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<document id="C359168F6CAD759A053CAB74E2FC3D01" ID-ISSN="1175-5326" ID-ZooBank="9913C60F-5F0D-42ED-A5E6-109EB5156B4F" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629832609958" checkinUser="felipe" docAuthor="Behounek, G., Han, H. L. &amp; V. S. Kononenko" docDate="2012" docId="0F360A170A14FFEBA8810EA0E619F9EB" docLanguage="en" docName="zt03587p088.pdf" docOrigin="Zootaxa 3587" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Xizanga Behounek, Han &amp; Kononenko 2012, gen. n." docType="treatment" docVersion="3" lastPageNumber="83" masterDocId="F30F726F0A11FFEEA8160C2FE415FFA7" masterDocTitle="Two new genera and three new species of the Pantheinae from East Asia and China (Lepidoptera, Noctuidae). Contribution to revision of Pantheinae VIII" masterLastPageNumber="88" masterPageNumber="78" pageNumber="83" updateTime="1699118637777" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="F87080F02059C0A18E1688C440436037">Two new genera and three new species of the Pantheinae from East Asia and China (Lepidoptera, Noctuidae). Contribution to revision of Pantheinae VIII</mods:title>
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<mods:name id="236C7EF46390A1A3B6F309CFDCFE32B0" type="personal">
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<mods:namePart id="1928009988C1CA52E130C747E2AD46D0">Behounek, G.</mods:namePart>
<mods:affiliation id="4AD6F19F12847390FED9B198A14EF86E">D- 85567 Grafing / Munich, Sudetenstrasse 6, Germany. E-mail: Gottfried. Behounek @ t-online. de</mods:affiliation>
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<mods:namePart id="ADDA9DDA02ACFC0B44ABA15860F95560">Han, H. L.</mods:namePart>
<mods:affiliation id="38745207512198BCBD02016187A80E55">School of Forestry, Northeast Forestry University, Harbin, CH- 150014 China. E-mail: hanhuilin @ yahoo. com. cn</mods:affiliation>
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<mods:namePart id="CD775978F2DDC5736FBEA5FD9266AC22">V. S. Kononenko</mods:namePart>
<mods:affiliation id="887BD3A260E92B968E1D1DD986152D07">Laboratory of Entomology, Institute of Biology and Soil Science Far Eastern Branch of Russian Academy of Sciences, RF- 690022 Vladivostok, Russia. E-mail: kononenko @ ibss. dvo. ru, vlad _ kononenko @ mail. ru</mods:affiliation>
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<paragraph id="8720BB010A14FFEBA8810EA0E739FD0E" blockId="5.[151,812,655,681]" box="[151,812,655,681]" pageId="5" pageNumber="83">
<heading id="DC680C6D0A14FFEBA8810EA0E739FD0E" bold="true" box="[151,812,655,681]" fontSize="11" level="1" pageId="5" pageNumber="83" reason="1">
<emphasis id="B5EB67130A14FFEBA8810EA0E739FD0E" bold="true" box="[151,812,655,681]" pageId="5" pageNumber="83">
Genus
<taxonomicName id="409FC0820A14FFEBA8FB0EA0E6C7FD0E" authority="Behounek, Han &amp; Kononenko" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[237,722,655,681]" class="Insecta" family="Noctuidae" genus="Xizanga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="83" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B5EB67130A14FFEBA8FB0EA0E545FD0E" bold="true" box="[237,336,655,681]" italics="true" pageId="5" pageNumber="83">Xizanga</emphasis>
Behounek, Han &amp; Kononenko
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<taxonomicNameLabel id="AED8DA680A14FFEBAACF0EA0E739FD0E" box="[729,812,655,681]" pageId="5" pageNumber="83" rank="genus">gen. n.</taxonomicNameLabel>
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<subSubSection id="CF85E88A0A14FFEBA8810EF8E6C0FD4B" box="[151,725,726,749]" pageId="5" pageNumber="83" type="type_taxon">
<paragraph id="8720BB010A14FFEBA8810EF8E6C0FD4B" blockId="5.[151,725,726,749]" box="[151,725,726,749]" pageId="5" pageNumber="83">
<emphasis id="B5EB67130A14FFEBA8810EF8E533FD4A" bold="true" box="[151,294,726,749]" pageId="5" pageNumber="83">
<typeStatus id="582405A30A14FFEBA8810EF8E4D9FD4B" box="[151,204,727,748]" pageId="5" pageNumber="83">Type</typeStatus>
species:
</emphasis>
<emphasis id="B5EB67130A14FFEBA93D0EF8E5F8FD4B" box="[299,493,727,748]" italics="true" pageId="5" pageNumber="83">
<taxonomicName id="409FC0820A14FFEBA93D0EF8E5FFFD4B" authority="Behounek &amp; Han &amp; V.S.Kononenko, 2012" authorityName="Behounek &amp; Han &amp; V.S.Kononenko" authorityYear="2012" box="[299,490,727,748]" class="Insecta" family="Noctuidae" genus="Xizanga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="83" phylum="Arthropoda" rank="species" species="mysterica" status="sp. nov.">Xizanga mysterica</taxonomicName>
,
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<taxonomicNameLabel id="AED8DA680A14FFEBA9E20EF8E63FFD4B" box="[500,554,727,748]" pageId="5" pageNumber="83" rank="species">sp. n.</taxonomicNameLabel>
designated here.
</paragraph>
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<subSubSection id="CF85E88A0A14FFEBA8810F36E792FCF2" pageId="5" pageNumber="83" type="diagnosis">
<paragraph id="8720BB010A14FFEBA8810F36E792FCF2" blockId="5.[151,1436,793,1612]" pageId="5" pageNumber="83">
<emphasis id="B5EB67130A14FFEBA8810F36E504FC95" bold="true" box="[151,273,793,818]" inLineHeading="true" pageId="5" pageNumber="83" reason="1">Diagnosis.</emphasis>
No genus of the
<taxonomicName id="409FC0820A14FFEBA9C00F36E659FC96" box="[470,588,793,817]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="83" phylum="Arthropoda" rank="family">Noctuidae</taxonomicName>
, and
<taxonomicName id="409FC0820A14FFEBAA9E0F36E716FC96" box="[648,771,793,817]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="83" phylum="Arthropoda" rank="subFamily" subFamily="Pantheinae">Pantheinae</taxonomicName>
in particular, in the Chinese-Himalayan fauna is similar to
<taxonomicName id="409FC0820A14FFEBA8810F11E4E6FCF2" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[151,243,830,853]" class="Insecta" family="Noctuidae" genus="Xizanga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B5EB67130A14FFEBA8810F11E4E6FCF2" box="[151,243,830,853]" italics="true" pageId="5" pageNumber="83">Xizanga</emphasis>
</taxonomicName>
in external and genitalic characters; see description below.
</paragraph>
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<subSubSection id="CF85E88A0A14FFEBA8D10F4EE034FAD4" pageId="5" pageNumber="83" type="description">
<paragraph id="8720BB010A14FFEBA8D10F4EE034FAD4" blockId="5.[151,1436,793,1612]" pageId="5" pageNumber="83">
<emphasis id="B5EB67130A14FFEBA8D10F4EE54DFCDD" bold="true" box="[199,344,865,890]" inLineHeading="true" pageId="5" pageNumber="83" reason="1">Description.</emphasis>
Adult (
<figureCitation id="1FA4A7840A14FFEBA9A60F4EE642FCDE" box="[432,599,865,889]" captionStart="FIGURES 915" captionStartId="7.[151,263,1841,1864]" captionTargetBox="[152,1436,193,1813]" captionTargetId="figure-68@7.[151,1436,193,1821]" captionTargetPageId="7" captionText="FIGURES 915. Imagines, wing venation, male genitalia. 9. Xizanga mysterica, male, holotype, China, Xizang; 10. X. mysterica, female, paratype, genit. prep. HHL-2449-2; 11. X. mysterica, wing venation; 12. X. mysterica, male genitalia, paratype, genit. prep. HHL-1056; 13. Anacronicta caliginea, Russia, Primorye, genit. prep. 272 VK IBSS." pageId="5" pageNumber="83">Figs. 9, 10, 11</figureCitation>
). Frons bulged, covered by short scales; eyes rounded, naked, surrounded with “lashes”; ocelli present; antennae of male filiform; proboscis well developed, short; labial palps moderate, not compressed, extending to mid of frons; 3
<superScript id="70EA16490A14FFEBAA6D0F86E69DFC10" attach="left" box="[635,648,937,951]" fontSize="6" pageId="5" pageNumber="83">rd</superScript>
segment of palps small, about two times smaller than 2
<superScript id="70EA16490A14FFEBAD090F86E13BFC10" attach="none" box="[1311,1326,937,951]" fontSize="6" pageId="5" pageNumber="83">nd</superScript>
, pointed; Thorax rather stout, covered with broad scales; hindwing venation with M2 well developed, unlike to other
<taxonomicName id="409FC0820A14FFEBA8810FDBE507FBAB" box="[151,274,1012,1036]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="83" phylum="Arthropoda" rank="subFamily" subFamily="Pantheinae">Pantheinae</taxonomicName>
genera arising from the middle of discal cell. Male genitalia (
<figureCitation id="1FA4A7840A14FFEBABDA0FDBE00AFBAB" box="[972,1055,1012,1036]" captionStart="FIGURES 915" captionStartId="7.[151,263,1841,1864]" captionTargetBox="[152,1436,193,1813]" captionTargetId="figure-68@7.[151,1436,193,1821]" captionTargetPageId="7" captionText="FIGURES 915. Imagines, wing venation, male genitalia. 9. Xizanga mysterica, male, holotype, China, Xizang; 10. X. mysterica, female, paratype, genit. prep. HHL-2449-2; 11. X. mysterica, wing venation; 12. X. mysterica, male genitalia, paratype, genit. prep. HHL-1056; 13. Anacronicta caliginea, Russia, Primorye, genit. prep. 272 VK IBSS." pageId="5" pageNumber="83">Fig. 12</figureCitation>
). Uncus short, relatively massive, extended basally, tegumen equal to vinculum, with broad lateral lobes; paratergal sclerites narrow, recurved; juxta moderate, anchor-like; vinculum rather massive with quadrangular saccus; valva short, broad basally, tapering distally and extended apically, costa very strong, heavily sclerotised, extended apically to abrupt tip of valva; basal lobe of sacculus very broad, elongate; clasper positioned longitudinally, as sclerotised plate; harpe missing; ventral margin of valva with distal sclerotised extension (saccular extension); presence of costal and ventral extension forms bifurcated ending of valva. Aedeagus straight, slightly curved basally; carina with scobinated ring-like band and strong, claw-like subbasal cornutus; vesica tubular, long, not armed. Female genitalia (
<figureCitation id="1FA4A7840A14FFEBACA008DFE118FAAF" box="[1206,1293,1264,1288]" captionStart="FIGURES 1723" captionStartId="9.[151,263,1877,1900]" captionTargetBox="[151,1436,203,1844]" captionTargetId="figure-65@9.[151,1436,193,1856]" captionTargetPageId="9" captionText="FIGURES 1723. Female genitalia, distribution map. 17. Anacronicta caliginea, Russia, Primorye, genit. prep. 273VK; 18. Flavala flavala, Nepal, genit. prep. 4152GB; 19. Xizanga mysterica, China, paratype, genit. prep. HHL-2449-2. Distribution map of Flavala flavala (20), F. crypta (21), F. secunda (22) and Xizanga mysterica (23)." pageId="5" pageNumber="83">Fig. 19</figureCitation>
). Ovipositor rather small; papillae anales quadrangular, covered with hairs; apophyses anteriores and posteriores short, equal in length; antrum funnel-like, with two lateral lobes with twin cut in centre; ductus bursae short, rounded, sclerotised; corpus bursae membranous, narrow, tube-like, extended to pyriform bottom part.
</paragraph>
</subSubSection>
<subSubSection id="CF85E88A0A14FFEBA8D109AFE619F9EB" pageId="5" pageNumber="83" type="etymology">
<paragraph id="8720BB010A14FFEBA8D109AFE520FA1B" blockId="5.[151,1436,793,1612]" pageId="5" pageNumber="83">
<emphasis id="B5EB67130A14FFEBA8D109AFE558FA3F" bold="true" box="[199,333,1408,1432]" inLineHeading="true" pageId="5" pageNumber="83" reason="1">Etymology.</emphasis>
The generic name
<taxonomicName id="409FC0820A14FFEBAA2209AEE685FA3F" authorityName="Behounek, Han &amp; Kononenko" authorityYear="2012" box="[564,656,1409,1432]" class="Insecta" family="Noctuidae" genus="Xizanga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B5EB67130A14FFEBAA2209AEE685FA3F" box="[564,656,1409,1432]" italics="true" pageId="5" pageNumber="83">Xizanga</emphasis>
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refers to the
<typeStatus id="582405A30A14FFEBAB2209AEE771FA3E" box="[820,868,1409,1433]" pageId="5" pageNumber="83">type</typeStatus>
locality of the type-species,
<collectingRegion id="455B75E30A14FFEBACAD09AFE16CFA3F" box="[1211,1401,1408,1432]" country="China" name="Xizang" pageId="5" pageNumber="83">Xizang province</collectingRegion>
of
<collectingCountry id="FF88FB910A14FFEBA881098BE4CFFA1B" box="[151,218,1444,1468]" name="China" pageId="5" pageNumber="83">China</collectingCountry>
(
<collectingRegion id="455B75E30A14FFEBA8FE098BE53CFA1B" box="[232,297,1444,1468]" country="China" name="Xizang" pageId="5" pageNumber="83">Tibet</collectingRegion>
).
</paragraph>
<paragraph id="8720BB010A14FFEBA8D109E7E619F9EB" blockId="5.[151,1436,793,1612]" pageId="5" pageNumber="83">
<emphasis id="B5EB67130A14FFEBA8D109E7E504FA47" bold="true" box="[199,273,1480,1504]" inLineHeading="true" pageId="5" pageNumber="83" reason="1">Notes.</emphasis>
The systematic position of this genus is unclear. We did not find any genus related to the new one among
<taxonomicName id="409FC0820A14FFEBA8FC09C3E570F9A3" box="[234,357,1516,1540]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="83" phylum="Arthropoda" rank="subFamily" subFamily="Pantheinae">Pantheinae</taxonomicName>
and other subfamilies of the “lower trifinae”. We tentatively place the new genus in
<taxonomicName id="409FC0820A14FFEBAD3709C3E189F9A3" box="[1313,1436,1516,1540]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="83" phylum="Arthropoda" rank="subFamily" subFamily="Pantheinae">Pantheinae</taxonomicName>
due to the quadrifid venation of the hindwing with developed vein M2, arising from the middle of discal cell, the bulged frons and short proboscis.
</paragraph>
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