84D56FFCC166A799AFD8B20EC7029" authorityName="Tschitscherine" authorityYear="1898" box="[252,399,529,550]" pageId="7" pageNumber="22" rank="subGenus" subGenus="Neohaptoderus">
|
||
<emphasis id="959CEAC7FFCC166A799AFD8B20EC7029" box="[252,399,529,550]" italics="true" pageId="7" pageNumber="23">Neohaptoderus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="60E84D56FFCC166A78D8FD8B231B7029" authorityName="Tschitscherine" authorityYear="1898" box="[446,632,529,550]" pageId="7" pageNumber="17" rank="subGenus" subGenus="Morphohaptoderus">
|
||
<emphasis id="959CEAC7FFCC166A78D8FD8B231B7029" box="[446,632,529,550]" italics="true" pageId="7" pageNumber="23">Morphohaptoderus</emphasis>
|
||
</taxonomicName>
|
||
. I do not share this hypothesis because many features combined, especially the short metepisternum (following apterous condition of the body),
|
||
<emphasis id="959CEAC7FFCC166A7864FDFB20747079" box="[258,279,609,630]" italics="true" pageId="7" pageNumber="23">cx</emphasis>
|
||
3 trisetose, and a peculiar internal sac of aedeagus, suggest rather isolated position of
|
||
<taxonomicName id="60E84D56FFCC166A7B6FFDE123A0709F" authorityName="Tschitscherine" authorityYear="1898" box="[521,707,635,656]" pageId="7" pageNumber="17" rank="subGenus" subGenus="Morphohaptoderus">
|
||
<emphasis id="959CEAC7FFCC166A7B6FFDE123A0709F" box="[521,707,635,656]" italics="true" pageId="7" pageNumber="23">Morphohaptoderus</emphasis>
|
||
</taxonomicName>
|
||
within
|
||
<taxonomicName id="60E84D56FFCC166A7A4AFEA122C6735F" authorityName="Bonelli" authorityYear="1810" box="[812,933,315,336]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFCC166A7A4AFEA122C6735F" box="[812,933,315,336]" italics="true" pageId="7" pageNumber="23">Pterostichus</emphasis>
|
||
</taxonomicName>
|
||
lineage D sensu Sasakawa and Kubota [2007].
|
||
<taxonomicName id="60E84D56FFCC166A7A4AFECC22DC7364" authorityName="Tschitscherine" authorityYear="1898" box="[812,959,342,363]" pageId="7" pageNumber="22" rank="subGenus" subGenus="Neohaptoderus">
|
||
<emphasis id="959CEAC7FFCC166A7A4AFECC22DC7364" box="[812,959,342,363]" italics="true" pageId="7" pageNumber="23">Neohaptoderus</emphasis>
|
||
</taxonomicName>
|
||
may instead be closer to the lineage or lineages defined by the combination of
|
||
<emphasis id="959CEAC7FFCC166A7D1AFEEB25F27389" box="[1148,1169,369,390]" italics="true" pageId="7" pageNumber="23">cx</emphasis>
|
||
3 bisetose and
|
||
<emphasis id="959CEAC7FFCC166A7C41FEEB24547389" box="[1319,1335,369,390]" italics="true" pageId="7" pageNumber="23">tr</emphasis>
|
||
3 with seta, among them
|
||
<taxonomicName id="60E84D56FFCC166A7A88FE11258173AF" authority="Reitter, 1892" authorityName="Reitter" authorityYear="1892" box="[1006,1250,395,417]" class="Insecta" family="Carabidae" genus="Agastillus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFCC166A7A88FE11252C73AF" box="[1006,1103,395,416]" italics="true" pageId="7" pageNumber="23">Agastillus</emphasis>
|
||
Reitter, 1892
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="60E84D56FFCC166A7D95FE1122CF73B3" authority="Reitter, 1896" authorityName="Reitter" authorityYear="1896" class="Insecta" family="Carabidae" genus="Haplomaseus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFCC166A7D95FE11241573AF" box="[1267,1398,395,416]" italics="true" pageId="7" pageNumber="23">Haplomaseus</emphasis>
|
||
Reitter, 1896
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="60E84D56FFCC166A7AD3FE3C25BA73B4" authority="Jakobson, 1907" authorityName="Jakobson" authorityYear="1907" box="[949,1241,422,444]" class="Insecta" family="Carabidae" genus="Oreoplatysma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFCC166A7AD3FE3C255D73B4" box="[949,1086,422,443]" italics="true" pageId="7" pageNumber="23">Oreoplatysma</emphasis>
