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<document id="CC95317B3DB594EA7CFCB22D1ABAC847" ID-CLB-Dataset="281027" ID-DOI="10.11646/zootaxa.5383.1.6" ID-GBIF-Dataset="6574f0d6-27c3-4e8b-ae8a-e37d774f5971" ID-ISSN="1175-5326" ID-Zenodo-Dep="10301472" ID-ZooBank="DF7AA4C1-DE85-4D6D-A6DE-5D07DB1B0F47" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1702023627589" checkinUser="plazi" docAuthor="Singh, Pratima, Chowdhury, Basudhara Roy, Gurumayum, Shantabala Devi &amp; Kosygin, Laishram" docDate="2023" docId="FB3787AA590C850F869DFDBAFEAD8631" docLanguage="en" docName="zootaxa.5383.1.6.pdf" docOrigin="Zootaxa 5383 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.1.6/52441" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glyptothorax heokheei Singh &amp; Chowdhury &amp; Gurumayum &amp; Kosygin 2023, new species" docType="treatment" docVersion="4" lastPageNumber="79" masterDocId="070EFFD2590D850B860AFFFDFFAC8351" masterDocTitle="Glyptothorax heokheei, a new species of catfish (Teleostei: Sisoridae) from the Brahmaputra River drainage, Arunachal Pradesh, Northeast India" masterLastPageNumber="82" masterPageNumber="75" pageNumber="76" updateTime="1702036731562" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="969C735BF90D2E4001F023908F4CD905" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CD5B425CD6BC72E7B9634FF01D5C2745">
<mods:title id="8DFFE25CA64C7DC984717BD37443E494">Glyptothorax heokheei, a new species of catfish (Teleostei: Sisoridae) from the Brahmaputra River drainage, Arunachal Pradesh, Northeast India</mods:title>
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<mods:name id="2699A9314E02C41637322BAF6A524EC7" type="personal">
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<mods:namePart id="3DF09910DD652A5D2D929D88BF3A746D">Singh, Pratima</mods:namePart>
<mods:nameIdentifier id="DB20E4936B5F8CB8A3FA39E7F179AA03" type="ORCID">0000-0002-3785-3918</mods:nameIdentifier>
<mods:affiliation id="803636C839A848105C1C730A856A758D">Zoological Survey of India, Freshwater Fish Section, 27 J. L. Nehru Road, Kolkata, India &amp; singhpratimakol @ gmail. com; https: // orcid. org / 0000 - 0002 - 3785 - 3918</mods:affiliation>
<mods:nameIdentifier id="61EC4E56B28DAC6556B11248EAD57968" type="email">singhpratimakol@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="ADC8D4A58A9AF5C97AF96F5333724FE9" type="personal">
<mods:role id="8F4D8D923572871A02FCAC1F6C55E1EB">
<mods:roleTerm id="3B62B82406143E52391BF7B7406900E6">Author</mods:roleTerm>
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<mods:namePart id="14691A2B528B5AD006FABF207290BEDB">Chowdhury, Basudhara Roy</mods:namePart>
<mods:nameIdentifier id="0EBFBD102A5D7402FF34C385295D28D0" type="ORCID">0000-0001-7356-6441</mods:nameIdentifier>
<mods:affiliation id="FAA4362E0E3E8D3B9C65E55499943AAA">Zoological Survey of India, Freshwater Fish Section, 27 J. L. Nehru Road, Kolkata, India &amp; basudhararoychowdhury @ rocketmail. com; https: // orcid. org / 0000 - 0001 - 7356 - 6441</mods:affiliation>
<mods:nameIdentifier id="3DBD717D9ECF69BF6964140E57CC1D8D" type="email">basudhararoychowdhury@rocketmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="D3B359706777E237A7D13AE27CF4042E" type="personal">
<mods:role id="229C6343C1A6231BD342326AD48C42A8">
<mods:roleTerm id="4D3EFB284BF669D610BAB8127ED091C2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="324807FF98120C9733342C02CC30C7C9">Gurumayum, Shantabala Devi</mods:namePart>
<mods:nameIdentifier id="351842988154571F7F6A5E9C9ACCB34F" type="ORCID">0000-0003-0917-8158</mods:nameIdentifier>
<mods:affiliation id="41B8E89D15720D0CC22406DF29D42E8F">Zoological Survey of India, Arunachal Pradesh Regional Centre, Senki Valley, Itanagar, Arunachal Pradesh, India &amp; santaguru @ refiffmail. com; https: // orcid. org / 0000 - 0003 - 0917 - 8158</mods:affiliation>
<mods:nameIdentifier id="3B0B67B217BE305E9DB4933B29FDF435" type="email">santaguru@refiffmail.com</mods:nameIdentifier>
</mods:name>
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<mods:roleTerm id="E5F047606F58726F497DDBDE227B2025">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2C081A24797EBE02D81E7FE06FD3EDA2">Kosygin, Laishram</mods:namePart>
<mods:nameIdentifier id="CC7B64BF0391996A95F354AD1C413819" type="ORCID">0000-0002-9566-2247</mods:nameIdentifier>
<mods:affiliation id="D2BAE4EECE584D96C19A362C608D09FC">Zoological Survey of India, Freshwater Fish Section, 27 J. L. Nehru Road, Kolkata, India &amp; lkzsi 5 @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9566 - 2247</mods:affiliation>
<mods:nameIdentifier id="3AA23C1C6F0BFE44EF2A6B16DC615B68" type="email">lkzsi5@yahoo.com</mods:nameIdentifier>
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<mods:typeOfResource id="1178A9AE10A3E3B15AA7701227331068">text</mods:typeOfResource>
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<mods:title id="31A8D02968E802B46BDB0C7A172B3B99">Zootaxa</mods:title>
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<mods:part id="60204D07F9F1B918E9C45EB1619836CB">
<mods:date id="DBDD2A4B59CC8B13C80C15E30F1C9E70">2023</mods:date>
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<mods:number id="9D4E0514D17E3ED12AB066D58E6B531C">2023-12-08</mods:number>
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<mods:number id="B88B48AB5880B460C2101D1D0E13167C">5383</mods:number>
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<mods:classification id="26868725C2C7BFCCD6E97365068DCB95">journal article</mods:classification>
<mods:identifier id="307B51A1A23A1C034E10B9F3000A6850" type="CLB-Dataset">281027</mods:identifier>
<mods:identifier id="A76FB01152864E875B4FC610070F322B" type="DOI">10.11646/zootaxa.5383.1.6</mods:identifier>
<mods:identifier id="B1AF81793E880E50EE58618524B3C909" type="GBIF-Dataset">6574f0d6-27c3-4e8b-ae8a-e37d774f5971</mods:identifier>
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<treatment id="FB3787AA590C850F869DFDBAFEAD8631" ID-GBIF-Taxon="219596767" LSID="urn:lsid:plazi:treatment:FB3787AA590C850F869DFDBAFEAD8631" httpUri="http://treatment.plazi.org/id/FB3787AA590C850F869DFDBAFEAD8631" lastPageId="4" lastPageNumber="79" pageId="1" pageNumber="76">
<subSubSection id="3B846537590C850A869DFDBAFDE68130" box="[151,586,583,610]" pageId="1" pageNumber="76" type="nomenclature">
<paragraph id="732136BC590C850A869DFDBAFDE68130" blockId="1.[151,586,583,645]" box="[151,586,583,610]" pageId="1" pageNumber="76">
<heading id="286981D0590C850A869DFDBAFDE68130" bold="true" box="[151,586,583,610]" fontSize="11" level="1" pageId="1" pageNumber="76" reason="1">
<emphasis id="41EAEAAE590C850A869DFDBAFDE68130" bold="true" box="[151,586,583,610]" pageId="1" pageNumber="76">
<taxonomicName id="B49E4D3F590C850A869DFDBAFE078133" authority="Singh &amp; Chowdhury &amp; Gurumayum &amp; Kosygin, 2023" authorityName="Singh &amp; Chowdhury &amp; Gurumayum &amp; Kosygin" authorityYear="2023" box="[151,427,583,610]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="76" phylum="Chordata" rank="species" species="heokheei" status="sp. nov.">
<emphasis id="41EAEAAE590C850A869DFDBAFE078133" bold="true" box="[151,427,583,610]" italics="true" pageId="1" pageNumber="76">Glyptothorax heokheei</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5AD957D5590C850A87B0FDBAFDE68130" box="[442,586,583,609]" pageId="1" pageNumber="76" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3B846537590C850A869DFD96FEB281D4" box="[151,286,619,645]" pageId="1" pageNumber="76" type="description">
<paragraph id="732136BC590C850A869DFD96FEB281D4" blockId="1.[151,586,583,645]" box="[151,286,619,645]" pageId="1" pageNumber="76">
(
<figureCitation id="EBA52A39590C850A8695FD96FF4181D4" box="[159,237,619,645]" captionStart="FIGURE 1" captionStartId="3.[151,250,1941,1965]" captionTargetBox="[151,1436,181,1916]" captionTargetId="figure-17@3.[151,1436,181,1916]" captionTargetPageId="3" captionText="FIGURE 1. Glyptothorax heokheei, ZSI FF 9555, holotype, 152.0 mm SL: a, dorsal; b, lateral; and c, ventral aspects." figureDoi="http://doi.org/10.5281/zenodo.10301474" httpUri="https://zenodo.org/record/10301474/files/figure.png" pageId="1" pageNumber="76">Figs. 1</figureCitation>
,
<figureCitation id="EBA52A39590C850A86F1FD96FEBA81D4" box="[251,278,619,645]" captionStart="FIGURE 2" captionStartId="4.[151,250,1063,1087]" captionTargetBox="[161,1426,191,1029]" captionTargetId="figure-19@4.[151,1436,181,1039]" captionTargetPageId="4" captionText="FIGURE 2. Thoracic adhesive apparatus of a. Glyptothorax heokheei sp. nov., ZSI FF 9556, paratype, 124.1 mm SL; b. G. dikrongensis, ZSI V/APFS/P-515, holotype, 94.2 mm SL. Illustrations not to scale." figureDoi="http://doi.org/10.5281/zenodo.10301480" httpUri="https://zenodo.org/record/10301480/files/figure.png" pageId="1" pageNumber="76">2a</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3B846537590C850A869DFD4EFBBA8068" pageId="1" pageNumber="76" type="materials_examined">
<paragraph id="732136BC590C850A869DFD4EFE9E8044" blockId="1.[151,1437,691,2014]" pageId="1" pageNumber="76">
<emphasis id="41EAEAAE590C850A869DFD4EFEE3819C" bold="true" box="[151,335,691,717]" pageId="1" pageNumber="76">Type materials.</emphasis>
<materialsCitation id="C3F63CE1590C850A875FFD4EFE9E8044" ID-GBIF-Occurrence="4455028301" collectingDate="2020-02-25" collectorName="S. D. Gurumayum" country="India" county="East Siang District" latitude="28.153889" location="Mebo" longLatPrecision="20" longitude="95.37972" municipality="Siku" pageId="1" pageNumber="76" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
<typeStatus id="AC25881E590C850A875FFD4EFE6C819C" box="[341,448,691,717]" pageId="1" pageNumber="76" type="holotype">Holotype</typeStatus>
, ZSI FF 9555, 152.0 mm SL,
<collectingCountry id="0B89762C590C850A8501FD4EFCE5819C" box="[779,841,691,717]" name="India" pageId="1" pageNumber="76">India</collectingCountry>
:
<collectingRegion id="B15AF85E590C850A855BFD4EFB85819C" box="[849,1065,691,717]" country="India" name="Arunachal Pradesh" pageId="1" pageNumber="76">Arunachal Pradesh</collectingRegion>
:
<collectingCounty id="9A404E30590C850A8233FD49FAA2819C" box="[1081,1294,691,718]" pageId="1" pageNumber="76">East Siang District</collectingCounty>
:
<collectingMunicipality id="9345ACC6590C850A8312FD4EFAE7819C" box="[1304,1355,691,717]" pageId="1" pageNumber="76">Siku</collectingMunicipality>
stream near
<location id="76416067590C850A86C6FD2AFEBC81A0" LSID="urn:lsid:plazi:treatment:FB3787AA590C850F869DFDBAFEAD8631:76416067590C850A86C6FD2AFEBC81A0" box="[204,272,727,753]" country="India" county="East Siang District" latitude="28.153889" longLatPrecision="20" longitude="95.37972" municipality="Siku" name="Mebo" pageId="1" pageNumber="76" stateProvince="Arunachal Pradesh">Mebo</location>
, a tributary of the
<location id="76416067590C850A87DEFD2AFDFB81A0" LSID="urn:lsid:plazi:treatment:FB3787AA590C850F869DFDBAFEAD8631:76416067590C850A87DEFD2AFDFB81A0" box="[468,599,727,753]" country="India" county="East Siang District" latitude="28.153889" longLatPrecision="20" longitude="95.37972" municipality="Siku" name="Siang River" pageId="1" pageNumber="76" stateProvince="Arunachal Pradesh">Siang River</location>
(
<location id="76416067590C850A8469FD2AFC9781A0" LSID="urn:lsid:plazi:treatment:FB3787AA590C850F869DFDBAFEAD8631:76416067590C850A8469FD2AFC9781A0" box="[611,827,727,753]" country="India" county="East Siang District" latitude="28.153889" longLatPrecision="20" longitude="95.37972" municipality="Siku" name="Brahmaputra River" pageId="1" pageNumber="76" stateProvince="Arunachal Pradesh">Brahmaputra River</location>
drainage);
<geoCoordinate id="16AA507B590C850A85BCFD24FB9F81A5" box="[950,1075,729,756]" degrees="28" direction="north" minutes="9" orientation="latitude" pageId="1" pageNumber="76" precision="15" seconds="14" value="28.153889">28°9’14″N</geoCoordinate>
,
<geoCoordinate id="16AA507B590C850A8237FD2AFB6B81A5" box="[1085,1223,727,756]" degrees="95" direction="east" minutes="22" orientation="longitude" pageId="1" pageNumber="76" precision="15" seconds="47" value="95.37972">95°22’ 47″E</geoCoordinate>
,
<collectorName id="DE6B536A590C850A82DBFD2AFA3481A0" box="[1233,1432,727,753]" pageId="1" pageNumber="76">S.D. Gurumayum</collectorName>
,
<date id="0720107C590C850A869DFD06FE828044" box="[151,302,763,790]" pageId="1" pageNumber="76" value="2020-02-25">
<collectingDate id="1764E994590C850A869DFD06FE828044" box="[151,302,763,790]" pageId="1" pageNumber="76" value="2020-02-25">25 Feb. 2020</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="732136BC590C850A86CDFCDDFBBA8068" blockId="1.[151,1437,691,2014]" box="[199,1046,799,826]" pageId="1" pageNumber="76">
<materialsCitation id="C3F63CE1590C850A86CDFCDDFBBE8068" ID-GBIF-Occurrence="4455028302" box="[199,1042,799,826]" collectingDate="2020-02-25" collectorName="S. D. Gurumayum" country="India" county="East Siang District" latitude="28.153889" location="Mebo" longLatPrecision="20" longitude="95.37972" municipality="Siku" pageId="1" pageNumber="76" specimenCount="4" stateProvince="Arunachal Pradesh" typeStatus="paratype">
<typeStatus id="AC25881E590C850A86CDFCDDFE94806B" box="[199,312,800,826]" pageId="1" pageNumber="76" type="paratype">Paratypes</typeStatus>
. ZSI FF 9556,
<specimenCount id="6598FD35590C850A87D5FCE2FDB4806B" box="[479,536,799,826]" count="4" pageId="1" pageNumber="76" type="generic">4 ex.</specimenCount>
,
<quantity id="B4669B59590C850A842EFCE2FD4F806B" box="[548,739,799,826]" metricMagnitude="-1" metricUnit="m" metricValue="1.1324999999999998" metricValueMax="1.241" metricValueMin="1.024" pageId="1" pageNumber="76" unit="mm" value="113.25" valueMax="124.1" valueMin="102.4">102.4–124.1 mm</quantity>
SL, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B846537590C850A86CDFCBEFB838708" pageId="1" pageNumber="76" type="diagnosis">
<paragraph id="732136BC590C850A86CDFCBEFB838708" blockId="1.[151,1437,691,2014]" pageId="1" pageNumber="76">
<emphasis id="41EAEAAE590C850A86CDFCBEFEEE800C" bold="true" box="[199,322,835,861]" pageId="1" pageNumber="76">Diagnosis.</emphasis>
<taxonomicName id="B49E4D3F590C850A8747FCBEFDFD800C" authorityName="Singh &amp; Chowdhury &amp; Gurumayum &amp; Kosygin" authorityYear="2023" box="[333,593,835,861]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="76" phylum="Chordata" rank="species" species="heokheei">
<emphasis id="41EAEAAE590C850A8747FCBEFDFD800C" box="[333,593,835,861]" italics="true" pageId="1" pageNumber="76">Glyptothorax heokheei</emphasis>
</taxonomicName>
is diagnosed from all congeners in having a unique combination of the following characters: a chevron-shaped thoracic adhesive apparatus extending anteriorly onto gular region with more or less pointed anterior apex at isthmus, unculiferous striae prominent anteriorly from tip of median depression (
<figureCitation id="EBA52A39590C850A8695FC52FF4C8098" box="[159,224,943,969]" captionStart="FIGURE 2" captionStartId="4.[151,250,1063,1087]" captionTargetBox="[161,1426,191,1029]" captionTargetId="figure-19@4.[151,1436,181,1039]" captionTargetPageId="4" captionText="FIGURE 2. Thoracic adhesive apparatus of a. Glyptothorax heokheei sp. nov., ZSI FF 9556, paratype, 124.1 mm SL; b. G. dikrongensis, ZSI V/APFS/P-515, holotype, 94.2 mm SL. Illustrations not to scale." figureDoi="http://doi.org/10.5281/zenodo.10301480" httpUri="https://zenodo.org/record/10301480/files/figure.png" pageId="1" pageNumber="76">Fig 2</figureCitation>
); body depth at anus 12.8–14.1% SL; head length 25.1–28.8 % SL; interorbital width 12.0–20.0 % HL; post-adipose distance 16.0–18.4 % SL; posterior margin of pectoral-fin spine with 14–18 serrae; nasal barbel not reaching anterior margin of eye; arrow-shaped anterior nuchal plate element; ventral surface of pectoral spine and first pelvic-fin ray non-plaited; 9–10 branched anal-fin rays; vertical through pelvic-fin origin posterior to base of last dorsal-fin ray, head and body mottled with dark brown spots and smooth skin.
</paragraph>
</subSubSection>
<subSubSection id="3B846537590C850986CDFB9EFD2A84B6" lastPageId="2" lastPageNumber="77" pageId="1" pageNumber="76" type="description">
<paragraph id="732136BC590C850A86CDFB9EFC9186CC" blockId="1.[151,1437,691,2014]" pageId="1" pageNumber="76">
<emphasis id="41EAEAAE590C850A86CDFB9EFEF5872C" bold="true" box="[199,345,1123,1149]" pageId="1" pageNumber="76">Description.</emphasis>
Morphometric data presented in
<tableCitation id="3E1C0307590C850A84D7FB9EFC98872C" box="[733,820,1123,1149]" captionStart="TABLE 1" captionStartId="2.[151,239,296,320]" captionText="TABLE 1. Morphometric data of holotype (ZSI FF 9555) and four paratypes (ZSI FF 9556) of Glyptothorax heokheei." pageId="1" pageNumber="76">Table 1</tableCitation>
. Body elongate. Head depressed with almost conical snout when viewed dorsally. Dorsal profile rising evenly from tip of snout to dorsal fin origin, then sloping gently to end of caudal peduncle. Occipital process not in contact with anterior nuchal plate. Anterior nuchal plate broad, arrow-shaped extensions when viewed dorsally. Ventral profile almost flat up to posterior end of pelvic-fin base, then slightly convex to end of caudal-fin base. Caudal peduncle moderately shallow. Mouth inferior, lips papillate, teeth on upper jaw as a continuous band, its anterior margin almost rounded; teeth on lower jaw in two patches, separated by wide partition. Eyes small, round, located on dorsal surface of head. Anus and urogenital opening located at vertical through posterior half of adpressed pelvic-fin. Skin smooth. Lateral line complete and mid-lateral. Gills on first branchial arch long, bearing 4+8 or 5+8 rakers.
</paragraph>
<paragraph id="732136BC590C850A86CDFA5AFCB78500" blockId="1.[151,1437,691,2014]" pageId="1" pageNumber="76">
Thoracic adhesive apparatus consisting of unculiferous ridges present over chevron-shaped field and pointed anteriorly at isthmus. Unculiferous striae orientated longitudinally, except for posteriormost lateral ridge, which is slightly tilted towards midline of body, prominent anteriorly from tip of central median depression transversely. Median longitudinal striae extending anteriorly along ventral midline of body to cover gular region (
<figureCitation id="EBA52A39590C850A82EEF9EEFA98857C" box="[1252,1332,1555,1581]" captionStart="FIGURE 2" captionStartId="4.[151,250,1063,1087]" captionTargetBox="[161,1426,191,1029]" captionTargetId="figure-19@4.[151,1436,181,1039]" captionTargetPageId="4" captionText="FIGURE 2. Thoracic adhesive apparatus of a. Glyptothorax heokheei sp. nov., ZSI FF 9556, paratype, 124.1 mm SL; b. G. dikrongensis, ZSI V/APFS/P-515, holotype, 94.2 mm SL. Illustrations not to scale." figureDoi="http://doi.org/10.5281/zenodo.10301480" httpUri="https://zenodo.org/record/10301480/files/figure.png" pageId="1" pageNumber="76">Fig. 2a</figureCitation>
). Median depression poorly developed, without unculiferous striae.
</paragraph>
<paragraph id="732136BC590C850A86CDF9A6FE3085B0" blockId="1.[151,1437,691,2014]" pageId="1" pageNumber="76">Barbels in four pairs. Maxillary barbel shorter than head length, slender, extending to middle of pectoral-fin base. Outer mandibular barbel longer than inner, extending to anterior margin of pectoral fin base. Nasal barbel slender, not reaching anterior margin of orbit when adpressed, extending to half distance between its base and anterior orbital margin.</paragraph>
<paragraph id="732136BC590C850986CDF916FD2983A8" blockId="1.[151,1437,691,2014]" lastBlockId="2.[151,1437,151,249]" lastPageId="2" lastPageNumber="77" pageId="1" pageNumber="76">Dorsal-fin located equidistant between snout tip and adipose fin origin, with ii, 6 (5) rays; its height greater than body depth at dorsal-fin origin. Dorsal-fin spine strong, almost straight, strongly rugose posteriorly with 11–17 ridges, and smaller than depth of body at its origin. Adipose-fin low, with anterior margin straight and posterior margin slightly convex, deeply incised with the dorsoposterior margin uniformly convex and attached to body. Pectoral fin shorter than head, with i, 8 (2) or i, 9 (2) or 10 (1) rays; its posterior margin concave. Pectoral-fin spine broad, long; its anterior margin smooth, and its posterior margin with 14–18 serrae. Pelvic-fin with i, 5 (4) or i, 6 (1) rays, its tip not reaching anal-fin origin when adpressed. Vertical through pelvic-fin origin posterior to vertical through posterior end of dorsal-fin base. Ventral surface of pectoral-fin spine and first pelvic-fin ray not plaited. Anal fin long, with iv, 9 (4), or iv, 10 (1) rays; its origin almost vertically opposite to adipose-fin origin. Caudal fin deeply forked, with 8 + 9 (5) branched rays.</paragraph>
<caption id="27E16634590F8509869DFED5FA208211" box="[151,1420,296,321]" pageId="2" pageNumber="77" startId="2.[151,239,296,320]" targetType="table">
<paragraph id="732136BC590F8509869DFED5FA208211" blockId="2.[151,1420,296,321]" box="[151,1420,296,321]" pageId="2" pageNumber="77">
<emphasis id="41EAEAAE590F8509869DFED5FEA68211" bold="true" box="[151,266,296,320]" pageId="2" pageNumber="77">TABLE 1.</emphasis>
Morphometric data of holotype (ZSI FF 9555) and four paratypes (ZSI FF 9556) of
<taxonomicName id="B49E4D3F590F8509829BFED5FA298211" authorityName="Singh &amp; Chowdhury &amp; Gurumayum &amp; Kosygin" authorityYear="2023" box="[1169,1413,296,320]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="77" phylum="Chordata" rank="species" species="heokheei">
<emphasis id="41EAEAAE590F8509829BFED5FA298211" box="[1169,1413,296,320]" italics="true" pageId="2" pageNumber="77">Glyptothorax heokheei</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="732136BC590F850984A6FEACFA9485B8" pageId="2" pageNumber="77">
<table id="019EC41C590F7AF486A9FEACFA9485B8" box="[163,1336,337,1769]" gridcols="5" gridrows="36" pageId="2" pageNumber="77">
<tr id="CDAE34FE590F7AF486A9FEACFA948238" box="[163,1336,337,361]" gridrow="0" pageId="2" pageNumber="77" rowspan-0="1">
<th id="8E7F5D82590F7AF484A6FEACFCA68238" box="[684,778,337,361]" gridcol="1" gridrow="0" pageId="2" pageNumber="77">Holotype</th>
<th id="8E7F5D82590F7AF48576FEACFC578238" box="[892,1019,337,361]" gridcol="2" gridrow="0" pageId="2" pageNumber="77">Range</th>
<th id="8E7F5D82590F7AF4827CFEACFB1C8238" box="[1142,1200,337,361]" gridcol="3" gridrow="0" pageId="2" pageNumber="77">Mean</th>
<th id="8E7F5D82590F7AF48312FEACFA948238" box="[1304,1336,337,361]" gridcol="4" gridrow="0" pageId="2" pageNumber="77">sd</th>
</tr>
<tr id="CDAE34FE590F7AF486A9FE84FA9482C0" box="[163,1336,377,401]" gridrow="1" pageId="2" pageNumber="77" rowspan-3="1" rowspan-4="1">
<th id="8E7F5D82590F7AF486A9FE84FE4D82C0" box="[163,481,377,401]" gridcol="0" gridrow="1" pageId="2" pageNumber="77">Standard length (in mm)</th>
<td id="8E7F5D82590F7AF484A6FE84FCA682C0" box="[684,778,377,401]" gridcol="1" gridrow="1" pageId="2" pageNumber="77">152.0</td>
<td id="8E7F5D82590F7AF48576FE84FC5782C0" box="[892,1019,377,401]" gridcol="2" gridrow="1" pageId="2" pageNumber="77">102.4–152.0</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FE5CFA9482E8" box="[163,1336,417,441]" gridrow="2" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FE5CFA9482E8" box="[163,1336,417,441]" colspan="5" colspanRight="4" gridcol="0" gridrow="2" pageId="2" pageNumber="77">In percent of standard length</th>
</tr>
<tr id="CDAE34FE590F7AF486A9FE34FA9482B3" box="[163,1336,457,482]" gridrow="3" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FE34FE4D82B3" box="[163,481,457,482]" gridcol="0" gridrow="3" pageId="2" pageNumber="77">Head length</th>
<td id="8E7F5D82590F7AF484A6FE34FCA682B3" box="[684,778,457,482]" gridcol="1" gridrow="3" pageId="2" pageNumber="77">28.8</td>
<td id="8E7F5D82590F7AF48576FE34FC5782B3" box="[892,1019,457,482]" gridcol="2" gridrow="3" pageId="2" pageNumber="77">25.1–28.8</td>
<td id="8E7F5D82590F7AF4827CFE34FB1C82B3" box="[1142,1200,457,482]" gridcol="3" gridrow="3" pageId="2" pageNumber="77">27.1</td>
<td id="8E7F5D82590F7AF48312FE34FA9482B3" box="[1304,1336,457,482]" gridcol="4" gridrow="3" pageId="2" pageNumber="77">1.4</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FE0CFA94815B" box="[163,1336,497,522]" gridrow="4" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FE0CFE4D815B" box="[163,481,497,522]" gridcol="0" gridrow="4" pageId="2" pageNumber="77">Body depth at dorsal-fin origin</th>
<td id="8E7F5D82590F7AF484A6FE0CFCA6815B" box="[684,778,497,522]" gridcol="1" gridrow="4" pageId="2" pageNumber="77">14.3</td>
<td id="8E7F5D82590F7AF48576FE0CFC57815B" box="[892,1019,497,522]" gridcol="2" gridrow="4" pageId="2" pageNumber="77">11.4–15.1</td>
<td id="8E7F5D82590F7AF4827CFE0CFB1C815B" box="[1142,1200,497,522]" gridcol="3" gridrow="4" pageId="2" pageNumber="77">13.3</td>
<td id="8E7F5D82590F7AF48312FE0CFA94815B" box="[1304,1336,497,522]" gridcol="4" gridrow="4" pageId="2" pageNumber="77">1.4</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FDE7FA948163" box="[163,1336,538,562]" gridrow="5" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FDE7FE4D8163" box="[163,481,538,562]" gridcol="0" gridrow="5" pageId="2" pageNumber="77">Body depth at anus</th>
<td id="8E7F5D82590F7AF484A6FDE7FCA68163" box="[684,778,538,562]" gridcol="1" gridrow="5" pageId="2" pageNumber="77">12.8</td>
<td id="8E7F5D82590F7AF48576FDE7FC578163" box="[892,1019,538,562]" gridcol="2" gridrow="5" pageId="2" pageNumber="77">12.8–14.1</td>
<td id="8E7F5D82590F7AF4827CFDE7FB1C8163" box="[1142,1200,538,562]" gridcol="3" gridrow="5" pageId="2" pageNumber="77">13.5</td>
<td id="8E7F5D82590F7AF48312FDE7FA948163" box="[1304,1336,538,562]" gridcol="4" gridrow="5" pageId="2" pageNumber="77">0.6</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FDBFFA94810B" box="[163,1336,578,602]" gridrow="6" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FDBFFE4D810B" box="[163,481,578,602]" gridcol="0" gridrow="6" pageId="2" pageNumber="77">Predorsal length</th>
<td id="8E7F5D82590F7AF484A6FDBFFCA6810B" box="[684,778,578,602]" gridcol="1" gridrow="6" pageId="2" pageNumber="77">38.7</td>
<td id="8E7F5D82590F7AF48576FDBFFC57810B" box="[892,1019,578,602]" gridcol="2" gridrow="6" pageId="2" pageNumber="77">33.9–38.7</td>
<td id="8E7F5D82590F7AF4827CFDBFFB1C810B" box="[1142,1200,578,602]" gridcol="3" gridrow="6" pageId="2" pageNumber="77">36.2</td>
<td id="8E7F5D82590F7AF48312FDBFFA94810B" box="[1304,1336,578,602]" gridcol="4" gridrow="6" pageId="2" pageNumber="77">1.8</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FD97FA9481D3" box="[163,1336,618,642]" gridrow="7" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FD97FE4D81D3" box="[163,481,618,642]" gridcol="0" gridrow="7" pageId="2" pageNumber="77">Prepectoral length</th>
<td id="8E7F5D82590F7AF484A6FD97FCA681D3" box="[684,778,618,642]" gridcol="1" gridrow="7" pageId="2" pageNumber="77">23.2</td>
<td id="8E7F5D82590F7AF48576FD97FC5781D3" box="[892,1019,618,642]" gridcol="2" gridrow="7" pageId="2" pageNumber="77">19.6-23.2</td>
<td id="8E7F5D82590F7AF4827CFD97FB1C81D3" box="[1142,1200,618,642]" gridcol="3" gridrow="7" pageId="2" pageNumber="77">21.2</td>
<td id="8E7F5D82590F7AF48312FD97FA9481D3" box="[1304,1336,618,642]" gridcol="4" gridrow="7" pageId="2" pageNumber="77">1.5</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FD6FFA9481FA" box="[163,1336,658,683]" gridrow="8" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FD6FFE4D81FA" box="[163,481,658,683]" gridcol="0" gridrow="8" pageId="2" pageNumber="77">Prepelvic length</th>
<td id="8E7F5D82590F7AF484A6FD6FFCA681FA" box="[684,778,658,683]" gridcol="1" gridrow="8" pageId="2" pageNumber="77">54.3</td>
<td id="8E7F5D82590F7AF48576FD6FFC5781FA" box="[892,1019,658,683]" gridcol="2" gridrow="8" pageId="2" pageNumber="77">46.2–54.3</td>
<td id="8E7F5D82590F7AF4827CFD6FFB1C81FA" box="[1142,1200,658,683]" gridcol="3" gridrow="8" pageId="2" pageNumber="77">49</td>
<td id="8E7F5D82590F7AF48312FD6FFA9481FA" box="[1304,1336,658,683]" gridcol="4" gridrow="8" pageId="2" pageNumber="77">3.3</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FD46FA948182" box="[163,1336,699,723]" gridrow="9" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FD46FE4D8182" box="[163,481,699,723]" gridcol="0" gridrow="9" pageId="2" pageNumber="77">Preanal length</th>
<td id="8E7F5D82590F7AF484A6FD46FCA68182" box="[684,778,699,723]" gridcol="1" gridrow="9" pageId="2" pageNumber="77">71.4</td>
<td id="8E7F5D82590F7AF48576FD46FC578182" box="[892,1019,699,723]" gridcol="2" gridrow="9" pageId="2" pageNumber="77">61.6–71.4</td>
<td id="8E7F5D82590F7AF4827CFD46FB1C8182" box="[1142,1200,699,723]" gridcol="3" gridrow="9" pageId="2" pageNumber="77">65.6</td>
<td id="8E7F5D82590F7AF48312FD46FA948182" box="[1304,1336,699,723]" gridcol="4" gridrow="9" pageId="2" pageNumber="77">3.6</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FD1EFA9481AA" box="[163,1336,739,763]" gridrow="10" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FD1EFE4D81AA" box="[163,481,739,763]" gridcol="0" gridrow="10" pageId="2" pageNumber="77">Dorsal-fin height</th>
<td id="8E7F5D82590F7AF484A6FD1EFCA681AA" box="[684,778,739,763]" gridcol="1" gridrow="10" pageId="2" pageNumber="77">17.8</td>
<td id="8E7F5D82590F7AF48576FD1EFC5781AA" box="[892,1019,739,763]" gridcol="2" gridrow="10" pageId="2" pageNumber="77">15.8–17.8</td>
<td id="8E7F5D82590F7AF4827CFD1EFB1C81AA" box="[1142,1200,739,763]" gridcol="3" gridrow="10" pageId="2" pageNumber="77">16.6</td>
<td id="8E7F5D82590F7AF48312FD1EFA9481AA" box="[1304,1336,739,763]" gridcol="4" gridrow="10" pageId="2" pageNumber="77">0.8</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FCF6FA948072" box="[163,1336,779,803]" gridrow="11" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FCF6FE4D8072" box="[163,481,779,803]" gridcol="0" gridrow="11" pageId="2" pageNumber="77">Dorsal-fin spine length</th>
<td id="8E7F5D82590F7AF484A6FCF6FCA68072" box="[684,778,779,803]" gridcol="1" gridrow="11" pageId="2" pageNumber="77">12.9</td>
<td id="8E7F5D82590F7AF48576FCF6FC578072" box="[892,1019,779,803]" gridcol="2" gridrow="11" pageId="2" pageNumber="77">10.4–12.9</td>
<td id="8E7F5D82590F7AF4827CFCF6FB1C8072" box="[1142,1200,779,803]" gridcol="3" gridrow="11" pageId="2" pageNumber="77">11.5</td>
<td id="8E7F5D82590F7AF48312FCF6FA948072" box="[1304,1336,779,803]" gridcol="4" gridrow="11" pageId="2" pageNumber="77">1.2</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FCC9FA94801D" box="[163,1336,820,844]" gridrow="12" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FCC9FE4D801D" box="[163,481,820,844]" gridcol="0" gridrow="12" pageId="2" pageNumber="77">Dorsal-fin base length</th>
<td id="8E7F5D82590F7AF484A6FCC9FCA6801D" box="[684,778,820,844]" gridcol="1" gridrow="12" pageId="2" pageNumber="77">9.8</td>
<td id="8E7F5D82590F7AF48576FCC9FC57801D" box="[892,1019,820,844]" gridcol="2" gridrow="12" pageId="2" pageNumber="77">9.8–10.9</td>
<td id="8E7F5D82590F7AF4827CFCC9FB1C801D" box="[1142,1200,820,844]" gridcol="3" gridrow="12" pageId="2" pageNumber="77">10.4</td>
<td id="8E7F5D82590F7AF48312FCC9FA94801D" box="[1304,1336,820,844]" gridcol="4" gridrow="12" pageId="2" pageNumber="77">0.4</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FCA1FA948025" box="[163,1336,860,884]" gridrow="13" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FCA1FE4D8025" box="[163,481,860,884]" gridcol="0" gridrow="13" pageId="2" pageNumber="77">Dorsal-to-adipose fin distance</th>
<td id="8E7F5D82590F7AF484A6FCA1FCA68025" box="[684,778,860,884]" gridcol="1" gridrow="13" pageId="2" pageNumber="77">24.5</td>
<td id="8E7F5D82590F7AF48576FCA1FC578025" box="[892,1019,860,884]" gridcol="2" gridrow="13" pageId="2" pageNumber="77">19.8–25.0</td>
<td id="8E7F5D82590F7AF4827CFCA1FB1C8025" box="[1142,1200,860,884]" gridcol="3" gridrow="13" pageId="2" pageNumber="77">22.2</td>
<td id="8E7F5D82590F7AF48312FCA1FA948025" box="[1304,1336,860,884]" gridcol="4" gridrow="13" pageId="2" pageNumber="77">2.3</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FC79FA9480CD" box="[163,1336,900,924]" gridrow="14" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FC79FE4D80CD" box="[163,481,900,924]" gridcol="0" gridrow="14" pageId="2" pageNumber="77">Post-adipose distance</th>
<td id="8E7F5D82590F7AF484A6FC79FCA680CD" box="[684,778,900,924]" gridcol="1" gridrow="14" pageId="2" pageNumber="77">16.8</td>
<td id="8E7F5D82590F7AF48576FC79FC5780CD" box="[892,1019,900,924]" gridcol="2" gridrow="14" pageId="2" pageNumber="77">16.0–8.4</td>
<td id="8E7F5D82590F7AF4827CFC79FB1C80CD" box="[1142,1200,900,924]" gridcol="3" gridrow="14" pageId="2" pageNumber="77">17.2</td>
<td id="8E7F5D82590F7AF48312FC79FA9480CD" box="[1304,1336,900,924]" gridcol="4" gridrow="14" pageId="2" pageNumber="77">0.9</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FC51FA948095" box="[163,1336,940,964]" gridrow="15" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FC51FE4D8095" box="[163,481,940,964]" gridcol="0" gridrow="15" pageId="2" pageNumber="77">Adipose-fin base length</th>
<td id="8E7F5D82590F7AF484A6FC51FCA68095" box="[684,778,940,964]" gridcol="1" gridrow="15" pageId="2" pageNumber="77">10.0</td>
<td id="8E7F5D82590F7AF48576FC51FC578095" box="[892,1019,940,964]" gridcol="2" gridrow="15" pageId="2" pageNumber="77">10.0–13.1</td>
<td id="8E7F5D82590F7AF4827CFC51FB1C8095" box="[1142,1200,940,964]" gridcol="3" gridrow="15" pageId="2" pageNumber="77">12.2</td>
<td id="8E7F5D82590F7AF48312FC51FA948095" box="[1304,1336,940,964]" gridcol="4" gridrow="15" pageId="2" pageNumber="77">1.3</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FC29FA9480BC" box="[163,1336,980,1005]" gridrow="16" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FC29FE4D80BC" box="[163,481,980,1005]" gridcol="0" gridrow="16" pageId="2" pageNumber="77">Pectoral-fin length</th>
<td id="8E7F5D82590F7AF484A6FC29FCA680BC" box="[684,778,980,1005]" gridcol="1" gridrow="16" pageId="2" pageNumber="77">21.2</td>
<td id="8E7F5D82590F7AF48576FC29FC5780BC" box="[892,1019,980,1005]" gridcol="2" gridrow="16" pageId="2" pageNumber="77">16.7–21.2</td>
<td id="8E7F5D82590F7AF4827CFC29FB1C80BC" box="[1142,1200,980,1005]" gridcol="3" gridrow="16" pageId="2" pageNumber="77">18.8</td>
<td id="8E7F5D82590F7AF48312FC29FA9480BC" box="[1304,1336,980,1005]" gridcol="4" gridrow="16" pageId="2" pageNumber="77">1.9</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FC00FA948744" box="[163,1336,1021,1045]" gridrow="17" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FC00FE4D8744" box="[163,481,1021,1045]" gridcol="0" gridrow="17" pageId="2" pageNumber="77">Pelvic-fin length</th>
<td id="8E7F5D82590F7AF484A6FC00FCA68744" box="[684,778,1021,1045]" gridcol="1" gridrow="17" pageId="2" pageNumber="77">15.2</td>
<td id="8E7F5D82590F7AF48576FC00FC578744" box="[892,1019,1021,1045]" gridcol="2" gridrow="17" pageId="2" pageNumber="77">12.2–15.2</td>
<td id="8E7F5D82590F7AF4827CFC00FB1C8744" box="[1142,1200,1021,1045]" gridcol="3" gridrow="17" pageId="2" pageNumber="77">14.4</td>
<td id="8E7F5D82590F7AF48312FC00FA948744" box="[1304,1336,1021,1045]" gridcol="4" gridrow="17" pageId="2" pageNumber="77">1.2</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FBD8FA94876C" box="[163,1336,1061,1085]" gridrow="18" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FBD8FE4D876C" box="[163,481,1061,1085]" gridcol="0" gridrow="18" pageId="2" pageNumber="77">Anal-fin length</th>
<td id="8E7F5D82590F7AF484A6FBD8FCA6876C" box="[684,778,1061,1085]" gridcol="1" gridrow="18" pageId="2" pageNumber="77">17.9</td>
<td id="8E7F5D82590F7AF48576FBD8FC57876C" box="[892,1019,1061,1085]" gridcol="2" gridrow="18" pageId="2" pageNumber="77">18.5–22.2</td>
<td id="8E7F5D82590F7AF4827CFBD8FB1C876C" box="[1142,1200,1061,1085]" gridcol="3" gridrow="18" pageId="2" pageNumber="77">19.8</td>
<td id="8E7F5D82590F7AF48312FBD8FA94876C" box="[1304,1336,1061,1085]" gridcol="4" gridrow="18" pageId="2" pageNumber="77">1.8</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FBB0FA948734" box="[163,1336,1101,1125]" gridrow="19" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FBB0FE4D8734" box="[163,481,1101,1125]" gridcol="0" gridrow="19" pageId="2" pageNumber="77">Anal-fin base length</th>
<td id="8E7F5D82590F7AF484A6FBB0FCA68734" box="[684,778,1101,1125]" gridcol="1" gridrow="19" pageId="2" pageNumber="77">11.8</td>
<td id="8E7F5D82590F7AF48576FBB0FC578734" box="[892,1019,1101,1125]" gridcol="2" gridrow="19" pageId="2" pageNumber="77">11.8–14.8</td>
<td id="8E7F5D82590F7AF4827CFBB0FB1C8734" box="[1142,1200,1101,1125]" gridcol="3" gridrow="19" pageId="2" pageNumber="77">13</td>
<td id="8E7F5D82590F7AF48312FBB0FA948734" box="[1304,1336,1101,1125]" gridcol="4" gridrow="19" pageId="2" pageNumber="77">1.2</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FB88FA9487DF" box="[163,1336,1141,1166]" gridrow="20" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FB88FE4D87DF" box="[163,481,1141,1166]" gridcol="0" gridrow="20" pageId="2" pageNumber="77">Caudal-peduncle length</th>
<td id="8E7F5D82590F7AF484A6FB88FCA687DF" box="[684,778,1141,1166]" gridcol="1" gridrow="20" pageId="2" pageNumber="77">17.4</td>
<td id="8E7F5D82590F7AF48576FB88FC5787DF" box="[892,1019,1141,1166]" gridcol="2" gridrow="20" pageId="2" pageNumber="77">17.4–22.9</td>
<td id="8E7F5D82590F7AF4827CFB88FB1C87DF" box="[1142,1200,1141,1166]" gridcol="3" gridrow="20" pageId="2" pageNumber="77">19.9</td>
<td id="8E7F5D82590F7AF48312FB88FA9487DF" box="[1304,1336,1141,1166]" gridcol="4" gridrow="20" pageId="2" pageNumber="77">2.1</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FB63FA9487E7" box="[163,1336,1182,1206]" gridrow="21" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FB63FE4D87E7" box="[163,481,1182,1206]" gridcol="0" gridrow="21" pageId="2" pageNumber="77">Caudal-peduncle depth</th>
<td id="8E7F5D82590F7AF484A6FB63FCA687E7" box="[684,778,1182,1206]" gridcol="1" gridrow="21" pageId="2" pageNumber="77">6.6</td>
<td id="8E7F5D82590F7AF48576FB63FC5787E7" box="[892,1019,1182,1206]" gridcol="2" gridrow="21" pageId="2" pageNumber="77">6.6–8.6</td>
<td id="8E7F5D82590F7AF4827CFB63FB1C87E7" box="[1142,1200,1182,1206]" gridcol="3" gridrow="21" pageId="2" pageNumber="77">8.08</td>
<td id="8E7F5D82590F7AF48312FB63FA9487E7" box="[1304,1336,1182,1206]" gridcol="4" gridrow="21" pageId="2" pageNumber="77">0.8</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FB3BFA94878F" box="[163,1336,1222,1246]" gridrow="22" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FB3BFE4D878F" box="[163,481,1222,1246]" gridcol="0" gridrow="22" pageId="2" pageNumber="77">Caudal-fin length</th>
<td id="8E7F5D82590F7AF484A6FB3BFCA6878F" box="[684,778,1222,1246]" gridcol="1" gridrow="22" pageId="2" pageNumber="77">24.1</td>
<td id="8E7F5D82590F7AF48576FB3BFC57878F" box="[892,1019,1222,1246]" gridcol="2" gridrow="22" pageId="2" pageNumber="77">24.1–29.1</td>
<td id="8E7F5D82590F7AF4827CFB3BFB1C878F" box="[1142,1200,1222,1246]" gridcol="3" gridrow="22" pageId="2" pageNumber="77">25.9</td>
<td id="8E7F5D82590F7AF48312FB3BFA94878F" box="[1304,1336,1222,1246]" gridcol="4" gridrow="22" pageId="2" pageNumber="77">2.1</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FB13FA948657" box="[163,1336,1262,1286]" gridrow="23" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FB13FE4D8657" box="[163,481,1262,1286]" gridcol="0" gridrow="23" pageId="2" pageNumber="77">Adhesive apparatus length</th>
<td id="8E7F5D82590F7AF484A6FB13FCA68657" box="[684,778,1262,1286]" gridcol="1" gridrow="23" pageId="2" pageNumber="77">9.7</td>
<td id="8E7F5D82590F7AF48576FB13FC578657" box="[892,1019,1262,1286]" gridcol="2" gridrow="23" pageId="2" pageNumber="77">9.5–10.3</td>
<td id="8E7F5D82590F7AF4827CFB13FB1C8657" box="[1142,1200,1262,1286]" gridcol="3" gridrow="23" pageId="2" pageNumber="77">9.8</td>
<td id="8E7F5D82590F7AF48312FB13FA948657" box="[1304,1336,1262,1286]" gridcol="4" gridrow="23" pageId="2" pageNumber="77">0.3</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FAEBFA94867E" box="[163,1336,1302,1327]" gridrow="24" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FAEBFE4D867E" box="[163,481,1302,1327]" gridcol="0" gridrow="24" pageId="2" pageNumber="77">Adhesive apparatus width</th>
<td id="8E7F5D82590F7AF484A6FAEBFCA6867E" box="[684,778,1302,1327]" gridcol="1" gridrow="24" pageId="2" pageNumber="77">10.6</td>
<td id="8E7F5D82590F7AF48576FAEBFC57867E" box="[892,1019,1302,1327]" gridcol="2" gridrow="24" pageId="2" pageNumber="77">10.6–12.1</td>
<td id="8E7F5D82590F7AF4827CFAEBFB1C867E" box="[1142,1200,1302,1327]" gridcol="3" gridrow="24" pageId="2" pageNumber="77">11.7</td>
<td id="8E7F5D82590F7AF48312FAEBFA94867E" box="[1304,1336,1302,1327]" gridcol="4" gridrow="24" pageId="2" pageNumber="77">0.6</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FAC2FA948606" box="[163,1336,1343,1367]" gridrow="25" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FAC2FA948606" box="[163,1336,1343,1367]" colspan="5" colspanRight="4" gridcol="0" gridrow="25" pageId="2" pageNumber="77">In percent of head length</th>
</tr>
<tr id="CDAE34FE590F7AF486A9FA9AFA94862E" box="[163,1336,1383,1407]" gridrow="26" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FA9AFE4D862E" box="[163,481,1383,1407]" gridcol="0" gridrow="26" pageId="2" pageNumber="77">Head depth at occiput</th>
<td id="8E7F5D82590F7AF484A6FA9AFCA6862E" box="[684,778,1383,1407]" gridcol="1" gridrow="26" pageId="2" pageNumber="77">39</td>
<td id="8E7F5D82590F7AF48576FA9AFC57862E" box="[892,1019,1383,1407]" gridcol="2" gridrow="26" pageId="2" pageNumber="77">35–43</td>
<td id="8E7F5D82590F7AF4827CFA9AFB1C862E" box="[1142,1200,1383,1407]" gridcol="3" gridrow="26" pageId="2" pageNumber="77">40.1</td>
<td id="8E7F5D82590F7AF48312FA9AFA94862E" box="[1304,1336,1383,1407]" gridcol="4" gridrow="26" pageId="2" pageNumber="77">3.3</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FA72FA9486F6" box="[163,1336,1423,1447]" gridrow="27" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FA72FE4D86F6" box="[163,481,1423,1447]" gridcol="0" gridrow="27" pageId="2" pageNumber="77">Head width (maximum)</th>
<td id="8E7F5D82590F7AF484A6FA72FCA686F6" box="[684,778,1423,1447]" gridcol="1" gridrow="27" pageId="2" pageNumber="77">74</td>
<td id="8E7F5D82590F7AF48576FA72FC5786F6" box="[892,1019,1423,1447]" gridcol="2" gridrow="27" pageId="2" pageNumber="77">67–81</td>
<td id="8E7F5D82590F7AF4827CFA72FB1C86F6" box="[1142,1200,1423,1447]" gridcol="3" gridrow="27" pageId="2" pageNumber="77">72.3</td>
<td id="8E7F5D82590F7AF48312FA72FA9486F6" box="[1304,1336,1423,1447]" gridcol="4" gridrow="27" pageId="2" pageNumber="77">5.4</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FA4AFA948681" box="[163,1336,1463,1488]" gridrow="28" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FA4AFE4D8681" box="[163,481,1463,1488]" gridcol="0" gridrow="28" pageId="2" pageNumber="77">Orbital diameter</th>
<td id="8E7F5D82590F7AF484A6FA4AFCA68681" box="[684,778,1463,1488]" gridcol="1" gridrow="28" pageId="2" pageNumber="77">6</td>
<td id="8E7F5D82590F7AF48576FA4AFC578681" box="[892,1019,1463,1488]" gridcol="2" gridrow="28" pageId="2" pageNumber="77">6–11</td>
<td id="8E7F5D82590F7AF4827CFA4AFB1C8681" box="[1142,1200,1463,1488]" gridcol="3" gridrow="28" pageId="2" pageNumber="77">8.28</td>
<td id="8E7F5D82590F7AF48312FA4AFA948681" box="[1304,1336,1463,1488]" gridcol="4" gridrow="28" pageId="2" pageNumber="77">1.9</td>
</tr>
<tr id="CDAE34FE590F7AF486A9FA1DFA9486A9" box="[163,1336,1504,1528]" gridrow="29" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9FA1DFE4D86A9" box="[163,481,1504,1528]" gridcol="0" gridrow="29" pageId="2" pageNumber="77">Snout length</th>
<td id="8E7F5D82590F7AF484A6FA1DFCA686A9" box="[684,778,1504,1528]" gridcol="1" gridrow="29" pageId="2" pageNumber="77">52</td>
<td id="8E7F5D82590F7AF48576FA1DFC5786A9" box="[892,1019,1504,1528]" gridcol="2" gridrow="29" pageId="2" pageNumber="77">44–52</td>
<td id="8E7F5D82590F7AF4827CFA1DFB1C86A9" box="[1142,1200,1504,1528]" gridcol="3" gridrow="29" pageId="2" pageNumber="77">46.5</td>
<td id="8E7F5D82590F7AF48312FA1DFA9486A9" box="[1304,1336,1504,1528]" gridcol="4" gridrow="29" pageId="2" pageNumber="77">3.4</td>
</tr>
<tr id="CDAE34FE590F7AF486A9F9F5FA948571" box="[163,1336,1544,1568]" gridrow="30" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9F9F5FE4D8571" box="[163,481,1544,1568]" gridcol="0" gridrow="30" pageId="2" pageNumber="77">Interorbital width</th>
<td id="8E7F5D82590F7AF484A6F9F5FCA68571" box="[684,778,1544,1568]" gridcol="1" gridrow="30" pageId="2" pageNumber="77">20</td>
<td id="8E7F5D82590F7AF48576F9F5FC578571" box="[892,1019,1544,1568]" gridcol="2" gridrow="30" pageId="2" pageNumber="77">12–20</td>
<td id="8E7F5D82590F7AF4827CF9F5FB1C8571" box="[1142,1200,1544,1568]" gridcol="3" gridrow="30" pageId="2" pageNumber="77">15.3</td>
<td id="8E7F5D82590F7AF48312F9F5FA948571" box="[1304,1336,1544,1568]" gridcol="4" gridrow="30" pageId="2" pageNumber="77">2.9</td>
</tr>
<tr id="CDAE34FE590F7AF486A9F9CDFA948519" box="[163,1336,1584,1608]" gridrow="31" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9F9CDFE4D8519" box="[163,481,1584,1608]" gridcol="0" gridrow="31" pageId="2" pageNumber="77">Nasal barbel length</th>
<td id="8E7F5D82590F7AF484A6F9CDFCA68519" box="[684,778,1584,1608]" gridcol="1" gridrow="31" pageId="2" pageNumber="77">31</td>
<td id="8E7F5D82590F7AF48576F9CDFC578519" box="[892,1019,1584,1608]" gridcol="2" gridrow="31" pageId="2" pageNumber="77">31–51</td>
<td id="8E7F5D82590F7AF4827CF9CDFB1C8519" box="[1142,1200,1584,1608]" gridcol="3" gridrow="31" pageId="2" pageNumber="77">40.3</td>
<td id="8E7F5D82590F7AF48312F9CDFA948519" box="[1304,1336,1584,1608]" gridcol="4" gridrow="31" pageId="2" pageNumber="77">8.3</td>
</tr>
<tr id="CDAE34FE590F7AF486A9F9A5FA948520" box="[163,1336,1624,1649]" gridrow="32" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9F9A5FE4D8520" box="[163,481,1624,1649]" gridcol="0" gridrow="32" pageId="2" pageNumber="77">Maxillary barbel length</th>
<td id="8E7F5D82590F7AF484A6F9A5FCA68520" box="[684,778,1624,1649]" gridcol="1" gridrow="32" pageId="2" pageNumber="77">65</td>
<td id="8E7F5D82590F7AF48576F9A5FC578520" box="[892,1019,1624,1649]" gridcol="2" gridrow="32" pageId="2" pageNumber="77">65–89</td>
<td id="8E7F5D82590F7AF4827CF9A5FB1C8520" box="[1142,1200,1624,1649]" gridcol="3" gridrow="32" pageId="2" pageNumber="77">78.8</td>
<td id="8E7F5D82590F7AF48312F9A5FA948520" box="[1304,1336,1624,1649]" gridcol="4" gridrow="32" pageId="2" pageNumber="77">10</td>
</tr>
<tr id="CDAE34FE590F7AF486A9F97CFA9485C8" box="[163,1336,1665,1689]" gridrow="33" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9F97CFE4D85C8" box="[163,481,1665,1689]" gridcol="0" gridrow="33" pageId="2" pageNumber="77">Inner mandibular barbel length</th>
<td id="8E7F5D82590F7AF484A6F97CFCA685C8" box="[684,778,1665,1689]" gridcol="1" gridrow="33" pageId="2" pageNumber="77">19</td>
<td id="8E7F5D82590F7AF48576F97CFC5785C8" box="[892,1019,1665,1689]" gridcol="2" gridrow="33" pageId="2" pageNumber="77">19–30</td>
<td id="8E7F5D82590F7AF4827CF97CFB1C85C8" box="[1142,1200,1665,1689]" gridcol="3" gridrow="33" pageId="2" pageNumber="77">26.4</td>
<td id="8E7F5D82590F7AF48312F97CFA9485C8" box="[1304,1336,1665,1689]" gridcol="4" gridrow="33" pageId="2" pageNumber="77">4.4</td>
</tr>
<tr id="CDAE34FE590F7AF486A9F954FA948590" box="[163,1336,1705,1729]" gridrow="34" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9F954FE4D8590" box="[163,481,1705,1729]" gridcol="0" gridrow="34" pageId="2" pageNumber="77">Outer mandibular barbel length</th>
<td id="8E7F5D82590F7AF484A6F954FCA68590" box="[684,778,1705,1729]" gridcol="1" gridrow="34" pageId="2" pageNumber="77">37</td>
<td id="8E7F5D82590F7AF48576F954FC578590" box="[892,1019,1705,1729]" gridcol="2" gridrow="34" pageId="2" pageNumber="77">37–63</td>
<td id="8E7F5D82590F7AF4827CF954FB1C8590" box="[1142,1200,1705,1729]" gridcol="3" gridrow="34" pageId="2" pageNumber="77">52.7</td>
<td id="8E7F5D82590F7AF48312F954FA948590" box="[1304,1336,1705,1729]" gridcol="4" gridrow="34" pageId="2" pageNumber="77">11</td>
</tr>
<tr id="CDAE34FE590F7AF486A9F92CFA9485B8" box="[163,1336,1745,1769]" gridrow="35" pageId="2" pageNumber="77">
<th id="8E7F5D82590F7AF486A9F92CFE4D85B8" box="[163,481,1745,1769]" gridcol="0" gridrow="35" pageId="2" pageNumber="77">Mouth width</th>
<td id="8E7F5D82590F7AF484A6F92CFCA685B8" box="[684,778,1745,1769]" gridcol="1" gridrow="35" pageId="2" pageNumber="77">46</td>
<td id="8E7F5D82590F7AF48576F92CFC5785B8" box="[892,1019,1745,1769]" gridcol="2" gridrow="35" pageId="2" pageNumber="77">31–46</td>
<td id="8E7F5D82590F7AF4827CF92CFB1C85B8" box="[1142,1200,1745,1769]" gridcol="3" gridrow="35" pageId="2" pageNumber="77">35.8</td>
<td id="8E7F5D82590F7AF48312F92CFA9485B8" box="[1304,1336,1745,1769]" gridcol="4" gridrow="35" pageId="2" pageNumber="77">6.1</td>
</tr>
</table>
</paragraph>
<paragraph id="732136BC590F850986CDF8C1FD2A84B6" blockId="2.[151,1437,1852,2023]" pageId="2" pageNumber="77">
<emphasis id="41EAEAAE590F850986CDF8C1FEE38407" bold="true" box="[199,335,1852,1878]" pageId="2" pageNumber="77">Coloration.</emphasis>
In 70% alcohol, dorsal surface of head and body yellowish brown; lateral surface light brown, fading to yellowish cream ventrally. Dorsal and lateral surfaces of head and body are mottled with dark brown spots. Anterior nuchal plate element is visible as a distinct yellowish arrow when viewed dorsally. Adipose fin pale yellow with dark brown base. Dorsal fin with an indistinct black band on distal half. Pectoral, pelvic, anal and caudal fins are dusky or mottled with dark brown spots.
</paragraph>
</subSubSection>
<caption id="27E16634590E8508869DF868FAEE84FD" ID-DOI="http://doi.org/10.5281/zenodo.10301474" ID-Zenodo-Dep="10301474" box="[151,1346,1941,1965]" httpUri="https://zenodo.org/record/10301474/files/figure.png" pageId="3" pageNumber="78" startId="3.[151,250,1941,1965]" targetBox="[151,1436,181,1916]" targetPageId="3" targetType="figure">
<paragraph id="732136BC590E8508869DF868FAEE84FD" blockId="3.[151,1346,1941,1965]" box="[151,1346,1941,1965]" pageId="3" pageNumber="78">
<emphasis id="41EAEAAE590E8508869DF868FEBF84FC" bold="true" box="[151,275,1941,1965]" pageId="3" pageNumber="78">FIGURE 1.</emphasis>
<taxonomicName id="B49E4D3F590E85088713F868FDFA84FD" authority=", ZSI FF" authorityName="ZSI FF" box="[281,598,1941,1965]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="78" phylum="Chordata" rank="species" species="heokheei">
<emphasis id="41EAEAAE590E85088713F868FDAD84FC" box="[281,513,1941,1965]" italics="true" pageId="3" pageNumber="78">Glyptothorax heokheei</emphasis>
, ZSI FF
</taxonomicName>
9555, holotype, 152.0 mm SL: a, dorsal; b, lateral; and c, ventral aspects.
</paragraph>
</caption>
<caption id="27E166345909850F869DFBDAFC4C8732" ID-DOI="http://doi.org/10.5281/zenodo.10301480" ID-Zenodo-Dep="10301480" httpUri="https://zenodo.org/record/10301480/files/figure.png" pageId="4" pageNumber="79" startId="4.[151,250,1063,1087]" targetBox="[161,1426,191,1029]" targetPageId="4" targetType="figure">
<paragraph id="732136BC5909850F869DFBDAFC4C8732" blockId="4.[151,1437,1063,1124]" pageId="4" pageNumber="79">
<emphasis id="41EAEAAE5909850F869DFBDAFEB9876E" bold="true" box="[151,277,1063,1087]" pageId="4" pageNumber="79">FIGURE 2.</emphasis>
Thoracic adhesive apparatus of a.
<taxonomicName id="B49E4D3F5909850F8475FBDAFCC5876E" authority="Singh &amp; Chowdhury &amp; Gurumayum &amp; Kosygin, 2023" authorityName="Singh &amp; Chowdhury &amp; Gurumayum &amp; Kosygin" authorityYear="2023" box="[639,873,1063,1087]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="79" phylum="Chordata" rank="species" species="heokheei" status="sp. nov.">
<emphasis id="41EAEAAE5909850F8475FBDAFCC5876E" box="[639,873,1063,1087]" italics="true" pageId="4" pageNumber="79">Glyptothorax heokheei</emphasis>
</taxonomicName>
<emphasis id="41EAEAAE5909850F857BFBD5FC6F8711" bold="true" box="[881,963,1064,1088]" pageId="4" pageNumber="79">
<taxonomicNameLabel id="5AD957D55909850F857BFBD5FC6F8711" box="[881,963,1064,1088]" pageId="4" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ZSI FF 9556, paratype, 124.1 mm SL; b.
<taxonomicName id="B49E4D3F5909850F838EFBD5FECE8732" authority=", ZSI V" authorityName="ZSI V" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="79" phylum="Chordata" rank="species" species="dikrongensis">
<emphasis id="41EAEAAE5909850F838EFBD5FEB48732" italics="true" pageId="4" pageNumber="79">G. dikrongensis</emphasis>
, ZSI V
</taxonomicName>
/APFS/P-515, holotype, 94.2 mm SL. Illustrations not to scale.
</paragraph>
</caption>
<subSubSection id="3B8465375909850F86CDFB48FA9787A5" pageId="4" pageNumber="79" type="distribution">
<paragraph id="732136BC5909850F86CDFB48FA9787A5" blockId="4.[151,1436,1205,1376]" pageId="4" pageNumber="79">
<emphasis id="41EAEAAE5909850F86CDFB48FECD879E" bold="true" box="[199,353,1205,1231]" pageId="4" pageNumber="79">Distribution.</emphasis>
<taxonomicName id="B49E4D3F5909850F876DFB4BFDC48781" authorityName="Singh &amp; Chowdhury &amp; Gurumayum &amp; Kosygin" authorityYear="2023" box="[359,616,1206,1232]" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="79" phylum="Chordata" rank="species" species="heokheei">
<emphasis id="41EAEAAE5909850F876DFB4BFDC48781" box="[359,616,1206,1232]" italics="true" pageId="4" pageNumber="79">Glyptothorax heokheei</emphasis>
</taxonomicName>
is presently known only from the
<typeStatus id="AC25881E5909850F85EDFB4BFBBB8781" box="[999,1047,1206,1232]" pageId="4" pageNumber="79">type</typeStatus>
locality, Siku stream near Mebo, a tributary of the Siang River, Brahmaputra drainage, East Siang District,
<collectingRegion id="B15AF85E5909850F85B4FB27FB3A87A5" box="[958,1174,1242,1268]" country="India" name="Arunachal Pradesh" pageId="4" pageNumber="79">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="0B89762C5909850F82AAFB27FB7687A5" box="[1184,1242,1242,1268]" name="India" pageId="4" pageNumber="79">India</collectingCountry>
(
<figureCitation id="EBA52A395909850F82E3FB27FA8187A5" box="[1257,1325,1242,1268]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[159,1436,193,1783]" captionTargetId="figure-17@5.[151,1436,181,1783]" captionTargetPageId="5" captionText="FIGURE 3. Map showing the type locality of Glyptothorax heokheei." figureDoi="http://doi.org/10.5281/zenodo.10301482" httpUri="https://zenodo.org/record/10301482/files/figure.png" pageId="4" pageNumber="79">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3B8465375909850F86CDFB03FEAD8631" pageId="4" pageNumber="79" type="etymology">
<paragraph id="732136BC5909850F86CDFB03FEAD8631" blockId="4.[151,1436,1205,1376]" pageId="4" pageNumber="79">
<emphasis id="41EAEAAE5909850F86CDFB03FEE28649" bold="true" box="[199,334,1278,1304]" pageId="4" pageNumber="79">Etymology.</emphasis>
The species is named after Heok Hee Ng of the Lee Kong Chian Natural History Museum, National University of
<collectingCountry id="0B89762C5909850F8732FADFFE06866D" box="[312,426,1314,1340]" name="Singapore" pageId="4" pageNumber="79">Singapore</collectingCountry>
(NUS), honouring his valuable contributions to the taxonomy and systematics of Asian catfishes.
</paragraph>
</subSubSection>
</treatment>
</document>
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