|
||
Jakobson, 1907
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="60E84D56FFCC166A7C6CFE3C254973D9" authority="Kryzhanovskij, 1968" authorityName="Kryzhanovskij" authorityYear="1968" class="Insecta" family="Carabidae" genus="Asioplatysma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFCC166A7C6CFE3C222A73D9" italics="true" pageId="7" pageNumber="23">Asioplatysma</emphasis>
|
||
Kryzhanovskij, 1968
|
||
</taxonomicName>
|
||
. Moreover
|
||
<taxonomicName id="60E84D56FFCC166A7DC3FE5B223C73FF" authority="Sciaky, 1994" authorityName="Sciaky" authorityYear="1994" class="Insecta" family="Carabidae" genus="Sinosteropus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFCC166A7DC3FE5B244173D9" box="[1189,1314,449,470]" italics="true" pageId="7" pageNumber="23">Sinosteropus</emphasis>
|
||
<bibRefCitation id="C3794B24FFCC166A7C48FE5B223C73FF" author="Sciaky R." pageId="7" pageNumber="23" pagination="1 - 19" refId="ref20291" refString="Sciaky R. 1994. Revision of Pterostichus subg. Morphohaptoderus Tschitscherine, 1898 with description of ten new species from China (Coleoptera: Carabidae) // Koleopt. Rundsch. Bd. 64. P. 1 - 19." type="book chapter" year="1994">Sciaky, 1994</bibRefCitation>
|
||
</taxonomicName>
|
||
, seems to be the closest ally or only a group of
|
||
<taxonomicName id="60E84D56FFCC166A7C45FE4122177004" authorityName="Tschitscherine" authorityYear="1898" pageId="7" pageNumber="22" rank="subGenus" subGenus="Neohaptoderus">
|
||
<emphasis id="959CEAC7FFCC166A7C45FE4122177004" italics="true" pageId="7" pageNumber="23">Neohaptoderus</emphasis>
|
||
</taxonomicName>
|
||
, since the two taxa share many significant characters, including very similar reproductive tract, genitalia, and pregenital abdominal urites in female. Many other features of
|
||
<taxonomicName id="60E84D56FFCC166A7A4AFDDC22CA7054" authorityName="Sciaky" authorityYear="1994" box="[812,937,582,603]" class="Insecta" family="Carabidae" genus="Sinosteropus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFCC166A7A4AFDDC22CA7054" box="[812,937,582,603]" italics="true" pageId="7" pageNumber="23">Sinosteropus</emphasis>
|
||
</taxonomicName>
|
||
such as pronotal basal angles rounded, the outer basolateral sulcus lost, and the parascutellar seta often missing, are by comparison derived characters. Plesiomorphic states of some of these characters could be seen within the
|
||
<emphasis id="959CEAC7FFC3166579DBFECC20497364" box="[189,298,342,363]" italics="true" pageId="8" pageNumber="24">perhoplites</emphasis>
|
||
-group that was provisionally assigned to
|
||
<taxonomicName id="60E84D56FFC316657BACFECC20667389" authorityName="Sciaky" authorityYear="1994" class="Insecta" family="Carabidae" genus="Sinosteropus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="959CEAC7FFC316657BACFECC20667389" italics="true" pageId="8" pageNumber="24">Sinosteropus</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<bibRefCitation id="C3794B24FFC316657875FEEB20857389" author="Schmidt J. & Tian Ming-Yi." box="[275,486,368,391]" pageId="8" pageNumber="24" pagination="175 - 181" refId="ref20252" refString="Schmidt J., Tian Ming-Yi. 2011. Three remarkable new Pterostichus species from Southern Tibet (Insecta: Coleoptera: Carabidae: Pterostichini) // Vernate. Bd. 30. S. 175 - 181." type="book chapter" year="2011">Schmidt, Tian, 2011</bibRefCitation>
|
||
]. This group is distinctive in having pronotal basal angles sharp, the parascutellar seta constantly missing, and an undescribed species of this group from
|
||
<collectingCountry id="DFFF7645FFC316657994FE5B204F73D8" box="[242,300,449,471]" name="China" pageId="8" pageNumber="24">China</collectingCountry>
|
||
has the pronotum bisulcate on each side.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |